0
10
20
30
40
50
60
MM-CO
MECO
EECO
0 10-10
Age
(Ma) Panama
Acre
0 80RM genera
0 40RM fam.
100 300
III
II
I
0
taxa
0 10
FM spp.
0 60
PM spp.
III
II
I
PETM
Pozo System
Pebas System
TEE
periods ofintensi�ed Andean uplift
AlpineMontanePre−montaneLowlandMixed elevations
1 10 20 30 40 50 60 70 80 90 100 110 120 130 140 150 160
Molluscs
CrocodyliansMammalsPollenGlobal
temp. (°C)Geological
events
(a)
(b)
nMSP
GAAR
PlantsInsects MammalsAmphib.F R Birds
. . . . . . . . . . .. . . . . .
0 10 0 10
taxa f/p