References Bailey·Brock J. H. & C. C. Emig, 2000. Hawaiian Phoronida
(Lophophorata) and their d istribution in the Pacific region. Pacific Science 54 (2), 1 1 9- 1 26.
Cohen B. L., 2000. Monophyly of brachiopods and phoronids: reconciliation of molecular evidence with Linnaean classification (the subphylum Phoroniformea nov.). Proceedings o/the Royal Society. London, Series B 267, 225·2 3 1 .
Emig C. C. & A . N . Golikov, 1990. O n Phoronids ofthe Far Eastern Seas of the USSR and their distribution in the Pacific Ocean (in Russian). Zool. Zh. 69, 22-30.
Emig C. C. & C. Roldan, 1 992. The occurrence in Australia of three species of Phoronida (Lopbophorata) and their distribution in the Pacific area. Rec. S. Aust. Mus. 26 ( I ), I · 8.
Emig C. C., 1 982. The biology of Phoronida. Advances in Marine Biology 1 9, 1 ·89.
Emig C. C., 1 984. On the origin of the Lophopborates. Z. zool. System. Evolut.-forsch. 22 (2), 9 1 -94.
Emig C. C, 1 985. Phylogenetic systematics in Phoronida (Lophophorata). Z. zool. System. Evolul.-forsch. 23 (3), 1 84- 1 93.
Emig C. C., 1 997. Les Lopbophorates constituent-ils un embranchement? Bull. Soc. zool. Fr. 1 22 (3), 279-288.
Emig C. C., Garcia Carrascosa A. M., Roldan C. & J. M. Vietiez, 1 999. The occurrence in the Chafarinas Islands (S.E. Alboran Sea, western Mediterranean) of four species ofPhoronida (Lophophorata) and their distribution in the north-eastern Atlantic and Mediterranean areas. Cahiers de Biologie Marine 40, 1 29. 1 33.
Emig C. C., Roldan C. & J. M. Vieitez, 2000. Foronldeos del Iitoral luso-espaiiol. Boletin de la Real Sociedad Espailola de Historia Natural (Secci6n Biologia), 96 ( \ -2), in press.
Vitiitez J. M., C. C. Emig, C. Rodriguez Bablo & A. M. Garcia Carrascosa, 1 987. Foronldeos de las costas de la Peninsula Iberica e Islas Baleares: sistematica, ecologla, distribuci6n y estados larvarios. Bol. Insf. esp. Oceanogr. 4 (2), 63-78.
PHYLUM PHORONIDA Phoronis
ovalis Wright. 1 856 hippocrepia Wright, 1 856 australis Haswell, 1 883 muelleri Selys-Lonchamps, 1 903 psammphila Cori, 1 889 pa/lida Si len, 1 95 2
Phoronopsis albomaculala Gilchrist, 1 907 harmeri Pixell, 1 9 1 2 californica Hilton, 1 930
European Register of Marine Species
BRYOZOA (ECTOPROCTA)
Compiled by Peter J. Hayward
This catalogue of European Bryozoa incorporates the checklists prepared by Hayward ( 1 997) and Brattegard ( 1 995, also checked by Hayward), the species lists in Hayward and Ryland ( 1998, 1 999), Zabala and Maluquer ( 1 988), and relevant primary sources published after 1 985. All faunistic and systematic papers relating to shelf and coastal European Bryozoa listed in the Zoological Record from 1 985 to date have been consulted, but only those including new species or new European records are referenced here. Synonyms of most species included are to be found in the sources cited above; an unsourced entry indicates either a recently described species or unpublished records held by the compiler. The ERMS list was checked by J. Harmelin.
Taxonomic and faunistic research on European Bryozoa has been continuous through the last three decades, but geographical coverage and frequency of investigation have been extremely variable. The British Isles, Ireland and the Atlantic coasts of north·west Europe have attracted most research effort and the faunas of these regions are reasonably well known, although completely new species continue to be discovered even in such well studied areas as the English Channel (Reverter-Gil and Fernandez-Pulpeiro, 1 996; Hayward and Hansen, 1 999). The Faeroe Islands have been the subject of two surveys (Kramp, 1 934; Hayward, 1 994), the latter resulting in 1 5 new species and 1 3 new records for the temperate northeast Atlantic. The faunas of Iceland and west Greenland are similarly poorly known but are currently under review. The synoptic key of Zabala and Maluquer ( 1988) was founded on numerous sources relating to the western Mediterranean and Adriatic faunas, on the few available accounts of Aegean Bryozoa and the even sparser records for the southern coasts of the Mediterranean. Mediterranean Bryozoa are sti l l only incompletely known, at best, and i t is certain that many more undescribed species will be d iscovered as research continues.
References Alvarez, J. A. 1 987. Notas sobre la fauna briozoologica marina
Iberica. II. Especies de la Costa Vasea: estudio zoogeographico. KOBlE (Serie Cieneias Naturales) Bilbao, 1 6: 2 1 5 - 222
Alvarez, J. A. 1990. Una coleccion de briozoos procedentes de la costa de Alicante. Boletin Insl. Esp. Oceanog., 6: 21 • 40
Alvarez, J. A. 1 992. Sobre algunas especies de la familia Lichenoporidae Smitt, 1 866 (Bryozoa, Cyc!ostomida) en la region Atlantico-Mediterranea. Parte I: genero Dispore/la Gray, 1 848. Cahiers de Biologie Marine, 33: 20 1 - 243
Alvarez, J. A. 1993, Feneslrulina barrosoi sp. nov. (Bryozoa: Cheilostomida) with a review of the genus Feneslrulina on the Iberian Peninsula. Journal o/the Marine Biological Association a/the U. K., 73: 83 1 · 835
Alvarez, J. A. 1 994. La famille des L ichenoporidae (Bryozoa, Cyclostomida) dans les provinces Atlantico· Mediterraneenne et boreale Partie III: etude des collections du Museum National d'Histoire Naturelle de Paris. Cahiers de Biologie Marine, 35: 49 1 - 509
Alvarez, J. A. 1 995. New data on the family Lichenoporidae Smill (Bryozoa: Cyclostomida) from the Mediterranean region. Journal o/Natural His/Dry, 29: 1 067 - 1 079.
Aristegui. 1. 1 984. Briozoos Quilostomados (Ecloprocta. Cheiloslomata) de Canarias: esfudio sistematiico,
/aunislico y biogeogrqfico. Tesis doctoral, Universidad de La Laguna. 524 pp.
- 325 -
Costello, Emblow and White (eds), 200 1
Balduzzi, A., Barbieri, M. and Grislina, M. 1 99 1 . Morphology and life strategies of Aetea (Bryozoa: Cheilostomata) living on some western Mediterranean Posidonia oceanica meadows. Bulletin de la Societe des Sciences Naturelles de [ 'Ouest de la France. Memoire HS I : 1 - 1 2.
Brattegard, T. 1 995.BryozoB. Pp. 278 - 295 in Brattegard, T. and Holthe, T. (eds), Distribution of marine. benthic macro-organisms in NO/way. Research Report for DN 1 997- 1 . Directorate for Nature Management: Trondheim.
Fernandez-Pulpeiro, E . • Reverter-Gil, O. and Rami!, F. 1990. Inventario de los briozoos de Galicia. Thalassas. 8: 57 - 67
Hannelin, J.-G. 1 997. Especes affines microsympalriques chez Puellina (Bryozoa, CheiJostomala) et description d'especes nouvelles. Zoologica Scripta, 1 7: 25 - 38
Hannelin, J.-G. and Aristegui, J. 1988. New Cribrilinidae (Bryozoa, Cheilostomata) from the upper bathyal of the Atlanto-Mediterranean region. Journal of Natural History. 22: 507 - 535
Hannelin, I.-G. and d'Hondt, 1.-L. I 992a. Bryozoaires des parages de Gibraltar (campagne oceanographique BALGIM, ( 984) 1 - Cheilostomes.Bulletin du Museum National d'Histoire Naturelle, Paris, serie 4, 14: 23 - 67
Harmelin, J.-G. and d'Hondt, J.-1. 1 992b. Bryozoaires des parages de Gibraltar (campagne oceanographique Balgim, 1 984) 2 - Ctenostomes et Cyclostomes. Bulletin du Museum National d'Histoire NalUrelle, Paris, serie 4, 14: 605 - 621
Hannelin, I.-G., Borona!, 1. , Moisette, P. and Rosso, A. 1 989. Distansescharella seguenzai Cipolla, 1921 (Bryozoa, Cheilostomata), nouvelles donnees morphologiques et ecologiques tiTtles de specimens fossiles (Miocene, Pliocene) et actuels de Mediterranee. Geobios, 22: 484 -501
Hayward, P. J. 1 994. New species and new records of cheilostomatous Bryozoa from the Faroe Islands, collected by BIOFAR. Sarsia, 79: 1 8 1 - 206
Hayward, P. J. 1 997. Bryozoa. Pp. 273 - 282, in Howson, C. M. and Picton, B. E. (eds), The species directory of the marine
fauna andjlora of the British Isles and surrounding seas. Belfast and Ross-on- Wye: Ulster Museum and the Marine Conservation Society.
Hayward, P. J. and Hansen, K. B. 1 999. Three newly recognised cheilostomate bryozoans from tbe British sea area. Journal of the Marine Biological Association o/the u.K. 79: 9 1 7 -92 1
Hayward, P. 1. and Ryland, J. S. 1 998. Cheilostomatous Bryozoa. Part I , Aeteoidea - Cribrilinoidea. Synopses o/the British Fauna (new series), 1 0 (second edition): I 366
Hayward, P. J. and Ryland, J. S . 1 999. Cbeilostomatous Bryozoa. Part 2, Hippothooidea - Celleporoidea. Synopses of the British Fauna (new series), 1 4 (second edition): I -4 1 6
Hondt, 1.-L. d ' 1 984. U n nouvel immigrant dans I e Bassin d' Arcachon, Watersipora aterrima (Ortmann, 1 890) (Bryozoaire Cheilostome). Congres national des Societes savantes, Difon. J 984, sciences, fasc. II: 237 - 245
Hondt, J.-L. d' 1 988. Bryozoa from the coast ofIsrael. Boll. Zool. 3: 1 9 1 - 203
Kramp, P. L. 1 934. Bryozoa. Zoology of the Faroes, 58: I - 39 Reverter-Gil, O. 1 994. Briozoos de la Ria de Ferrol. Tesis
doctoral, Universidad de Santiago de Compostela. Reverter-Gil, O. and Femandez-Pulpeiro, E. 1 996. Cribrilinidae
from the Ria de Ferrol (NW Spain). Journal of Natural History, 30: 1 247 - 1 260
Reverter-Gil, O. and Femandez-Pulpeiro, E. 1 997. Two new species of Schizomavella (Bryozoa, Cheiloslomatida). Cahiers de Biologie Marine, 38: I - 6
Reverter- Gil, 0., d'Hondt, 1.-L. and Femandez-Pulpeiro, E. 1 995. Mise a jour de I'inventaire des bryozoaires de Roscoff publie par Echalier et Prenant ( 1 95 1 ). Cahiers de Biologie Marine, 36: 1 23 - 1 3 1
Rosso, A . 1 996. Popolamenti e Tanatocenosi a briozoi d i fondi mobili circalitorali del Golfo di Noto (Sicilia, Italia). Naturalista ski!. S. IV, 30: 1 89 - 225
Rosso, A. 1 999. Recent and fossil species of Characodama Mapiestone, 1 900 (Bryozoa) from the Mediterranean, with
description of two new species. Journal of Natural History, 33: 4 1 5 - 437
Zabala, M. and Maluquer, P. 1 988. Illustrated keys for the classification of Mediterranean Bryozoa. Treballs del Museu de Zoolagia, Barcelona, 4 : I - 294
Zabala, M., Maluquer, P. and Hannelin, I.-G. 1 993. Epibiotic bryozoans on deep-water scleractinian corals from the Catalonia slope (western Mediterranean, Spain, France). Scientia Marina, 57: 65 - 78
PHYLUM BRYOZOA
Class Gymnolaemata
Order Cheilostomatida
Family Adeonidae Adeonella
calveti (Canu & Bassler, 1 930) polystomella (Reuss, 1 847)
Adeonellopsis distoma (Busk, 1 858) multiporosa Aristegui, 1 98 5
Anarthropora monodon (Busk, 1 860)
Reptadeonella insidiosa (Jullien, 1 903) violacea (Johnston, 1 847)
Family Aeteidae Aetea
anguina (Linnaeus, 1 758) lepadiformis Waters, 1 906 ligulata Busk, 1 852 longicollis (Jullien, 1903) sica (Couch, 1 844) truncala (Landsborough, 1 852)
Family Ascosiidae Ascm'ia
pandora Jullien, 1 882 Fedora
edwardsi Jullien, 1 882
Family Beaniidae Beania
hirtissima (Heller, 1 967) hirtissima subsp. cylindrica (Hincks, 1 886) magellanica (Busk, 1 852) mirabilis Johnston, 1 840 robusta (Hinch, 1 8 8 1 ) thula Hayward, 1 994
Family Bitecfiporidae Codonellina
lacunala Hayward & Hansen, 1 999 Hippomonavella
borealis (Waters, 1 900) Hippoporina
harmsworlhi (Waters, 1 900) lineolifera (Hincks, 1 886) pertusa (Esper, 1 796) reticulafopunctala (Hincks, 1 877)
Metroperiella lepralioides (Calvet, 1 903)
Pentapora fascialis (Pallas, 1 766)
Pentapora folicacea (see Hayward & Ryland 1 999) ottomulleriana (Moll, 1 803)
Pseudojlustra anderssoni Kluge, 1 946 hincksi Kluge, 1 946 minima Hayward, 1 994 sinuosa (Andersson, 1 902) solid a (Stimpson, 1 854) virgula Hayward, 1 994
Schizomavella arrogata (Waters, 1 879) auriculala (Hassall, 1 842) auriculata subsp. asymetrica (Calvet, 1 927)
- 326 -
auricu/ata subsp. hirsuta (Calvet, J 927) aarica/ala subsp. inordinala Canu & Bassler, 1 930 aariculata subsp. leontiniensis (Waters, J 878) aarica/ala subsp. lineala (Nordgaard, 1 896) cristala (Hincks, J 879) cuspidata (Hincks, 1 880) discoidea (Busk, 1 859) jischeri (lullien, 1 882) gautier; Reverter Gil & Fernandez Pulpeiro, 1 997 hastala (Hincks, 1 862) hondti Reverter Gil & Fernandez Pulpeiro, 1 996 linearis (Hassall, 1 84 1 ) linearis subsp. profunda Harmelin & d'Hondt, 1 992 mamillala (Hincks, 1 880) marsupifera (Busk, 1 884) monoecensis (Calve!, 1 927) nepluni (lullien, 1 882) ochracea (Hincks, 1 880) porifera (Smitt, 1 868) rudis (Manzoni, 1 869) sarniensis Hayward & Thorpe, 1 995 teresae Reverter Gil & Fernandez Pulpeiro, 1996 triangularis Reverter Gil & Fernandez Pulpeiro, 1 997
Family BryocryptelJidae Bryocryptella
koehleri (Cal vet, J 896) torquala (Jullien, 1 903)
Cyslisella saccala (Busk, 1 856)
Marguella pulchra lullien, 1 903
Palmicellaria cf. aviculifera (Canu & Bassler, 1 928)
Palmiskenea aquilonia Hayward, 1 994
faroensis Hayward, 1 994 lorea (Alder, 1 864) plana (Hincks, 1 888) skenei (Ellis & Solander, 1 786)
Porella aculiroslris Smitt, 1 868 alba Nordgaard, 1 906 belli Dawson, 1 859 concinna subsp. tubulala Calvet, 1 927 compressa (J. Sowerby, 1 805) concinna (Busk, 1 854) cymosa Hayward, 1 994 groenlandica (Norman, 1 894) /aevis (Fleming, 1 828) minula (Norman, 1 868) palula (M. Sars, 1 85 1 ) perislomala (Nordgaard, 1 905) princeps Norman, 1 903 proboscidea Hincks, 1 888 smilli Kluge, 1 907 slruma (Norman, 1 868) tabulala (Busk, 1 86 1 ) targidula Hayward, 1994
Rhamphos/ome/la argenlea (Hincks, 1 88 1 ) bilaminata (Hincks, 1 877) coslata Lorenz, 1 886 hindcsi Nordgaard, 1 906 ovata (Smitt, 1 868) plicala (Smitt, 1 868) radialufa (Hincks, 1 877) scabra (0. Fabricius, 1 780) spinigera Lorenz, 1 886
Family Bugulidae Bicellarie/la
ciliala (Linnaeus, 1 758) Bicellarlna
alderi (Busk, 1 859) Bugula
aperla (Hincks, 1 886)
European Register of Marine Species
avicularia (Linnaeus, 1 758) cala/hus Norman, 1 864 cala/has subsp. cala/has Ryland, 1962 calalhus subsp. minor Ryland, 1 962 flabe/la/a (J V Thompson, in Gray, 1 848) fulva Ryland, 1 960 gaulier/ Ryland, 1 96 1 germanae Calvet, 1 902 gracilis (Busk, 1 852) harmswor/hi Waters, 1 900 neritina (Linnaeus, 1 758) plamosa (Pallas, 1 766) prenanli Castric-Fey, 197 1 purpurotincta Norman, 1 868 simplex Hincks, 1 886 spicata (Hincks, 1 880) Iflolonifera Ryland, 1 960 Iricuspis Kluge, 1 9 1 5 furbinala Alder, 1 857
Bugulella elegans Hayward, 1 978 fragilis Verril l , 1 879
Corynopore/la /enuis Hincks, 1 888
Dendrobeania decorala (Verrill, 1 879) = Bagula elongala Nordgaard elongala (Nordgaard, 1 906) fessa Kluge, 1 955 frulicosa (Packard, 1 863) murrayana (Bean, in Johnston, 1 847) pseudomurrayana Kluge, 1 955 quadr/denlala (Loven, 1 834) sessilis (d'Hondt, 1 974)
Kineloskias arborescens Danielssen, 1 868 smilli Danie1ssen, 1 868
Nordgaardia cornucopioides d'Hondt, 1 983 pusilla (Nordgaard, 1 907)
Uschakovia gorbunovi Kluge, 1946
Family Calescharldae Coronellina
fagei (Gautier, 1 962)
Family Calioporldae Alder/na
imbellis (Hincks, 1 860) Ammatophora
nodulosa (Hincks, 1 877) Amphibles/rum
aari/um (Hincks, 1 877) jiemingii (Busk, 1 854) Iyra/alam (Calvet, 1 907) sepfenlrionalis (Kluge, 1 906) solidum (Packard, 1 860)
- 327 -
Ap/ousina capriensis (Waters, 1 898) jilam (Jullien, 1 903)
Callopora crallcula (Alder, 1 856) depressa Cook, 1 968 discreta (Hincks, 1 862) dumerilil (Audouin, 1 826) dumerilil subsp. pouilleli (Alder, 1 857) lineala (Linnaeus, 1 767) minuta Harmelin, 1 973 rylandi Bobin & Prenant, 1 965 whileavesi Norman, 1 903
Cauloramphus cymbaefarmis (Hincks, 1 877) spiniferum (Johnston, 1 832)
Clavodesia biradiculala Harmelin & d'Hondt, 1 992
Copidazoam
Costello, Emblow and White (eds), 200 1
exiguum (Barroso, 1 920) maeilenta (Jullien, 1 882) planum (Hincks, 1 880) tenuiroslre (Hincks, 1 880)
Crassimarginalelfa erassimarginala (Hincks, 1 880) maderensis (Waters, 1 898) so/Mula (Hincks, 1 860) lensa (Norman, 1 903)
Doryporella spathulifera (Smitt, 1 868) reticulata Ryland, 1963
Ellisina antarctica Hastings, 1945 cr. antarclica gautieri Fernandez Pulpeiro & Reverter Gil, 1993
Megapora ringens (Busk, 1856)
Parellisina curviroslris (Hincks, 1 862)
Ramphonotus minax (Busk, 1 860) gorbunovi Kluge, 1 946
Relevirgula akdenizae Chimenz e/ al., 1997
Tegella arctica (d'Orbigny, 1 8 5 l ) armifera (Hincks, 1 880) kildinensis Kluge, 1 955 norvegica Nordgaard, 1 9 1 8 spitzbergensis (Bidenkap, 1 897) unicornis (Fleming, 1828)
Family Candidae Caberea
boryi (Audouin, 1 826) ellisii (Fleming, 1 8 1 4)
Notoplites damicornis Hayward & Ryland, 1 978 evocala (Jullien, 1 882) = Notopliles norman; harmeri Ryland, 1 963 jejJreysii (Norman, 1 868) sibirica (Kluge, 1 929) smilli (Norman, 1 868)
Scrupocellaria aegeensis Hannelin, 1969 arc/iea (Busk. 1 855) bertholleti (Audouin, J 826) delilii (Audouin, 1826) elongata (Smitt, 1 868) incurvata Waters, 1 896 inermis Norman, 1 867 in/ermedia Norman, 1 893 jolfoisi (Audouin, 1 826)
Scupocellaria macrorhyncha Gautier, 1 962
Scl'upoeeilaria madel'ensis Busk, 1 860 reptans (Linnaeus, 1 767) scabra (van Beneden, 1848) scrupea Busk, 1 852 scruposa (Linnaeus, 1 758) Spallilata d'Orbigny, 1 85 1
Tricellaria gracilis (van Beneden, 1 848) inopinata d'Bondt et al., 1 985 peachii (Busk, 1 85 1 ) fernala (Ellis & Solander, 1 786)
Family Cellariidae Cel/aria
jislu/osa (Linnaeus, 1 758) normani Hastings, 1 946 salicornioides Lamouroux, J 8 1 6 sinuosa (Hassall, 1 840)
Euginoma
relicu/ata d'Hondt, 1 98 1 vermiformis Jullien, 1 882
Family Cel\eporidac BI¢}'oneliaria
divergens (Smitt, 1873) Buskea
billardi (Calve!, 1906) dichotoma (Bincks, 1 862) nitida (Heller, 1 867) quincuncialis (Norman, 1 867)
Cel/epora pumicosa Waters, 1877 pumicosa (Pallas, 1 766)
Celieporaria sardonica (Waters, 1879)
Cei/epol'ina caminata (Waters, 1 879) canariensis Aristegui, 1 989 decipiens Hayward, 1 976 hassallii (Johnston, 1 847) lucida (Hincks, 1 880) pygmaea (Norman, J 868) surcularis (Packard, 1 863) lubu/osa (Hincks, 1 880) ventricosa (Lorenz, 1 886)
Lagenipora lepralioides (Norman, 1 868)
Omalosecosa ramu/osa (Linnaeus, 1 767)
Palmicellaria elegans Alder, J 864
Rhamphostomellina posidoniae Hayward, 1975
Turbicellepora avicu/aris (Hincks, 1 860) borea/e Hayward & B ille Hansen, 1 999 camera Hayward, 1 978 canaliculata (Busk, 1 886) canlabra (Barroso, 1 9 1 9) eoronopus (VVood, 1 844) coronopusoida Calvet, 1931 crenulala Hayward, 1 978 magnicosfata (Barroso, 1 9 1 9) nodulosa Lorenz, 1 886 smilti (Kluge, 1 962) torquata Hayward, 1 978 tubigera (Busk, 1 859)
Family Chaperiidae Chaperiopsis
annulus (Manzoni, 1 870) Larnacicus
eorniger (Busk, 1 859)
Family Cheiloporinidae Cheilopora
sincera (Smitt, 1 868) Cheiloporina
circumcincfa Neviani, 1 896)
Family Chlidoniidae Chlidonia
pyriformis (Bertoloni, 18 1 0) Crepis
longipes Jullien, 1 882
Family Chorizoporidae Chorizopora
brongniarlii (Audouin, 1 826)
Family Cleidochasmatidae Characodoma
- 328 -
mamilla/urn (Seguenza, 1 880) Cleidochasma
oranense (VVaters, 1 9 1 8) porcel/anum (Busk, 1 860) slrangnlalUm (Calvet, 1 906)
Cleidochasmidra
canakkalense Unsal & d'Hondt, 1979
Family Cribrilinidae Collarina
balzaci (Audouin, 1826) Cribrilina
alcicornis Jullien, 1882 annulata (0. Fabricius, ( 780) cryptooecium Norman, 1903 punctata (Hassall, 1841) spicuiifera Kluge, 1955 watersi Andersson, 1902
Dislansescharella seguenzai Cipolla, 1921
Figularia flgularis (Johnston, 1847)
Gephyrotes nilidopunctala (Smin, 1868)
Membraniporella nitida (Johnston, \838)
Puellina arrecla Bishop & Househam. 1987 bathyalis Harmelin & Aristegui. 1988 bijida (d'Hondt, 1970) cassidainsis (Harmelin, 1984) corbula Bishop & Househam, 1987 denliculala Harmelin & Aristegui, 1988 directa Bishop & Househam, 1987 gattyae (Landsborough, 1852) hincksi (Friedl, 1917) innominata (Couch. 1844) minima (Harmel in, 1984) modica Bishop & Househam, 1987 orientalis subsp. azoren.is Harmelin, 1988 orien/alis subsp. lusitanica (Harmelin, 1988) orien/alis subsp. orientalis Harmelin & Aristegui, 1987
pscudoradiata subsp. canaricnsis Harmelin & Aristegui, 1988
pseudoradiala subsp. pseudoradiata Harmelin & Aristegui, 1988
parva Reverter Gil & Fernandez Pulpeiro, 1996 pedunculala Gautier, 1956 picardi Harmelin, 1987 praecox Bishop & Househam, 1987 pseudoradiala Harmelin & Aristegui, 1988 radiata (Moll, 1803) seliformis Harmelin & Aristegui, 1988 setiformis subsp. romana Harrnelin & Aristegui, 1988 setiformis subsp. setiformis Harmelin & Aristegui, 1988
selosa (Waters, 1899) venusla Canu & Bassler. 1925
Family Cryptosulidae Cryptosula
pallasiana (Moll. 1803) Harmeria
scutelala (Busk, 1855)
Family Electridae Conopeum
reticulum (Linnaeus, 1767) seurati (Canu, 1928)
Family Cupuladriidae Cupufadria
biporosa (Canu & Bassler, 1923) canariensis (Busk, 1859) doma(d'Orbigny, 1851) mult/spinata (Canu & Bassler, 1923)
Family Cyclicoporidlle Lepralioides
nordlandica (Nordgaard, 1 905)
Family Electridlle Aspidelectra
melolontha (Landsborough, 1852)
European Register of Marine Species
Electra eruslulenla (Pallas, 1766) monoslachys (Busk, 1854) pi/osa (Linnaeus, 1767) posidoniae Gautier, 1957 verlicillata (Ellis & Solander, 1786)
Pyripora calenularia (Fleming, 1828)
Family Epistomiidae Epistomia
bursarla (Linnaeus. 1758) Synnolum
aegypliacum (Audouin, 1826)
Family Eucratcidae Eucralea
foricala (Linnaeus, 1758)
Family Flustridae Carbasea
carbasea (Ellis & Solander. 1786) Char/ella
barleel (Busk. 1860) membranaceolrllncata (Smitt. 1868) papyracea (Ellis & Solander, 1786) tenella (Hincks, 1887)
Fluslra Jofiacea (Linnaeus. 1758)
Gregarinidra gregaria (Heller, 1867)
Hincksina Jlustroides subsp. crassispinala Cal vet long/spinosa Harmelin & d'Hondt. 1992 see/elos (Busk. 1858)
HineksinoJluslra oClodon (Busk. 1852)
Sarsiflustra abyssicola (Sars, 1872)
Securiflustra securifrons (Pallas, 1766)
TerminoJlustra ca/veti (Guerin-Ganivet, 1911)
Family Gigllntoporidlle Coscin/opsis
ambita Hayward, 1974 Cyfindroporella
tubulosa (Norman, 1868)
Family Haplopomidae Haplopoma
bimucrona/um (Moll. 1803) bimucronatum subsp. occiduum (Waters, 1877) graniferum (Johnston, 1847) graniforllm subsp. carina/um (Calvet, 1902) impressum (Audouin, 1826) planum Ryland, 1963 sciaphilllm Sill!n & Harmelin, 1976
Family Heliodomidae Seloselfina
capriensis (Waters, 1926) eonslric/a Harmer, 1926 roulei Calvet, 1906
Family Hincksiporidac Hincksipora
spinulifera (Hincks, 1889)
Family Hippaliosinidac Hippaliosina
depressa (Busk. 1854)
Family Hippopodinidae Hippomenelfa
mucronelliformis (Waters. 1899) Hippopodina
Jeegeensis (Busk, 1884)
Family Hippoporidridae Hagiosynodos
- 329 -
M
Costello, Emblow and White (eds), 200 1
latus (Busk, 1 856) kirchenpaueri (Heller, 1 867) kirchenpaueri subsp. Iregoubovii Gautier, 1 962
Hippopore/la obesa (Waters, 1900)
Hippoporidra lusitania Taylor & Cook, 198 1 picardi Gautier, 1962
Family Hippothoidae Celleporella
angusla Alvarez, 1991 galaica Cesar-Aldariz el al., 1 999 hya/ina (Linnaeus, 1 767)
Haplola r:lavala (Hincks, 1 857)
Hippothoa divaricala Lamouroux, 1 82 1
flagellum Manzoni, 1 870 Plesiothoa
gigerium (Ryland & Gordon, 1977)
Family Laeernidae Cribellopara
trieholoma (Waters, 1 9 1 8)
Family Lllneeoporidae Calyplolheca
obseura Harmelin et al., 1 989 rugosa Hayward, 1 974 Iriarmata Hayward, 1 974
Family Lepraliellidae Buchneria
layalensis (Waters, 1 888) Lepra/iel/a
cont/gua (Smitt, 1 868) hippopus (Smitt, 1 867)
Family Mllrgarettidae Margarella
cereoides (Ellis & Solander, 1 786)
Family Membraniporidae Membranipora
arborescens (Canu & Bassler, 1 928) membranacea (Linnaeus, 1 767) tenuis Desor, 1 848
Family Microporellidae Calloporina
decorata (Reuss, 1 848) Diparula
veaueosa (Peach, 1 868) F eneslrulina
asturiasensis Alvarez, 1 992 barrosoi Alvarez, 1993 joannae (Cal vet, 1 902) mailisii (Audouin, 1 826)
Microporella appendiclilata (Heller, 1 867) arctica Norman, 1 903 ciliata (Pallas, 1 766) coronala (Audouin, 1 826) orienlalis Harmer, 1 957 umbracula (Audouin, 1 826)
Family Mlcroporidae Ca/pensia
nobilis (Esper, 1 796) Micropora
coriacea (Johnston, 1 847) normani Levinsen, 1909
Mollia eireumeinela (Heller, 1 867) patel/aria (Moll, 1 8 1 6)
Monoporella nodulifera (Hincks, 1 88 1 )
Rosseliana rosselii (Audouin, 1 826)
Steraechmella
buski Lagaaij, 1 952
Family Myriaporidae Myriapora
coarctata (M. Sars, 1 863) sllbgraei/e (d'Orbigny, 1 852) truncata (Pallas, 1 766)
Family Onychocellidae Onychocella
angu/osa (Reuss, 1 847) marioni (Jul l ien, 1 88 1 ) vibraculifera Neviani, 1 895
Family Calloporidae Rectonychoeella
di!ljunela Canu & Bassler, 1 930
Family Onychocellidae Smitlipora
solida (Nordgaard. 1907)
Family Pasytheidae Gemellipora
ebllrnea (Smitt. 1 873)
Family Phidoloporidlle Hippellozoon
mediterraneum (Waters, 1 894) Jaculina
blanchardi lullien, 1 903 paral/elala (Waters, 1 895) tessel/ata Hayward, 1979
Phidolopora elongala (Smitt, 1 868)
Reteparella aporosa (Waters, 1 894) atlantica (Busk, 1 884) beaniana (King, 1 846) camp/anata (Waters, 1 894) couchU (Hincks, 1 878) elegans Harmelin, 1 976 leuerbornii Hass, 1 948 grimaldii (Jullien, 1903) harmeri Hass, 1948 incognita Hayward & Ryland, 1 996 jermanensis (Waters, 1 909) mediterranea Hass, 1948 septentrionalis Harmer, 1 933 sparteli (Calvet, 1 906) sudbourniensis Gautier, 1962 walersi (Nordgaard, 1 907)
Releporellina delicatula Hayward, 1974 hyperborea Hayward, 1 994
Rhynchozoon bispinosum (Johnston, 1 847) globosum Harmer, 1957 neapolitanum Gautier, 1962 pseudodigilalum Zabala & Maluquer, 1 988 quadrispinalum Zabala & Maluquer, 1 988
Schizoretepora imperati (Busk, 1 884) longiselae (Canu & Bassler, 1 928) solanderia (Risso, 1 826)
Schizolheea divisa (Norman, 1 864) fissa (Busk, 1 856) serralimargo Hincks, 1 886 tubigera (Jullien, 1903)
Stephanollana armala (Hincks, 1 862)
Family Phoceanidae Phoceana
columnaris lullien, 1 903 Sertuiipora
gullata Harmelin & d'Hondt, 1 992
Family Romancheinidae Arclonula
- 330 -
M
M
aretiea (M. Sars, 1 85 I ) Eseharella
abyssicola (Norman, 1 869) acula Zabala el al., 1 993 connectens (Ridley, 1 88 1 ) hexaspinoso Aristegui, 1 986 immerso (Fleming, 1 828) indiviso Levinsen, 1 9 1 6 klugei Hayward, 1 979 labiola (Boeck MS, in Smitt, 1 868 labioso (Busk, 1 856) laqueata (Norman, 1 864) levinseni Hayward, 1 994 longicollis (Ju\lien, 1 882) octodenlata (Hincks. 1 880) praealta (Calvet, 1 906) ryland; Geraci, 1 994 variolosa (Johnston, 1 838) ventricosa (Hassall, 1 842)
Escharoides bidenkap; (Kluge, 1 946) coccinea (Abildgaard, 1 806) jacksoni (Waters, 1 900) mamillata (Wood, 1 844) megaroslris (Canu & Bassler, 1928
Exechonella antillea (Osbum, 1 927)
Hemicyclopora discrepans (Jullien, 1 903 emucronala (Smitt, 1 872) labrato Hayward, 1 994 microstoma (Norman, 1 864) multispinala (Busk, 1 86 1 ) polita (Norman, 1 864)
Neolagenipora collaris (Norman, 1 867) cximia (Hincks, 1 860) rugosa Hayward, 1 994
Family Savignyellidae Savignyella
lafontii (Audouin, 1 826)
Family Schizoporellidae Arlhropoma
cecilii (Audouin, 1 826) Escharina
alderi (Busk, 1 856) boreale Hayward, 1994 dutertrei (Audouin, 1 826) dutertrei subsp. haywardi Zabala et al., 1 993 dutel'trei subsp. prolecra Zabala et al., 1 993 hyndmanni (Johnston, 1 847) johnstoni (Quelch, 1 884) porosa (Smitt, 1 973) vulgaris (Moll, 1 803)
Phaeostachys spinifera (Johnston, 1 847)
Schizobrachiella sanguinea (Norman, 1 868)
Schizoporella cornualis Hayward & Ryland, 1 995 crustacea (Smitt, 1 868) dunkeri (Reuss, 1 848) = Schizoporella longirostris Hincks, 1 886 elmwoodiae (Waters, 1 900) errata (Waters, 1 878) hesperia Hayward & Ryland, 1 995 hexagona Nordgaard, 1 905 Iimbata Lorenz, 1 886 magnifica Hincks, 1 886 magniporata Nordgaard, 1 906 mutabilis Calvet, 1 927 neptuni (Jullien, 1 882) patula Hayward & Ryland, 1 995 smitti Kluge, 1962 tefragona (Reuss, 1 848)
European Register of Marine Species
unicornis (Johnston, in Wood, 1 844)
Family Scrupariidae Scruparia
ambigua (d'Orbigny, 1 84 1 ) chelata (Linnaeus, 1 758)
Family Selosellidae Sefosella
cavernicola Harmelin, 1 977 Jolini (Jullien, 1 882) vulnerala (Busk, 1 860)
Family SmiUinidae Parasmiftina
jejJreysi (Norman, 1 903) parsevalii (Audouin, 1 826) raigii (Audouin, 1 826) Irispinosa (Johnston, 1 838) tropica (Waters, 1 909)
Smitlina ajJinis (Hincks, 1 862) bella (Busk, 1 860) cervicornis (Pallas, 1 766) cheilostoma (Manzoni, 1 870) colleti (Jullien, 1 903) crysfailina (Norman, 1 867) inerma (Calve!, 1 906) jordii Reverter Gil & Femandez Pulpeiro, 1 999 jeJJreysi Norman, 1 903 landsborovii (Johnston, 1 847) majuscula (Smitt, 1 868) malleolus (Hincks, 1 884) minuscula (Smitt, 1 868) rem% rostrala Canu & Bassler, 1 928 rigid a Lorenz, 1 886 smitti (Kirchenpauer, 1 874) tubulifera (Heller, 1 867)
Smilfoidea amplissima Haywasd, 1 979 exilis Hayward, t 994 glaciala Waters, 1 900 marmorea (Hincks, 1 877) microoecia Hayward, 1 994 ophidiana (Waters, 1 879) propinqua (Smitt, 1 868) reliculata (MacGillivray, 1 842)
Family StomachetoseJlidae Cigclisula
turrila (Smitt, 1 873) Stomachetosella
cruenta (Busk, 1 854) normani Hayward, 1 994 sinuosa (Busk, 1 860)
Family Tessaradomidae Tessaradoma
boreale (Busk, 1 860)
Family Tcuchoporidae Phylaclella
labrosa (Busk, 1 854) Teuchopora
edwardsi (Jullien, 1 882)
Family Trypostegidae Trypos/ega
claviculala (Hincks, 1 884) venusta (Norman, \ 864)
Family Umbonulidae Hippoplcurifera
- 3 3 1 -
pulchra (Manzoni, t 870) Posterula
sarsi (Smitt, 1 868) Ragionl/la
rosacea (Busk, 1 856) Umbonula
littoralis Hastings, 1 944 oviceliata Hastings, 1 944
Costello. Emblow and White (eds). 2001
Family Watersiporidae Wafersipora
alerrima (Ortmann, 1 890) comp/anala (Nonnan, 1 864) cucullala (Busk, 1 854) subovoidea (d'Orbigny, 1 852) sublarquala (d'Orbigny, 1 852)
Order Ctenostomatida
Family Aethozoontidae Aelhozoon
pellucidum Hayward, 1 978
Family Alcyonidiidae Alcyonidium
albidum Alder, 1 85 7 candidum Ryland, 1 963 cellariaides Calve!, 1 900 diaphanum (Hudson, 1 778) disci/orme Smitt, 1 872 duplex Prouho, 1 892 gelalinosum ( Linnaeus, 1 76 1 ) hirsutum (Fleming, 1 828)
several species mamillatum Alder, 1 857 mytili Dalyell, 1 84 8 parasilicum (Fleming, 1 828) variegalum (Prouho, 1 892)
Family Arachnidiidae Arachnidium
clavatum Hincks, 1 877 ftbrosum Hincks, 1 880 hippo/hooides Hincks, 1 859
Arachnoidea annosciae d'Hondt &. Geraci, 1 976 thalassae (d'Hondt, 1 978)
Tanganella muelleri Kraeplin, 1 887
Family Benedeniporidae Benedenipora
calenala Fergens, 1 889 delicalula d'Hondt &. Geraci, 1975
Family Buskiidae Buskia
nilens Alder, 1 856 socialis Hincks, 1 887
Family Clavoporldae Ascarhiza
mowatarii d'Hondt, 1 983 Clavopora
hyslricis (Busk, 1 874) Melalcyonidium
gautieri d'Hondt, 1 976
Family Claviporidae Pseudalcyonidium
bobinae d'Hondt, 1 976
Family Flustrellldridae Flustrellidra
corniaulata (Smitt, 1 872) hispida (0. Fabricius, 1 780)
Family HypophorelJidae Hypophorella
expansa Ehlers, 1 876
Family Immergentiidae /mmergentia
suecica Silen, 1 946
Family Loblancoporldae Lobiancopora
hyalina Pergens, 1 889
Family Mlmosellidae Bantariella
verticil/ala (Heller, 1 867) Mimosella
crosslandi d'Hondt. 1 983 gracilis Hincks, 1851 verlicillala (Heller, 1 867)
Family MonohryozooDtidae Monobryozoon
ambulans RemaDe, 1 936
Family Nolellidae Anguinella
pa/mala van Beneden, 1 845 No/ella
dilalata (Hincks, 1 860)
Family Nollelidae gigantea (Busk, 1 856)
Family Nolellidae Iimicola (Franzen, 1 960) pusilla (Hincks, 1 8g0) slipata (Gosse, 1 8 55)
Panollcella nllians Jebrarn, 1 985
Family Penetrantiidae Penelrantia
brevis Sih:n, 1 946 concharllm Sileo, 1 946
Family Pherusellidae Pherusella
tubulosa (Ellis & Solander, 1 786)
Family Spathiporidae Spathipora
:rertum (Fischer, 1 866)
Family Terebriporidae /mmergentia
orbignyona Fischer, 1 866 Spathipora
comma (Soule, 1 950)
Family Tritlcellidae Farrella
repens (Farre, 1 837) Trilicella
flava Dalyell. 1 848 = Triticella wrenii G.O. Sars pedicellata (Alder, 1 857)
Triticel/opsis tissieri Gautier, 1 962
Family Vesiculariidae Amathia
distam Busk, 1 886 guermeii Chimonides, 1 987 inlermedis Chimonides, 1 987 lendigera (Linnaeus, 1 758) pruvoti Calvet, 1 9 1 1 semiconlloillta (Larnouroux, 1 824) torfuosa Tenison-Woods, 1 880 vidoviei (Heller, 1 867)
Bowerbankia citrina (Hincks, 1 877) gracilis Leidy, 1 855 gracillima (Hincks, 1 877) imbricata (Adams, 1 798) Pl/stu/osa (Ellis & Solander, 1 786)
Vesiclliaria spinosa (Linnaeus, 1 967)
Zoobotryon verticil/atum (delle Chiaje, 1 822)
Family Victorellidae Bulbefla
abscondita Braem, 1 9 5 1 Victorella
muelleri (Kraepelin, 1 887) pavida Saville Kent, 1 870
Family Walkeriidae Walkeria
- 332 -
atlanliea Busk, J 886 tuberosa Heller, 1 867 uva (Linnaeus, 1 758)
Class Stenolaemata
Order Cyc/ostomatida
Family Annectocymidae Anneetoeyma
arcuata (Hannelin, 1 976) indistineta Canu & Bassler, 1929 major (Johnston, 1 847) lubulosa (Busk, 1 875)
Enlalophoroecia balgimae Harmelin & d'Hondt, 1 992 clavafa (Busk, 1 859) elegans (Nann an, 1 909) gracilis Hannelin, 1 976 robusla Hannelin, 1 976 dejlexa (Couch, 1 842) produela (M. Sars, 1 869)
Family Crisiidac Bicrisia
abyssicola Kluge, 1 862 gibraltarensis Hannelin, 1990
Crisia aculeala Hassall, 1 84 1 calyplostoma Hayward & Ryland, 1 978 conferta Busk, 1 875 conslans Kluge, 1 946 cuneata Maplestone, 1905 denticulala (Lamarck, 1 8 1 6) denticulata var. arctica M. Sars, 1 863 eburnea (Linnaeus, 1 758) elongala (Milne Edwards, 1 838)
flstulosa Heller, 1 867 klugei Ryland, 1 967 occidenlalis (Trask, 1 857) oranensis Waters, 1 9 1 6 pyrula Hannelin, 1 990 ramosa Hanner, 1 89 1 sigmoidea Waters, 1 9 1 6 lenella Calvet, 1 906
Crisidia cornula (Linnaeus, 1 758)
Crisiella producla (Smitt, 1 865)
Fillerisia genieulala (Milne Edwards, 1 838) smilli (Kluge, 1 946)
Family Diastoporidae Cardioecia
watersi (O'Donoghue & de Watervil le, 1939) Diplosolen
obelia (Johnston, 1 838) inlriearius (Smitt, 1 872)
Eurysfrolos compacla (Norman, 1 866) occulla (Hannelin, 1 976)
Liripara amphorae (Hannelin, 1 974) violacea (Hannelin, 1 976)
Plagioecia dorsalis (Waters, 1 879) inoedijica/a (Jullien, 1 882) palina (Lamarck, 1 8 1 6) platydiscus Hannelin sarniensis (Nonnan, 1 864)
Family Horneridae Hornera
frondieulala Lamouroux, 1 82 1 lichenoides (Linnaeus, 1 758)
Family Lichenoporidae Coronopora
truneala (Fleming, 1 928)
European Register of Marine Species
Defrancia lucernaria M. Sars, 1 85 1
Disporella alboranensis Alvarez, 1992 borgi Alvarez, 1 995 boutani Alvarez, 1 995 harmeri (Neviani, 1939) hispida (Fleming, 1 828) piramidala Alvarez, 1 992 rob usia Alvarez, 1 992 smitti (Calvet, 1 906) spinulosa Jullien, 1 888 zurigneae Alvarez, 1 992
Lichenopora aZliensis Alvarez, 1 994 dislinola Alvarez, 1 993 jloseulus Hincks, 1 862 medUerranea de Blainville radiala (Audouin, 1 826) verrucaria (0. Fabricius, 1 780)
Family Mecynoeeiidae Mecynaeeia
defiealula (Busk, 1 875)
Family Oncousoeciidae Oncousoecia
canadensis (Osborn, 1 933) diastoporides (Nonnan, 1 869) dUalans (Johnston, 1 847) polygonalis (Kluge, 1 9 1 5)
Siomalopora semierecla Jullien, in Jullien & Calve!, 1 903
Stomaloporina incurvala (Hincks, 1 859)
Family SlegohorncridBe Slegohornera
arcliea (Kluge, 1 946) vialaeea (M. Sars, 1 863)
Family Terviidae Tervia
irregularis (Meneghini, 1 844)
Family Tubuliporidae Idmidronea
- 333 -
allanlica (Forbes, in Johnston, 1 847) bidenkapi (Kluge, 1 955) coerulea Hannelin, 1 976
jlexuosa (Pourtales, 1 867) Plalonea
sloeehas Hannelin, 1976 Tubulipora
aperla (Hanner, 1 898) jlabellaris (0. Fabricius, 1 780) hemiphragmala Hannelin, 1 976 liIiacea (Pallas, 1766) lobifera Hastings, 1 963 nOlomale (Busk, 1 875) penicillala (0. Fabricius, 1 780) phalangea Couch, 1 844 plumosa Thomson, in Hanner, 1 898 ziczac Harmelin, 1 976