Key to the checklistThe layout and acceptance of families, genera, and species is given alphabetically and follows that of the most recent species list (Germishuizen & Meyer 2003) published by the National Botanical Institute, Preto-ria.
• Accepted genus and species names are in bold type. THEMEDA Forssk. triandra Forssk.
• Names that are not accepted are in roman type and are followed by a cross-reference to the accepted name.
triandra Forssk. var. burchellii (Hack.) Stapf = T. triandra
• Synonyms are in italics. Not all synonyms for a species are listed. (T. triandra Forssk. var. burchellii (Hack.) Stapf)
• The names of the provinces indicate the distribution of species within Angola, as shown on the map below.
Bengo, Benguela, Bié, Cabinda, Cuando-Cubango, Cunene, Huambo, Huíla, Kuanza Norte, Kuanza Sul, Luanda, Lubango, Lunda Norte, Lunda Sul, Malanje, Moxico, Namibe, Uíge, Zaire.
• An asterisk (*) precedes the names of alien taxa.
A checklist of Angola grasses
Checklist das Poaceae de Angola
A checklist of Angola grasses
Checklist das Poaceae de Angola
Esperança Costa
Teresa Martins
Francisca Monteiro
2004
Southern African Botanical Diversity Network Report No. 28
Recommended citation formatCosta, E., Martins, T. & Monteiro, F. 2004. A checklist of Angola grasses—Checklist das Poaceae de Angola. Southern African Botanical Diversity Network Report No. 28. SABONET, Pretoria.
Produced byLuanda HerbariumUniversidade Agostinho NetoRua Fernando Pessoa No. 103Villa AliceCaixa Postal 3244Tel: (244) 2 336 168Fax: (244) 2 336 168
Published bySouthern African Botanical Diversity Network (SABONET)c/o National Botanical Institute, Private Bag X101, 0001, Pretoria
Printed in 2004 in the Republic of South Africa by Capture Press, Pretoria, (27) 12 349-1802
ISBN 1-919976-09-4
© 2004 SABONET. All rights reserved. No part of this publication may be reproduced or transmitted in any form or by any means without the permission of the copyright holder.
Editor-in-chief: Marthina MössmerScientific editor: Lyn FishSubeditor: Lidia GibsonText design and layout: Antoinette BurkhardtCover design: Rolf Burkhardt, Pretoria, South Africa (27) 83 635-1446.Front cover: Mangrove vegetation (top); Serra da Namphande, Huíla (bottom left); degraded vegetation in Namibe (bottom right).
SABONET website: www.sabonet.org
This report is a product of the Southern African Botanical Diversity Network (SABONET) and was made possible through support provided by the Global Environment Facility (GEF)/United Nations Development Programme (UNDP) and the United States Agency for International Development (USAID)/World Conservation Union-Regional Office for southern Africa (IUCN ROSA) (Plot no. 14818 Lebatlane Road, Gaborone West, Extension 6 Gaborone, Botswana), under the terms of Grant No. 690-0283-A-00-5950. The opinions expressed herein are those of the authors and do not necessarily reflect the views of USAID, the SABONET Steering Committee or SABONET National Working Groups.
Contents
Acknowledgements .....................................................................................................................................vi
Agradecimentos ...........................................................................................................................................vi
Introduction ................................................................................................................................................ 3
Key to the checklist ..................................................................................................................................... 4
Introdução .................................................................................................................................................. 6
Método de consulta do catálogo ................................................................................................................. 8
References/Bibliografia ................................................................................................................................ 9
Checklist ................................................................................................................................................... 10
Mangrove vegetation in Angola.
Acknowledgements
The present checklist was only made possible through the assistance of the staff of the Luanda Herbarium to whom we are greatly indebted.
We would also like to acknowledge the Dean’s office of the Faculty of Science of the University Agostinho Neto who granted us the means necessary for the internal compilation of the data, thus making the task possible.
Our gratitude also goes to Professor Patrick Phiri of the Lusaka Herbarium of the University of Zambia, who spent a week with us investigating Poaceae.
We would also like to express our thanks to the Main Researcher of the Botanical Centre of Lisbon (Centro de Botânica de Lisboa), Eurico Sampaio Martins, and Lyn Fish of the National Herbarium in Pretoria, South Africa, for correcting the manuscript.
The SABONET Programme is thanked for providing the financial resources necessary for the compilation of the checklist.
Our thanks also go to all those who, in a direct or indirect manner, took part in and made this work pos-sible.
Agradecimentos
A presente “checklist” foi apenas possível compilar com a ajuda de todo o pessoal do Herbário de Luanda aos quais estamos imensamente gratos.
Os nossos agradecimentos são extensivos à Direcção da Faculdade de Ciências que nos possibilitou e facilitou os meios necessários para a compilação interna dos dados.
Ao Professor Patrick Phiri do Herbário de Lusaka da Universidade da Zâmbia que connosco permaneceu uma semana a investigar sobre Poaceae, os nossos agradecimentos.
Ao Investigador Principal do Centro de Botânica de Lisboa Eurico Sampaio Martins os nossos agradecimentos pela correcção do manuscrito.
Ao Programa SABONET que providenciou os recursos financeiros para a sua elaboração, os nossos agradeci-mentos.
A todos aqueles que de uma forma directa ou indirecta participaram e tornaram possível a realização da pre-sente obra, os nossos agradecimentos.
Welwitschia mirabilis growing in Namibe.
1A checklist of Angola grasses
ANGOLA: Provincial Distribution
2 A checklist of Angola grasses
Serra de Namphande, Huíla.
Cissus uter, Namibe.
3A checklist of Angola grasses
Introduction
The Luanda Herbarium (Herbário de Luanda), known by the abbreviation LUAI, is an institution which keeps collections of dried plants which were collected in various parts of the country and of the world. These plants are prepared and kept in proper conditions with respect to light, temperature, humidity, and ventilation.
The first botanical specimens of Angola were collected by British medical doctors towards the end of the 18th century. Amongst the various scientific explorations carried out in the 18th and 19th centuries, we would like to highlight the participation of the botanists Friedrich Welwitsch, John Gossweiler, Joachim Monteiro, José Anchieta, Joaquim da Silva, Max Buchner, and a number of others.
Among the various present day local researchers who have collected extensively in various regions of the country in the areas of phytogeographic and ecological studies, we would like to highlight the names of Azancot de Menezes and Manuela Batalha in the taxonomy and identification of Angolan flora, with particular emphasis on plants that are traditionally used.
The Luanda Herbarium is part of the Lubango Herbarium which was transferred to Luanda and established in 1968 under the tutelage of the Faculty of Sciences of the University Agostinho Neto, and which is situated in the building of the National Centre for Scientific Research (in Portuguese, “Centro Nacional de Investigação Científica”, CNIC), in Avenida Revolução de Outubro (“Revolução de Outubro” Avenue). This Herbarium houses 35,000 botanical samples which are representative of approximately 5 000 species.Since 1995, this Herbarium also houses, on a temporary ba-sis, 45,000 botanical samples of the Institute for Agronomic Investigation (Instituto de Investigação Agronómica, IIA) of
the Ministry for Agriculture and Rural Development. These specimens belong to the Huambo Herbarium.
The main objectives of the Herbarium are the following:• The correct identification of the species according to the
rules and norms of the International Code of Botanical Nomenclature.
• Species conservation.• Contributing to the compilation of local and national
floras and checklists.• Mapping the country’s vegetation.• Making an inventory and a study of economically
important plants: medicinal, wild, edible, timber, or others which are potentially important for the sustain-able development of Angola.
Among the main areas of activity the following can be highlighted:• The creation of a database of Angola’s flora, with
special relevance given to endemic species.• The creation of pollen collections for the correct iden-
tification of species by means of pollen.• The study of economic botany and of the ecology of
botanical species.• The study of sensitive ecosystems and the study of
methods of regeneration of areas of degraded vegeta-tion.
The Herbarium maintains strong ties with Portuguese her-baria, mainly with the Botanical Centre of Lisbon (Centro de Botânica de Lisboa), and it is also linked to the Southern African Regional Network for the study of vegetation and for the identification of threatened plants aimed at compiling a national list of species threatened with extinction.
Euphorbia dinteri, Namibe.
4 A checklist of Angola grasses
General description of the Angolanvegetation
Angola is an African country situated in the southern hemi-sphere, and has a surface area of 1,246,700 km2. Angola’s coastline is 1,650 km long. Its geographical characteristics and position make Angola a country with some of the richest botanical diversity in sub-Saharan Africa. Its flora, however, is poorly known, as the studies done on the botanical diver-sity of the country are few (for example, Conspectus florae angolensis). From the existing bibliography we highlight the Carta Fitogeográfica by Gossweiler & Mendonça (1939), which was later expanded by Barbosa (1970).
According to Barbosa (1970), Angola’s vegetation may be subdivided into 32 types, which range from the tropical forests in Cabinda to the semiarid zones of the Namib. Other authors have also stood out for their important work, such as Monteiro (1970), who describes in detail the flora of Bié with special relevance given to the Miombo woodlands.
Based on statistical analysis, it has been estimated that there are 1,260 species endemic to Angola (Excell & Gonçalves 1973, Bamps 1975). Even though this estimation refers to the whole country, the North has been insufficiently studied.
With regard to Cabinda, the types that include perennially green forests are described as being the predominant vegeta-tion type. The most important species in Cabinda include Oxystigma oxyphyllum, Terminalia superba, Gilletioden-dron oogouense, Gossweilerodendron balsamiferum, and Entandrophragma. This type of vegetation is also found in regions of the Uíge Bengo and of the Kwanza Norte prov-inces where Celtis and Morus species occur frequently.
Savannas cover the greatest part of the territory and they are often described as various types of vegetation such as
mosaic with thickets and forests. In the savannas one finds tall growing grasses mainly connected to open forests.
In the savanna forest mosaics it is a common occurrence to find low growing grasses where one can find species of vari-ous genera, such as Hyparrhenia, Andropogon, Pennisetum, and Panicum. In these formations one may also find woody shrubs such as Hymenocardia acida, Erythrina abyssinica, Piliostigma thonningii, and Cussonia angolensis.
One particular type of vegetation consists of the mangrove communities. In Angola, mangroves reach their greatest development in the provinces of Cabinda and Zaire; as one moves South they lose their importance progressively, both in area as well as in size of the dominant species until they are reduced to small formations in the Benguela province and they then practically disappear in the desert-like areas of the South (Costa 1996). In these mangroves one finds an arboreal strata with zoning consisting of Rhizophora mangle, R. racemosa, R. harrisonii, Avicennia germinans, and Laguncularia racemosa. On the fringe of the mangrove we find an area consisting in its greatest part of Sesuvium portulacastrum and S. mesembrianthemoides.
Compiling the checklist
Although there are various studies pertaining to the Poaceae family, namely in the Flora zambeziaca and others which refer to Angola, we are not aware of any checklist pertain-ing to this family.
For this reason, the present compilation was based initially on local research and focuses on all the existing Angolan and Portuguese publications.
On the other hand, we also used Flora zambeziaca, Flora of Tropical West Africa, Flora of southern Africa, and some talks or articles on the subject.
Serra de Namphande, Huíla.
5A checklist of Angola grasses
Key to the checklist
The layout and acceptance of families, genera, and species is given alphabetically and follows that of the most recent spe-cies list (Germishuizen & Meyer 2003) published by the National Botanical Institute, Pretoria.
• Accepted genus and species names are in bold type. THEMEDA Forssk. triandra Forssk.
• Names that are not accepted are in roman type and are followed by a cross-reference to the accepted name. triandra Forssk. var. burchellii (Hack.) Stapf = T. triandra
• Synonyms are in italics. Not all synonyms for a species are listed. (T. triandra Forssk. var. burchellii (Hack.) Stapf)
• The names of the provinces indicate the distribution of species within Angola, as shown on the map on page 1. Bengo, Benguela, Bié, Cabinda, Cuando-Cubango, Cunene, Huambo, Huíla, Kuanza Norte, Kuanza
Sul, Luanda, Lubango, Lunda Norte, Lunda Sul, Malanje, Moxico, Namibe, Uíge, Zaire.
• An asterisk (*) precedes the names of alien taxa.
Phragmites mauritianus in Namibe.
6 A checklist of Angola grasses
IntroduçãoO Herbário de Luanda designado LUAI, é uma instituição que alberga colecções de plantas secas colhidas em várias partes do País e do Mundo. Estas plantas são preparadas e conservadas em condições adequadas de luminosidade, temperatura, humidade e ventilação.
Os primeiros espécimes botânicos em Angola, foram col-hidos por médicos Britânicos nos finais do século XVIII. Entre as diversas explorações científicas realizadas nos séculos I, XVIII e XIX, destacamos a participação dos botânicos Friedrich Welwitsch, John Gossweiler, Joachim Monteiro, José Anchieta, Joaquim da Silva, Max Buchner e tantos outros.
Dentre os vários investigadores contemporâneos nacionais que têm efectuado importantes colecções em várias regiões do País no âmbito de estudos fitogeográficos e ecológicos, destacam-se nomes como Azancot de Menezes e Manuela Batalha na taxonomia e identificação da flora de Angola, realçando-se a de uso tradicional.
O Herbário de Luanda (LUAI) é uma parte do Herbário do Lubango que foi transferido para Luanda e estabelecido em 1968 sendo tutelado pela Faculdade de Ciências da Universidade Agostinho Neto, funcionando no prédio do Centro Nacional de Investigação Científica (CNIC), situado na Avenida Revolução de Outubro. Este Herbário alberga 35,000 exemplares botânicos distribuídos por cerca de 5,000 espécies.
Encontram-se ainda temporariamente conservados neste Herbário desde 1995 45,000 exemplares botânicos do In-stituto de Investigação Agronómica (IIA) do Ministério da Agricultura e do Desenvolvimento Rural. Estes espécimes pertencem ao Herbário do Huambo.
O Herbário tem como principais objectivos:
• Identificação correcta das espécies de acordo as regras e normas do código internacional de nomenclatura botânica;
• Conservação das espécies;• Contribuição para a elaboração de floras locais, na-
cionais e “check-lists• Mapeamento da vegetação do país;• Inventariação e estudo das plantas económicamente im-
portantes quer medicinais, quer selvagens, comestívéis, madeiravéis, ou outras potencialmente importantes de modo a contribuir para o desenvolvimento sustentável de Angola.
De entre as principais áres de actividade salientam-se:• Criação de uma base de dados sobre a flora existente em
Angola com especial realce das espécies endémicas;• Criação de palinotecas para a correcta identificação
dos espécimes vegetais com base no polén;• Estudo da botânica económica e da ecologia das es-
pécies botânicas;• Estudo de ecossistemas sensíveis e estudo de métodos
de regeneração de áreas de vegetação degradada.
O Herbário que mantém uma forte ligação com os Herbários Portugueses, nomeadamente com o Centro de Botânica de Lisboa, está também ligado à Rede Regional da África Aus-tral de estudos da vegetação para a identificação de plantas ameaçadas com vista à elaboração de uma lista nacional de espécies ameaçadas de extinção.
Aspectos gerais sobre a vegetação emAngola
Angola é um país que se situa no hemisfério Sul do conti-nente Africano, com uma superfície de 1,246,700 km2. A fronteira marítima de Angola é de 1,650 km de costa. As características geográficas e a sua localização tornam-no um
Euphorbia dinteri, Namibe.
7A checklist of Angola grasses
país com uma das mais ricas diversidade vegetal na África subshariana, porém, insuficientemente conhecida pois pou-cos são os estudos efectuados sobre a diversidade botânica do país (por exemplo, Conspectus Florae Angonlense). Da bibliografia existente destaca-se a Carta Fitogeográfica de Gossweiler & Mendonça (1939), melhorada mais tarde por Barbosa (1970).
Segundo Barbosa (1970), cerca de 32 tipos de vegetação podem ser considerados em Angola, desde as florestas húmidas de Cabinda até as zonas semiáridas do Namibe. Entretanto outros autores foram se destacando em obras relevantes como sendo Monteiro (1970) que descreve ao pormenor a vegetação do Bié com relevancia na vegetação do tipo Miombo.
Baseado em análises estatísticas, foi estimado que 1,260 espécies endémicas ocorrem em Angola (Excell & Gonçalves 1973, Bamps 1975). Embora os tipos de vegetação incluam todo país, a parte Norte está mal estudada.
Em Cabinda são descritos os tipos que incluem florestas sem-pre verdes como tipo fisionómico predominante. As espécies mais importantes em Cabinda incluem Oxystigma oxy-phyllum, Terminalia superba, Gilletiodendron oogouense, Gossweilerodendron balsamiferum e Entandrophragma.Também este tipo de vegetação é encontrado em regiões da província do Uíge Bengo e Kuanza-Norte onde as espécies de Celtis sp. e Morus sp. são frequentes.
As savanas cobrem a maior parte do território muitas vezes descritas em vários tipos de vegetação como mosaico com bosques e florestas. Nassavanas é frequente a presença de gramíneas de altura elevada maioritariamente ligadas a florestas abertas.
Em moisacos florestas savanas é frequente as gramínias de baixo porte onde se encontram espécies de vários géneros como: Hyparrhenia, Andropogon, Pennisetum e Panicum.Também é notório nestas formações a presença de arbustos lenhosos como Hymenocardia acida, Erythrina abyssinica, Piliostigma thonningii e Cussonia angolensis.
Um tipo de vegetação particular são as comunidades de mangal. Em Angola os mangais atingem o seu maoir desen-volvimento nas províncias de Cabinda, Zaire, e a medida que caminham para o Sul do país, vão perdendo progressi-vamente importância tanto em áera ocupada como em porte das dominantes, até se esbaterem em pequenas formações na província de Benguela e desaparecerem praticamente nas áreas desérticas do sul (Costa 1996). Nestes mangais, destacam-se principalmente um estrato árboreo com zon-agem constituído por Rhizophora mangle, R. racemosa, R. harrisonii, Avicennia germinans e Laguncularia racemosa. Constituindo a orla do mangal encontramos um estrato constituído na sua maioria por Sesuvium portulacastrum e S. mesembriamthemoides.
A checklist
Embora haja alguns estudos referentes à família Poaceae, nomeadamente na Flora Zambezíaca e outros que referem Angola, na verdade não é conhecida nenhuma checklist sobre esta família.
Assim, a presente compilação baseou-se inicialmente numa pesquisa e concentração local de tudo o que existe em Angola em publicações feitas em Angola ou Portugal.
Por outro lado, serviu-nos ainda por base a pesquisa bibli-ográfica em Flora zambesíaca, Flora of West Tropical Africa, Flora of southern Africa e algumas comunicações ou artigos sobre a matéria.
Serra de Namphande, Huíla.
8 A checklist of Angola grasses
Método de consulta do catálogoEste catálogo é uma lista provisória visto não incluir as muitas incertae que se encontram nos herbários. O layout e a aceitação de famílias, géneros, e espécies é apresentada por ordem alfabética e segue o modelo da última lista de espécies (Germishuizen & Meyer 2003) publicada pelo National Botanical Institute, em Pretória.
• Os nomes de géneros e espécies aceites aparecem em tipo forte. THEMEDA Forssk. triandra Forssk.
• Nomes não aceites aparecem em tipo Romano, seguidos por uma remissiva ao nome aceito. triandra Forssk. var. burchellii (Hack.) Stapf = T. triandra
• Os sinónimos são grifados. A lista não inclui todos os sinónimos para uma determinada espécie. (T. triandra Forssk. var. burchellii (Hack.) Stapf)
• Os nomes das províncias indicam a distribuição das espécies dentro de Angola. O mapa apresenta as divisões geográ-ficas. Bengo, Benguela, Bié, Cabinda, Cuando-Cubango, Cunene, Huambo, Huíla, Kuanza Norte, Kuanza
Sul, Luanda, Lubango, Lunda Norte, Lunda Sul, Malanje, Moxico, Namibe, Uíge, Zaire.
• Um asterisco (*) precede os nomes de géneros e espécies estrangeiros à região.
Phragmites mauritianus, Namibe.
9A checklist of Angola grasses
References BibliografiaBARBOSA, L.A.B. 1970. Carta fitogeográfica de Angola. Instituto
de Investigação Científica de Angola.
BRUMMITT, R.K. & POWELL, C. E. (eds). 1992. Authors of plant
names. Royal Botanic Gardens, Kew.
CHIPPENDALL, L.K.A. 1955. A guide to the identification of
grasses in South Africa. In D. Meredith (ed.), The grasses
and pastures of South Africa, Part 1. Central News Agency,
Johannesburg.
CLAYTON, W.D. 1970. In E. Milne-Redhead & R. M. Polhill
(eds), Flora of Tropical East Africa. Gramineae (Part 1). Crown
Agents, London.
CLAYTON, W.D. 1972. Gramineae. In F. N. Hepper, Flora of West
Tropical Africa, Vol. 3.2. Crown Agents. London.
CLAYTON, W.D., PHILLIPS, S.M. & RENVOIZE, S.A. 1974.
In: R.M. Polhill (ed.), Flora of Tropical East Africa, Graminae
(Part 2). Crown Agents. London.
CLAYTON, W.D. & RENVOIZE, S.A. 1982. In: R.M. Polhill
(ed.), Flora of Tropical East Africa, Graminae (Part 3). A.A.
Balkema, Rotterdam.
COSTA, E.M.E.F. 1996. Taxonomia de géneros da tribo Genisteae.
Genista, Echinospartum em Portugal e Adenocarpus e Agryo-
lobium em Angola. Dissertação de Doutoramento. Inst. Sup.
Agron., Univ. Técn. Lisboa.
DYER, A.R., 1976. The genera of southern African flowering
plants:Vol. 2. Gymnosperms and Monocotyledons. Depart-
ment of Agricultural Technical Services, Botanical Research
Institute, Pretoria.
EXCELL, A.W. & GONÇALVES, M.L. 1973. A statistical analy-
sis of a sample of flora of Angola. Garcia de Orta, Sér. Bot.
1 (1-2).
FERNANDES, A., LAUNERT, E. & WILD, H. (eds). 1971. Flora
zambesiaca, Vol. 10: Part 1. Royal Botanic Gardens, Kew,
London.
GIBBS RUSSELL, G.E., WATSON, L., KOEKMOER, M., SMOOK,
L., BARKER, N.P., ANDERSON, H.M. & DALLWITZ, M.J.
1990. Grasses of southern Africa. Memoirs of the Botanical
Survey of South Africa no. 58.
GOSSWEILER, J. & MENDONÇA, F.A. 1939. Carta Fitogeográfica
de Angola. Memória descritiva dos principais tipos de vegetação
da Colónia determinados pelos seus aspectos fisiográficos e
caracteres ecológicos segundo a nomenclatura de Ruben. Edição
do Governo-General de Angola.
JACOT GUILLARMOD, A. 1971. Flora of Lesotho. Cramer,
Germany.
LAUNERT, E. & POPE, G.V. (eds). 1989. Flora zambesiaca, Vol.
10: Part 3. Royal Botanic Gardens, Kew, London.
LEISTNER, A. (ed.). 2000. Seed plants of southern Africa: families
and genera. Strelitzia 10. National Botanical Institute, Preto-
ria.
MÜLLER, M.A.N. 1983. Grasses of South West Africa/Namibia.
Directorate of Agriculture and Forestry, Windhoek.
POPE, G.V. (ed.). 1999. Flora zambesiaca, Vol. 10: Part 2. Royal
Botanic Gardens, Kew, London.
POPE, G.V. & MARTINS, E.S. (eds). 2002. Flora zambesiaca Vol.
10: Part 4. Royal Botanic Gardens, Kew, London.
SCHMITZ, M. 1984. An illustrated key for the identification of the
grasses of Lesotho. National University of Lesotho, Roma.
VAN OUDTSHOORN, F. 1999. Guide to grasses of southern
Africa. Briza Publications, Pretoria.
ZIZKA, G. 1988. Revision der Melinidae Hitchcock (Poaceae).
Bibliotheca Botanica 138.
10 A checklist of Angola grasses
ACRACHNE ARISTIDA
ChecklistACRACHNE Wight & Arn. ex Chiov. racemosa (B.Heyne ex Roem. & Schult.) Ohwi Distrib.: Huíla; Namibe
(A. verticillata (Roxb.) Chiov.)
(Eleusine verticillata Roxb.)
ACROCERAS Stapf gabunense (Hack.) Clayton Distrib.: Kuanza Norte
macrum Stapf Distrib.: Cunene; Huíla
AGROSTIS L. huttoniae (Hack.) C.E.Hubb. = A. lachnantha lachnantha Nees Distrib.: Huíla; Luanda
(A. huttoniae (Hack.) C.E.Hubb.)
semiverticillata (Forssk.) C.Chr. = Polypogon viridis
ALLOEOCHAETE C.E.Hubb. andongensis (Rendle) C.E.Hubb. Distrib.: Malanje
(Danthonia andogensis Rendle)
ALLOTEROPSIS J.Presl angusta Stapf Distrib.: Benguela
paniculata (Benth.) Stapf Distrib.: Moxico
semialata (R.Br.) Hitchc. subsp. eckloniana (Nees) Gibbs Russ.
Distrib.: Huíla
semialata (R.Br.) Hitchc. subsp. semialata Distrib.: Cunene; Huíla; Malanje; Uíge
ANDROPOGON L. abyssinicus Fresen. Distrib.: Cunene; Moxico
africanus Franch. Distrib.: Moxico
(A. prolixus Stapf)
amplectens Nees var. catangensis Chiov. = Dihetero-pogon amplectens var. catangensis
brazzae Franch. Distrib.: Huíla; Moxico
chinensis (Nees) Merr. Distrib.: Malanje
densiflorus (Steud.) Stapf = Cymbopogon densiflorus eucomus Nees Distrib.: Cuando-Cubango; Huambo; Huíla; Lunda Sul;
Namibe
fastigiatus Sw. Distrib.: Huíla
(Diectomis fastigiata (Sw.) Kunth)
festuciformis Rendle Distrib.: Moxico
gabonensis Stapf Distrib.: Zaire
gayanus Kunth Distrib.: Huíla; Luanda; Lunda Sul; Moxico
grandiflorus Hack. = Diheteropogon filifolius huillensis Rendle Distrib.: Huíla; Moxico
lacunosus J.G.Anderson Distrib.: Huíla
laxatus Stapf Distrib.: Huíla
lopollensis Rendle = Schizachyrium lopollense nyassae Rendle = Hyparrhenia nyassae perligulatus Stapf. Distrib.: Cunene; Moxico
prolixus Stapf = A. africanus roseus Napper = A. textilis schirensis Hochst. ex A.Rich. Distrib.: Cunene; Huíla
textilis Welw. ex Rendle Distrib.: Moxico
(A. roseus Napper)
ANTHEPHORA Schreb. *cristata (Döll) Hack. ex De Wild. & T.Durand Distrib.: Cabinda; Luanda
elongata De Wild. Distrib.: Cuando-Cubango; Huíla
(A. burttii Stapf & C.E.Hubb.)
pubescens Nees Distrib.: Cuando-Cubango; Cunene; Huíla; Namibe
(A. ramosa Gooss.)
schinzii Hack. Distrib.: Cunene; Huambo; Huíla
APOCHAETE (C.E.Hubb.) J.B.Phipps hispida (L.f.) J.B.Phipps = Tristachya leucothrix
ARISTIDA L. adscensionis L. Distrib.: Cunene; Huíla; Namibe
(A. submucronata Schumach.)
alopecuroides Hack. = A. congesta subsp. congesta angolensis C.E.Hubb. = Sartidia angolensis bipartita (Nees) Trin. & Rupr. Distrib.: Huíla
congesta Roem. & Schult. subsp. congesta Distrib.: Cunene; Huíla; Cuando-Cubango
(A. alopecuroides Hack.)
contractinodis Stent & J.M.Rattray = A. junciformis subsp. macilenta
dewinteri Giess Possible distribution: Cunene; Namibe
dinteri Hack. = Stipagrostis dinteri gossweileri Pilg. = A. recta graciliflora Pilg. = A. stipitata subsp. graciliflora hordeacea Kunth Distrib.: Cuando-Cubango; Cunene; Huíla; Moxico;
Namibe
junciformis Trin. & Rupr. subsp. macilenta (Henrard)
11A checklist of Angola grasses
ARUNDINELLA BRACHYACHNE
Melderis Distrib.: Moxico
(A. contractinodis Stent & J.M.Rattray)
junciformis Trin. & Rupr. subsp. welwitschii (Rendle) Melderis
Distrib.: Cuando-Cubango; Cunene; Huíla; Moxico
leucophaea Henrard Distrib.: Moxico
meridionalis (Stapf) Henrard Distrib.: Cuando-Cubango; Cunene; Huíla; Moxico;
Namibe
pilgeri Henrard Distrib.: Cuando-Cubango
recta Franch. Distrib.: Malanje; Moxico
(A. gossweileri Pilg.)
rhiniochloa Hochst. Distrib.: Cuando-Cubango; Cunene; Huíla; Moxico;
Namibe
scabrivalvis Hack. subsp. scabrivalvis Distrib.: Cuando-Cubango
stipitata Hack. subsp. graciliflora (Pilg.) Melderis Distrib.: Huíla
(A. graciliflora Pilg.)
stipitata Hack. subsp. robusta (Stent & J.M.Rattray) Melderis
Distrib.: Moxico
stipitata Hack. subsp. stipitata Distrib.: Cuando-Cubango; Moxico
stipoides Lam. Distrib.: Cuando-Cubango; Huíla; Moxico
subacaulis (Nees) De Winter = Stipagrostis subacaulis submucronata Schumach. = A. adscensionis L.
ARUNDINELLA Raddi nepalensis Trin. Distrib.: Huíla
AXONOPUS P.Beauv. flexuosus (Peter) C.E.Hubb. ex Troupin Distrib.: Moxico
BAMBUSA Schreb. abyssinica A.Rich. = Oxytenanthera abyssinica
BECKEROPSIS K.Schum. uniseta (Nees) K.Schum. = Pennisetum unisetum
BEWSIA Gooss. biflora (Hack.) Gooss. Distrib.: Cuando-Cubango; Moxico
(Diplachne bifora Hack.)
BOTHRIOCHLOA Kuntze bladhii (Retz.) S.T.Blake Distrib.: Cunene; Huíla; Moxico
(B. glabra (Roxb.) A.Camus)
glabra (Roxb.) A.Camus = B. bladhii insculpta (Hochst. ex A.Rich.) A.Camus Distrib.: Benguela; Cunene; Huíla
radicans (Lehm.) A.Camus
Distrib.: Cunene; Namibe
BRACHIARIA (Trin.) Griseb. andongensis (Rendle) Stapf Distrib.: Malanje
arrecta (T.Durand & Schinz) Stent Distrib.: Benguela; Huíla
(B. latifolia Stapf)
bovonei (Chiov.) Robyns Distrib.: Malanje; Moxico
brachylopha Stapf = B. serrata brevispicata (Rendle) Stapf Distrib.: Moxico
brizantha (A.Rich.) Stapf Distrib.: Benguela; Bié; Cuando-Cubango; Cunene;
Huambo; Huíla; Malanje; Uíge;
clavipila (Chiov.) Robyns Distrib.: Moxico
deflexa (Schumach.) C.E.Hubb. ex Robyns Distrib.: Benguela; Huíla; Luanda; Namibe
(Pseudobrachiaria deflexa (Schumach.) Launert)
dura Stapf var. dura Distrib.: Moxico
eruciformis (J.E.Smith) Griseb. Distrib.: Cunene; Huíla
(B. isachne (Roem. & Schult.) Stapf)
filifolia Stapf = B. subulifolia glomerata (Stapf) A.Camus Distrib.: Cuando-Cubango
grossa Stapf Distrib.: Moxico
humidicola (Rendle) Schweick. Distrib.: Cunene; Huíla
(Panicum humidicola Rendle)
jubata (Fig. & De Not.) Stapf Distrib.: Huambo; Malanje
latifolia Stapf = B. arrecta malacodes (Mez & K.Schum.) Scholz Distrib.: Cunene; Huíla; Namibe
(B. poaeoides Stapf)
melanotyla (Hack.) Henrard = B. nigropedata mutica (Forssk.) Stapf nigropedata (Ficalho & Hiern) Stapf Distrib.: Cunene; Huíla; Moxico; Namibe
(B. melanotyla (Hack.) Henrard)
poaeoides Stapf = B. malacodes psammophila (Welw. ex Rendle) Launert Distrib.: Cuando-Cubango; Namibe
sadinii Vanderyst = Panicum sadinii serrata (Thunb.) Stapf Distrib.: Cuando-Cubango; Cunene; Huíla; Moxico
(B. brachylopha Stapf)
subulifolia (Mez) Clayton Distrib.: Huíla; Malanje
(B. filifolia Stapf)
xantholeuca (Schinz) Stapf Distrib.: Cunene; Moxico
(Panicum xantholeucum Schinz)
BRACHYACHNE (Benth.) Stapf fulva Stapf
12 A checklist of Angola grasses
BRACHYACHNE DACTYLOCTENIUM
BRACHYACHNE (Benth.) Stapf (cont.) Distrib.: Moxico
obtusiflora (Benth.) C.E.Hubb. Distrib.: Moxico
patentiflora (Stent & J.M.Rattray) C.E.Hubb. Distrib.: Moxico
BRACHYPODIUM P.Beauv. flexum Nees Distrib.: Cabinda
BROMUNIOLA Stapf & C.E.Hubb. gossweileri Stapf & C.E.Hubb. Distrib.: Moxico
CENCHRUS L. *biflorus Roxb. Distrib.: Cuando-Cubango; Cunene; Moxico
ciliaris L. Distrib.: Bengo; Cuando-Cubango; Cunene; Huambo;
Huíla; Luanda; Namibe
CENTROPODIA Reichenb. mossamedensis (Rendle) Cope Distrib.: Namibe
(Danthonia mossamedensis Rendle)
CHASMOPODIUM Stapf caudatum (Hack.) Stapf Distrib.: Malanje
CHLORIDION Stapf cameronii Stapf = Stereochlaena cameronii
CHLORIS Sw. flaballata (Hack.) Launert Distrib.: Benguela; Kuanza Sul; Luanda; Namibe
(C. pubescens Wawra & Peyr.)
gayana Kunth Distrib.: Huíla
myriostachya Hochst. = C. roxburghiana notocoma Hochst. = C. virgata pilosa Schumach. Distrib.: Namibe
pubescens Wawra & Peyr. = C. flabellata pycnothrix Trin. Distrib.: Bengo; Huíla
roxburghiana Schult. Distrib.: Bengo; Huíla
(C. myriostachya Hochst.)
virgata Sw. Distrib.: Huíla; Luanda
(C. notocoma Hochst.)
CHRYSOPOGON Trin. nigritanus ( Benth.) Veldkamp Distrib.: Huíla
(Vetiveria nigritana (Benth.) Stapf)
(Vetiveria zizanioides (L.) Nash var. nigritana (Benth.)
A.Camus)
CLADORAPHIS Franch. cyperoides (Thunb.) S.M.Phillips Distrib.: Cunene; Namibe
(Eragrostis cyperoides (Thunb.) Beauv.)
COELORACHIS Brongn. afraurita (Stapf) Stapf Distrib.: Huíla
COIX L. *lacryma-jobi L. Distrib.: Uíge
CORTADERIA Stapf *selloana (Schult. & Schult. f.) Asch. & Graebn. Distrib.: Huambo
CRASPEDORHACHIS Benth. rhodesiana Rendle Distrib.: Cuando-Cubango; Cunene; Moxico
sarmentosa (Hack.) Pilg. = Willkommia sarmentosa
CTENIUM Panz. concinnum Nees Distrib.: Moxico
newtonii Hack. Distrib.: Moxico
somalense (Chiov.) Chiov. Distrib.: Moxico
CYMBOPOGON Spreng. caesius (Hook. & Arn.) Stapf Distrib.: Huíla
(C. excavatus (Hochst.) Stapf ex Burtt Davy)
(C. giganteus Chiov.)
*citratus (DC.) Stapf Distrib.: Huíla
densiflorus (Steud.) Stapf Distrib.: Huíla; Moxico
(Andropogon densiflorus Steud.)
excavatus (Hochst.) Stapf ex Burtt Davy = C. caesius giganteus Chiov. = C. caesius nardus (L.) Rendle Distrib.: Huambo; Huíla
(C. validus (Stapf) Stapf ex Burtt Davy)
validus (Stapf) Stapf ex Burtt Davy = C. nardus
CYNODON Rich. bradleyi Stent = C. incompletus dactylon (L.) Pers. Distrib.: Bié; Cuando-Cubango; Cunene; Huíla; Luanda;
Moxico
*incomplectus Nees Distrib.: Bié; Huíla
(C. bradleyi Stent)
nlemfuensis Vanderyst var. nlemfuensis Distrib.: Moxico
DACTYLOCTENIUM Willd. aegyptium (L.) Willd. Distrib.: Cuando-Cubango; Cunene; Huíla; Luanda;
13A checklist of Angola grasses
DANTHONIA ECHINOCHLOA
Moxico; Namibe
(D. mpuetensis De Wild.)
giganteum Fisher & Schweick. Distrib.: Cuando-Cubango; Cunene; Huíla; Moxico
mpuetensis De Wild. = D. aegyptium
DANTHONIA DC. andongensis Rendle = Alloeochaete andongensis mossamedensis Rendle = Centropodia mossamedensis
DANTHONIOPSIS Stapf dinteri (Pilg.) C.E.Hubb. Distrib.: Cunene
lignosa C.E.Hubb. Distrib.: Cunene; Huíla
intermedia C.E.Hubb. = D. viridis viridis (Rendle) C.E.Hubb. Distrib.: Moxico
(D. intermedia C.E.Hubb.)
DIANDROCHLOA De Winter namaquensis (Nees ex Schrad.) De Winter = Eragrostis
japonica pusilla (Hack.) De Winter = Eragrostis pusilla
DICHANTHIUM Willemet annulatum (Forssk.) Stapf var. papillosum (Hochst. ex
A.Rich.) de Wet & Harlan Distrib.: Cunene
DIECTOMIS P.Beauv. fastigiata (Sw.) Kunth = Andropogon fastigiatus
DIGITARIA Haller angolensis Rendle Distrib.: Huíla; Malanje; Moxico
(D. yokoensis Vanderyst)
adscendens (Kunth) Henrard = D. ciliaris brazzae (Franch.) Stapf Distrib.: Cuando-Cubango; Moxico
ciliaris (Retz.) Koeler Distrib.: Moxico
(D. adscendens (Kunth) Henrard)
comifera Pilg. Distrib.: Moxico
debilis (Desf.) Willd. Distrib.: Cuando-Cubango; Moxico
diagonalis (Nees) Stapf Distrib.: Moxico
eriantha Steud. Distrib.: Moxico
(D. livida Henrard)
eylesii C.E.Hubb. Distrib.: Moxico
friesii Pilg. = D. longiflora gayana (Kunth) Stapf ex A.Chev. Distrib.: Moxico
gazensis Rendle Distrib.: Cuando-Cubango; Huíla; Moxico
livida Henrard = D. eriantha Steud.
longiflora (Retz) Pers. Distrib.: unknown
maniculata Stapf Distrib.: Moxico
milanjiana (Rendle) Stapf Distrib.: Moxico
monodactyla (Nees) Stapf Distrib.: Cuando-Cubango; Cunene; Moxico
nitens Rendle Distrib.: Moxico
pearsonii Stapf Distrib.: Kuanza Norte
perrottetii (Kunth) Stapf Distrib.: Huíla; Moxico
phaeotricha (Chiov.) Robyns Distrib.: Moxico
redheadii (C.E.Hubb.) Clayton Distrib.: Moxico
seriata Stapf Distrib.: Cuando-Cubango
setifolia Stapf Distrib.: Huíla
ternata (A.Rich.) Stapf Distrib.: Moxico
velutina (Forssk.) P.Beauv. Distrib.: Huíla; Huambo
ventriosa Van der Veken Distrib.: Moxico
yokoensis Vanderyst = D. angolensis
DIHETEROPOGON Stapf amplectens (Nees) Clayton var. catangensis (Chiov.)
Clayton Distrib.: Cuando-Cubango; Huíla; Moxico
(Andropogon amplectens Nees var. catangensis Chiov.)
filifolius (Nees) Clayton Distrib.: Huíla; Moxico
(D. grandiflorus (Hack.) Stapf)
grandiflorus (Hack.) Stapf = D. filifolius
DINEBRA Jacq. guineensis Franch. = Heteranthoecia guineensis retroflexa (Vahl) Panz. var. condensata S.M.Phillips Distrib.: Huíla; Luanda
DIPLACHNE P.Beauv. biflora Hack. = Bewsia biflora fusca (L.) P.Beauv. ex Roem. & Schult. = Leptochloa
fusca gigantea Launert = Leptochloa gigantea
ECHINOCHLOA P.Beauv. colona (L.) Link. Distrib.: Cuando-Cubango; Cunene; Namibe
crus-pavonis (Kunth) Schult. Distrib.: Huíla; Malanje
haploclada (Stapf) Stapf Distrib.: Moxico
jubata Stapf Distrib.: Cuando-Cubango; Moxico
14 A checklist of Angola grasses
ECHINOCHLOA ERAGROSTIS
ECHINOCHLOA P.Beauv. (cont.) polystachya Kunth = Pseudechinolaena polystachya pyramidalis (Lam.) Hitchc. & Chase Distrib.: Cuando-Cubango; Cunene; Huíla; Moxico
stagnina (Retz.) P.Beauv. Distrib.: Cunene; Huíla
ELEUSINE Gaertn. africana Kenn.-O’Byrne Distrib.: Cunene; Malanje; Moxico
*coracana (L.) Gaertn. Distrib.: Huambo; Huíla; Moxico
indica (L.) Gaertn. Distrib.: Cunene; Malanje; Moxico
verticillata Roxb. = Acrachne racemosa
ELIONURUS Willd. argenteus Nees = E. muticus muticus (Spreng.) Kuntze Distrib.: Cunene; Huíla; Malanje
(E. argenteus Nees)
tripsacoides Willd. Distrib.: Cuando-Cubango
ELYMANDRIA Stapf androphilla (Stapf) Stapf Distrib.: Huambo
grallata (Stapf) Clayton Distrib.: Benguela; Bié
(Hyparrhenia grallata Stapf)
gossweileri (Stapf) Clayton Distrib.: Zaire
ELYTROPHORUS P.Beauv. globularis Hack. Distrib.: Cunene; Huíla; Moxico
ENNEAPOGON P.Beauv. cenchroides (Licht. ex Roem. & Schult.) C.E.Hubb. Distrib.: Cuando-Cubango; Cunene; Namibe
desvauxii P.Beauv. Distrib.: Cuando-Cubango
scaber Lehm. Distrib.: Namibe
scoparius Stapf Distrib.: Namibe
ENTEROPOGON Nees macrostachyus (Hochst. ex A.Rich.) Munro ex Benth. Distrib.: Huíla; Luanda; Moxico; Namibe
ENTOLASIA Stapf imbricata Stapf
ENTOPLOCAMIA Stapf aristulata (Hack. & Rendle) Stapf Distrib.: Benguela; Cunene; Huíla
(Tetrachne aristulata Hack. & Rendle)
ERAGROSTIS Wolf acamptoclado Cope
Distrib.: Benguela
aethiopica Chiov. Distrib.: Huíla
amabilis (L.) Hook. & Arn. (E. tenella (L.) P.Beauv. ex Roem. & Schult.)
andongensis Rendle = E. nindensis angolensis Rendle Distrib.: Huíla
angusta Hack. = E. rotifer annulata Rendle ex Scott-Elliot Distrib.: Cunene; Huíla; Namibe
arenicola C.E.Hubb. Distrib.: Moxico
aspera (Jacq.) Nees Distrib.: Cuando-Cubango; Cunene; Huíla; Moxico;
Namibe
atrovirens (Desf.) Steud. Distrib.: Cunene; Moxico
aurorae Launert Distrib.: Bié
blepharolepis Hack. = E. hispida bicolor Nees Distrib.: Huíla
biflora Hack. ex Schinz Distrib.: Huíla
brainii (Stent) Launert Distrib.: Moxico
buchananii K.Schum. = E. nindensis caniflora Rendle Distrib.: Huíla
capensis (Thunb.) Trin. Distrib.: Huambo; Huíla; Luanda; Malanje
chalcantha Trin. = E. racemosa chapelieri (Kunth) Nees Distrib.: Huíla; Kuanza Norte
cilianensis (All.) Vignolo ex Janch. Distrib.: Cuando-Cubango; Moxico
ciliaris (L.) R.Br. Distrib.: Luanda
cimicina Launert Distrib.: Cuando-Cubango
curvula (Schrad.) Nees Distrib.: Cunene; Huíla; Namibe
(E. robusta Stent)
cylindriflora Hochst. Distrib.: Cuando-Cubango; Huíla; Kuanza Norte;
Namibe
(E. horizontalis Peter)
cyperoides (Thunb.) P.Beauv. = Cladoraphis cyperoides dinteri Stapf Distrib.: Benguela; Cuando-Cubango; Cunene; Huíla;
Namibe
exasperata Peter Distrib.: Huíla; Kuanza Norte
fascicularis Trin. = E. prolifera flavicans Rendle Distrib.: Huíla; Kuanza Norte; Moxico
friesii Pilg. Distrib.: Said to occur
glandulosipedata De Winter Distrib.: Huíla
15A checklist of Angola grasses
habrantha Rendle Distrib.: Huíla
henrardii Jansen = E. trichophora heteromera Stapf Distrib.: Huíla; Kuanza Norte
hierniana Rendle Distrib.: Cunene
(E. uniglumis Hack.)
hispida K.Schum. Distrib.: Huíla; Kuanza Norte; Lubango; Malanje
(E. blepharolepis Hack.)
homomalla Nees Distrib.: Huíla
(E. hygrophila C.E.Hubb. & Schweick.)
horizontalis Peter = E. cylindriflora. huillensis Rendle Distrib.: Huíla
hygrophila C.E.Hubb. & Schweick. = E. homomalla inamoena K.Schum. Distrib.: Cuando-Cubango
japonica (Thunb.) Trin. Distrib.: Huíla; Moxico; Namibe
(Diandrochloa namaquensis (Nees ex Schrad.) De
Winter)
(E. namaquensis Nees)
lappula Nees Distrib.: Cuando-Cubango; Cunene; Huíla
lehmanniana Nees var. lehmanniana Distrib.: Huíla; Kuanza Norte
lepidobasis Cope Distrib.: Known only from Mwinilunga District, Zambia,
but may be expected to occur in Angola. Flora zambesiaca
10,2: 119 (1999)
macilenta (A.Rich.) Steud. Distrib.: Huíla
margaritacea Stapf = E. rotifer membranacea Hack. ex Schinz Distrib.: Cunene; Huíla; Kuanza Norte
micrantha Hack. Distrib.: Cunene
microsperma Rendle Distrib.: Huíla; Malanje
milnei Launert ex Cope Distrib.: Known only from Mwinilunga District, Zambia,
but may be expected to occur in Angola. Flora zambesiaca
10,2: 61–62 (1999)
namaquensis Nees = E. japonica nindensis Ficalho & Hiern Distrib.: Cunene; Kuanza Norte; Malanje; Namibe
(E. andongensis Rendle)
pallens Hack. Distrib.: Cuando-Cubango
patens Oliv. Distrib.: Huíla; Malanje; Namibe
patentipilosa Hack. Distrib.: Huíla
(E. pseudosclerantha Chiov.)
pilgeriana Dinter ex Pilg. Distrib.: Namibe
pilosa (L.) P.Beauv. Distrib.: Cunene; Huíla; Namibe
plana Nees Distrib.: Huíla
plurigluma C.E.Hubb. Distrib.: Lunda Sul
porosa Nees Distrib.: Cunene; Huíla; Moxico; Namibe
prolifera (Sw.) Steud. Distrib.: Bengo; Luanda; Namibe
(E. fascicularis Trin.)
pseudosclerantha Chiov. = E. patentipilosa pusilla Hack. Distrib.: Huíla; Kuanza Norte
(Diandrochloa pusilla (Hack.) De Winter)
racemosa (Thunb.) Steud. Distrib.: Bengo; Cuando-Cubango; Cunene; Huíla;
Kuanza Norte; Malanje; Moxico
(E. chalcantha Trin.)
rejuvenescens Rendle Distrib.: Bié; Huíla
rigidior Pilg. Distrib.: Cuando-Cubango; Cunene; Huíla
robusta Stent = E. curvula rogersii C.E.Hubb. Distrib.: Namibe
rotifer Rendle Distrib.: Cuando-Cubango; Huíla; Malanje; Namibe
(E. angusta Hack.)
(E. margaritacea Stapf)
sarmentosa (Thunb.) Trin. Distrib.: Cuando-Cubango
sclerantha Nees subsp. sclerantha Distrib.: Huíla
sclerantha Nees subsp. villosipes (Jedwabn.) Launert Distrib.: Huíla; Kuanza Norte
stapfii De Winter Distrib.: Cunene
superba Peyr. Distrib.: Bengo; Benguela; Cuando-Cubango; Cunene;
Huíla; Luanda
tenella (L.) P.Beauv. ex Roem. & Schult. = E. amabilis tenuifolia (A.Rich.) Hochst. ex Steud. Distrib.: Benguela; Huíla; Uíge
thollonii Franch. Distrib.: Bié
tremula Hochst. ex Steud. Distrib.: Malanje
trichophora Coss. & Durieu Distrib.: Cuando-Cubango; Cunene; Huíla
(E. henrardii Jansen)
uniglumis Hack. = E. hierniana vacillans Rendle Distrib.: Huíla
venustula Launert ex Cope Distrib.: Huíla
viscosa (Retz.) Trin. Distrib.: Cunene; Huíla; Kuanza Norte
welwitschii Rendle Distrib.: Malanje; Moxico
ERIOCHLOA Kunth fatmensis (Hochst. & Steud.) Clayton
ERAGROSTIS ERIOCHLOA
16 A checklist of Angola grasses
ERIOCHLOA Kunth (cont.) Distrib.: Moxico
(E. nubica (Steud.) Thell.)
meyeriana (Nees) Pilg. Distrib.: Moxico
(Panicum meyerianum Nees var. grandeglume Stent &
J.M.Rattray)
nubica (Steud.) Thell. = E. fatmensis
ERIOCHRYSIS P.Beauv. pallida Munro Distrib.: Huíla; Lubango; Lunda Sul; Malanje
purpurata (Rendle) Stapf Distrib.: Benguela; Malanje
EUCLASTA Franch. condylotricha (Hochst. ex Steud.) Stapf Distrib.: Kuanza Norte
EULALIA Kunth aurea (Bory) Kunth Distrib.: Cuando-Cubango
villosa (Thunb.) Nees Distrib.: Huíla; Kuanza Norte
EUSTACHYS Desv. paspaloides (Vahl) Lanza & Mattei Distrib.: Cuando-Cubango; Cunene; Huíla; Kuanza
Norte
FESTUCA L. abyssinica A.Rich. Distrib.: Huíla
costata Nees Distrib.: Huíla
FINGERHUTHIA Nees africana Nees Distrib.: Cunene; Huíla; Kuanza Norte
GUADUELLA Franch. densiflora Pilg. Distrib.: Cabinda
dichroa Cope Distrib.: Cabinda
HACKELOCHLOA Kuntze granularis (L.) Kuntze Distrib.: Cuando-Cubango; Cunene; Malanje; Moxico
HARPOCHLOA Kunth pseudoharpechloa (Chiov.) Clayton Distrib.: Huíla
HEMARTHRIA R.Br. altissima (Poir.) Stapf & C.E.Hubb. Distrib.: Moxico
(H. fasciculata (Lam.) Kunth)
fasciculata (Lam.) Kunth = H. altissima natans Stapf Distrib.: Moxico
HETERANTHOECIA Stapf guineensis (Franch.) Robyns Distrib.: Lunda Sul
(Dinebra guineensis Franch.)
HETEROPHOLIS C.E.Hubb. sulcata (Stapf) C.E.Hubb. Distrib.: Moxico
HETEROPOGON Pers. contortus (L.) P.Beauv. ex Roem. & Schult. Distrib.: Benguela; Cuando-Cubango; Cunene; Huíla;
Kuanza Norte; Luanda; Moxico
melanocarpus (Elliott) Benth. Distrib.: Huíla; Kuanza Norte
HOMOZEUGOS Stapf conciliatum G.F.Guala Distrib.: Huambo
fragilis Stapf Distrib.: Bié
gossweileri Stapf Distrib.: Bié
huillense (Rendle) Stapf Distrib.: Huíla
(Pogonatherum huillense (Rendle) Roberty)
katakton Clayton Distrib.: Bié; Moxico
HYPARRHENIA E.Fourn. altissima Stapf = H. rufa anamesa Clayton Distrib.: Moxico
andongensis Rendle Distrib.: Malanje
bracteata (Humb. & Bonpl. ex Willd.) Stapf Distrib.: Benguela; Huíla; Moxico
cymbaria (L.) Stapf Distrib.: Huíla; Kuanza Norte
dichroa (Steud.) Stapf Distrib.: Moxico
diplandra (Hack.) Stapf Distrib.: Benguela; Huambo; Huíla; Kuanza Sul; Moxico
(H. mutica Clayton)
dissoluta (Nees ex Steud.) C.E.Hubb. = Hyperthelia dis-soluta
dregeana (Nees) Stapf ex Stent (H. pilosissima (Hack.) J.G.Anderson)
eylesii C.E.Hubb. = Elymandra grallata familiaris (Steud.) Stapf Distrib.: Moxico
filipendula (Hochst.) Stapf var. filipendula Distrib.: Malanje; Moxico
filipendula (Hochst.) Stapf var. pilosa (Hochst.) Stapf Distrib.: Cunene; Huíla; Kuanza Norte; Moxico; Uíge
finitima (Hochst.) Andersson ex Stapf Distrib.: Moxico
gossweileri Stapf Distrib.: Cuando-Cubango
grallata Stapf = Elymandra grallata hirta (L.) Stapf
ERIOCHLOA HYPARRHENIA
17A checklist of Angola grasses
Distrib.: Moxico
(H. modica (De Wild.) Robyns)
modica (De Wild.) Robyns = H. hirta newtonii (Hack.) Stapf var. macra Stapf Distrib.: Lunda Sul
newtonii (Hack.) Stapf var. newtonii Distrib.: Cuando-Cubango; Huambo; Huíla
niariensis (Franch.) Clayton Distrib.: Malanje
nyassae (Rendle) Stapf Distrib.: Cuando-Cubango; Moxico
(Andropogon nyassae Rendle)
pilosissima (Hack.) J.G.Anderson = H. dregeana poecilotricha (Hack.) Stapf Distrib.: Huíla; Moxico
quarrei Robyns Distrib.: Moxico
rudis Stapf Distrib.: Cuando-Cubango
rufa (Nees) Stapf Distrib.: Luanda; Moxico
(H. altissima Stapf)
schimperi (Hochst. ex A.Rich.) Andersson ex Stapf Distrib.: Moxico
subplumosa Stapf Distrib.: Namibe
variabilis Stapf Distrib.: Benguela; Huambo; Huíla; Malanje; Moxico
welwitschii (Rendle) Stapf Distrib.: Huambo; Huíla; Kuanza Norte; Lunda Sul;
Malanje; Moxico
HYPERTHELIA Clayton dissoluta (Nees ex Steud.) Clayton Distrib.: Cuando-Cubango; Cunene; Huíla; Kuanza Norte;
Malanje; Moxico
(Hyparrhenia dissoluta (Nees ex Steud.) C.E.Hubb.)
HYPOGYNIUM Nees ceresiiforme ( Nees ) Roberty = Monocymbium ceresii-
forme
IMPERATA Cirillo cylindrica (L.) Raeusch. Distrib.: Cuando-Cubango; Huíla; Kuanza Norte; Kuanza
Sul; Luanda; Lunda Sul; Malanje
ISACHNE R.Br. angolensis Rendle Distrib.: Moxico
buettneri Hack. Distrib.: Moxico
gossweileri Stapf & C.E.Hubb. Distrib.: Kuanza Norte
nervata Franch = Panicum nervatum
ISCHAEMUM L. afrum (J.F.Gmel.) Dandy Distrib.: Huíla; Kuanza Norte
(I. brachyatherum (Hochst.) Fenzl ex Hack.)
arcuatum (Nees) Stapf = I. fasciculatum
angolensis (Rendle) Stapf Distrib.: Huambo; Huíla
brachyatherum (Hochst.) Fenzl ex Hack. = I. afrum fasciculatum Brongn. Distrib.: Huíla
(I. arcuatum (Nees) Stapf)
JARDINEA Steud. angolensis (Rendle) Stapf = Phacelurus gabonensis
KAMPOCHLOA Clayton brachyphylla Clayton Distrib.: Moxico
KAOKOCHLOA De Winter nigrirostris De Winter Distrib.: Possibly occurring in Cunene; Namibe
LEERSIA Sw. friesii Melderis Distrib.: Moxico
hexandra Sw. Distrib.: Moxico
(Oryza hexandra (Sw.) Doell.)
nematostachya Launert Distrib.: Moxico
LEPTOCARYDION Stapf vulpiastrum (De Not.) Stapf Distrib.: Huíla
LEPTOCHLOA P.Beauv. caerulescens Steud. Distrib.: Malanje
fusca (L.) Kunth Distrib.: Moxico
(Diplachne fusca (L.) P.Beauv. ex Roem. & Schult.)
gigantea (Launert) Cope & N.Snow Distrib.: Moxico
(Diplachne gigantea Launert)
obtusifolia Hochst. Distrib.: Luanda
uniflora Hochst. ex A.Rich. Distrib.: Kuanza Norte
LEUCOPHRYS Rendle mesocoma (Nees) Rendle Distrib.: Namibe
LOPHACME Stapf digitata Stapf Distrib.: Moxico
LOUDETIA Steud. angolensis C.E.Hubb. Distrib.: Cunene; Huíla; Kuanza Norte; Kuanza Sul
arundinacea (A.Rich.) Steud. Distrib.: Kuanza Norte; Luanda; Malanje; Moxico
cuanzensis Lubke & J.B.Phipps Distrib.: Kuanza Norte
densispica (Rendle) C.E.Hubb.
HYPERTHELIA LOUDETIA
18 A checklist of Angola grasses
LOUDETIA Steud. (cont.) Distrib.: Benguela; Huíla
lanata (Stent & J.M.Rattray) C.E.Hubb. Distrib.: Cuando-Cubango; Huambo; Namibe
(Trichopteryx lanata Stent & J.M.Rattray)
phragmitoides (Peter) C.E.Hubb. Distrib.: Moxico
simplex (Nees) C.E.Hubb. Distrib.: Cuando-Cubango; Cunene; Huíla; Kuanza Norte;
Kuanza Sul; Lunda Sul
superba De Not. = Tristachya superba
LOXODERA Launert rhytachnoides (Launert) Clayton Distrib.: Uíge
MEGALOPROTACHNE C.E.Hubb. albescens C.E.Hubb. Distrib.: Huíla
(M. glabrescens Roiv.)
glabrescens Roiv. = M. albescens
MEGASTACHYA P.Beauv. mucronata (Poir.) P.Beauv. Distrib.: Benguela; Malanje
MELINIS P.Beauv. ambigua Hack. subsp. ambigua Distrib.: Moxico
ambigua Hack. subsp. longicauda (Mez) Zizka Distrib.: Huambo; Luanda; Moxico
(M. longicauda (Mez) Stapf & C.E.Hubb.)
amethystea (Franch.) Zizka Distrib.: Lunda Norte; Lunda Sul
(Rhynchelytrum amethysteum (Franch.) Chiov.)
angolensis Rendle Distrib.: Huíla
ascendens Mez Distrib.: Huíla
(Rhynchelytrum ascendens (Mez) Stapf & C.E.Hubb.)
biaristata (Rendle) Stapf & C.E.Hubb. Distrib.: Malanje
effusa (Rendle) Stapf Distrib.: Moxico
gossweileri C.E.Hubb. Distrib.: Huíla
kallimorpha (Clayton) Zizka Distrib.: Cuando-Cubango
(Rhynchelytrum kallimorphon Clayton)
longicauda (Mez) Stapf & C.E.Hubb. = M. ambigua subsp. longicauda
longiseta (A.Rich.) Zizka subsp. bellespicata (Rendle) Zizka
Distrib.: Cuando-Cubango; Cunene; Huíla; Namibe
(Rhynchelytrum bellespicatum (Rendle) Stapf &
C.E.Hubb.)
longiseta (A.Rich.) Zizka subsp. longiseta Distrib.: Cuando-Cubango
(Rhynchelytrum minutiflorum (Rendle) Stapf &
C.E.Hubb.)
macrochaeta Stapf & C.E.Hubb.
Distrib.: Huíla
minutiflora P.Beauv. Distrib.: Uíge
nerviglumis (Franch.) Zizka Distrib.: Moxico
(Rhynchelytrum nerviglume (Franch.) Chiov.)
repens (Willd.) Zizka subsp. grandiflora (Hochst.) Zizka
Distrib.: Bengo; Cunene; Namibe
(Rhynchelytrum grandiflorum Hochst.)
repens (Willd.) Zizka subsp. nigricans (Mez) Zizka Distrib.: Huambo; Malanje
(Rhynchelytrum nigricans (Mez) Stapf & C.E.Hubb.)
repens (Willd.) Zizka subsp. repens Distrib.: Cuando-Cubango; Cunene; Huíla; Malanje;
Moxico; Namibe; Uíge
(Rhynchelytrum repens (Willd.) C.E.Hubb.)
rupicola (Rendle) Zizka Distrib.: Malanje
(Rhynchelytrum rupicola (Rendle) Stapf & C.E.Hubb.)
subglabra Mez Distrib.: Cuando-Cubango
tanatricha (Rendle) Zizka Distrib.: Luanda; Malanje
(Rhynchelytrum tanatrichum (Rendle) Stapf &
C.E.Hubb.)
tenuissima Stapf Distrib.: Namibe
tomentosa Rendle Distrib.: Malanje
(Rhynchelytrum tomentosum (Rendle) Stapf &
C.E.Hubb.)
welwitschii Rendle Distrib.: Malanje
(Rhynchelytrum welwitschii (Rendle) Stapf &
C.E.Hubb.)
MICROCHLOA R.Br. caffra Nees Distrib.: Cunene; Cuando-Cubango; Huíla; Lunda Sul
ensifolia Rendle Distrib.: Malanje
indica (L.f.) P.Beauv. Distrib.: Cuando-Cubango; Cunene; Moxico
kunthii Desv. Distrib.: Cuando-Cubango; Cunene; Huíla; Moxico
MISCANTHIDIUM Stapf junceum (Stapf) Stapf = Miscanthus junceus sorghum (Nees) Stapf = Miscanthus sorghum
MISCANTHUS Andersson junceus (Stapf) Pilg. Distrib.: Cuando-Cubango; Kuanza Norte
(Miscanthidium junceum (Stapf) Stapf)
sorghum (Nees) Pilg. Distrib.: Said to occur, Flora zambesiaca 10,4: 10 (2002)
(Miscanthidium sorghum (Nees) Stapf)
MONELYTRUM Hack. annuum Gooss. = M. luederitzianum
LOUDETIA MONELYTRUM
19A checklist of Angola grasses
luederitzianum Hack. Distrib.: Namibe
(M. annuum Gooss.)
MONOCYMBIUM Stapf ceresiiforme (Nees) Stapf Distrib.: Cuando-Cubango; Cunene; Huambo; Huíla;
Kuanza Norte; Malanje
(Hypogynium ceresiiformis (Nees) Roberty)
ODYSSEA Stapf paucinervis (Nees) Stapf Distrib.: Namibe
OLYRA L. latifolia L. Distrib.: Kuanza Norte
OPLISMENUS P.Beauv. hirtellus (L.) P.Beauv. Distrib.: Bié; Huíla; Lunda Norte; Lunda Sul
OROPETIUM Trin. capense Stapf Distrib.: Namibe
ORYZA L. barthii A.Chev. Distrib.: Cunene; Huíla
hexandra (Sw.) Doell. = Leersia hexandra longistaminata A.Chev. & Roehr. Distrib.: Cuando-Cubango; Moxico
ORYZIDIUM C.E.Hubb. & Schweick. barnardii C.E.Hubb. & Schweick. Distrib.: Cuando-Cubango; Huíla
OXYTENANTHERA Munro abyssinica (A.Rich.) Munro Distrib.: Luanda; Malanje
(Bambusa abyssinica A.Rich.)
PANICUM L. adenophorum K.Schum. Distrib.: Moxico
brazzavillense Franch. Distrib.: Malanje
(P. ianthum Stapf)
catumbense Rendle = Sacciolepis catumbensis coloratum L. var. coloratum Distrib.: Bié; Cunene; Cuando-Cubango; Huíla; Kuanza
Norte
dregeanum Nees Distrib.: Cunene; Cuando-Cubango
ecklonii Nees Distrib.: Bié; Kuanza Norte; Malanje
(P. pectinatum Rendle)
fluviicola Steud. Distrib.: Malanje
glabrescens Steud. = P. subalbidum gracilicaule Rendle
Distrib.: Moxico
(P. verruciferum Mez)
graciliflorum Rendle Distrib.: Kuanza Norte
graniflorum Stapf Distrib.: Moxico
griffonii Franch. Distrib.: Malanje
heterostachyum Hack. Distrib.: Benguela; Cabinda; Cunene; Huambo; Huíla;
Kuanza Norte; Namibe
humidicola Rendle = Brachiaria humidicola hymeniochilum Nees Distrib.: Moxico
ianthum Stapf = P. brazzavillense infestum Andersson ex Peters Distrib.: Moxico
juncifolium Stapf = P. natalense kalaharense Mez Distrib.: Cuando-Cubango
laetum Kunth Distrib.: Moxico
laevifolium Hack. var. laevifolium = P. schinzii lindleyanum Nees Distrib.: Malanje
margariteferum (Chiov.) Robyns Distrib.: Moxico
maximum Jacq. Distrib.: Cuando-Cubango; Cunene
merkeri Mez Distrib.: Luanda
meyerianum var. grandeglume Stent & J.M.Rattray = Eriochloa meyeriana
natalense Hochst. Distrib.: Huíla; Lunda Sul
(P. juncifolium Stapf)
nervatum (Franch.) Stapf Distrib.: Moxico
(Isachne nervata Franch.)
pansum Rendle Distrib.: Malanje
parvifolium Lam. Distrib.: Huíla
pectinatum Rendle = P. ecklonii phragmitoides Stapf Distrib.: Malanje
pectinellum Stapf Distrib.: Said to occur, Flora zambesiaca 10,3: 12 (1989)
repens L. Distrib.: Cuando-Cubango; Cunene; Huíla
sadinii (Vanderyst) Renvoize Distrib.: Moxico
(Brachiaria sadinii Vanderyst)
schinzii Hack. Distrib.: Namibe
(P. laevifolium var. laevifolium Hack.)
seslerioides Rendle = Sacciolepis seslerioides stapfianum Fourc. Distrib.: Benguela
subalbidum Kunth Distrib.: Moxico
MONOCYMBIUM PANICUM
20 A checklist of Angola grasses
PANICUM L. (cont.) (P. glabrescens Steud.)
verruciferum Mez = P. gracilicaule xantholeucum Schinz = Brachiaria xantholeuca
PARATHERIA Griseb. prostrata Griseb. Distrib.: Moxico
PASPALIDIUM Stapf geminatum (Forssk.) Stapf Distrib.: Huíla; Moxico; Namibe
PASPALUM L. *dilatatum Poir. Distrib.: Huíla
conjugatum Berg. Distrib.: Cabinda; Luanda
orbiculare G.Forst. = P. scrobiculatum scrobiculatum L. Distrib.: Cabinda; Cunene; Huíla; Kuanza Norte
(P. orbiculare G.Forst.)
*urvillei Steud. Distrib.: Huambo
vaginatum Sw. Distrib.: Kuanza Sul
PENNISETUM Rich. angolense Rendle = P. macrourum glabrum Steud. = P. thunbergii glaucocladum Stapf & C.E.Hubb. = P. macrourum glaucum (L.) R.Br. Distrib.: unknown
(P. typhoides (Burm.) Stapf & C.E.Hubb.)
gossweileri Stapf & C.E.Hubb. = P purpureum macrourum Trin. Distrib.: Bengo; Kuanza Norte; Kuanza Sul; Malanje
(P. angolense Rendle)
polystachion (L.) Schult. subsp. polystachion Distrib.: Benguela; Cabinda; Huíla; Kuanza Sul; Lubango;
Namibe; Uíge
purpureum Schumach. Distrib.: Bengo; Huambo; Huíla; Uíge
(P. gossweileri Stapf & C.E.Hubb.)
thunbergii Kunth Distrib.: Huíla
(P. glabrum Steud.)
typhoides (Burm.) Stapf & C.E.Hubb. = P. glaucum unisetum (Nees) Benth. Distrib.: Moxico
(Beckeropsis uniseta (Nees) K.Schum.)
PEROTIS Aiton huillensis (Rendle) Clayton Distrib.: Cunene; Huíla; Kuanza Norte
leptopus Pilg. Distrib.: Huambo; Huíla; Malanje; Moxico; Namibe
patens Gand. Distrib.: Cunene; Huambo; Huíla; Kuanza Norte;
Lubango; Uíge
vaginata Hack.
Distrib.: Cuando-Cubango; Cunene; Namibe
PHACELURUS Griseb. gabonensis (Steud.) Clayton Distrib.: Malanje
(Jardinea angolensis (Rendle) Stapf)
(Rottboellia angolensis Rendle)
huillensis (Rendle) Clayton Distrib.: Huíla
(Rottboellia huillensis Rendle)
PHRAGMITES Adans. australis (Cav.) Steud. Distrib.: Cuando-Cubango; Cunene
(P. communis Trin.)
communis Trin. = P. australis mauritianus Kunth Distrib.: Cuando-Cubango; Cunene; Huíla; Lunda Sul;
Moxico; Namibe
PHYLLORHACHIS Trimen sagittata Trimen Distrib.: Kuanza Norte; Malanje
PIPTOPHYLLUM C.E.Hubb. welwitschii (Rendle) C.E.Hubb. Distrib.: Kuanza Norte
POGONARTHRIA Stapf falcata (Hack.) Rendle = P. squarrosa fleckii (Hack.) Hack. Distrib.: Cunene; Huíla; Namibe
(P. tuberculata Pilg.)
squarrosa (Roem. & Schult.) Pilg. Distrib.: Cuando-Cubango; Cunene; Huíla; Namibe
(P. falcata Rendle)
tuberculata Pilg. = P. fleckii
POGONATHERUM P.Beauv. huillense (Rendle) Roberty = Homozeugos huillense
POLYPOGON Desf. viridis (Gouan) Breistr. Distrib.: Huíla
(Agrostis semiverticillata (Forssk.) C.Chr.)
PSEUDECHINOLAENA Stapf. polystachya (Kunth) Stapf Distrib.: Moxico
(Echinochloa polystachya (Kunth) Roberty)
PSEUDOBRACHIARIA Launert deflexa (Schumach.) Launert = Brachiaria deflexa
RHYNCHELYTRUM Nees amethysteum (Franch.) Chiov. = Melinis amethystea ascendens (Mez) Stapf & C.E.Hubb. = Melinis ascen-
dens bellispicatum (Rendle) Stapf & C.E.Hubb. = Melinis
longiseta subsp. bellispicata grandiflorum Hochst. = Melinis repens subsp. grandi-
PANICUM RHYNCHELYTRUM
21A checklist of Angola grasses
flora kallimorphon Clayton = Melinis kallimorpha minutiflorum (Rendle) Stapf & C.E.Hubb. = Melinis
longiseta subsp. longiseta nerviglume (Franch.) Chiov. = Melinis nerviglumis nigricans (Mez) Stapf & C.E.Hubb. = Melinis repens
subsp. nigricans repens (Willd.) C.E.Hubb. = Melinis repens subsp.
repens rupicola (Rendle) Stapf & C.E.Hubb. = Melinis ru-
picola tanatrichum (Rendle) Stapf & C.E.Hubb. = Melinis
tanatricha tomentosum (Rendle) Stapf & C.E.Hubb. = Melinis
tomentosa welwitschii (Rendle) Stapf & C.E.Hubb. = Melinis
welwitschii
RHYTACHNE Desv. robusta Stapf Distrib.: Type: see Flora zambesiaca 10,4: 169 (2002)
(Rottboellia robusta (Stapf) Keng)
rottboellioides Desv. Distrib.: Huíla
ROTTBOELLIA L.f. angolensis Rendle = Phacelurus gabonensis cochinchinensis ( Lour.) Clayton Distrib.: Cuando-Cubango; Cunene; Moxico
(R. exaltata L.f.)
exaltata L.f. = R. cochinchinensis huillensis Rendle = Phacelurus huillensis robusta (Stapf) Keng = Rhytachne robusta
SACCHARUM L. *officinarum L. Distrib.: Benguela
SACCIOLEPIS Nash africana C.E.Hubb. & Snowden Distrib.: Cunene; Huíla; Kuanza Norte
barbiglandularis Mez = S. transbarbata catumbensis (Rendle) Stapf Distrib.: Moxico
(Panicum catumbense Rendle)
chevalieri Stapf Distrib.: Benguela; Huíla
(S. strictula Pilg.)
claviformis B.K.Simon = S. indica glaucescens Stapf = S. typhura huillensis (Rendle) Stapf = S. spiciformis indica (L.) Chase Distrib.: Huíla; Moxico
(S. claviformis Simon)
leptorhachis Stapf = S. rigens micrococca Mez Distrib.: Malanje
rigens (Mez) A.Chev. Distrib.: Moxico
(S. leptorhachis Stapf)
seslerioides (Rendle) Stapf
Distrib.: Moxico
(Panicum seslerioides Rendle)
spiciformis (A.Rich.) Stapf Distrib.: Huíla
(S. huillense (Rendle) Stapf)
strictula Pilg. = S. chevalieri transbarbata Stapf Distrib.: Huíla
(S. barbiglandularis Mez)
typhura (Stapf) Stapf Distrib.: Huíla; Kuanza Sul; Malanje; Moxico; Namibe
(S. glaucescens Stapf)
SARTIDIA De Winter angolensis (C.E.Hubb.) de Winter Distrib.: Huíla
(Aristida angolensis C.E.Hubb.)
SCHIZACHYRIUM Nees brevifolium (Sw.) Nees ex Büse Distrib.: Huíla; Kuanza Norte; Moxico
exile (Hochst.) Pilg. Distrib.: Moxico
(S. inclusum Stent)
inclusum Stent = S. exile jeffreysii (Hack.) Stapf Distrib.: Cuando-Cubango; Cunene; Huíla; Malanje
lopollense (Rendle) Sales Distrib.: Huíla
(Andropogon lopollensis Rendle)
sanguineum (Retz.) Alston Distrib.: Cuando-Cubango; Cunene; Huíla
(S. semiberbe Nees)
semiberbe Nees = S. sanguineum
SCHMIDTIA Steud. ex J.A.Schmidt kalahariensis Stent Distrib.: Huíla; Luanda; Namibe
pappophoroides Steud. ex J.A.Schmidt Distrib.: Cunene; Huíla; Kuanza Norte; Namibe
SEHIMA Forssk. galpinii Stent Distrib.: Cuando-Cubango
ischaemoides Forssk. Distrib.: Cuando-Cubango
SETARIA P.Beauv. angustissima Stapf = S. lindenbergiana atrata Hack. Distrib.: Moxico
chevalieri Stapf = S. megaphylla homonyma (Steud.) Chiov. Distrib.: Cuando-Cubango; Moxico
(S. microprolepis Stapf)
incrassata (Hochst.) Hack. Distrib.: Cuando-Cubango; Cunene; Kuanza Sul; Namibe
lindenbergiana (Nees) Stapf Distrib.: Huíla; Moxico
(S. angustissima Stapf)
longiseta P.Beauv.
RHYTACHNE SETARIA
22 A checklist of Angola grasses
SETARIA P.Beauv. (cont.) Distrib.: Moxico
(S. paniciformis Rendle)
megaphylla (Steud.) T.Durand & Schinz Distrib.: Cunene; Kuanza Norte; Moxico
(S. chevalieri Stapf)
(S. phillipsii De Wit)
(S. plicatilis (Hochst.) Engl.)
microprolepis Stapf = S. homonyma pallide-fusca (Schumach.) Stapf & C.E.Hubb. = S.
pumila paniciformis Rendle = S. longiseta phillipsii De Wit = S. megaphylla plicatilis (Hochst.) Engl. = S. megaphylla pseudaristata (Peter) Pilg. Distrib.: Cuando-Cubango
pumila (Poir.) Roem. & Schult. Distrib.: Cuando-Cubango; Cunene; Huíla; Huambo;
Kuanza Norte; Moxico
(S. pallide-fusca (Schumach.) Stapf & C.E.Hubb.)
petiolata Stapf & C.E.Hubb. Distrib.: Malanje; Moxico
sagittifolia (A.Rich.) Walp. Distrib.: Cuando-Cubango; Cunene; Huíla; Luanda;
Lunda Sul
sphacelata (Schumach.) Moss var. sericea (Stapf) Clay-ton
Distrib.: Cuando-Cubango; Huíla; Kuanza Norte; Moxico
sphacelata (Schumach.) Moss var. sphacelata Distrib.: Cuando-Cubango; Cunene; Moxico
verticillata (L.) P.Beauv. Distrib.: Cuando-Cubango; Cunene; Huíla; Luanda;
Namibe
welwitschii Rendle Distrib.: Luanda
SORGHASTRUM Nash nudipes Nash Distrib.: Huíla
(Sorghum nutans (L.) Nash var. angolense Rendle)
pogonostachyum (Stapf) Clayton Distrib.: Benguela
SORGHUM Moench arundinaceum (Desv.) Stapf Distrib.: Cuando-Cubango; Cunene; Moxico
*bicolor (L.) Moench Distrib.: Cuando-Cubango; Cunene; Huíla
(S. caffrorum P.Beauv.)
caffrorum P.Beauv. = S. bicolor nutans (L.) Nash var. angolense Rendle = Sorghastrum
nudipes versicolor Andersson Distrib.: Cunene; Huíla
SPOROBOLUS R.Br. africanus (Poir.) A.Robyns & Tournay Distrib.: Cuando-Cubango; Moxico
barbigerus Franch. = S. subtilis consimilis Fresen. Distrib.: Namibe
congoensis Franch. Distrib.: Moxico
(S. eylesii Stent & J.M.Rattray)
conrathii Chiov. = S. welwitschii coromandelianus (Retz.) Kunth Distrib.: Cunene
eylesii Stent & J.M.Rattray = S. congoensis festivus Hochst. ex A.Rich. Distrib.: Cuando-Cubango; Cunene; Namibe
(S. menyharthii Hack.)
fimbriatus (Trin.) Nees Distrib.: Cuando-Cubango; Cunene; Moxico; Namibe
ioclados (Trin.) Nees Distrib.: Cuando-Cubango; Cunene
kentrophyllus (K.Schum.) Clayton Distrib.: Cuando-Cubango
mauritianus (Steud.) T.Durand & Schinz Distrib.: Moxico
mayumbensis Franch. = S. molleri menyharthii Hack. = S. festivus molleri Hack. Distrib.: Moxico
(S. mayumbensis Franch.)
myrianthus Benth. Distrib.: Moxico
natalensis (Steud.) T.Durand & Schinz Distrib.: Cuando-Cubango; Cunene
panicoides A.Rich. Distrib.: Cuando-Cubango; Cunene
piliferus (Trin.) Kunth Distrib.: Huíla
pyramidalis P.Beauv. Distrib.: Cuando-Cubango; Cunene; Moxico
sanguineus Rendle Distrib.: Huíla; Moxico
schlechteri Schweick. = S. centrifugus spicatus (Vahl) Kunth Distrib.: Cuando-Cubango
stapfianus Gand. Distrib.: Huíla
subtilis Kunth Distrib.: Moxico
(S. barbigerus Franch.)
virginicus (L.) Kunth Distrib.: Namibe
welwitschii Rendle Distrib.: Cuando-Cubango; Cunene; Moxico
(S. conrathii Chiov.)
STEREOCHLAENA Hack. cameronii (Stapf) Pilg. Distrib.: Moxico
(Chloridion cameronii (Stapf) Pilg.)
STIPAGROSTIS Nees damarensis (Mez) De Winter Distrib.: Possibly in Angola in Namibe
dinteri (Hack.) De Winter Distrib.: Namibe
(Aristida dinteri Hack.)
giesssii Kers
SETARIA STIPAGROSTIS
23A checklist of Angola grasses
Distrib.: Namibe
hermannii (Mez) De Winter Distrib.: Possibly in Angola in Namibe
hirtigluma (Steud. ex Trin. & Rupr.) De Winter subsp. patula (Hack.) De Winter
Distrib.: Cunene
hirtigluma (Steud. ex Trin. & Rupr.) De Winter subsp. pearsonii (Henrard) De Winter
Distrib.: Cunene; Namibe
hochstetteriana (Beck ex Hack.) De Winter var. secalina (Henrard) De Winter
Distrib.: Namibe
prodigiosa (Welw.) De Winter Distrib.: Cunene
subacaulus (Nees) De Winter Distrib.: Namibe
(Aristida subacaulus Nees ex Steud.)
uniplumis (Licht.) De Winter Distrib.: Namibe
vexillifera Kers Distrib.: Namibe
STREPTOGYNA P.Beauv. crinata P.Beauv. Distrib.: Cabinda
STREPTOLOPHUS Hughes sagittifolius Hughes Distrib.: Namibe
STYPPEIOCHLOA De Winter catheriniana Cope & Ryves Distrib.: Type: In Kew Bulletin 58: 739 (2003)
Distrib.: Type only; no locality
TETRACHNE Nees aristulata Hack. & Rendle = Entoplocamia aristulata
TETRAPOGON Desf. tenellus (Roxb.) Chiov. Distrib.: Said to occur, see Flora zambesiaca 10,2:
207–209 (1999)
THELEPOGON Roth elegans Roth ex Roem. & Schult. Distrib.: Cuando-Cubango
THEMEDA Forssk. triandra Forssk. Distrib.: Cuando-Cubango; Huíla; Kuanza Sul; Moxico
TRACHYPOGON Nees capensis (Thunb.) Trin. = T. spicatus spicatus (L.f.) Kuntze Distrib.: Cuando-Cubango; Cunene; Huíla; Lunda Sul;
Moxico
(T. capensis (Thunb.) Trin.)
TRAGUS Haller berteronianus Schult. Distrib.: Cuando-Cubango; Cunene; Huíla
racemosus (L.) All. Distrib.: Huíla
TRICHOLAENA Schrad. ex Schult. delicatula Stapf & C.E.Hubb. = T. monachne monachne (Trin.) Stapf & C.E.Hubb. Distrib.: Benguela; Cuando-Cubango; Cunene; Huíla;
Luanda; Lunda Sul; Moxico; Namibe
(T. delicatula Stapf & C.E.Hubb.)
TRICHONEURA Andersson eleusinoides (Rendle) Ekman Distrib.: Namibe
grandiglumis (Nees) Ekman Distrib.: Benguela; Cuando-Cubango; Huíla
TRICHOPTERYX Nees dregeana Nees Distrib.: Cuando-Cubango
fruticulosa Chiov. Distrib.: Moxico
(T. katangensis De Wild.)
gracillima C.E.Hubb. = T. marungensis katangensis De Wild. = T. fruticulosa lanata Stent & J.M.Rattray = Loudetia lanata marungensis Chiov. Distrib.: Moxico
(T. gracillima C.E.Hubb.)
stolziana Henrard Distrib.: Moxico
TRIPOGON Roem. & Schult. minimus (A.Rich.) Hochst. ex Steud. Distrib.: Cuando-Cubango
modestus S.M.Phillips & Launert Distrib.: Said to occur, see Flora zambesiaca 10,2: 32
(1999)
TRIRAPHIS R.Br. fleckii Hack. = T. purpurea pumila R.Br. Distrib.: Namibe
purpurea Hack. Distrib.: Cunene; Namibe
(T. fleckii Hack.)
ramosissima Hack. Distrib.: Cunene; Namibe
(T. elliottii Rendle)
schinzii Hack. Distrib.: Cuando-Cubango; Huíla
(T. schlechteri Pilg. ex Stent)
schlechteri Pilg. ex Stent = T. schinzii
TRISTACHYA Nees bequaertii De Wild. Distrib.: Moxico
hispida (L.f.) K.Schum. = T. leucothrix hitchcockii (C.E.Hubb.) Conert = T. lualabaensis hubbardiana Conert Distrib.: Said to occur, see Flora zambesiaca 10,3: 206
(1989)
STREPTOGYNA TRISTACHYA
24 A checklist of Angola grasses
TRISTACHYA Nees (cont.) huillensis Rendle Distrib.: Huíla
leucothrix Nees Distrib.: Malanje
(Apochaete hispida (L.f.) J.B.Phipps)
(T. hispida (L.f.) K.Schum.)
lualabaensis (De Wild.) J.B.Phipps Distrib.: Cuando-Cubango; Moxico
(T. hitchcockii (C.E.Hubb.) Conert)
nodiglumis K.Schum. Distrib.: Cuando-Cubango; Huíla; Malanje; Moxico;
Uíge
rehmannii Hack. Distrib.: Benguela; Cuando-Cubango; Moxico
superba (De Not.) Schweinf. & Asch. Distrib.: Cuando-Cubango; Cunene; Huambo; Huíla;
Moxico
(Loudetia superba De Not.)
thollonii Franch. Distrib.: Moxico
URELYTRUM Hack. agropyroides (Hack.) Hack. Distrib.: Cuando-Cubango
digitatum K.Schum. Distrib.: Malanje
(U. stapfianum C.E.Hubb.)
stapfianum C.E. Hubb = U. digitatum
UROCHLOA P.Beauv. bolbodes (Steud.) Stapf = U. oligotricha brachyura (Hack.) Stapf Distrib.: Cuando-Cubango; Cunene
mosambicensis (Hack.) Dandy Distrib.: Cuando-Cubango; Namibe
(U. pullulans Stapf)
oligotricha (Fig. & De Not.) Henrard Distrib.: Cuando-Cubango; Cunene; Huíla; Namibe
(U. bolbodes (Steud.) Stapf)
pullulans Stapf = U. mosambicensis trichopus (Hochst.) Stapf Distrib.: Cuando-Cubango; Cunene
VETIVERIA Bory nigritana (Benth.) Stapf = Chrysopogon nigritanus zizanioides (L.) Nash var. nigritana (Benth.) A.Camus =
Chrysopogon nigritanus
VOSSIA Wall. & Griff. cuspsidata (Roxb.) Griff. Distrib.: Cuando-Cubango
WILLKOMMIA Hack. annua Hack. Distrib.: Cunene; Namibe; Huíla
newtonii Hack. Distrib.: Cunene
sarmentosa Hack. Distrib.: Cuando-Cubango; Cunene; Huíla; Moxico
(Craspedorhachis sarmentosa (Hack.) Pilg.)
ZONOTRICHE (C.E.Hubb.) J.B.Phipps decora (Stapf) J.B.Phipps Distrib.: Moxico
inamoena (K.Schum.) Clayton Distrib.: Moxico
*ZEA L. *mays L. Distrib.: Bengo; Benguela; Bié; Cuando-Cubango;
Cunene; Huambo; Huíla; Kuanza Norte; Kuanza Sul;
Luanda; Lunda Norte; Lunda Sul; Malanje; Moxico;
Namibe; Uíge
TRISTACHYA ZEA
25A checklist of Angola grasses
This publication is a product of the Southern African Botanical Diversity Network (SABONET), a programme aimed at strengthening the level of botanical expertise, expanding and improving herbarium and botanic garden collections, and fostering closer collaborative links among botanists in the southern African subcontinent.
The main objective of SABONET is to develop a strong core of professional botanists, taxonomists, horticulturists, and plant diversity specialists within the ten countries of southern Africa (Angola, Botswana, Lesotho, Malawi, Mozambique, Namibia, South Africa, Swaziland, Zambia, and Zimbabwe). This core group will be competent to inventory, monitor, evaluate, and conserve the botanical diversity of the region in the face of specific development challenges, and to respond to the technical and scientific needs of the Convention on Biological Diversity.
To enhance the human resource capacity and infrastructure available in the region, SABONET offers training courses, workshops, and collaborative expeditions in un-der-collected areas. The programme produces a newsletter, SABONET News, and a series of occasional publications, the Southern African Botanical Diversity Network Report Series, of which this publication is part.
SABONET is co-funded by:• The United States Agency for International Development (USAID/World Conservation Union—Regional Office for southern Africa (IUCN-ROSA)• The Global Environment Facility (GEF)/United Nations Development Programme (UNDP)
For more information about our projects in southern Africa contact one of the following addresses:
General enquiries about SABONETSABONET Coordinatorc/o National Botanical InstitutePrivate Bag X101Pretoria 0001South AfricaTel: (27) 12 804 3200Fax: (27) 12 804 3211/5979E-mail: [email protected]://www.sabonet.org
ANGOLALuanda Herbarium (Prof. Esparança Costa)Universidade Agostinho NetoRua Fernando Pessoa No. 103Villa AliceCaixa Postal 3244Tel: (244) 2 336 168Fax: (244) 2 336 168E-mail: [email protected]
BOTSWANABotswana National MuseumNational Herbarium (Mr Nonofo Mosesane)Private Bag 00114GaboroneTel: (267) 3973860/3974616Fax: (267) 311186/3902797E-mail: [email protected]
LESOTHONational Environment Secretariat (Mr Thulo Qhotsokoane)Ministry of EnvironmentPrivate Bag A23Maseru 100
Tel: (266) 311 767Fax: (266) 310 506/321505E-mail: [email protected]
MALAWINational Herbarium and Botanic Gardens of Malawi (Dr Zacharia Magombo)P.O. Box 528ZombaTel: (265) 525 388/118/145Fax: (265) 524164/108E-mail: [email protected]
MOZAMBIQUELMA Herbarium (Ms Samira Izidiné)Instituto Nacional de Investigaçáo AgronómicaCaixa Postal 3658MavalaneMaputoTel: (258) 1 460 255/130/190/097Fax: (258) 1 460 074E-mail: [email protected]
NAMIBIANational Herbarium (Dr Gillian Maggs-Kölling)National Botanical Research InstitutePrivate Bag 13184WindhoekTel: (264) 61 202 2020Fax: (264) 61 258 153E-mail: [email protected]
SOUTH AFRICANational Herbarium (Prof. Gideon Smith)National Botanical InstitutePrivate Bag X101
Pretoria 0001Tel: (27) 12 804 3200Fax: (27) 12 804 3211/5343E-mail: [email protected]
SWAZILANDNational Herbarium (Mr Titus Dlamini)Malkerns Agricultural Research StationP.O. Box 4MalkernsTel: (268) 52 82111/83017/83038Fax: (268) 52 83360/83490E-mail: [email protected]
ZAMBIAHerbarium (Dr Patrick Phiri)Department of Biological SciencesUniversity of ZambiaP.O. Box 32379LusakaTel: (260) 1 293 158Fax: (260) 1 294806/253952E-mail: [email protected]
ZIMBABWENational Herbarium and Botanic Garden (Ms Nozipo Nobanda)P.O Box A889AvondaleHarareTel: (263) 4 708 938/744170/745230Fax: (263) 4 708 938E-mail: [email protected]
About SABONET
1. *Southern African national herbaria: status reports, 1996. C.K. Willis (ed.). 1997. 59 pp. ISBN 1-874907-36-6.
2. *Index herbariorum: southern African supplement. G.F. Smith & C.K. Willis (eds). 1997. 55 pp. ISBN 1-874907-37-4.
3. *PRECIS Specimen database user guide. C.A. Pren-tice & T.H. Arnold. 1998. 130 pp. ISBN 1-874907-39-0.
4. *†Inventory, evaluation and monitoring of botanical diversity in southern Africa: a regional capacity and institution building network (SABONET). B.J. Hunt-ley, E.M. Matos, T.T. Aye, U. Nermark, C.R. Nagen-dran, J.H. Seyani, M.A.C. da Silva, S. Izidine, G.L. Maggs, C. Mannheimer, R. Kubirske, G.F. Smith, M. Koekemoer, G.M. Dlamini, P.S.M. Phiri, N. Nobanda & C.K. Willis. 1998. 73 pp. ISBN 1-919795-36-7.
5. *Plant taxonomic and related projects in southern Africa. T.H. Arnold & M. Mössmer (compilers). 1998. 101 pp. ISBN 1-919795-34-0.
6. *†Southern African herbarium needs assessment. G.F. Smith, C.K. Willis & M. Mössmer. 1999. 88 pp. ISBN 1-919795-45-6.
7. *A checklist of Namibian plant species. P. Craven (ed.). 1999. 206 pp. ISBN 1-919795-37-5.
8. †Index herbariorum: southern African supplement. Second edition. G.F. Smith & C.K. Willis. 1999. 181 pp. ISBN 1-919795-47-2.
9. *Making your garden come alive! Environmental interpretation in botanical gardens. M. Honig. 2000. 96 pp. ISBN 1-919795-50-2.
10. †Plant taxonomic expertise: An inventory for south-ern Africa. M. Mössmer & C.K. Willis. 2000. 350 pp. ISBN 1-919795-53-7.
11. *Southern African botanical gardens needs assess-ment. D.J. Botha, C.K. Willis & J.H.S. Winter. 2000. 156 pp. ISBN 1-919795-54-5.
12. *Action plan for southern African botanical gardens. C.K. Willis & S. Turner (eds). 2001. 35 pp. ISBN 1-919795-61-8.
13. Conspectus of southern African Pteridophyta. J.P. Roux. 2001. 223 pp. ISBN 1-919795-58-8.
14. *Southern African plant Red Data Lists. J.S. Golding (ed.). 2002. 256 pp. ISBN 1-919795-64-2.
Other publications in this series15. *Addressing the needs of the users of botanical infor-
mation. Y. Steenkamp & G.F. Smith. 2002. 56 pp. ISBN 1-919795-65-0.
16. *A checklist of Zimbabwean grasses. C. Chapano. 2002. 28 pp. ISBN 1-919795-66-9.
17. *A checklist of Lesotho grasses. K. Kobisi & L.E. Kose. 2002. 28 pp. ISBN 1-919795-68-5.
18. Trees of Botswana: names and distribution. M.P. Setshogo & F. Venter. 2003. 160 pp. ISBN 1-919795-69-3.
19. *Swaziland ferns and fern allies. J.P. Roux. 2003. 242 pp. ISBN 1-919795-97-9.
20. Checklist of grasses in Namibia. E.S.Klaassen & P. Craven. 2003. 130 pp. ISBN 99916-63-16-9.
21. A checklist of Zimbabwean bryophytes. P. Manyanga & S.M. Perold. 2004. 22 pp. ISBN 1-919976-02-7.
22. African Botanic Gardens Congress ‘Partnerships and Linkages’: proceedings of a congress held at Durban Botanic Gardens, South Africa, 24–29 November 2002. / Congrès des Jardins Botaniques Africains ‘Relations et Partenariats’: compte rendu d’un congrès tenu dans les Jardins Botaniques de Durban, Afrique du Sud, 24–29 Novembre 2002. C.K. Willis (ed.). 2004. 96 + 96 pp. ISBN 1-919976-04-3.
23. Integration of Red Data List concepts into the policy framework in Mozambique: proceedings of a work-shop held in Kaya-Kwanga, Maputo, Mozambique, 29–31 August 2001. S.A. Izidine, I. Nhantumbo & J. Golding (eds). 2004. 19 + 19 pp. ISBN 1-919976-05-1.
24. A checklist of Botswana grasses. M. Kabelo & D. Mafokate. 2004. 18 pp. ISBN 1-919976-06-X.
25. Herbarium essentials: the southern African herbar-ium user guide. J.E. Victor, M. Koekemoer, L. Fish, S.J. Smithies & M. Mössmer. 2004. ISBN 1-919795-54-5.
27. Swaziland Flora Checklist. K.P. Braun, S.D.V. Dlamini, D.R. Mdladla, N.P. Methule, P.W. Dlamini & M.S. Dlamini. 2004. ISBN 1-919976-10-8.
29. Herbaria in SABONET countries: building botani-cal capacity and meeting end-user expectations. T.J. Smith, G.F. Smith & Y. Steenkamp. 2004. ISBN 1-919976-11-6.
* Out of print.
† Available in PDF format on the SABONET web site: http://www.sabonet.org/publications/download.htm
Most of the books in this series can also be ordered in PDF format on CD. Contact [email protected]