Anomala Samouelle, 1819 (Rutelinae: Scarabaeidae) of Buxa ...€¦ · World Scientific News 118...

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( Received 16 December 2018; Accepted 06 January 2019; Date of Publication 07 January 2019 )

WSN 118 (2019) 17-42 EISSN 2392-2192

Anomala Samouelle, 1819 (Rutelinae: Scarabaeidae) of Buxa Tiger Reserve, Dooars, West Bengal, India.

Part – II

Subhankar Kumar Sarkar1,a, Sumana Saha2,b, Dinendra Raychaudhuri3,c

1Department of Zoology, University of Kalyani, Kalyani, Nadia, West Bengal, PIN – 741235, India

2Department of Zoology, Barasat Govt. College, 10 K.N.C Road, Barasat (N 24 Parganas), Kolkata - 700124, West Bengal, India

3Department of Agricultural Biotechnology, IRDM Faculty Centre, Ramakrishna Mission Vivekananda University, Narendrapur, Kolkata - 700103, West Bengal, India

a-cE-mail address: [email protected] , [email protected] , [email protected]

ABSTRACT

In continuation to Part-I of the discourse, detailed taxonomic account of the remaining 10 species

of Anomala Samouelle, 1819 out of the total 20 species recorded from Buxa Tiger Reserve is presented

herewith. An identification key of all the species dealt in both Part I & Part II is provided in Part-I. Each

of the 10 species dealt in this paper is redescribed, illustrated and supplemented by digital images. A

note on the distribution of the recorded taxa is also provided.

Keywords: Anomala, Rutelinae, Scarabaeidae, Redescription, Buxa Tiger Reserve, Dooars, India

1. INTRODUCTION

As stated in Part-I of the treatise, genus Anomala, though have one of the largest species

representation and cosmopolitan distribution, lacks comprehensive taxonomic exposition

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World Scientific News 118 (2019) 17-42

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particularly from this part of the world. The genus is known to be composed of 1000 species

globally of which 800 are reported from the old world (Jameson et al. 2003). Indian Anomala

however is known by the regional works of Mittal (1999, 2005), Chatterjee & Biswas (1995),

Saha & Raychaudhuri (1998), Raychaudhuri & Saha (2014), Sarkar et al. (2010, 2017),

Chandra (1988, 2005), Chandra & Ahirwar (2005), Chandra & Uniyal (2007), Chandra et al.

(2012), Chandra & Gupta (2012a, 2012b, 2012c).

The study area, Buxa Tiger Reserve (BTR) represents one of the tropical rain forests of

Eastern Himalayas which is one of the 35 biodiversity hotspots of the world and one of the four

of India and is located in West Bengal, India.

In an attempt to provide detailed taxonomic account of the insect fauna of the hyper

diverse BTR, the authors have recorded 76 scarab species (Saha & Raychaudhuri 1998; Sarkar

et al. 2010, 2014, 2015, 2016a, 2016b, 2016c, 2017, 2018) of which 20 belong to the genus

Anomala. In Part-I of the article, taxonomic account of 10 species was presented and the

remaining 10 are dealt here in this paper. Each of the taxa is redescribed and illustrated nearly

after a century since Arrow (1917). A key for identification of all the 20 recorded species is

presented in Part-I of the discourse. A note on the distribution of the recorded species is also

provided in both Part-I & Part-II. The taxa are considered sensu Arrow (1917) and Potts (1974,

1977).

Study area

Detailed information about the study area and a location map is provided in Part–I (Sarkar

et al. 2017) of the discourse.

2. MATERIALS AND METHODS

Material and methods used in the work is discussed in Part–I (Sarkar et al. 2017) of the

monologue.

3. RESULTS

Family Scarabaeidae Latreille, 1802

Subfamily Rutelinae MacLeay, 1819

Tribe Anomalini Streubel, 1839

Subtribe Anomalina Streubel, 1839

Genus Anomala Samouelle, 1819

Anomala Samouelle, 1819, The Entomologist’s Useful Companion, p. 191.

Diagnosis: Body short, broad or elongate; opaque or shiny; labrum horizontal; antennae 9-10

segmented; head short and broad, transverse; clypeus semicircular or rectangular, front margin

straight, curved or medially notched; pronotum transverse, sides medially round or medially

angulate, base round or sinuate; elytra punctate striate or sulcate or punctured in longitudinal

rows, apical and sutural margins often with membranous fringe; pygidial apex round; fore tibia

bi or tridentate, mid and hind tibiae carinate externally and truncate at extremity.


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Type species: Melolontha aenea De Geer, 1775

Distribution: Worldwide (Redtenbacher 1848; Blanchard 1851; Arrow 1917; Machatschke

1957, 1972, 1974; Chatterjee & Biswas 1995; Moron et al. 1997; Saha & Raychaudhuri 1998;

Jameson 2002; Chandra 1988, 2005; Chandra & Ahirwar 2005; Chandra & Uniyal 2007;

Chandra et al. 2012; Chandra & Gupta 2012a, 2012b, 2012c; Raychaudhuri & Saha 2014;

Sarkar et al. 2010, 2017; Ratcliffe & Jameson 2017; GBIF 2017).

Key to species:

Key to all the 20 species recorded from the study area is provided in Part-I (Sarkar et al. 2017).

Anomala bilobata Arrow, 1912

(Figures 1A, 2A-B, 3A-J)

Anomala bilobata Arrow, 1912: 329.

Anomala bilobata Arrow, 1917: 156.

Description: ♂

Length 13.62 mm, humeral width 6 mm. Body elongate, cylindrical, compact and convex.

Colour and markings (Figures 1A, 3A): Testaceous with clypeus, anterior part of head, hind

tibiae and all tarsi brownish red; shiny; venter scantily pubescent.

Head (Figures 3A, 3B): Small and transverse; minutely and densely punctured.

Clypeus (Figures 3A, 3B): Short and transverse; minutely and densely punctured; front margin

strongly reflexed, medially notched and bilobed.

Fronto-clypeal suture (Figures 3A, 3B): Marked by a transverse sinuate carina.

Interocular width: 3.11 x transverse eye diameter.

Antenna (Figure 3C): 10 segmented, club 0.82 x stem.

Pronotum (Figures 3A, 3B): Transverse; minutely and densely punctured; sides round; base

finely margined, sinuate and roundly produced into a lobe at middle; front angles acute and

hind angles round.

Scutellum (Figure 3A): Short; obtusely triangular; minutely punctured; sides gradually round;

apex conical.

Elytra (Figure 3A): Finely punctured in longitudinal and irregular lines; intervals irregularly

punctured, 2nd wide anteriorly; angles nearly right angle; humps elevated.

Pygidium (Figure 3D): Coarsely punctured forming rugosity.

Mesosternum (Figure 3E): Minutely and densely punctured; medially marked by a longitudinal

line.

Fore tibia (Figure 3F): Tridentate; subterminally with 1 sharp and pointed spur.

Mid and hind tibiae (Figures 3G, 3H): Carinate externally; truncate and fringed with long spines

at extremity; terminally with 2 unequal spurs.

Tarsi (Figures 3F, 3G, 3H): 5 segmented.

Claws (Figures 3F, 3G, 3H): Fore unequal, with the longer one cleft; mid and hind equal.

Parameres (Figures 2A, 2B, 3I, 3J): Short and elongate; base broad; sides straight; inner margin

sinuate; apex blunt.


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Material examined: 1♂, Damanpur, BTR, 02.ix.02, coll. S.K. Sarkar; 1♂, Nimati, BTR,

13.iii.03, coll. S.K. Sarkar.

Distribution: India: West Bengal; Myanmar (Arrow 1917; Chatterjee & Biswas 1995; Saha &

Raychaudhuri 1998).

Anomala bilunata Fairmarie, 1888

(Figures 1B, 4A-H)

Anomala bilunata Fairmarie, 1888: 341.

Anomala bilunata Arrow, 1917: 147.

Description: ♀

Length 10.9 mm, humeral width 5.6 mm. Body elongate, cylindrical, compact and moderately

convex.

Colour and markings (Figures 1B, 4A): Testaceous with clypeus and head red brown; vertex of

head, a faint midlongitudinal and a small anterolateral patch on each side of pronotum, elytral

suture, hind tibiae and all tarsi dark brown; shiny except opaque lateral areas of elytra

Head (Figures 4A, 4B): Short and transverse; rugose.

Clypeus (Figures 4A, 4B): Short; rugose; front margin medially straight and round and at

angles.

Fronto-clypeal suture (Figures 4A, 4B): Medially marked by a transverse sinuate line and

laterally by a carina on each side.



Pronotum (Figure 4A): Coarsely and densely punctured; with a mid-longitudinal line at its

anterior half; sides gradually round; base sinuate and roundly produced at middle; front angles

acute and hind angles obsolete.

Scutellum (Figure 4A): Short; scantily and minutely punctured; sides round; apex conical.

Elytra (Figure 4A): Deeply sulcate, with coarse confluent punctures in the sulci; intervals

minutely and scantily punctured; humeral and apical angles nearly right angle; humps indistinct.

Pygidium (Figure 4D): Coarsely and densely punctured.

Mesosternum (Figure 4E): Medially smooth, laterally coarsely and densely punctured.

Fore tibia (Figure 4F): Tridentate; subterminally with 1 short spur.




Claws (Figures 4F, 4G, 4H): Fore and mid unequal, with the longer one cleft; hind equal.

Material examined: 1♀, Rajabhatkhawa, BTR, 19.v.95, coll. S. Saha; 1♀, Raimatang, BTR,

30.v.96, coll. S. Saha; 1♀, Nimati, BTR, 24.vi.03, coll. S.K. Sarkar.

Distribution: India: Assam, Manipur, Sikkim and West Bengal; Bhutan; China (Arrow 1917;

Chatterjee & Biswas 1995; Saha & Raychaudhuri 1998; Chandra & Ahirwar 2005).


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Anomala pusilla Arrow, 1912

(Figures 1C, 5A-H)

Anomala pusilla Arrow, 1912: 82.

Anomala pusilla Arrow, 1917: 263.

Description: ♀

Length 11 mm, humeral width 4.36 mm. Body short, oval, narrow in front, compact and convex.

Colour and markings (Figures 1C, 5A): Pale yellow, with golden suffusion above; clypeus,

forehead and pronotumred brown; a faint mid-longitudinal patch on each side of pronotum

black; opaque except shiny pygidium.

Head (Figures 5A, 5B): Short and transverse; rugosely punctured.

Clypeus (Figures 5A, 5B): Short; rugosely punctured; front margin round.

Fronto-clypeal suture (Figures 5A, 5B): Marked by a transverse straight line.



Pronotum (Figure 5A): Coarsely and densely punctured at sides, medially nearly smooth; sides

angulate at middle; base round; front angles acute and hind angles obtuse.

Scutellum (Figure 5A): Short; scantily and minutely punctured; sides sinuate; apex conical.

Elytra (Figure 5A): Deeply punctate striate, striae 10 on each disc; intervals minutely and

scantily punctured; humeral and apical angles nearly right angle; humps indistinct.

Pygidium (Figure 5D): Coarsely rugose.

Mesosternum (Figure 5E): Rugose.

Fore tibia (Figure 5F): Sharply bidentate; subterminally with 1 short spur.





Material examined: 1♀, Panbari, BTR, 20.iii.04, coll. S.K. Sarkar.

Distribution: India: Assam, Manipur, Sikkim and West Bengal (Arrow 1917; Chatterjee &

Biswas 1995; Sarkar et al. 2010).

Anomala birmana Heller, 1891

(Figures 1D-E, 2C-D, 6A-J)

Anomala birmana Heller, 1891: 297.

Anomala birmana Arrow, 1917: 181.

Description: ♂

Length 9.8 mm, humeral width 4.66 mm. Body short, compact and convex.

Colour and markings (Figures 1D, 6A): Pale testaceous yellow with head, median part of

pronotum, basal and lateral margins of elytra dark brown; shiny.


Clypeus (Figures 6A, 6B): Short; semicircular; rugose; front margin medially straight and round

at angles.


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Fronto-clypeal suture (Figures 6A, 6B): Marked by a transverse curved line.



Pronotum (Figure 6A): Minutely and densely punctured; with a mid-longitudinal line; sides

round; base gently round; front angles acute and hind angles obtuse.

Scutellum (Figure 6A): Broader than long; minutely punctured; sides round; apex conical.

Elytra (Figure 6A): Coarsely punctured in uniform longitudinal rows; intervals minutely and


Pygidium (Figure 6D): Coarsely and confluently punctured.

Mesosternum (Figure 6E): Coarsely and densely punctured.

Fore tibia (Figure 6F): Bluntly bidentate; subterminally with 1 short spur.





Parameres (Figures 2C, 2D, 6I, 6J): Long; base broad; sides sinuate; inner margin nearly

straight; apex conical, inner angle acute.

Material examined: 2♂1♀, Nimati, BTR, 21.v.97, coll. S. Saha.

Distribution: India: West Bengal; Myanmar (Arrow 1917; Saha & Raychaudhuri 1998).

Anomala (Singhala) polymorpha Arrow, 1911

(Figures 1F, 7A-H)

Anomala (Singhala) polymorpha Arrow, 1911: 486.

Anomala polymorpha Arrow, 1917: 134.

Description: ♀

Length 13.3 mm, humeral width 5.71 mm. Body narrow, oval, compact and moderately convex.

Colour and markings (Figures 1F, 7A): Pale testaceous yellow; metallic; with clypeus, head

and pygidium brown; moderately shiny.

Head (Figures 7A, 7B): Short; coarsely and densely punctured.

Clypeus (Figures 7A, 7B): Short; coarsely and densely punctured; front margin medially

straight; angles sharp.

Fronto-clypeal suture (Figures 7A, 7B): Medially marked by a transverse curved line.



Pronotum (Figure 7A): Minutely and sparingly punctured; sides and base gradually round, latter

narrowly margined; front angles acute and hind angles obsolete.

Scutellum (Figure 7A): Short; scantily and minutely punctured; sides straight; apex conical.

Elytra (Figure 7A): Deeply punctured in longitudinal rows; intervals minutely and scantily

punctured; humeral and apical angles nearly right angle; humps indistinct.


Mesosternum (Figure 7E): Coarsely and densely punctured.



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Material examined: 2♀, Rajabhatkhawa, BTR, 04.iv.03, coll. S.K. Sarkar.

Distribution: India: Karnataka, Madhya Pradesh, Maharashtra, Tamil Nadu and West Bengal

(Arrow 1917; Sarkar et al. 2010).

Anomala fissilabris Arrow, 1912

(Figures 1G, 8A-H)

Anomala fissilabris Arrow, 1912:331.

Anomala fissilabris Arrow, 1917:157.

Description: ♀

Length 13 mm, humeral width 5.6 mm. Body elongate, compact and convex.

Colour and markings (Figures 1G, 8A): Testaceous with head and clypeus dark brown; margins

of pronotum, scutellum and elytral suture black; pronotumanterolaterally on both side with a

faint and broken black spot; hind tibiae and tarsi reddish black; venter scantily setose.

Head (Figures 8A, 8B): Short and transverse; coarsely and densely punctured.

Clypeus (Figures 8A, 8B): Short and transverse; coarsely and densely punctured; front margin

reflexed, medially notched and bilobed.

Fronto-clypeal suture (Figures 8A, 8B): Marked medially by a transverse sinuate line and

laterally by a carina on each side.



Pronotum (Figure 8A): Transverse; roughly hexagonal; nearly smooth except few coarse

punctures; anterolaterally with an indication of a little elevation on each side; sides roughly

angulate at middle; base round; front angles acute and hind angles obtuse.

Scutellum (Figure 8A): Obtusely triangular; minutely punctured; sides gradually round; apex

blunt.

Elytra (Figure 8A): Finely punctured in longitudinal rows; intervals scatteredly punctured;

lateral margin elevated and reflexed; apex and suture with a fine membranous flange; humeral

angles obtuse and apical angles not formed; humeral hump elevated and apical hump evident.

Pygidium (Figure 8D): Scantily and minutely punctured.

Mesosternum (Figure 8E): Finely punctured; medially marked by a longitudinal line.

Fore tibia (Figure 8F): Tridentate, third smallest; subterminally with 1 short spur.




Claws (Figures 8F, 8G, 8H): Fore unequal, with the longer one cleft; mid and hind equal.

Material examined: 1♀, Nimati, BTR, 24.vi.03, coll. S.K. Sarkar.

Distribution: India: West Bengal; Myanmar (Arrow 1917; Sarkar et al. 2010).


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Anomala biharensis Arrow, 1917

(Figures 1H, 9A-H)

Anomala biharensis Arrow, 1917: 166.

Description: ♀

Length 13.85 mm, humeral width 6.28 mm. Body moderately elongate, oval, compact and

convex.

Colour and markings (Figures 1H, 9A): Pale testaceous yellow with head, margins of clypeus,

pronotum, scutellum and elytra, extremities of tibiae and tarsi brown; smooth and shiny.

Head (Figures 9A, 9B): Short and transverse; minutely and densely punctured.

Clypeus (Figures 9A, 9B): Short; transversely rectangular; rugose; front margin strongly

reflexed, nearly straight and round at angles.


Interocular width: 3 x transverse eye diameter.


Pronotum (Figure 9A): Minutely and densely punctured; margins elevated; sides gradually

round; base round and margined; front angles acute and hind angles obtuse.

Scutellum (Figure 9A): Short; scantily and minutely punctured; sides round; apex blunt.

Elytra (Figure 9A): Punctate striate, striae 7 on each disc; intervals scantily punctured; humeral

angles obtuse and apical angles acute; humps evident.

Pygidium (Figure 9D): Rugose.

Mesosternum (Figure 9E): Laterally rugose, medially smooth and marked by a longitudinal

line.

Fore tibia (Figure 9F): Tridentate, third very feeble; subterminally with 1 short spur.




Claws (Figures 9F, 9G, 9H): Fore and mid unequal, with the longer one of fore cleft; hind equal.

Material examined: 1♀, South Volka, BTR, 19.v.97, coll. S. Saha.

Distribution: India: Bihar and West Bengal (Arrow 1917; Saha & Raychaudhuri 1998).

Anomala variegata Hope, 1831

(Figures 1 I, 10A-H)

Anomala variegata Hope, 1831: 24.

Anomala variegata Arrow, 1917: 192.

Description: ♀

Length 12.88 mm, humeral width 5.55 mm. Body short, oval, compact and convex.

Colour and markings (Figures 1I, 10A): Testaceous with clypeus and head red brown; pronotum

basally and marginally, hind tibiae and tarsi black brown; pronotum decorated with a black,

longitudinal and irregular mark extending from front to near base on each side of middle; venter

scantily setose.

Head (Figures 10A, 10B): Elongate and transverse; coarsely and densely punctured.


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Clypeus (Figures 10A, 10B): Short and transverse; depressed; coarsely and densely punctured;

front margin reflexed and gently round at angles.

Fronto-clypeal suture (Figures 10A, 10B): Marked by a transverse sinuate carina.



Pronotum (Figure 10A): Transverse; minutely and densely punctured; sides gradually round;

base gently round; front angles acute and hind angles obtuse.

Scutellum (Figure 10A): Short; obtusely triangular; nearly smooth; margins with a membranous

fringe; sides gradually round; apex bluntly pointed.

Elytra (Figure 10A): Punctate striate, striae 13 on each disc; intervals smooth; margins elevated

and laterally reflexed; apex and suture with a fine membranous fringe; humeral and apical

angles nearly right angles; humps evident.

Pygidium (Figure 10D): Transversely rugose.

Mesosternum (Figure 10E): Laterally coarsely punctured; medially smooth, depressed and

marked by a longitudinal line.


Mid and hind tibiae (Figures 10G, 10H): Carinate externally; truncate and fringed with long

spines at extremity; terminally with 2 unequal spurs.



Material examined: 1♀, Rajabhatkhawa, BTR, 18.v.95, coll. D. Raychaudhuri; 1♀,

Rajabhatkhawa, BTR, 19.v.95, coll. S.Saha; 1♀, Gadadhar, BTR, 16.ix.96, coll. D.

Raychaudhuri; 1♀, Hatipota, BTR, 19.iv.02, coll. S.K. Sarkar; 1♀, Ghoramara, BTR, 30.iv.02,

coll. S.K. Sarkar.

Distribution: India: Sikkim and West Bengal; Nepal (Arrow 1917; Chatterjee & Biswas 1995;

Saha & Raychaudhuri 1998).

Anomala tincticeps Arrow, 1917

(Figures 1J-K, 2E-F, 11A-J)

Anomala tincticeps Arrow, 1917: 154.

Description: ♂


Colour and markings (Figures 1J, 11A): Pale testaceous yellow, with head, sutural margin of

elytra, all tibiae and tarsi blackish red; moderately smooth and shiny; sides of sternum scantily

setose.


Clypeus (Figures 11A, 11B): Short; rugose; front margin round and reflexed.


Interocular width: 3 x transverse eye diameter.


Pronotum (Figure 11A): Minutely and densely punctured; sides and base gradually round, latter

finely margined; front angles acute and hind angles obsolete.



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Elytra (Figure 11A): Coarsely punctate striate, striae 7 on each disc; intervals minutely and









Parameres (Figures 3E, 3F, 11I, 11J): Short and elongate; base broad; sides straight; inner

margin sinuate and medially bluntly toothed; apex conical.

♀: First tooth of fore tibia very sharp and long.

Material examined: 1♀, Rajabhatkhawa, BTR, 20.v.95, coll. S. Saha; 1♂, Raimatang, BTR,

30.v.96, coll. S. Saha.

Distribution: India: West Bengal; Myanmar (Arrow 1917; Saha & Raychaudhuri 1998).

Anomala siliguria Arrow, 1917

(Figures 1L, 12A-H)

Anomala siliguria Arrow, 1917: 188.

Description: ♀


Colour and markings (Figures 1L, 12A): Testaceous, with clypeus, head, pronotum, scutellum,

elytral suture, extremities of all tibiae and tarsi red; suffused with a metallic lusture except on

elytra and lower surface; moderately shiny.


Clypeus (Figures 12A, 12B): Short; rugose; front margin round and strongly reflexed.




Pronotum (Figure 12A): Coarsely, densely and deeply punctured; sides and base gradually

round; front angles acute and hind angles obsolete.


Elytra (Figure 12A): Coarsely and deeply punctured in longitudinal rows; intervals smooth;

humeral and apical angles nearly right angle; humps evident.

Pygidium (Figure 12D): Coarsely, densely and very distinctly punctured.







Material examined: 1♀, Damanpur, BTR, 02.v.02, coll. S.K. Sarkar.

Distribution: India: Sikkim, Uttarakhand and West Bengal (Arrow 1917; Sarkar et al. 2010).


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4. CONCLUSIONS

The present discourse provides a first ever comprehensive taxonomic account of beetles

comprising the genus Anomala Samouelle, 1819 under the subfamily Rutelinae (Coleoptera:

Scarabaeidae) occurring in Buxa Tiger Reserve. The survey for insect fauna of the study area

yielded in the recognition of 76 scarab species, of which 36 belong to the subfamily Rutelinae.

The genus Anomala Samouelle, 1819 dominates the subfamily Rutelinae as well as the family

Scarabaeidae with 20 species representation and speaks high in terms of species richness.

Photographic documentation, illustrations, key for identification and redescription of all the

recorded species will be of immense value for future researchers while undertaking the

taxonomic study of scarab beetles in India.

Acknowledgement

The authors are grateful to MOEF, GOI for financing the research work (Sanction no. – 14/34/2000-ERS/RE,

Dt.17/10/01), Hon’ble Vice chancellor, Ramakrishna Mission Vivekananda University, Kalyani University and

Calcutta University for providing necessary laboratory facilities. Thanks are extended to all officials and field

staffs of Buxa Tiger Reserve for their cooperation during field work. Thanks are due to all colleagues for their

assistance during the entire work.

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Figures 1A-L. A - Anomala bilobata Arrow, 1912, ♂ dorsal habitus; B - Anomala bilunata

Fairmarie, 1888, ♀ dorsal habitus; C - Anomala pusilla Arrow, 1912, ♀ dorsal habitus; D -

Anomala birmana Heller, 1891, ♂ dorsal habitus; E - Anomala birmana Heller, 1891, ♀dorsal

habitus; F - Anomala (Singhala) polymorpha Arrow, 1911, ♀ dorsal habitus; G - Anomala

fissilabris Arrow, 1912, ♀ dorsal habitus; H - Anomala biharensis Arrow, 1917, ♀ dorsal

habitus; I - Anomala variegata Hope, 1831, ♀ dorsal habitus; J - Anomala tincticeps Arrow,

1917, ♂ dorsal habitus; K - Anomala tincticeps Arrow, 1917, ♀ dorsal habitus; L - Anomala

siliguria Arrow, 1917, ♀dorsal habitus.


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Figures 2 A-F. A - Genital capsule of Anomala bilobata Arrow, 1912; B - Parameres of

Anomala bilobata Arrow, 1912; C - Genital capsule of Anomala birmana Heller, 1891; D -

Parameres of Anomala birmana Heller, 1891; E - Genital capsule of Anomala tincticeps Arrow,

1917; F - Parameres of Anomala tincticeps Arrow, 1917.


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Figures 3A-J. Anomala bilobata Arrow, 1912, Male: A - Dorsal habitus; B - Head & Clypeus,

dorsal view; C - Antenna, lateral view; D - Pygidium, dorsal view; E - Mesosternum, ventral

view; F - Fore tibia, tarsi & claws; G - Mid tibia, tarsi & claws; H - Hind tibia, tarsi & claws; I

- Genital capsule; J - Parameres. Scale bar – 1 mm.


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Figures 4A-H. Anomala bilunata Fairmarie, 1888, Female: A - Dorsal habitus; B - Head &

Clypeus, dorsal view; C - Antenna, lateral view; D - Pygidium, dorsal view; E - Mesosternum,

ventral view; F - Fore tibia, tarsi & claws; G - Mid tibia, tarsi & claws; H - Hind tibia, tarsi &

claws. Scale bar – 1 mm.


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Figures 5A-H. Anomala pusilla Arrow, 1912, Female: A - Dorsal habitus; B - Head & Clypeus,


view; F - Fore tibia, tarsi & claws; G - Mid tibia, tarsi & claws; H - Hind tibia, tarsi & claws.

Scale bar – 1 mm.


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Figures 6A-J. Anomala birmana Heller, 1891, Male: A - Dorsal habitus; B - Head & Clypeus,


view; F - Fore tibia, tarsi & claws; G - Mid tibia, tarsi & claws; H - Hind tibia, tarsi & claws; I

- Genital capsule; J - Parameres. Scale bar – 1 mm.


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Figures 7A-H. Anomala (Singhala) polymorpha Arrow, 1911, Female: A - Dorsal habitus; B -

Head & Clypeus, dorsal view; C - Antenna, lateral view; D - Pygidium, dorsal view; E -

Mesosternum, ventral view; F - Fore tibia, tarsi & claws; G - Mid tibia, tarsi & claws; H - Hind

tibia, tarsi & claws. Scale bar – 1 mm.


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Figures 8A-H. Anomala fissilabris Arrow, 1912, Female: A - Dorsal habitus; B - Head &





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Figures 9A-H. Anomala biharensis Arrow, 1917, Female: A - Dorsal habitus; B - Head &





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Figures 10A-H. Anomala variegata Hope, 1831, Female: A - Dorsal habitus; B - Head &





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Figures 11A-J. Anomala tincticeps Arrow, 1917, Male: A - Dorsal habitus; B - Head &



claws; I - Genital capsule; J - Parameres. Scale bar – 1 mm.


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Figures 12A-H. Anomala siliguria Arrow, 1917, Female: A - Dorsal habitus; B - Head &




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