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G. Tyler Miller’sLiving in the Environment13th Edition
G. Tyler Miller’sLiving in the Environment13th Edition
Ecosystems: Components, Energy Flow, and Matter Cycling
Chapter 4
Ecosystems: Components, Energy Flow, and Matter Cycling
Chapter 4
Key ConceptsKey Concepts
• What is ecology?What is ecology?• What basic processes keep us and What basic processes keep us and
other organisms alive?other organisms alive?• What are the major components of What are the major components of
the ecosystem?the ecosystem?• What happens to energy in the What happens to energy in the
ecosystem?ecosystem?• How do scientists study the How do scientists study the
ecosystem?ecosystem?• What are ecosystem services?What are ecosystem services?
What is ecology?What is ecology?
The study of how organisms interact The study of how organisms interact with one another and with their non-with one another and with their non-living environment.living environment.
((oikos oikos “place to live”“place to live” logoslogos “study “study of”)of”)
How nature is connected.How nature is connected.
Communities
Subatomic Particles
Atoms
Molecules
Protoplasm
Cells
Tissues
Organs
Organ systems
Organisms
Populations
Populations
Communities
Ecosystems
Biosphere
Earth
Planets
Solar systems
Galaxies
Universe
Organisms
Realm of ecology
Ecosystems
Biosphere
DNA(informationstorage, nonucleus)
Cell membrane(transport ofraw materials)
Protein constructionand energy conversionoccur without specializedinternal structures
Prokaryotic Cell
Proteinconstruction
Nucleus(information storage)
Energyconversion
Cell membrane(transport of raw
materials andfinished products)Packaging
Eukaryotic Cell
Figure 4-3 (1)Page 67Figure 4-3 (1)Page 67
The Nature of EcologyThe Nature of Ecology
OrganismOrganismAny form of lifeAny form of life
Ecosystem OrganizationEcosystem Organization
The Nature of EcologyThe Nature of Ecology
• OrganismOrganism– Any form of lifeAny form of life– SpeciesSpecies
• Group of organisms Group of organisms that resemble one that resemble one anotheranother
• Actually or potentially Actually or potentially breed with one breed with one anotheranother
• Produce live, fertile Produce live, fertile offspringoffspring
Ecosystem OrganizationEcosystem Organization
The Nature of EcologyThe Nature of Ecology
• CommunitiesCommunities– Populations of the different Populations of the different
species occupying a species occupying a particular placeparticular place
– Biological communityBiological community
• PopulationsPopulations– Group of interacting Group of interacting
individual of the same individual of the same species that occupy a species that occupy a specific area a the same specific area a the same time.time.
• OrganismsOrganisms– Any living organismAny living organism
Ecosystem OrganizationEcosystem Organization
The Nature of EcologyThe Nature of Ecology
•BiosphereBiosphere
All of the earth’s ecosystemsAll of the earth’s ecosystems
•EcosystemEcosystem
A community of different species A community of different species interacting with one another and interacting with one another and their nonliving environmenttheir nonliving environment
Ecosystem OrganizationEcosystem Organization
The Earth’s Life-Support SystemsThe Earth’s Life-Support Systems
Fig. 4-6 p. 68Fig. 4-6 p. 68
•AtmosphereAtmosphere
TroposphereTroposphere
StratosphereStratosphere
•HydrosphereHydrosphere
•LithosphereLithosphere
•BiosphereBiosphere
Sustaining Life of EarthSustaining Life of Earth
One-way flow of One-way flow of energyenergy
SunSun
Living materials and Living materials and living thingsliving things
Into the environmentInto the environment
Cycling of matterCycling of matter
Atoms, ions, Atoms, ions, molecules needed for molecules needed for survivalsurvival
GravityGravity
Ecosystem Concepts and ComponentsEcosystem Concepts and Components
• BiomesBiomes– ““By-ohms”By-ohms”– Land Land
ecosystemsecosystems– Distinct climate Distinct climate
and specific and specific life-formslife-forms
• Role of climateRole of climate– Long term Long term
patterns of patterns of weatherweather
– Determines Determines what type of what type of life will thrivelife will thrive
• Aquatic life Aquatic life zoneszones– freshwater freshwater – ocean or ocean or
marine lifemarine life Fig. 4-9 p. 70Fig. 4-9 p. 70
Sun
Producers (rooted plants)
Producers (phytoplankton)
Primary consumers (zooplankton)
Secondary consumers (fish)
Dissolvedchemicals Tertiary consumers
(turtles)
Sediment
Decomposers (bacteria and fungi)
Figure 4-11 Page 72Figure 4-11 Page 72
Major components of freshwater ecosystemMajor components of freshwater ecosystem
Sun
Producer
PrecipitationFalling leaves
and twigs
Producers
Primary consumer(rabbit)
Secondary consumer(fox)
Carbon dioxide (CO2)
Oxygen (O2)
Water
Soil decomposers
Soluble mineral nutrients
Figure 4-12 page 72Figure 4-12 page 72
Major components of a terrestrial ecosystemMajor components of a terrestrial ecosystem
Terrestrial Ecosystems Aquatic Life Zones
• Sunlight
• Temperature
• Precipitation
• Wind
• Latitude (distance from equator)
• Altitude (distance above sea level)
• Fire frequency
• Soil
• Light penetration
• Water currents
• Dissolved nutrient concentrations (especially N and P)
• Suspended solids
• Salinity
Principles of Ecological FactorsPrinciples of Ecological Factors
Abiotic factors
Figure 4-13Page 73
Principles of Ecological FactorsPrinciples of Ecological Factors•Law of toleranceLaw of tolerance- - the existence, abundance, and the existence, abundance, and distribution of a species in an ecosystem are distribution of a species in an ecosystem are determined by whether the levels ofdetermined by whether the levels of one or more one or more physical or chemical factors fall within the range of physical or chemical factors fall within the range of tolerance.tolerance.
OptimumRange
Zone ofPhysiological
Stress
Zone ofIntolerance
Principles of Ecological FactorsPrinciples of Ecological Factors
Fig. 4-14 p. 73; Refer to Fig. 4-13 p. 73Fig. 4-14 p. 73; Refer to Fig. 4-13 p. 73
•Limiting factors principleLimiting factors principle – – too much or too little of too much or too little of any abiotic factor can limit or prevent growth of a any abiotic factor can limit or prevent growth of a population, even if all other factors are at or near population, even if all other factors are at or near the optimum range of tolerance.the optimum range of tolerance.
The Biotic Components of EcosystemsThe Biotic Components of Ecosystems
• Producers Producers (autotrophs)(autotrophs)– Transform energy by Transform energy by
PhotosynthesisPhotosynthesis
• Consumers Consumers (heterotrophs)(heterotrophs)– Transform energy by Transform energy by
Aerobic RespirationAerobic Respiration
• DecomposersDecomposers
Fig. 4-16 p. 75Fig. 4-16 p. 75
Conservation of Matter and Energy CaptureConservation of Matter and Energy Capture
PhotosynthesisPhotosynthesis
6 CO6 CO2 2 + 6 H+ 6 H220 0 CC66HH1212OO66 + 6 O + 6 O22
RespirationRespiration
CC66HH1212OO66 + 6 O + 6 O22 6 CO6 CO2 2 + 6 H+ 6 H220 0
Trophic “Feeding” LevelsTrophic “Feeding” Levels
• First Trophic LevelFirst Trophic Level
• Second Trophic LevelSecond Trophic Level
• Third Trophic LevelThird Trophic Level
• Fourth Trophic LevelFourth Trophic Level
ProducersProducers (plants) (plants)
Primary consumersPrimary consumers (herbivores)(herbivores)
– Feed directly on Feed directly on producersproducers
Secondary consumerSecondary consumer (carnivores)(carnivores)
– Feed on Primary Feed on Primary ConsumersConsumers
Tertiary consumerTertiary consumer– Feed on other Feed on other
carnivorescarnivores
Trophic LevelsTrophic Levels
• OmnivoreOmnivore– Eat plants and animalsEat plants and animals
• Detritivores and ScavengersDetritivores and Scavengers– Feed on Feed on detritusdetritus, dead organisms, and waste, dead organisms, and waste
• DecomposersDecomposers– Break down dead organic materialBreak down dead organic material– Release the resulting simpler compounds into the Release the resulting simpler compounds into the
soilsoil– Anaerobic respiration (absence of oxygen)Anaerobic respiration (absence of oxygen)
• Methane, ethyl alcohol, acetic acid, hydrogen sulfideMethane, ethyl alcohol, acetic acid, hydrogen sulfide
MushroomWoodreduced
to powder
Long-hornedbeetle holes
Bark beetleengraving
Carpenterant
galleries
Termite andcarpenter
antwork
Dry rot fungus
Detritus feeders Decomposers
Time progression Powder broken down by decomposersinto plant nutrients in soil
Figure 4-15 page 75Figure 4-15 page 75
Biodiversity:What is it and why is it important?Biodiversity:What is it and why is it important?
The different life-forms and The different life-forms and life-sustaining processes.life-sustaining processes.
Biodiversity:What is it and why is it important?Biodiversity:What is it and why is it important?
Kinds of biodiversity include:Kinds of biodiversity include:• Genetic diversityGenetic diversity
– Variety in the genetic makeup among Variety in the genetic makeup among individuals within a speciesindividuals within a species
• Species diversitySpecies diversity– Variety among species found in different Variety among species found in different
habitats of the planethabitats of the planet
• Ecological diversityEcological diversity– Variety of biological communitiesVariety of biological communities
• Functional diversityFunctional diversity– Biological and chemical processes or Biological and chemical processes or
functions needed for survivalfunctions needed for survival
Connections: Food Webs and Energy Flow in EcosystemsConnections: Food Webs and Energy Flow in Ecosystems
Food chains – sequence of organisms each of which is a food source for the next.
Food chains – sequence of organisms each of which is a food source for the next.
Connections: Food Webs and Energy Flow in EcosystemsConnections: Food Webs and Energy Flow in Ecosystems
Food webs – a network of interconnected food chains
Food webs – a network of interconnected food chains
ECOLOGY
Ecological Pyramid
A food chain that shows the
relationship between the
organisms in each trophic level.
Ecological PyramidsEcological Pyramids
• Pyramid of Pyramid of energy flowenergy flow
• Ecological Ecological efficiencyefficiency– Range 5%-20%Range 5%-20%– Typically 10%Typically 10%
• Pyramid of Pyramid of biomassbiomass
• Pyramid of Pyramid of numbersnumbers
Fig. 4-20 p. 79
The figures represent number of individuals counted at each trophic level.
Ecological Pyramids of Numbers
• The total dry weight of organisms in a particular trophic level is referenced as “biomass”.
Ecological Pyramids of Biomass
BIOMASS=
# of organisms x
the weight of an average individual
Ecological Pyramids of EnergyEcological Pyramids of Energy
Energy in ecosystems flows from producers to consumers.
Energy is depicted in kilocalories.
Primary producers convert only about 1% of the energy in available sunlight.
The average amount of energy that is available to the next trophic level is about 10%.10%.
Primary Productivity of EcosystemsPrimary Productivity of Ecosystems
• Gross Primary ProductivityGross Primary Productivity (GPP) (GPP)– RateRate at which an ecosystem’s producers convert at which an ecosystem’s producers convert
solar energy into chemical energy as solar energy into chemical energy as biomassbiomass– kg/mkg/m22/year (kcal/m/year (kcal/m22/year)/year)
• Net Primary ProductivityNet Primary Productivity (NPP) (NPP)– Difference between the rate at which producers Difference between the rate at which producers
store energystore energy as biomass and the rate at which as biomass and the rate at which producers producers use chemical energyuse chemical energy stored as biomass stored as biomass
Connections: Matter Cycling in EcosystemsConnections: Matter Cycling in Ecosystems
• Biogeochemical (nutrient) cyclesBiogeochemical (nutrient) cycles– Hydrologic cycle (HHydrologic cycle (H22O)O)
– Atmospheric cycles (C,N)Atmospheric cycles (C,N)
– Sedimentary cycles (S,P)Sedimentary cycles (S,P)
Hydrologic (Water) CycleHydrologic (Water) Cycle
• Absolute humidityAbsolute humidity– The amount of The amount of
water vapor found water vapor found in a mass of air in a mass of air (g water/kg air)(g water/kg air)
• Relative humidityRelative humidity– The amount of The amount of
water vapor in a water vapor in a certain amount of certain amount of air, air, expressed as a expressed as a percentagepercentage the the maximum amount it maximum amount it could hold could hold at that at that temperaturetemperature
• Condensation nucleiCondensation nuclei– tiny particles on which tiny particles on which
droplets of water formdroplets of water form
• Dew pointDew point– Temperature at which Temperature at which
condensation occurscondensation occurs
Affects of Human Activity on the Water CycleAffects of Human Activity on the Water Cycle
• Withdrawing large quantities of water• Clearing vegetation
– Increased runoff– Reduced infiltration– Increased flooding– Soil erosion
• Modifying water quality– Adding nutrients– Other pollutants
The Carbon Cycle (Aquatic)The Carbon Cycle (Aquatic)
Fig. 4-28 p. 84-85Fig. 4-28 p. 84-85 http://www.mhhe.com/biosci/genbio/tlw3/eBridge/Chp29/animations/ch29/1_carbon_cycle.swf
Human Activities Affecting the Carbon CycleHuman Activities Affecting the Carbon Cycle
•Clearing treeClearing tree•Burning fossil fuels and woodBurning fossil fuels and wood
NO3 –
in soil
Nitrogen Fixation
by industry for agriculture
Fertilizers
Food Webs On Land
NH3, NH4+
in soil
1. Nitrification
bacteria convert NH4+ to
nitrite (NO2–)
loss by leaching
uptake by autotrophs
excretion, death,
decomposition
uptake by autotrophs
Nitrogen Fixationbacteria convert to ammonia
(NH3) ; this dissolves to form
ammonium (NH4+)
loss by leaching
Ammonificationbacteria, fungi convert the
residues to NH3 , this
dissolves to form NH4+
2. Nitrification
bacteria convert NO2- to
nitrate (NO3-)
Denitrificationby bacteria
Nitrogenous Wastes, Remains In Soil
Gaseous Nitrogen (N2)
In Atmosphere
NO2 –
in soil
Figure 4-29 Page 86Figure 4-29 Page 86The Nitrogen
Cycle
Human Activities Affecting The Nitrogen CycleHuman Activities Affecting The Nitrogen Cycle
• Burning fossil fuelsBurning fossil fuels– Acid rainAcid rain
• Animal wasteAnimal waste• Removing N from topsoilRemoving N from topsoil• Adding N to aquatic systemsAdding N to aquatic systems
Human Activities Affecting the Phosphorus CycleHuman Activities Affecting the Phosphorus Cycle
• MiningMining• Forest removalForest removal• Adding phosphorus to aquatic Adding phosphorus to aquatic
systemssystems– eutrophicationeutrophication
How Do Ecologists Learn About Ecosystems?How Do Ecologists Learn About Ecosystems?
Field researchField research
Remote sensingRemote sensing
Geographic information systems (GIS)Geographic information systems (GIS)
Laboratory researchLaboratory research
Systems analysisSystems analysis
GIS and Systems AnalysisGIS and Systems Analysis
Fig. 4-33 p. 91Fig. 4-33 p. 91Fig. 4-32 p. 91Fig. 4-32 p. 91
Ecosystem Servicesand SustainabilityEcosystem Servicesand Sustainability
Fig. 4-34 p. 92Fig. 4-34 p. 92
•Using renewable Using renewable solar energy as solar energy as an energy sourcean energy source
•Recycling the Recycling the chemical chemical nutrients nutrients organisms need organisms need for survival, for survival, growth, and growth, and reproduction.reproduction.