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Page 1: 6 U E S Z B F A O I G C D 8 7 B Z V W E F T S D I C A G J 9 R D B C 7 F 8 A G V k B …gene/papers/Buckley1997_manam_stress.pdf · 2010-11-15 · 8 7 f f r 9 < @ g >? b: s c

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� � � � � £ � � � � � � ¦ � � � � � � © � � �   ª � ¦ � � � �   « � � � ª � � � � � ¦ � � � �   � � � � � �   � © � � � � � � � � � ¥ , K L M 2 / -   � � � � �  � � �   � � � � � � � � �     � � � � � � � � � � « � � � � �   � � ¢ � � �   « � � � ª � � ª « � N O O P � / �   � �     � ª � � � > � ¨ � � � � � ? �   � � �   � � � � � � �

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� � � � � � � �   � � ¦ � � �   « � � � ª � � �   � � � � �   � � ª « � � « � ¢ � � � � �   � � � �   � ¡ � ¡ � � �Q ° + � � � � � � � � �   �   � � � � � � � � � � � � � � � � �   ¡ � � � ¨ �   � � ¦ � ¦ � «   F � « � � � � ¨ � � � � � � � � � ¬ � � £ � 5 ¥   ¡ � � � ¨ �   ­ � � �

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> 8 > < C 7 J ) A 8 7 F @ < G . 7 < Y @ A G < > : V6 F G 8 A F < J µ < 9 9 < 8 I { V # N N f V a ¢ Ù × Ò % Ñ Í Ð Ô Ð Ö Ò ß Ò Ó Ñ Ø Ò Õ Ò ° ÿ V ¹ C B > C A 8 9 F < G G @ A > 8 > < C 7 J H 6 W J \ 8 I S A < F = @ J H 8 G G V6 7 Z @ 9 8 G J a ? 8 A C 7 V # N N g V $ U B @ R > < C 7 8 9 G > A @ G G c 8 > > A 8 B > < 7 = G T D D < U @ G < 7 W T A Z < G ? X A @ R A @ G @ 7 > 8 > < C 7 G Y G V > ? @ = A 8 I c

I 8 A V W C 8 R R @ 8 A < 7 L 8 = @ A @ > 8 9 V # N N Î V6 > � J { T 7 Z C J 8 7 F j A I < 7 H @ G > @ A V # N N g V z @ 8 9 < = 7 I @ 7 > V W C 8 R R @ 8 A < 7 L 8 = @ A @ > 8 9 V # N N Î V6 > � J { T 7 Z C J 8 7 F j A I < 7 H @ G > @ A V # N N ^ V W ? @ B C A @ c R @ A < R ? @ A : G > A T B > T A @ C D > ? @ 9 @ U < B C 7 8 7 F B C 7 G > A 8 < 7 > G C 7

A @ A 8 7 Z < 7 = V 6 7 ¼ ß Ö ½ Ù Ø Õ Ö Ð ÿ Ò Ó £ Ô Õ Õ Ô × ¢ Í Õ Ù Ð Ð Õ Ï × × Ô Õ Ö Ò ß Ô Ú ² Ô Ñ Ù Ø Õ # ] J # ] # c f º N V6 > C J � T B < 9 9 @ 6 V # N ] N V H 8 7 8 I G > A @ G G X > ? @ B : B 9 @ 8 7 F @ U > A 8 I @ > A < B 8 9 < > : V 6 7 ² ¢ Ò ß Ò Ú Ò � ÿ Ô Ð d Ô ß Ð Ô Þ Ø Í � # J b ^ c ^ N V

a : 7 > 8 U z @ G @ 8 A B ? \ @ 7 > @ A J . 7 < Y @ A G < > : C D \ 8 9 < D C A 7 < 8 J a 8 7 > 8 \ A T ; VL 8 = @ A J z @ 7 � V # N ] N V º % Ù Ð Ø Ö × Ô Ú Ð ¢ Ù Ò Ø ÿ Ò Ó Õ Ð Ø Ù Õ Õ Ô ß ° ° Ù Õ Ð Ø Ù Õ Õ Ö ß � Ö ß ¶ ß � Ú Ö Õ ¢ Ô ß ° b Í Ð × ¢ V ¹ C B > C A 8 9 F < G G @ A c

> 8 > < C 7 J z @ � Z G T 7 < Y @ A G < > @ < > > @ . > A @ B ? > V ¹ < G > A < S T > @ F S : 6 \ T 4 A < 7 > < 7 = J ¹ C A F A @ B ? > VL 8 = @ A J z @ 7 � V # N N b V j 9 > @ A 7 8 > < Y @ G > C > ? @ < 8 I S < B c > A C B ? 8 < B 9 8 E V � Ô Ð Í Ø Ô Ú ¯ Ô ß � Í Ô � Ù Ô ß ° ¯ Ö ß � Í Ö Õ Ð Ö × a ¢ Ù Ò Ø ÿ

# # X b ] # c g b f VL 8 = @ A J z @ 7 � J K 8 A A : Y 8 7 F @ A K T 9 G > J 8 7 F µ < I ¾ C 7 7 @ Y @ 9 F J @ F G V # N N Î V a ¢ Ù Ñ Ø Ò Õ Ò ° ÿ æ % Ò Ø Ñ ¢ Ò Ú Ò � ÿ Ö ß Ð Ù Ø Ó Ô × Ù V

\ 8 I S A < F = @ X \ 8 I S A < F = @ . 7 < Y @ A G < > : 4 A @ G G V� 8 F @ D C = @ F J 4 @ > @ A J 8 7 F 6 8 7 H 8 F F < @ G C 7 V # N N k V a ¢ Ù Õ Ò Í ß ° Õ Ò Ó Ð ¢ Ù É Ò Ø Ú ° ¿ Õ Ú Ô ß � Í Ô � Ù Õ V \ 8 I S A < F = @ J H 8 G G V X

) 9 8 B Z E @ 9 9 V� < B ? > @ 7 S @ A Z J [ A 8 7 > < G @ Z V # N ] b V º � Ø Ô % % Ô Ø Ò Ó £ Ô ß Ô % V K C 7 C 9 T 9 T X . 7 < Y @ A G < > : C D K 8 E 8 < < 4 A @ G G VH B \ 8 A > ? : J { C ? 7 { V J 8 7 F j 9 8 7 a V 4 A < 7 B @ V # N N b 8 V T @ 7 @ A 8 9 < ; @ F 8 9 < = 7 I @ 7 > V 6 7 À Ù Ô Ø Ë Ò Ò X Ò Ó £ Ò Ø Ñ ¢ Ò Ú Ò � ÿ J @ F V

T @ @ A > ) C C < � 8 7 F { 8 8 R Y 8 7 H 8 A 9 @ J Î N c # ^ b V ¹ C A F A @ B ? > X L 9 T E @ A VH B \ 8 A > ? : J { C ? 7 { V J 8 7 F j 9 8 7 a V 4 A < 7 B @ V # N N b S V ² Ø Ò Õ Ò ° Ö × % Ò Ø Ñ ¢ Ò Ú Ò � ÿ ´ » × Ò ß Õ Ð Ø Ô Ö ß Ð Ö ß Ð Ù Ø Ô × Ð Ö Ò ß Ô ß ° Õ Ô Ð Ö Õ æ

Ó Ô × Ð Ö Ò ß V H G V J . 7 < Y @ A G < > : C D H 8 G G 8 B ? T G @ > > G J j I ? @ A G > J 8 7 F z T > = @ A G . 7 < Y @ A G < > : VH B \ 8 A > ? : J { C ? 7 { V J 8 7 F j 9 8 7 a V 4 A < 7 B @ V # N N g V j 7 C Y @ A Y < @ E C D R A C G C F < B I C A R ? C 9 C = : V W C 8 R R @ 8 A < 7 L 8 = @ A @ >

8 9 V # N N Î VH B \ 8 A > ? : J { C ? 7 { V J 8 7 F j 9 8 7 a V 4 A < 7 B @ V # N N ^ V [ 8 < > ? D T 9 7 @ G G 8 7 F A @ F T R 9 < B 8 > < Y @ < F @ 7 > < > : V 6 7 ¼ ß Ö ½ Ù Ø Õ Ö Ð ÿ Ò Ó

£ Ô Õ Õ Ô × ¢ Í Õ Ù Ð Ð Õ Ï × × Ô Õ Ö Ò ß Ô Ú ² Ô Ñ Ù Ø Õ # ] J f g N c b ] g VH B K T = ? J ) A < 8 7 V # N N b V ` R > < I 8 9 G 8 > < G D 8 B > < C 7 C D G T S B 8 > @ = C A < ; 8 > < C 7 < 7 K 8 T G 8 R 9 T A 8 9 G V 4 8 R @ A R A @ G @ 7 > @ F 8 >

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y @ G R C A J H 8 A < 7 8 J 8 7 F 6 A @ 7 @ x C = @ 9 V # N ] k V ² Ø Ò Õ Ò ° Ö × Ñ ¢ Ò ß Ò Ú Ò � ÿ V ¹ C A F A @ B ? > X [ C A < G V` A = T 7 J ` A ? 8 7 V # N N k V d Ö � ß æ Á Ô Õ Ù ° £ Ò Ø Ñ ¢ Ò Ú Ò � ÿ Ô ß ° ² ¢ Ò ß Ò Ú Ò � ÿ É Ö Ð ¢ Õ Ñ Ù × Ö Ô Ú Ø Ù Ó Ù Ø Ù ß × Ù Ð Ò Ï Ñ Ð Ö % Ô Ú Ö Ð ÿ

a ¢ Ù Ò Ø ÿ V ¹ C B > C A 8 9 F < G G @ A > 8 > < C 7 J . 7 < Y @ A G < > : C D \ 8 9 < D C A 7 < 8 J ) @ A Z @ 9 @ : V4 A < 7 B @ J j 9 8 7 a V # N ] b V z @ 9 8 > < 7 = > C > ? @ = A < F V ¯ Ö ß � Í Ö Õ Ð Ö × ´ ß µ Í Ö Ø ÿ # g X # N c # º º V4 A < 7 B @ J j 9 8 7 a V # N ] ^ V 6 I R A C Y < 7 = > A @ @ > ? @ C A : V 6 7 ² Ø Ò × Ù Ù ° Ö ß � Õ Ò Ó Ð ¢ Ù Ô ß ß Í Ô Ú % Ù Ù Ð Ö ß � Ò Ó Ð ¢ Ù Á Ù Ø X Ù Ú Ù ÿ

¯ Ö ß � Í Ö Õ Ð Ö × Õ d Ò × Ö Ù Ð ÿ # # J g Î # c g N º V ) @ A Z @ 9 @ : � < 7 = T < G > < B G a C B < @ > : J . 7 < Y @ A G < > : C D \ 8 9 < D C A 7 < 8 J ) @ A Z @ 9 @ : V4 A < 7 B @ J j 9 8 7 a V # N N # V ¹ T 8 7 > < > 8 > < Y @ B C 7 G @ _ T @ 7 B @ G C D A ? : > ? I < B C A = 8 7 < ; 8 > < C 7 V 6 7 ² Ô Ø Ô Õ Ù Õ Õ Ö Ò ß Ò ß Ð ¢ Ù

Õ ÿ Ú Ú Ô Ë Ú Ù Ö ß Ñ ¢ Ò ß Ù Ð Ö × Õ Ô ß ° Ñ ¢ Ò ß Ò Ú Ò � ÿ J b ^ ^ c b N ] V \ ? < B 8 = C � < 7 = T < G > < B a C B < @ > : J . 7 < Y @ A G < > : C D \ ? < B 8 = C J\ ? < B 8 = C J 6 9 9 V

4 A < 7 B @ J j 9 8 7 a V J 8 7 F 4 8 T 9 a I C 9 @ 7 G Z : V # N N # V ` R > < I 8 9 < > : V 4 8 R @ A R A @ G @ 7 > @ F 8 > > ? @ j A < ; C 7 8 4 ? C 7 C 9 C = : \ C 7 cD @ A @ 7 B @ J . 7 < Y @ A G < > : C D j A < ; C 7 8 J W T B G C 7 J j R A < 9 ^ c k V

4 A < 7 B @ J j 9 8 7 a V J 8 7 F 4 8 T 9 a I C 9 @ 7 G Z : V # N N b V Ï Ñ Ð Ö % Ô Ú Ö Ð ÿ a ¢ Ù Ò Ø ÿ » × Ò ß Õ Ð Ø Ô Ö ß Ð Ö ß Ð Ù Ø Ô × Ð Ö Ò ß Ö ß � Ù ß Ù Ø Ô Ð Ö ½ Ù� Ø Ô % % Ô Ø V H G V J z T > = @ A G . 7 < Y @ A G < > : 8 7 F . 7 < Y @ A G < > : C D \ C 9 C A 8 F C J ) C T 9 F @ A V

a @ 9 Z < A Z J $ 9 < G 8 S @ > ? V # N N ^ V W ? @ R A C G C F < B G > A T B > T A @ C D D T 7 B > < C 7 E C A F G V 6 7 ¼ ß Ö ½ Ù Ø Õ Ö Ð ÿ Ò Ó £ Ô Õ Õ Ô × ¢ Í Õ Ù Ð Ð ÕÏ × × Ô Õ Ö Ò ß Ô Ú ² Ô Ñ Ù Ø Õ # ] J g b N c g k N V

a R 8 @ 9 > < J 4 ? < 9 < R V # N N b V µ @ 8 Z @ F = @ G 8 7 F D < 7 8 9 = @ I < 7 8 > @ G < 7 a E < G G T @ A I 8 7 V 6 7 ² Ø Ò × Ù Ù ° Ö ß � Õ Ò Ó � ¶ ¯ d f g J Y C 9 Vf J ^ Î b c ^ ] ] V

$ T = @ 7 @ ) T B Z 9 @ :¹ @ R 8 A > I @ 7 > C D � < 7 = T < G > < B Gk # N µ < 9 9 < 8 I G K 8 9 9. 7 < Y @ A G < > : C D 4 @ 7 7 G : 9 Y 8 7 < 84 ? < 9 8 F @ 9 R ? < 8 J 4 j # N # º g c k b º ^

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