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APPROVED: Jesus Rosales-Ruiz, Major Professor Shahla Ala’i Rosales, Committee Member Richard G. Smith, Committee Member and Chair of the Department of Behavior Analysis David Hartman, Dean of the School of Community Service Sandra L. Terrell, Dean of the Robert B. Toulouse School of Graduate Studies THE EFFECTS OF CONCURRENT FIXED INTERVAL-FIXED RATIO SCHEDULES OF REINFORCEMENT ON HUMAN RESPONDING Teresa Camille Parsons, B.S. Thesis Prepared for the Degree of MASTER OF SCIENCE UNIVERSITY OF NORTH TEXAS August 2005
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APPROVED: Jesus Rosales-Ruiz, Major Professor Shahla Ala’i Rosales, Committee Member Richard G. Smith, Committee Member and

Chair of the Department of Behavior Analysis

David Hartman, Dean of the School of Community Service

Sandra L. Terrell, Dean of the Robert B. Toulouse School of Graduate Studies

THE EFFECTS OF CONCURRENT FIXED INTERVAL-FIXED RATIO SCHEDULES

OF REINFORCEMENT ON HUMAN RESPONDING

Teresa Camille Parsons, B.S.

Thesis Prepared for the Degree of

MASTER OF SCIENCE

UNIVERSITY OF NORTH TEXAS

August 2005

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Parsons, Teresa Camille. Effects of concurrent fixed interval-fixed ratio

schedules of reinforcement on human responding. Master of Science (Behavior

Analysis), August 2005, 65 pp., 18 figures, 9 tables, references, 57 titles.

The present study contributes an apparatus and research paradigm useful in

generating human performances sensitive to concurrent schedules of reinforcement.

Five participants produced performances observed to be under temporal and ratio

control of concurrent fixed interval-fixed ratio schedules. Two aspects of interaction

between FI and FR schedules were distinguishable in the data. First, interaction

between two schedules was observed in that changes in the value of one schedule

affected behavior reinforced on another schedule. Second, switching from one pattern

to the other functioned as an operant unit, showing stability during schedule

maintenance conditions and sensitivity to extinction. These effects are discussed in the

context of current views on behavior under concurrent schedules of reinforcement, and

some implications for the conceptualization, measurement, analysis, and treatment of

complex behavior are presented.

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Copyright 2005

by

Teresa Camille Parsons

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ACKNOWLEDGEMENTS

I thank my major professor Jesus Rosales-Ruiz for sharing my excitement over each

new cumulative record, and teaching me to choose reinforcers wisely. I am deeply honored by

his introducing me to Goldiamond (in writing), Morse (in a car), and Lindsley (in the Barrett lab),

each of whom influenced, informed, and broadened my perspective, while Dr. Rosales-Ruiz

helped me to focus it.

Contributions from past and present members of the UNT Physio Lab, other professors,

and friends have made the research presented here possible. Specifically, I thank Richard

Anderson for sharing technical expertise, theoretical discussions, and procedural guidance, and

Yuka Koremura for her helpful comments and encouragement in the laboratory. I have learned

much from and with Ruthie, whose deep understanding I consider to be one of the things that

stay. I thank Mike for his questions, support and conversations, Tanya for our friendship during

school, and the Behavior Analysis faculty for presenting diverse opportunities and providing

feedback on my progress.

I deeply appreciate that Drs. Richard Smith and Shahla Ala’i Rosales agreed to serve as

thesis committee members. They were both perfect choices, as each has played an

instrumental role in my education thus far.

Finally, I thank my family for their timely encouragement, especially that of my parents

Joe and Peggy Parsons. Daddy, I try always to imitate your practices of loving people and

discovering their stories. Mom, I value my past experiences watching you pursue science, and

listen to nature. If some of my behavior results in a valuable contribution, it should be

considered a product of the rich environment you provided to me and my brother Mark—who

was my very first participant.

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TABLE OF CONTENTS

Page

ACKNOWLEDGEMENTS .............................................................................................iii

LIST OF TABLES.......................................................................................................... v

LIST OF ILLUSTRATIONS............................................................................................vi

Chapter

1. INTRODUCTION ..................................................................................... 1

2. METHOD ............................................................................................... 10

3. RESULTS .............................................................................................. 17

4. DISCUSSION ........................................................................................ 28

Appendices

A. INFORMED CONSENT FORM.............................................................. 55

B. APPARATUS DIAGRAM ....................................................................... 57

C. EXPERIMENTAL DESIGN .................................................................... 59

REFERENCE LIST...................................................................................................... 61

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LIST OF TABLES

Page

1. Proportion of Responses and Consequences Obtained on each Schedule for Participant CCSP05.......................................................................................... 50

2. Proportion of Responses and Consequences Obtained on each Schedule for Participant SMSP05 ......................................................................................... 50

3. Proportion of Responses and Consequences Obtained on each Schedule for Participant MPFA04.......................................................................................... 51

4. Proportion of Responses and Consequences Obtained on each Schedule for Participant PWFA04 ......................................................................................... 51

5. Session Information for CCSP05...................................................................... 52

6. Session Information for SMSP05...................................................................... 52

7. Session Information for LNFA04....................................................................... 53

8. Session Information for MPFA04...................................................................... 53

9. Session Information for PWFA04 ..................................................................... 54

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LIST OF ILLUSTRATIONS

Page

1. Cumulative Records for Participant CCSP05 ................................................... 37

2. Histogram of Patterns Produced during Session 1 for Participant CCSP05 ..... 38

3. Cumulative Record of Session 5 for Participant CCSP05 ................................ 38

4. Switches for Participant CCSP05 ..................................................................... 39

5. Cumulative Records for Participant SMSP05 ................................................... 40

6. Histogram of Patterns Produced during Session 1 for Participant SMSP05..... 41

7. Switches for Participant SMSP05..................................................................... 41

8. Cumulative Records for Participant LNFA04 .................................................... 42

9. Histogram of Patterns Produced during Session 1 for Participant LNFA04...... 43

10. Switches for Participant LNFA04 ...................................................................... 43

11. Cumulative Records for Participant MPFA04 ................................................... 44

12. Histogram of Patterns Produced during Session 1 for Participant MPFA04 ..... 45

13. Switches for Participant MPFA04 ..................................................................... 45

14. Cumulative Records for Participant PWFA04................................................... 46

15. Histogram of Patterns Produced during Session 1 for Participant PWFA04..... 47

16. Switching for Participant PWFA04.................................................................... 47

17. Scatterplots of Keypresses for All Participants ................................................. 48

18. Left-to-Right Patterns and Keystrokes across Sessions for Participants CCSP05, PWFA04, and SMSP05 .................................................................................... 49

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CHAPTER 1

INTRODUCTION

Revealing the determinants of phenomena is the business of science. Among the

community concerned with the experimental analysis of behavior, schedules of

reinforcement are widely appreciated as substantial determinants of the behavior of

organisms (Morse & Kelleher, 1970; Ferster & Skinner, 1957; Schoenfeld, 1950).

Beginning in 1938, considerable attention has been paid in the literature to

performances of organisms under singly programmed schedules. However, while

characteristic behavior patterns have been well documented with respect to simple

schedules concerned with the single operant (e.g., Ferster & Skinner, 1957; Zeiler,

1984), there is doubt that studying one operant in isolation is sufficient for

understanding complex behavior. For example, Catania (1966) asserts: “The isolated

operant is, however, a special case, and attention inevitably must turn also to the

interaction of two or more operants.” (p. 213)

The term interaction, a familiar notion in science, is used to describe the

relationship between two separate entities. How that interaction is captured by

measurement deserves attention. Psychologist Roger Barker described the “behavior

stream” as the actual arrangement of the behavioral continuum in time, comprised of

natural units that occur without the investigator’s intervention (1963). Barker

distinguishes behavioral units, naturally occurring within the behavior stream, from

behavioral tesserae, “alien parts of the behavior stream in the sense that they are

formed when an investigator, ignoring or dismantling the existing stream of behavior,

imposes or chooses parts of it according to his own preconceptions and intentions.” (p.

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2) Barker’s distinction between experimenter-defined units and naturally occurring ones

can be understood given a geographical example: if a natural “unit” of geography

consists of a riverbed through a prairie, “tesserae” exist in the imaginary line separating

a body of land into two states. Interaction between the states (for example, commerce)

occurs as a function of the division, distinguished from the “natural” interaction of

sources of food and water with people and animals.

Carefully defining appropriate units of analysis helps to establish the subject

matter of a new science. In 1938 Skinner acknowledged that while the analysis of

behavior should take into account the natural lines of fracture between behavior and

environment, the prediction of behavior requires the isolation of a reproducible unit.

Rate of occurrence of the operant thus became a ubiquitous dependent variable for

understanding the effects and process of learning. Yet, an operant response actually

occurs as part of a complex act, or chain (Skinner, 1938).

Moving towards a more comprehensive analysis of the context surrounding the

response, Schoenfeld and Farmer (1970) applied the metaphor of the behavior stream

to behavior of a single organism:

We take it as axiomatic that behavior is a continuous stream. As Skinner noted, the stream may be divided for analytic purposes. …The continuousness of behavior means that the organism can be thought of as “always doing something.” (p. 222)

Discovering what behavior is emitted, besides the target responses under scrutiny,

is essential to understanding ongoing behavior. Viewing behavior of an organism under

concurrent schedules may constitute a step towards the analysis of ‘real-world’

contingencies. The available alternatives on a concurrent schedule include more than

simply to respond or not to respond; they include also the alternative of changing over

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from one alternative to another (Catania, 1966; Skinner, 1950; Ferster, 1959; Findley,

1958). Skinner (1950; 1957) suggested that analysis of such complex interaction should

include both the relative strengths of responding on different schedule components, and

the strength of other operants engendered by the schedule. He proposed that “changing

over” as an alternative is a separate operant under the control of the programming

devices, and suggested it be considered in the analysis of “choice” (Skinner, 1950).

Findley (1958) made changing over to another schedule explicit (and easy for the

experimenter to count) by programming a switching key that, when pecked, changed

the schedule in effect. Pigeons pecked a key that was illuminated either red or green,

associated with a different variable interval schedule of grain delivery, and changed the

reinforcement schedule by pecking a second key. Thus Findley’s procedure treated

switching as an operant, making possible greater stimulus control over the behaviors

specified by the concurrent schedule (1958).

Whereas previous research (Ferster & Skinner, 1957; Findley,1958; Skinner,

1950) had considered switching to be an separate operant under the control of the

experimental procedures and important in the analysis of choice, a different view

describes responding on separate components of concurrent schedules as

independent. Herrnstein (1997) described behavior of animals under concurrent

variable interval-variable interval schedules: “[A]nimals match relative frequency of

reinforcement… because they respond to the two keys independently.” In Herrnstein’s

concurrent variable-interval schedules paradigm, responses on two schedules are

separated by a changeover delay (COD) imposing a response requirement or a waiting

period. Instances of responding on one schedule or the other are framed as “choices”

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between alternatives (Baum, 1982; Crowley & Donahoe, 2004; Herrnstein & Loveland,

1976). In experiments with pigeons, the COD separated responding on two keys, each

associated with a different schedule of reinforcement, and functioned as a penalty for

switching between alternatives. That is, a peck on one key followed by a peck on the

other produced a period of 1.5 seconds during which no food was delivered Herrnstein,

1961).

Herrnstein’s matching law (1961; 1970) predicts that the rates of performance of

two operants maintained by concurrent schedules of reinforcement will approximate the

obtained reinforcement rates on the respective schedules. Experimenters programming

concurrent schedules use the changeover key procedure or a variation to insure

separation of the two performances observed (Findley, 1958; Herrnstein, 1961; Sidman,

1962; Catania, 1962). The performances generated by concurrent schedules are often

described in terms of matching (in which responses allocated to the two schedules

approximate the obtained rates of reinforcement for the two respective alternatives),

undermatching, or overmatching (Baum, 1974; Wearden, 1983).

Some studies have attempted to extend the use of the matching law to describe

human performance on concurrent schedules of reinforcement, with mixed results

(Horne & Lowe, 1993; Takahashi, 1997; Takahashi & Shimakura, 1998; Madden &

Perone, 1999; Neuringer, Deiss, & Imig, 2000). When the response distribution of

human concurrent schedule performances does not reflect proportions predicted by the

matching law, various aspects of the experiments such as instructions and schedule

correlated stimuli are manipulated that may facilitate matching (e.g., Takahashi, 1997;

Takahashi and Shimakura, 1998; Madden and Perone, 1999), while experimenters

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present hypotheses purporting to explain why human organisms do not “match” (Horne

& Lowe, 1993; Takahashi & Shimakura, 1998). For instance, Takahashi (1997)

arranged a vigilance task in which human participants detected signals coming from

either a left or right speaker on concurrent VI-VI schedules. The initial results were

explained as “undermatching.” After exposing the participants to training on the VI

schedule components separately, the participants produced performances more closely

aligned with matching law predictions; hence, the authors state that undermatching was

reduced (1997). Takahashi and Shimakura (1998) compared the effects of instructions

about the features of schedules of reinforcement on the choices of participants working

for points on concurrent VI-VI schedules. Results indicated that when participants

received instructions describing how to earn the highest scores, the participants’

responding closely matched rates of reinforcement available. Thus, Takahashi and

Shimakura reported that for humans, choice performance fits the generalized matching

law when participants state correct rules about concurrent VI schedules (1998).

Since Herrnstein’s assertion that all behavior can be described as choice (1961),

the matching law has been used to describe choices between cocaine and food,

basketball shots worth different point amounts, math problems, and different amounts of

money (Anderson, Velkey, & Woolverton, 2002; Bourret & Vollmer, 2003; Neef, Mace, &

Shade, 1992; Savastano & Fantino, 1994; see also Billington & DiTommaso, 2003, for

examples of how the matching law is applied to educational settings in which students

choose between teacher-specifed problems or activities). However, there is

disagreement that behavior is best depicted as binary. According to Catania (1966):

The reversible character of COs [changeovers from one schedule component to another] in concurrent performances also distinguishes these performances from

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those studied in choice experiments. The latter are usually arranged in discrete trials, and it may be that there are important differences between a procedure in which an organism is confronted with two or more alternatives on a given trial, and one in which an organism is engaging in one behavior while the opportunity for a CO to some other behavior is continuously available. This is reflected in the data analysis of choice and concurrent performances. (p. 227)

Recall the distinction made earlier between artificial, or arbitrary, divisions of the

behavior stream and naturally occurring ones (Barker, 1963). Perhaps the practice of

segmenting the behavior stream into two choices constitutes an arbitrary distinction

which necessarily generates interaction, which in turn is quantifiable by the matching

law. Sidman (1960) wrote that in typical free-operant procedures, “the lever is never

withdrawn from the experimental space to prevent the subject from responding at times

that would be inconvenient for the investigator’s theory” (p. 409). Withdrawing the lever

may be analogous to the programmed COD which the matching law typically requires to

correctly model data generated by concurrent schedules. While the COD successfully

maintains some independence on concurrent schedules (Findley, 1958; Herrnstein,

1961; Sidman, 1962; and Catania, 1962), Catania noted that a changeover is implicit,

even when not recorded, each time a response in one operant class follows a response

in another operant class (1966). In experiments with both rats and human participants,

Neuringer (2000) recorded but imposed no penalty for switches between components of

concurrent schedules. Rats pressed two levers for food on concurrent FI schedules, and

people pressed mouse buttons on concurrent VI schedules in a computer game. For the

human participants, pressing the space bar changed the schedule in effect for mouse

button pressing, and no differential consequences were contingent on changing over.

Neuringer found that for both rats and people, choices between alternatives were a

function of reinforcement obtained for those alternatives. The present experiment used

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a similar procedure in which changeovers were counted, but not penalized.

In contrast to Neuringer’s findings, previous work on concurrent schedules of

reinforcement suggests that the changeover delay may be important in producing a

matching relationship between response rates and obtained reinforcement rates

(Herrnstein, 1961). Mace, Neef, Shade, and Mauro (1994) arranged concurrent

variable-interval schedules for adolescents working different sets of math problems.

Mace et al. (1994) reported that changes in the schedule values were accompanied by

changes in patterns of allocation only when adjunct procedures such as the COD,

timers, and demonstrations were used. Variations of the COD such as blackout periods,

shock contingent on changing over, and requiring several responses on a changeover

key have also been found to facilitate the development of performances more closely

aligned with matching law predictions (Todorov, 1971; Pliskoff, Cicerone, & Nelson,

1978). Such results support the interpretation that the changeover responses can make

up an operant class, as suggested by Skinner (1950). Similarly, Baum compared the

effects of different travel requirements associated with switching between schedules,

providing further evidence that contingent punishment affected changeover responses.

When the travel requirement was large, pigeons’ rate of switching between two keys

decreased, in contrast to more switching maintained by a shorter travel requirement

(1982). But in contrast to Skinner’s description and analysis of choice, Baum (1982)

holds that the COD, travel, blackout, and changeover requirements in general should be

excluded from calculations used to measure choice.

The empirical questions surrounding the procedures best suited to understand

choice have not yet been answered. In their paper describing several different methods

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used to study the choice concept, Crowley and Donahue (2004) noted that choice, like

other nonscientific terms “encompasses a diverse set of behavioral processes whose

understanding requires a variety of experimental methods” (p. 157). They cautioned that

without a variety of methods to study such processes, repeatable findings could be

mistaken for evidence of valid principles.

If behavior is described as choice (e.g., Herrnstein, 1961) and responses of

changing from one alternative to another are excluded from calculations of choice

(Baum, 1982), part of the behavior stream may go unmeasured or uncounted. A

cumulative record depicting responses to discretely presented stimuli, in which the

learner can make only one response to each stimulus, would not record the learner’s

response, but the experimenter’s behavior (Lindsley, 1996). Likewise, Schoenfeld and

Cole (1972) observe, "Rate is a measure for digitized Rs, and dilemmas are created

when the continuity of behavior is neglected in favor of quantum events” (p. 147). In

terms of concurrent schedules, dilemmas are created by the specification of only two

responses. The behavior stream includes not only responses “allocated” to one or the

other schedule component, but also their interaction; switching between the

components; and any part of the stream that does not include the two or more specified

responses. Davison (2004) analyzed inter-response time distributions of behavior under

concurrent schedules. Based on the distributions, Davison found evidence that a major

portion of responding on concurrent schedules might be composed of behavior other

than either of the responses specified by the schedules (2004).

What unit of behavior responds to the independent variable cannot always be

determined prior to observation. To this end, Schoenfeld (1972) suggested that

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researchers define both the response of interest (R) as well as Not-R, other portions of

the behavior stream that could be affected. Similarly, Goldiamond (1968; 1974) called

for experimenters to specify alternative sets of behavior, outlining an approach that has

helped to explain some complex behavior (Layng, Andronis, & Goldiamond, 1999;

Andronis, Layng, & Goldiamond, 1997).

Crowley and Donahoe (2004) suggested that a range of experimental procedures

be used to study the diverse set of behavioral processes that may constitute choice.

The current study takes up that task, using a free-operant preparation to study

concurrent FI-FR schedules with no changeover delay. The present experiment tested

an apparatus to study the effects of concurrent fixed interval-fixed ratio schedules on

human performances during acquisition, schedule maintenance and extinction. Effects

of concurrent FI-FR schedules were examined on rates of switching between one

schedule component to another, switching between responses on a specified schedule

component to other behavior, and rates of target performances.

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CHAPTER 2

METHOD

Participants and Setting

One male (CCSP05) and four female (LNFA04, MPFA04, PWFA04, SMSP05)

college students (mean age=22) participated in the study. Participants were recruited

via an advertisement in the local university newspaper. A screening process identified

students who had not taken courses in behavior analysis and were not behavior

analysis majors. Each participant received five dollars per completed session, paid in a

cumulative sum at the end of his or her involvement in the study and not contingent on

performance (see Appendix A). Each experimental session lasted 20 minutes, and

participants each completed 12-15 sessions. Sessions were conducted in the

Department of Behavior Analysis at the University of North Texas. The experimental

room was furnished with a computer on a desk, a keyboard, mouse, headphones, and a

chair.

Apparatus

The present experiment used an apparatus developed by Flygler and Rosales-

Ruiz (1997) and modified by Anderson (2004). A computer, monitor, keyboard, mouse,

and headphones constituted the apparatus (see Appendix B). A participant sat at the

desk facing the computer monitor. Before each session, the experimenter said, “You

may press these keys” while gesturing to keys 1-9 on the numeric keypad and then

said, “If you see a button on the screen, you may click it with the mouse.” Targeted

response topographies included pressing the 3-key sequences 1-2-3 and 7-8-9. At the

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beginning of each session, the participant put on the headphones and sat at the desk

facing the monitor, which contained a single button onscreen. A mouse-click on this

button (labeled START) produced a three-by-three grid containing nine gray squares

resembling three-dimensional buttons. Each square button corresponded to a number

key on the numeric keypad. When the participant pressed a number key on the keypad,

the corresponding button onscreen immediately changed, now appearing pressed in.

Each occurrence of a targeted key press sequence (e.g., 1-2-3) produced a “click”

sound. A square was centered under the grid. In some conditions, a button appeared in

the square. The button was inscribed with a speaker icon and if pressed, produced a

sound clip heard through the headphones.

Dependent Variables

The effects of concurrent fixed interval-fixed ratio schedule arrangements were

assessed on rate of responding, number of switching patterns produced, frequency of

switching between schedule components and changes in the frequency of switching

between schedule components, distribution of keystrokes, number of keystrokes and

patterns, and the proportion of responses emitted and consequences obtained on each

schedule.

Rate of responding. Response rates were calculated for target patterns 1-2-3 and

7-8-9 during each session. The number of target patterns produced during a session

was divided by the number of minutes in the session (20) to derive rate per minute for

each target pattern.

Number of switch patterns. The number of different switching pattern topographies

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was calculated. A switch was defined as two consecutive responses that differed in

topography by one or more keystrokes.

Frequency of target switches. Sequences of responses in which one pattern was

followed by another were counted as switches. Only switches between schedule

components (e.g., switches between 1-2-3 and 7-8-9) were defined as target switches.

Target switches could occur in either direction (that is, from 7-8-9 to 1-2-3 or vice

versa).

Number of keystrokes. The number of keystrokes per session was counted by

recording the presses of each number key (1-9) on the keypad.

Number of patterns. A pattern consisted of 3 or more keystrokes moving from left

to right on the numeric keypad.

Proportion of responses allocated. For each schedule, the proportion of target

responses was calculated by dividing the number of one type of responses (e.g., 1-2-3)

performed on one schedule by the total number of 1-2-3 and 7-8-9 responses per

session.

Proportion of consequences obtained. The number of consequences obtained on

one schedule was divided by the total number of consequences delivered during the

session.

Independent Variables

Effects of concurrent FR-FR (fixed ratio 1-fixed ratio 1) schedules, concurrent fixed

interval-fixed ratio (FI30s-FR5 or FI30s-FR10) schedules, and concurrent extinction-

fixed ratio schedules (EXT-FR5 and EXT-FR10) were assessed. During a concurrent

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FR-FR schedule, either of two target responses was followed by a sound on a

continuous schedule of reinforcement. During a concurrent FI30s-FR5 schedule, the

target response (1-2-3) produced a sound if it occurred at least 30 s from the last

consequence delivered on that schedule, while every fifth 7-8-9 response produced a

sound. During EXT-FR schedules, 1-2-3 produced no consequences, while 7-8-9

produced consequences on a given (either FR5 or FR10) schedule.

Procedure

Participants experienced one session per day at about the same time each day.

Each session lasted 20 minutes.

Experimental Design

The experimental design was an A-B-C-D-C design. After an initial preference

assessment, acquisition of the left-to-right response was shaped by a continuous

schedule of reinforcement (A). Then, a concurrent FR1-FR1 schedule (B) selected two

different responses. Those responses were then exposed to concurrent FI-FR

schedules of reinforcement (C), followed by extinction (D). Finally, Condition C included

reinstatement of a previously experienced schedule of reinforcement for either one or

both target responses. The design included a multiple baseline across participants and

a partial replication across conditions (see Appendix C for diagram).

Preference assessment. The participant entered the room, sat at the computer,

and wore the headphones. The participant pressed a button labeled “Press to start” to

initiate the preference assessment. 345 sounds were sequentially presented in random

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order through the headphones. Participants answered the question “Do you like the

sound?” for each clip by pressing the “Yes” or “No” button. Clips (mean duration: 3 s)

included common sounds such as movie lines (e.g., “Go ahead, make my day”), familiar

quips by cartoon or sitcom characters (“Doh!) and clips of music or sound effects (e.g.,

the sound of a bell or children laughing). Each trial consisted of the presentation of one

sound, followed by the words “Do you like the sound?” on the screen. Then, the words

“Yes” and “No” appeared. Clicking either “Yes” or “No” sorted the sound clips into two

sound banks and reset the trial. A new sound was then presented. Each sound was

presented only once. Sounds to which the participant had answered “No” were no

longer used, while sounds labeled “Yes” were compiled into a bank. During the rest of

the experiment, a target response could produce a sound clip from his “Yes” sound

bank (depending on the schedule of reinforcement in effect during the condition).

Left-to-right pattern shaping. The participant sat at the computer. The

experimenter said, “Click the button when you are ready to start. You will see a ‘Thank

you’ message on the screen letting you know when you are finished. At that point, I will

return to release you.”

The experimenter said, “During the session, you may press any of these 9 keys”

while gesturing toward the numeric keypad. The experimenter then pointed to the

screen and said, “If you see a button appear here, you may click it with the mouse.”

The experimenter then left the room.

Keypresses of 1, 4, and 7 corresponded to the left column of the 3 x 3 grid (see

Appendix B). The middle column of the grid corresponded to the numbers 2, 5, 8 and

the grid’s right column corresponded to keys 3, 6, 9. When the first press of a key in the

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left column of the grid (either 1, 4, or 7) occurred, the corresponding button changed,

appearing pressed in. Further keypresses in the left column (1, 4, or 7) produced no

visible change, and the first press in the middle column changed the corresponding

button to appear pressed in. This continued until a left-to-right sequence of keypresses

had occurred, with at least one response in each column.

When a target response occurred (at least 3 keypresses moving from left-to-right,

such as 1-2-3, 1-5-9, 4-5-6 or 7-8-9), a chime sounded and a button appeared below

the grid. The button, when pressed, produced a clip from the “Yes” sound bank. The

trial then reset. Each instance of left-to-right keypress sequences was followed by a

consequence sound (e.g., a continuous reinforcement schedule was in effect for left-to-

right target responses).

Target response shaping. The apparatus operated exactly as during response

acquisition, except that only certain (targeted) responses produced a button and sound.

A concurrent fixed ratio-fixed ratio schedule (FR1-FR1) selected two different left-to-

right 3-key sequences. At any time during FR1-FR1, pressing 1-2-3 or 7-8-9 resulted in

a button, producing a sound clip. Participants CCSP05, SMSP05, and PWFA04

received one exposure to the FR1-FR1 condition, while Participants LNFA04 and

MPFA04 experienced two sessions under concurrent FR1-FR1 schedules.

Concurrent fixed interval-fixed ratio schedules. After initial exposure to concurrent

FR-FR schedules, each participant experienced concurrent fixed interval-fixed ratio

schedules. The sequence 1-2-3 produced a sound on a fixed-interval-30s schedule (the

first response produced after 30 seconds had elapsed from the previous consequence

delivery), while 7-8-9 sequences were followed by a sound on FR-5 (every 5th

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occurrence of 7-8-9 produced a sound). Participants CCSP05, SMSP05, LNFA04, and

PWFA04 were exposed to concurrent FI30s-FR5 schedules, and MPFA04 was exposed

to concurrent FI30s-FR5 and concurrent FI30s-FR10 schedules.

Target responses not meeting schedule requirements always produced a click

sound.

Extinction. The grid continued to change in response to key presses and target

responses still produced a click. No responses were followed by a button and sound.

Reversal. After the extinction phase, participants experienced either a concurrent

FI-FR or concurrent EXT-FR schedule for 1-2-3 and 7-8-9 responses. Participants

CCSP05 and SMSP05 experienced concurrent FI30s-FR5 schedules. Participants

LNFA04 and PWFA04 experienced EXT-FR5. MPFA04 experienced EXT-FR10.

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CHAPTER 3

RESULTS

CCSP05

Figure 1 depicts session cumulative records of 1-2-3 and 7-8-9 performances for

Participant CCSP05. A phase label is centered above each session in which a

contingency change occurred. Phase labels specify both the schedule and target

response (e.g., in the condition Concurrent FI30(123)-FR5(789), the pattern 1-2-3 was

followed by a sound on a Fixed Interval-30s schedule and the pattern 7-8-9 was

followed by a sound on a Fixed Ratio-5 schedule). Numbers positioned below the

cumulative records are session numbers. In Session 1, Participant CCSP05

experienced acquisition training in which left-to-right sequences of key presses were

followed by a sound on a continuous schedule of reinforcement. Left-to-right patterns

occurred at 12.2 per minute (rates reported in Tables 5-9). Next, CCSP05 experienced

one session of target shaping in which both 1-2-3 and 7-8-9 were followed by a sound

(concurrent FR1-FR1). During that session, rates of 1-2-3 and 7-8-9 were comparable

(11.9 rpm and 8.6 rpm). After stable baselines were established for both target

response sequences, the contingency changed to Concurrent FI30-FR5 for 1-2-3 and 7-

8-9, respectively. During Session 3, performance initially alternated between schedule

components and then differentiated. Low running rates on 1-2-3 (8.05 rpm and 7.45

rpm) and higher, constant levels of 7-8-9 (55.0 and 56.85 rpm) were observed during

Session 4 and 5. During Extinction, rates of 7-8-9 were initially higher (43.95 rpm,

compared to 12.9 for 1-2-3 in Session 6). Switching between schedule components

dropped dramatically during extinction. In Session 7, CCSP05 responded for several

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minutes on one schedule, then switched to the other component. Beginning in Session

9, long pauses were observed during both schedule components. Responding during

Session 11 consisted of bursts of responding on 7-8-9 (15.15 rpm overall) punctuated

by pauses of increasing duration, while 1-2-3 performance occurred at 2.25 rpm. A

return to the schedule maintenance phase reinstated high stable performance on 7-8-9

(52.75 rpm) and low response rates on 1-2-3 (3.2 rpm).

Figure 2 shows the distribution of left-to-right patterns CCSP05 produced during

Session 1 (Acquisition). Forty-two different patterns occurred, and the most frequent

patterns produced were 123, 456, and 789.

In Figure 3, two cumulative records produced during the first 10 minutes of

Session 5 for CCSP05 are enlarged to show detail. The top graph is a record of 1-2-3

responses and the bottom graph is 7-8-9 performance under the concurrent FI30-FR5

schedule. Triangles represent consequence (sound) deliveries. Initially, a burst of 7-8-9

responses occurred until a sound was delivered (bottom graph); next, a run of 1-2-3

patterns (top graph) ended in a sound delivery, followed by another 30-s run of 1-2-3.

Responding stabilized with regular 5-pattern sequences of 7-8-9, punctuated by

switching to the FI30 component every 30-40 seconds.

Figure 4 depicts the number of target patterns (1-2-3 and 7-8-9) and the number

of switches for Participant CCSP05 across sessions. Target patterns are marked with

lines; the open circle represents switches from 1-2-3 to 7-8-9, the closed circle

represents switching from 7-8-9 to 1-2-3; and all switches (including switching to a

different pattern that was not 1-2-3 or 7-8-9) are marked with small squares. During the

FR1-FR1 condition, switching between 1-2-3 and 7-8-9 occurred equally. Under

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Concurrent FI-FR schedules, switching from 7-8-9 to 1-2-3 decreased and stabilized at

a session average of 32 (recall that sounds could be delivered a maximum of 40 times

per session on the FI30s schedule). During the first 2 sessions of Extinction, the gap

between data paths for all switches versus the target switches widened. Switches from

1-2-3 to 7-8-9 or from 7-8-9 to 1-2-3 decreased across extinction sessions, while more

total switches occurred relative to the number of target switches. During Session 12

when the Concurrent FI30-FR5 schedule was again in effect for 1-2-3 and 7-8-9, the

number of switches returned to the levels maintained by the previous concurrent

schedule.

For Participant CCSP05, Table 1 shows the proportion of consequences

obtained on one schedule versus the proportion of responding allocated to that

schedule. During the Concurrent FI30-FR5 phase, CCSP05 spent more time

responding within the FR component and obtained a higher proportion of sounds there.

During the reversal to Concurrent FI30-FR5, CCSP05 allocated 94.28% of responding

to 7-8-9 while responding 5.72% of the time on 1-2-3. Thus, 91.74 % of all

consequences were obtained during 7-8-9 performance while the other 8.26% were

obtained during 1-2-3 responding.

SMSP05

Figure 5 shows cumulative records of 1-2-3 and 7-8-9 performances for

Participant SMSP05. During acquisition of the left-to-right response, patterns occurred

at a session running rate of 12.5 rpm. During Session 2, both 1-2-3 and 7-8-9

performances occurred consistently at 8.6 rpm under FR1-FR1 contingency. When the

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contingency changed to Concurrent FI30-FR5, performance on 1-2-3 was initially higher

than 7-8-9. Over the next two sessions 1-2-3 decelerated while 7-8-9 accelerated.

During Session 7, performances stabilized with 7-8-9 rates higher (41.6 rpm) than those

on 1-2-3 (12.85). Session 8 outcomes were similar to Session 7. Close inspection of the

cumulative records reveals a pattern of responding in which several cycles of 7-8-9

responses were closely followed by a run of 1-2-3 responses, culminating in a switch to

7-8-9. Under Extinction, SMSP05 initially produced 1-2-3 at high rates followed by

frequent switches to the other schedule component. Whereas responding had

previously been differentiated between schedules, responding under extinction

consisted primarily of similar response rates and fewer switches between the

components. Beginning in Session 10, rates of 1-2-3 versus 7-8-9 were 30 and 30.35,

38.9 and 33.65, 13.35 and 14.65, and finally (Session 13), 36.5 and 33.5. A return to the

Concurrent FI30-FR5 contingency re-established 7-8-9 performance higher (38.5 rpm)

than 1-2-3 performance (17.9 rpm).

Figure 6 shows the distribution of left-to-right patterns produced during Session 1

for SMSP05. Fifty-two different patterns occurred during Session 1. The most frequent

patterns produced were 1-5-6, 4-5-6, 1-2-3, 1-5-9, 7-5-3, and 7-8-9. During response

acquisition the operant levels of 1-2-3 and 7-8-9 were 1.1 and 0.95 rpm, respectively.

Figure 7 depicts the number of target patterns (1-2-3 and 7-8-9) per session,

compared to the number of switches between target patterns for Participant SMP05.

The number of target patterns produced during Concurrent FI30-FR5 was similar across

sessions. Switching stabilized during the last two sessions of schedule maintenance

(Sessions 7-8). Extinction began with a higher number of target patterns, and more

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switches from one pattern to another than during the concurrent schedule. Much

responding during extinction consisted of patterns other than 1-2-3 or 7-8-9. The

switching between components declined to near-zero levels during the last Extinction

session. A return to the Concurrent FI30-FR5 contingency reinstated switching to levels

previously seen during schedule maintenance.

Table 2 shows the proportion of 1-2-3 versus 7-8-9, and the proportion of

consequences obtained on the FI versus FR components for SMSP05. Responding

shifted between the two schedule components until Session 7, in which differentiation

between the components occurred and 7-8-9 was consistently performed at higher

rates. In Session 7, 7-8-9 constituted a higher proportion of responding (76.40%) than

responding on 1-2-3, and 7-8-9 responses produced 85.57% of the sounds obtained

during the session. The proportion of 1-2-3 versus 7-8-9 responses tended toward

indifference during Extinction. The contingency reversal in Session 14 produced a

distribution similar to Session 8, with a larger proportion of responding allocated to the

FR5 component, and a larger proportion of sounds obtained on the FR versus the FI

schedule.

LNFA05

Cumulative records of 1-2-3 and 7-8-9 performances are depicted for Participant

LNFA04 in Figure 8. In Session 1, a steady rate (11.65 rpm) of left-to-right patterns was

produced. Under the Concurrent FR1-FR1 contingency for 1-2-3 and 7-8-9 responses,

only 1-2-3 performances occurred. Changing the schedules to Concurrent FI30-FR5

resulted in low rates of responding on 7-8-9 that were constant across sessions 4, 5, 6,

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and 7 (9.3, 10.05, 10.00, and 10.25 rpm, consecutively). Rates of 1-2-3 performances

during those sessions were higher and stable (28.6, 34.6, 37.3, and 37.4). Across the

schedule maintenance condition, LNFA04 emitted a consistent pattern of responding

including several sequences of 1-2-3 followed by 5 instances of 7-8-9, followed by

switching back to 1-2-3 performance. During Extinction, this pattern was disrupted as

performance shifted to alternating on each component for longer and longer periods of

time. During Session 9 responding alternated during the first 7 min between 1-2-3 and

7-8-9, followed by a 3 min run of 7-8-9 responses. Performance then shifted to 1-2-3 for

the remainder of the session. Session 11 consisted of only 5 runs of responding on 7-8-

9. During the 3 pauses of increasing duration on the 7-8-9 component, runs of 1-2-3

performances occurred. Session 12 is similar to 11 and is not shown in Figure 10 due to

a computer error. During Session 13, LNFA05 experienced Concurrent EXT-FR5 in

which only 7-8-9 responses were followed by sounds. Only 13 1-2-3 responses

occurred, while 7-8-9 performance stabilized immediately at 37.3 per minute.

Performance was sensitive to the contingency change to FR5 from EXT, in that rates

during the last session changed immediately and were the highest of the entire study for

7-8-9, and the lowest (0.65 rpm) for 1-2-3.

Figure 9 shows the left-to-right response distribution for LNFA05’s Session 1.

Note that 25 different patterns were produced, with relatively high rates on 4-5-6, 7-5-3,

1-5-9, 7-8-9, and 1-2-3, although in Session 2 and 3 (see Figure 8) only 1-2-3

responses were produced.

In Figure 10 shows the number of switches and number of patterns for LNFA04.

No switches between 1-2-3 and 7-8-9 occurred during FR1-FR1 schedules, although

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some switches to other patterns occurred. During FI30-FR5, switching and target

patterns were stable. Beginning in Session 8, Extinction was accompanied by a drastic

separation between the high number of All Switches (small square marker) and the

number of switches between schedule components (circles). The number of patterns

produced increased during extinction, while the number of switches between schedule

components eventually decreased to near zero. Switches between schedule

components remained low during the last session, in which only 7-8-9 responses

produced sounds.

MPFA04

Figure 11 shows cumulative records of 1-2-3 and 7-8-9 performances for

MPFA04. During Session 1, 13.85 left-to-right responses occurred per minute. A

change to Concurrent FR1-FR1 schedules was accompanied by higher rates of 1-2-3,

whereas 7-8-9 occurred at 2.0 rpm (Session 2) and then 0.4 rpm (Session 3).

Performance of 7-8-9 responses had stopped by the second min of Session 3. When

Concurrent FI30-FR5 schedules for 1-2-3 and 7-8-9 were implemented, a majority of

responding shifted to 7-8-9, and by Session 5, 1-2-3 responding had diminished to 0.1

rpm (responding during the last 19.5 minutes occurred exclusively on 7-8-9). During

Session 7, the response requirement on the fixed ratio component was increased to 10.

This contingency change created low response rates on 1-2-3 and higher response

rates on 7-8-9. During the last session in schedule maintenance, 2.1 rpm occurred for 1-

2-3, and 72.5 7-8-9 responses occurred per minute. Extinction produced high and

undifferentiated rates of responding on both schedule components. 1-2-3 performance,

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maintained at low rates during Concurrent FI30-FR10 schedules, occurred at higher

rates than 7-8-9 during Sessions 11, 12, 13, and 14. Concurrent EXT-FR10 schedules

eliminated responding on 1-2-3 and shifted responding exclusively to 7-8-9

performance, maintained at 76 rpm for the last 19 minutes of Session 15.

Figure 12 shows that MPFA04 produced 46 different left-to-right key sequence

topographies during Session 1. 1-2-3 responses constituted the highest proportion of

patterns produced relative to other patterns.

Session frequencies of patterns and switches are depicted in Figure 13 for

MPFA04. Recall that 1-2-3 responding stopped during the Concurrent FI30-FR5

condition; this phenomenon can be seen in Figure 13 in the low frequencies of switches

from 123 to 789 and switches from 789 to 123. During the last two sessions of the FI30-

FR10 contingency, switching between 1-2-3 and 7-8-9 stabilized. In Extinction,

increasing numbers of patterns other than 1-2-3 or 7-8-9 were emitted. A downward

trend across Extinction sessions was seen in the number of all switches and the number

of target switches.

Table 3 shows the relative proportion of responding on 1-2-3 and the

consequences produced on that component, versus responding and consequences

produced for 7-8-9 patterns for Participant MPFA04. In Sessions 5 and 6, only 7-8-9

responses produced consequences. Increasing the FR schedule value shifted some

responding to the FI component during Sessions 7, 8 and 9. The proportion of

responding on 7-8-9 gradually decreased over the Extinction condition, and

reinstatement of the FR10 contingency for 7-8-9 resulted in 99.02 percent of responding

allocated to that schedule during the last session.

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PWFA04

Cumulative records of 1-2-3 and 7-8-9 are shown for Participant PWFA04 in

Figure 14. During Session 1, left-to-right patterns occurred at a stable 10.25 rpm.

Concurrent FR1-FR1 schedules produced comparable records of 1-2-3 and 7-8-9, with

respective rates of 6.05 and 5.45 rpm. Under concurrent FI30-FR5 schedules, rates

differentiated for 1-2-3 and 7-8-9 early in Session 3, remaining consistent over the next

4 sessions. 7-8-9 occurred at 28.2, 31.35, 31.25, 40.45, and 36.25 rpm for the

consecutive sessions 3, 4, 5, 6, and 7, while 1-2-3 occurred at the relatively lower rates

of 8.4, 8.7, 8.0, 1,8, and 6.5 rpm. During Extinction, rates of 7-8-9 decreased overall

relative to rates of 1-2-3. Performance during extinction generally consisted of periods

of production of one pattern followed by changing to the other pattern for an extended

period. Examples are seen in Session 11, where long runs of responding on 7-8-9

alternate with short bursts of responding on 1-2-3, and in Session 12 where long bouts

of 1-2-3 alternate with 3 short runs of 7-8-9 patterns. During Session 13, Concurrent

EXT(123) FR5(789) schedules were in effect. 1-2-3 was produced at 0.8 rpm while 7-8-

9 responses occurred at 43.95 per minute.

Left-to-right patterns (Session 1) are depicted in Figure 15 for Participant

PWFA04. Forty-seven different patterns occurred, with 9 topographies occurring at least

10 times.

Figure 16 shows across sessions the number of target patterns, switches from 1-

2-3 to 7-8-9, switches from 7-8-9 to 1-2-3, and the number of all switches for PWFA04.

The number of target patterns produced is consistent across sessions during concurrent

FI-FR schedules. During Extinction, the number of all switches separates from the

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number of switches between schedules components, indicating that relative to the

schedule maintenance condition, responding during extinction consisted of many

switches between patterns that were neither 1-2-3 nor 7-8-9.

Table 4 shows the proportion of responding on 1-2-3 versus the proportion of

consequences obtained via 1-2-3, and the proportion of responding and consequences

obtained for 7-8-9. Responding closely matched the proportion of obtained

consequences for PWFA04 during Concurrent FI30-FR5 schedules. For example, note

Session 4 in which 21.72% of consequences were obtained on 1-2-3 and 20.72% of the

total responses occurred on 1-2-3. Likewise, 7-8-9 responding constituted 78.28% of

total responding and obtained 79.62% of all consequences delivered. This relationship

between 1-2-3 and 7-8-9 responding was constant across subsequent sessions under

concurrent FI-FR schedules in which 7-8-9 consistently made up a majority of

responding and obtained a majority of consequent sounds.

General Results

Figure 17 shows scatterplots of keystrokes as they occurred within sessions.

Results for Participant CCSP05 are on top, followed by scatterplots for SMSP05,

LNFA04, MPFA04, and PWFA04. Each block is a scatterplot depicting one session.

Blocks in the left column show scatterplots from Session 1 for all participants, blocks in

the middle column show the last session in the schedule maintenance phase

(Concurrent FI-FR schedules), and blocks in the far right column show the first session

of Extinction. The Y-axis for each row of scatterplots is labeled from 1 to 9, representing

the 9 numbers on the numeric keypad. During Acquisition, much variation in the location

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of keystrokes was present for all participants. Scatterplots from the schedule

maintenance phase, in which only 1-2-3 and 7-8-9 patterns were followed by sounds,

show that the keys 1, 2, 3, 7, 8, and 9 constitute a majority of keystrokes. During

extinction, increased variability in the topography of left-to-right patterns occurred for all

participants. While each participant responded under schedule maintenance with a

slightly different pattern, each participant’s pattern is consistent throughout the entire

session. During extinction, the pattern of switching between 1-2-3 and 7-8-9, responding

on various keys, and allocation of a majority of responses to one schedule component

or the other are all disrupted and variable, compared to schedule maintenance.

Figure 18 depicts the number of left-to-right patterns and the number of

keystrokes across sessions for 3 participants. Plotted along the vertical axis is the

number of patterns and keystrokes, and sessions are listed across the X-axis. A left-to-

right pattern could include any number of keystrokes greater than three. During

schedule maintenance, the numbers of both keystrokes and left-to-right patterns are

maintained for each participant. The number of keystrokes to patterns was roughly 3:1

during schedule maintenance, with 3 keystrokes per pattern on average. During

extinction, the two data paths separate, showing that patterns during extinction were

made up of variable numbers of keystrokes. After extinction, when participants were

again exposed to concurrent schedules, the number of keystrokes and patterns

immediately changed, returning to the 3:1 ratio produced during the earlier schedule

maintenance condition.

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CHAPTER 4

DISCUSSION

The present study contributes an apparatus and research paradigm useful in

generating human performances sensitive to concurrent schedules of reinforcement.

Five participants produced performances observed to be under temporal and ratio

control of concurrent fixed interval-fixed ratio schedules. Two aspects of interaction

between FI and FR schedules were distinguishable in the data. First, interaction

between two schedules was observed in that changes in the value of one schedule

affected behavior reinforced on another schedule. Second, switching from one pattern

to the other functioned as an operant unit, showing stability during schedule

maintenance conditions and sensitivity to extinction. These effects are discussed in the

context of current views on behavior under concurrent schedules of reinforcement, and

some implications for the conceptualization, measurement, analysis, and treatment of

complex behavior are presented.

The current study utilized a technique for shaping a left-to-right pattern of free-

operant responding. For each subject, the left-to-right shaping contingency established

a baseline that was later shown to be sensitive to changing contingencies. That is,

stable rates of responding changed immediately upon introduction of the fixed interval-

fixed ratio contingency, and differential patterns emerged corresponding to the FI and

FR contingencies. Extinction produced variable pattern lengths and location of

keystrokes, effects similar to those documented in previous research on extinction

performances of humans (Anderson, 2000) and non-humans (Antonitis, 1951; Azrin,

Hutchinson, & Hake, 1966). Also, response rates rose during extinction and then fell,

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and upon reintroduction of the contingency, returned to previously seen levels under the

concurrent FI-FR schedules. For Participants CCSP05, SMSP05, MPFA04, and

PWFA04, the FI-30s schedule component produced lower rates of responding that

corresponded with consequences available every 30 seconds if a response occurred.

The FR component produced higher rates of ratio-controlled responding in which high

rates consisted of runs of 5 or 10 responses, depending on the schedule in effect. A

different effect was seen with Participant LNFA04, who produced higher rates of

responding on the FI contingency and lower rates on the FR schedule (see below for

further discussion of LNFA04’s performance).

Although the results show control by single schedules, interactions were also

seen between the schedules. The current study provides several examples of

interaction between concurrent schedules maintaining differentiated human

performances. One type of interaction was seen when the value of schedules were

changed. For example, under Concurrent FR1-FR1 schedules for patterns 1-2-3 and 7-

8-9, Participant LNFA04 produced only the pattern 1-2-3. When the schedule values

were changed to FI30s-FR5, LNFA04 produced both 1-2-3 and 7-8-9 at stable rates for

4 sessions. As opposed to the other participants, LNFA04 produced 1-2-3 at rates

significantly higher than 7-8-9 rates during those sessions. After experiencing 4

sessions of Extinction, re-introduction of an FR5 contingency for 7-8-9 was

accompanied by high rates on 7-8-9. Similarly, Participant MPFA04 initially performed

7-8-9 at very low rates, producing 1-2-3 almost exclusively, on Concurrent FR1-FR1

schedules. Changing the contingency to FI30 for 1-2-3 and FR5 for 7-8-9 was

accompanied by stable high rates on 7-8-9. At this point MPFA04 stopped producing 1-

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2-3 patterns. Increasing the FR value to 10 for 7-8-9 produced differentiated rates for 1-

2-3 and 7-8-9. This result is consistent with Findley’s (1958) finding that altering the

schedule arrangement can shift “preference” from one schedule to another.

Allocating more behavior to a given component than predicted may be a function

of the actual contingencies in effect, together with that organism’s previous experience.

Response distributions could be analyzed in descriptive terms rather than explained as

“preference”. Michael (2004) called attention to Skinner’s molecular analyses of

alternation on concurrent schedules, suggesting that the matching law could be

“molecularized” by considering the differential conditioned reinforcement strengths that

result from switching from right to left and left to right. Michael closes by arguing: “This

type of analysis would seem an appropriate replacement for explanations in terms of

choice or preference, which may well be functioning as explanatory fictions (one of

Skinner’s most useful pejoratives)” (p. 91, emphasis in the original).

A second type of interaction describes the interaction of consequences provided

on one schedule with the behavior on another. This has been termed in the literature

“local interaction.” Catania (1966) cited his 1963 study in which local interactions

between schedules were observed for combinations of ratio and interval schedules.

While both schedules maintained responding, responding on one schedule appeared to

be interacting with reinforcement deliveries obtained on the other. Catania described

pigeons responding on the FR component early in the FI interval, switching to the FI

component until a reinforcing stimulus was presented, and switching back to the FR

component (1966). The current study contributes similar evidence of interactions

between schedules. The sequence of 1-2-3 followed immediately by 7-8-9 (and the

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reciprocal) was consistently produced during schedule maintenance for each

participant. Each participant exhibited a variation of the switch pattern, and the rate of

switching between 123 and 789 after consequences were discontinued was observed to

follow a characteristic extinction curve different than that of other operant units. These

results contribute human evidence to a growing body of work in which switching

between schedules is shown to be an operant sensitive to reinforcement contingencies

(Crowley and Donohue, 2004; Machado, 1997; Shull & Pliskoff, 1967; Stubbs, Pliskoff,

& Reid, 1977).

Catania (1973) distinguished between the functional operant and descriptive

operant. During a concurrent schedule, an experimenter has designated consequences

to follow instances of one or the other of two response classes. The switching between

the defined schedule components may turn out to be a functional operant. Whether

switching should be eliminated using a paradigm that provides no special

consequences for it (Crowley and Donahoe, 2004), or explicitly studied as a result of a

concurrent schedule (Skinner, 1950; Skinner & Ferster, 1957; Michael, 2004) depends

on the strategic goals underlying research. One goal may include clarifying variables

controlling the strength of a response. In fact, the concurrent schedules studied by

Skinner and Ferster were preparations testing the relative strength of two operants

(1950). Crowley and Donahoe considered the acquisition of other operants (e.g.,

switching) to contaminate the measurement of the actual relative strength of two

(experimenter-defined) operants, and used several procedural variations to clarify the

role of switching in the acquisition of matching (2004).

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A different strategic goal of research, such as informing applied practice, may call

for examining the variables under which switching occurs regardless of its role in

maintaining a matching relation. Past research has imposed various penalties on

switching, requiring travel to the other alternative (Baum, 1982), instituting a blackout

period after a switch response (Todorov, 1971), or requiring a number of responses on

a changeover key (Pliskoff, Cicerone, & Nelson, 1978). In some cases, it could be

beneficial to increase the likelihood of switching to other alternatives. For example, a

common feature of disorders associated with autism is restricted activities and interests,

according to the Diagnostic and Statistical Manual of Mental Disorders (4th ed.; DSM-

IV; American Psychiatric Association, 1994). Sampling other activities could be

analogous to an operant class comprised of switches to alternatives on a concurrent

schedule. Developing procedures that instate and maintain switching in a behavioral

repertoire could benefit many populations.

Catania (1966) observed that on concurrent schedules although the emission of

independent responses is possible, the consequences scheduled by one schedule are

not independent of responses on the other. Thus, even if two keys are programmed

independently, interaction between them in a real-world setting may be impractical to

eliminate. It may be instructive to examine the actual interaction between more

schedules and different parameters without a changeover delay, to reveal interaction

effects that are present in classrooms, households and playgrounds, but may be

sterilized from laboratory settings. In this study interaction between the two schedules

was shown in that rates of switching between the components appeared sensitive to

changing contingencies and was especially affected by extinction.

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A practical use of concurrent schedules with human participants in the literature

is as a platform to study resurgence, in which behavior is observed to occur during

extinction (Epstein, 1985; Leitenberg, Rawson, & Mulick, 1975; Lieving & Lattal, 2003;

Lieving, Hagopian, Long, & O’Connor, 2004). These studies examine resurgence under

concurrent reinforcement-extinction schedules in which a response class is first

exposed to reinforcement, then placed on extinction while a new, concurrently available

alternative produces reinforcement. Using this paradigm, Lieving et al. (2004) found

responding during extinction to be a function of previous reinforcement (that is,

responding was not best described as induced variability). The authors emphasized the

importance of understanding the conditions under which the phenomenon identified as

resurgence occurs. This point is relevant to practitioners employed as change agents

working with humans engaging in complex behavior that results in deleterious side

effects. Much has been written on the importance of assessing the schedules of

reinforcement contributing to the maintenance of such behavior (see Lerman & Iwata

(1996) for a review of literature regarding variables that contribute to resistance to

extinction, with implications for application). The current study contributes evidence that

behavior previously reinforced can be extinguished quickly while an available alternative

is concurrently reinforced. CCSP05 and SMSP05 experienced extinction and then a

return to concurrent FI-FR schedules (accompanied by a return to FI-FR-controlled

performances). For the other three participants (LNFA04, MPFA04, and PWFA04),

extinction was followed by a concurrent EXT-FR condition. Each of these participants

exhibited performances during extinction in which neither operant decreased to zero-

levels of occurrence. When an alternative response (7-8-9) was reinforced concurrently

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with extinction contingency for (1-2-3), rates of responding on the 7-8-9 schedule

component responded dramatically and quickly, while responding on the EXT

component dropped to near-zero levels. Furthermore, these participants stopped

producing variable patterns and variable inter-response times as soon as an alternative

was reinforced. The results of the present study provide a laboratory example of how

practical it could be to provide alternatives while implementing extinction.

The metaphor of the behavior stream offers a potentially useful window through

which continuous behavior may be interpreted. Within the behavior stream, more than

one “alternative” could be measured, together with its interaction with other alternatives.

Still, isolation of a unit of behavior within the stream is recognized to be difficult, even in

descriptive analysis. Discussing the problem of quantifying interaction between a

mother and her infant, Brazelton, Kosolowski, and Main (1974) wrote, “The behavior of

any one member becomes a part of a cluster of behaviors which interact with a cluster

of behaviors from the other member of the dyad. No single behavior can be separated

from the cluster for analysis without losing its meaning in the sequence.” Attempting to

avoid this loss of meaning, Bakeman and Gottman described a time-series analysis of

the dependencies among events and sequences of events occurring in a stream of

behavior across time (1986). These researchers point out that in classical parametric

statistics, the assumption that observations are independent forms the basis of

distribution statistics. However, when researchers examine series of observational data

using sequential analysis, dependence in the data is the phenomenon of interest.

Whether complex behavior is conceptualized as choice between two

independent alternatives (Herrnstein, 1961; 1970) or as a behavior stream that includes

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interactions between the alternatives (Schoenfeld and Farmer, 1970; Goldiamond,

1974) has different implications for behavior analysis and related fields. The current

literature debate on this issue undercores the fact that the stimulus control of behavior

on concurrent schedules is not yet fully fleshed out. Consider a situation in which a

spouse engages in a sequence that includes many responses—perhaps including

eating, hitting, yelling, cooking, sexual behavior, and taking out the trash. While it may

be difficult and impractical to separate one operant completely from the sources of

control of the others, analysis of the stream of behavior could illuminate sequences of

responding that are typically initiated by similar circumstances and perhaps function as

units. The current study provides further evidence that switching, although not explicitly

targeted by the scheduled contingencies, behaved like a separate operant.

In summary, this study extends previous work by Crowley and Donahoe (2004)

and others (e.g., Machado, 1997; Shull & Pliskoff, 1967) in which switching responses

functioned as an operant unit under experimental control. Further research could use

the apparatus described here to maintain and provide consequences for the switching

response, extending previous research in which switching was part of an operant

contingency (Baum, 1982; Todorov, 1971; Pliskoff et al., 1978). No formal changeover

delay was programmed, replicating work by Skinner and Ferster (1957), Catania (1962;

1967), and Neuringer (2000) in which contingency changes were accompanied by

changes in the allocation of responding to schedule components without a COD.

Extinction produced new behaviors and variability in several dimensions of behavior,

consistent with work by Anderson (2000), Antonitis (1951), and Azrin et al. (1966) This

study is one of several recent experiments using an apparatus that generates schedule-

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controlled performances of human participants (Anderson, 2000; Koremura, 2001;

Rosales-Ruiz et al., 1999).

Examining comprehensively the interaction of two or more sources of

reinforcement, determining their respective influence on an individual’s behavior stream,

and viewing that behavior stream from a perspective that considers the alternatives

concurrently available to the organism (including ad libitum changing from one

alternative to another) has tremendous implications for teaching, reinstating, eliminating,

and changing behavior. The results of this study indicate that procedural progress

toward such a task is possible.

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Figure 1. Cumulative records for Participant CCSP05.

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0

5

10

15

20

25

30

35

40

45

123

126

129

153

156

159

183

186

189

423

426

429

453

456

459

483

486

489

723

726

729

753

756

759

783

786

789

1323

1326

2129

4156

1425

621

626

2312

357

353

5915

974

123

9878

974

3695

374

8124

5932

6598

429

2366

9852

1698

456

Left-to-Right Patterns

Cou

nt

CCSP05Session 1

Figure 2. Histogram of patterns produced during Session 1 for Participant CCSP05.

200

200Res

pons

es

1050

Figure 3. Cumulative record of Session 5 for Participant CCSP05.

Minutes

FR5 component (789 responses)

FI30s component (123 responses)

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CCSP05

1

10

100

1000

10000

1 2 3 4 5 6 7 8 9 10 11 12

Sessions

Num

ber

PatternsSw itch 123 to 789Sw itch 789 to 123All Sw itches

FR1

FR1-

FR1 FI30-FR5 EXTINCTION FI30-FR5

Figure 4. Switches for Participant CCSP05.

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1

123789

2

FRI(left-right) Concurrent FRI(123) FR1(789)

3

123

789

Concurrent FI30(123) FR5(789)123

789

4

123

789

5

123

789

6 7

123

789

123

789

8

123

789

10

123789

123789

11

EXTINCTION

1000

500

200

Res

pons

es

Minutes

SMSP05

12 13 14

123789

123

789123

789

Concurrent FI30(123) FR5(789)

9

20 minutes

Figure 5. Cumulative records for Participant SMSP05.

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0

5

10

15

20

25

30

Left-to-Right Patterns

Cou

ntSMSP05Session 1

Figure 6. Histogram of patterns produced during Session 1 for Participant SMSP05.

SM SP05

1

10

100

1000

10000

1 2 3 4 5 6 7 8 9 10 11 12 13 14

Sessions

Num

ber

Target PatternsSw itch 123 to 789Sw itch 789 to 123All Sw itches

FR1

FR1-

FR1 FI30-FR5 EXTINCTION FI30-FR5

Figure 7. Switches for Participant SMSP05.

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FR1 (left-right) Concurrent FR1 (123) FR1 (789)

Concurrent FI30(123) FR5(789)

EXTINCTION

Concurrent EXT(123) FR5(789)

123

789789

123123 123

123123

123123

123

123 123

789 789

789 789789

789

789

132

4 5 6 7

8 9 10 11

13

LNFA04

20 minutes

1500

750

200

Res

pons

es

Minutes

Figure 8. Cumulative records for Participant LNFA04.

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0

10

20

30

40

50

60C

ount

123

159

423

456

489

753

789

4756

4829

5423

5456

7453

7489

8159

8456

8789

4288

3

6548

9

8752

3

9195

9

9515

9

2744

89

4691

23

9845

286

8631

4752

9

LNFA04

Left-to-Right Patterns

Figure 9. Histogram of patterns produced during Session 1 for Participant LNFA04.

LNFA04

1

10

100

1000

10000

1 2 3 4 5 6 7 8 9 10 11 12 13

Sessions

Num

ber

PatternsSw itch 123 to 789Sw itch 789 to 123All Sw itches

FR1 FR1-FR1 FI30-FR5 EXTINCTION EXT-FR5

Figure 10. Switches for Participant LNFA04.

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FR1 (left-right) Concurrent FR1(123)-FR1(789)

Concurrent FI30(123)-FR5(789)

Concurrent FI30(123)-FR10(789)

EXTINCTION

Concurrent EXT(123)FR10(789)

20 minutes

1600

800

200

Res

pons

es

Minutes

123

789 789

789789 789

789789 789

789

789789

789789

789

123

123123 123

123123 123

123

123 123

123

123

123

1 2 3

4 5 6

7 8 9

10 11 12

13 14 15

MPFA05

Figure 11. Cumulative records for Participant MPFA04.

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0

10

20

30

40

50

60

70

Cou

nt

75.3

123.

012

6.0

129.

015

3.0

156.

015

9.0

183.

018

6.0

189.

042

3.0

426.

042

9.0

453.

045

6.0

459.

048

3.0

486.

048

9.0

723.

072

6.0

729.

075

3.0

756.

075

9.0

783.

078

6.0

789.

012

56.0

1323

.015

26.0

1659

.031

23.0

4123

.044

56.0

6159

.073

83.0

7489

.079

89.0

8786

.023

423.

053

753.

055

0123

.057

1983

.014

7442

3.0

9632

5874

89.0

MPFA04Session 1

Left-to-Right Patterns

Figure 12. Histogram of patterns produced during Session 1 for Participant MPFA04.

MPFA04

1

10

100

1000

10000

1 2 3 4 5 6 7 8 9 10 11 12 13 14 15

Sessions

Num

ber

PatternsSw itch 123 to 789Sw itch 789 to 123All Sw itches

FR1 FR1-FR1 FI30-FR5 FI30-FR10 EXTINCTION EXT-FR10

Figure 13. Switches for Participant MPFA04.

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PWFA04

FR1(left-right) Concurrent FR1(123)- FR1(789)

Concurrent FI30(123)- FR5(789)

EXTINCTION

Concurrent EXT(123)- FR5(789)

123789

123

789

123 123

123123

123 123 123

123

123

123

789 789

789789

789

789

789

789

789

789

1 2

3 4 5

6 7

8 9 10

11 12

13

20 minutes

1400

700

200

Res

pons

es

Minutes

Figure 14. Cumulative records for Participant PWFA04.

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02

46

810

1214

1618

2022

Cou

nt

123

126

153

156

159

423

426

453

456

459

486

489

753

756

759

786

789

1586

1589

2156

4589

4856

5456

7526

7789

7989

8756

1578

641

523

4522

345

879

6645

675

123

9915

925

8426

3591

5951

4586

7512

8975

2226

1474

589

9815

823

3265

9845

998

7144

856

9987

4151

396

3852

1586

3258

9641

526

3958

2641

7859

PWFA04Session 1

Left-to-Right Patterns

Figure 15. Histogram of patterns produced during Session 1 for Participant PWFA05.

PWFA04

1

10

100

1000

10000

1 2 3 4 5 6 7 8 9 10 11 12 13

Sessions

Num

ber

PatternsSw itch 123 to 789Sw itch 789 to 123All Sw itches

FR1

FR1-

FR1 FI30-FR5 EXTINCTION EXT-FR5

Figure 16. Switching for Participant PWFA04.

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123456789SMSP05

Key

Acquisition Schedule Maintenance

CCSP05

123456789

Extinction

123456789MPFA04

123456789PWFA04

20 min 20 min 20 min

123456789LNFA04

Figure 17. Scatterplots of keypresses for all participants.

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Figure 18. Left-to-right patterns and keystrokes across sessions for participants CCSP05, PWFA04, and SMSP05.

CCSP05

0

2000

4000

6000

8000

1 3 5 7 9 11

Keystrokes

Left-to-Right Patterns

PWSP05

0

2000

4000

6000

0 2 4 6 8 10 12 14

FR1 FI30-FR5 EXTINCTION

SMSP05

0

2000

4000

6000

0 2 4 6 8 10 12

Sessions

FR1 FI30-FR5 EXTINCTION

FR1 FR1-FR1

FI30-FR5 EXTINCTION FI30-FR5

EXT-FR5

FI30-FR5

Num

ber

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Table 1 Proportion of Responses and Consequences Obtained on each Schedule for Participant CCSP05 Proportion

Condition Session Responding

on 123 Consequences

Obtained on 123 Responding

on 789 Consequences

Obtained on 789 FR1 1 n/a n/a n/a n/a Concurrent FR1(123)-FR1(789)

2 58% 58% 42% 42%

3 39% 16% 61% 85% 4 14% 9% 86% 91%

Concurrent FI30(123)-FR5 (789) 5 14% 12% 87% 88%

6 23% 0 77% 0 7 55% 0 46% 0 8 67% 0 33% 0 9 26% 0 74% 0 10 39% 0 61% 0

EXTINCTION

11 13% 0 97% 0 Concurrent FI30(123)-FR5 (789)

12 6% 8 94% 92%

Table 2 Proportion of Responses and Consequences Obtained on each Schedule for Participant SMSP05

Proportion

Condition Session Responding

on 123 Consequences

Obtained on 123 Responding on

789 Consequences

Obtained on 789 FR1 1 n/a n/a n/a n/a Concurrent FR1(123) FR1(789)

2 50% 50% 50% 50%

3 59% 32% 41% 68% 4 67% 36% 33% 64% 5 40% 23% 60% 77% 6 63% 30% 37% 70% 7 24% 14% 76% 86%

Concurrent FI30(123) FR5(789)

8 31% 18% 69% 82% 9 58% 0 42% 0

10 48% 0 50% 0 11 54% 0 46% 0 12 48% 0 52% 0

EXTINCTION

13 52% 0 48% 0 Concurrent FI30(123) FR5(789)

14 32% 18% 68% 82%

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Table 3 Proportion of Responses and Consequences Obtained on each Schedule for Participant MPSP04

Condition SessionResponding on

123

ConsequencesObtained on

123Responding on

789

ConsequencesObtained on

789FR1 1 n/a n/a n/a n/a

2 87% 87% 13% 13%3 97% 97% 3% 3%4 24% 11% 76% 89%5 1% 0% 99% 100%6 1% 0% 99% 100%7 15% 23% 85% 77%8 3% 19% 97% 81%9 3% 19% 97% 81%

10 43% 0 57% 011 55% 0 45% 012 61% 0 39% 013 53% 0 47% 014 76% 0 24% 0

EXT- FR10(789) 15 1% 0% 99% 100%EXTINCTION

Proportion

ConcurrentFR1(123)

ConcurrentFI30(123)FR5(789)

ConcurrentFI30(123)Fr10(789)

Table 4 Proportion of Responses and Consequences Obtained on each Schedule for Participant PWFA04

Proportion

Condition Session Responding

on 123

Consequences Obtained on

123 Responding

on 789

Consequences Obtained on

789 FR1 1 n/a n/a n/a n/a Concurrent FR1(123) FR1(789)

2 53% 53% 47% 47%

3 23% 19% 77% 81% 4 22% 20% 78% 77% 5 20% 20% 80% 80% 6 4% 7% 96% 93%

Concurrent FI30(123) FR5(789)

7 15% 15% 85% 84% 8 24% 0 76% 0 9 16% 0 84% 0

10 41% 0 59% 0 11 14% 0 86% 0

EXTINCTION

12 83% 0 17% 0 EXT/ FR5 (789) 13 2% 0% 98% 100%

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Table 5 Session Information for CCSP05

Session Number Schedule Total

Responses Overall RPM 123 responses RPM

789 responses RPM

1 FR1 244 12.2 n/a n/a

2 Concurrent FR1-FR1 408 20.4 11.8 8.6

3 1079 53.95 21.2 32.75 4 1136 56.8 8.05 48.75 5

Concurrent FI30-FR5

1100 55 7.45 47.55 6 1137 56.85 12.9 43.95 7 1125 56.25 30.65 25.6 8 774 38.7 25.85 12.85 9 572 28.6 7.55 21.05

10 1323 66.15 25.8 40.35 11

EXTINCTION

348 17.4 2.25 15.15

12 Concurrent FI30-FR5 1119 55.95 3.2 52.75

Table 6 Session Information for Participant SMSP05

Session Number Schedule Total

Responses Overall RPM 123 responses RPM

789 responses RPM

1 FR1 250 12.5 n/a n/a

2 Concurrent FR1-FR1 343 17.15 8.55 8.6

3 912 45.6 27.1 18.5 4 1013 50.65 33.85 16.8 5 959 47.95 18.95 29 6 1149 57.45 36.45 21 7 1089 54.45 12.85 41.6 8

Concurrent FI30-FR5

1058 52.9 16.25 36.65 9 1547 77.35 44.65 32.7 10 1207 60.35 30 30.35 11 1451 72.55 38.9 33.65 12 560 28 13.35 14.65 13

EXTINCTION

1400 70 36.5 33.5

14 Concurrent FI30-FR5 1128 56.4 17.9 38.5

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Table 7 Session Information for Participant LNFA04

Session Number Schedule Total

Responses Overall RPM 123 responses RPM

789 responses RPM

1 FR1 233 11.65 n/a n/a 2 277 13.85 11.5 0 3

Concurrent FR1-FR1 303 15.15 15.15 0

4 758 37.9 28.6 9.3 5 893 44.65 34.6 10.05 6 946 47.3 37.3 10 7

Concurrent FI30-FR5

953 47.65 37.4 10.25 8 1252 62.6 41.65 20.95 9 1355 67.75 43.7 24.05 10 1401 70.05 58.8 11.25 11 1291 64.55 47.4 17.15 12

EXTINCTION

1388 69.4 44.5 24.9

13 Concurrent FI30-FR5 759 37.95 0.65 37.3

Table 8 Session Information for Participant MPFA04

Session Number Schedule Total

Responses Overall RPM 123 responses RPM

789 responses RPM

1 FR1 277 13.85 n/a n/a 2 306 15.3 13.3 2 3

Concurrent FR1-FR1 319 15.95 15.55 0.4

4 1058 52.9 12.95 39.95 5 1063 53.15 0.45 52.7 6

Concurrent FI30-FR5

1062 53.1 0.1 53 7 1370 68.5 10.55 57.95 8 1484 74.2 2.2 72 9

Concurrent FI30-FR10

1492 74.6 2.1 72.5 10 1818 90.9 38.6 52.3 11 1839 91.95 50.55 41.4 12 1523 76.15 46.45 29.7 13 1358 67.9 36.05 31.85 14

EXTINCTION

1598 79.9 60.65 19.25 15 EXT-FR10 1535 76.75 0.75 76

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Table 9 Session Information for Participant PWFA04 Session Number Schedule Total

Responses Overall RPM 123 responses RPM

789 responses RPM

1 FR1 205 10.25 n/a n/a

2 Concurrent FR1-FR1 230 11.5 6.05 5.45

3 732 36.6 8.4 28.2 4 801 40.05 8.7 31.35 5 785 39.25 8 31.25 6 845 42.25 1.8 40.45 7

Concurrent FI30-FR5

856 42.8 6.55 36.25 8 904 45.2 11 34.2 9 1248 62.4 9.8 52.6 10 681 34.05 14.1 19.95 11 882 44.1 6.15 37.95 12

EXTINCTION

957 47.85 39.5 8.35 13 EXT-FR5 895 44.75 0.8 43.95

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APPENDIX A

INFORMED CONSENT FORM

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Informed Consent Form

______ I understand that participation in this study requires me to schedule a time to work for approximately 20 minutes each week day, and I can expect to participate in 10-20 sessions. _____ I understand that pay is $5 per each session (approximately 20 minutes) and that I will be paid at the end of my involvement in the study. ______ My participation in this study is voluntary. There are no known risks associated with the procedures used in this study. It has been explained to me that I can withdraw my participation at any time without penalty by contacting the research assistant and informing her that I will no longer participate. ______ I have been given written contact information for the research assistant for this study, and will contact her at least 24 hours in advance if I must re-schedule a session. ______ I understand that this study is experimental in nature and that I will be fully debriefed at the end of my involvement in the study, when the experimenter will answer my questions about the experiment. I will not discuss the experiment with anyone else participating in this study. Results from this study may be presented at conferences or events relating to the research, and I understand that if my results are used, I will never be identified by my name. _____ I have read the conditions outlined above, and have agreed to participate in this study on these conditions. _____ I understand that I will receive a copy of this consent form when signed/dated. Participant Name (Printed) Signature Date Experimenter Name (Printed) Signature Date Principle Investigator (Printed) Signature Date

CONTACT INFORMATION (Investigator Name/Phone/Email):

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APPENDIX B

APPARATUS DIAGRAM

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1 2 3

4 5 6

7 98

2. 3X3 grid with 9boxes correspondingto the 9 keys on thenumeric keypad.

START

1. Participant facedcomputer monitorwithin reach of key-board and mouse,wearing audio head-phones.

4. The first key pressin the middle columnoccurs. The corre-sponding squareappears “pressed in”.

3. After the first keypress correspondingto a number in theLEFT column (1, 4,or 7), the corre-sponding squareappears “pressed in”.

5. Participant pressesa key in the right col-umn; the correspond-ing square appears“pressed in”.

6. After targetedsequence occurs, but-ton appears belowgrid. A mouse-click onthe button produces ashort sound clip.

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APPENDIX C

EXPERIMENTAL DESIGN

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Parti

cipa

nts

1 2 3 4 5 6 7 8 9 10 11 12 13 14 15

CCFI30-FR5

PWEXT-FR5

SMFI30-FR5

LNEXT-FR5

MPEXT-FR10

Pre

f Ass

mt

FR1

(L-R

)

FR1-

FR1 FI30-FR5

FI30-FR10

EXTINCTION

A B C D C

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