CHECKLIST OF THE

DYNASTINAE OF

HONDURAS,

NICARAGUA, AND

ELSALVADOR

W Ratcliffe, B. C. and R. D. Cave W Ratcliffe, B. C. and R. D. Cave W Ratcliffe, B. C. and R. D. Cave W Ratcliffe, B. C. and R. D. Cave

Listado de los escarabajos Dynastinae de

Honduras, Nicaragua, y El Salvador,

tomados del documento original de

W Ratcliffe, B. C. and R. D. Cave del Bulletin Univ.

Nebraska State Museum 21: 1-500. Año2006

Rubén Ernesto López Sorto

20/09/2010

Se elaboró un listado y una revisión comprensiva de las 116 especies de escarabajos dinástinos que ocurren en Honduras, Nicaragua, y El Salvador. Publicadas por W Ratcliffe, B. C. y R. D. Cave en el Boletín de la Universidad de Nebraska 21: 1-500. del Año 2006.

CHECKLIST OF THE DYNASTINAE OF HONDURAS, NICARAGUA, AND ELSALVADOR

Tribe Cyclocephalini

Ancognatha sellata Arrow, 1911

Aspidolea fuliginea Burmeister, 1847

Aspidolea singularis Bates, 1888

Aspidolea texana Höhne, 1922 (synonym)

Aspidolea similis Höhne, 1922 (synonym)

Aspidolea cevallosi Martínez, 1975 (synonym)

Cyclocephala abrelata Ratcliffe and Cave, 2002

Cyclocephala alexi Ratcliffe and Delgado, 1992

Cyclocephala amblyopsis Bates, 1888

Cyclocephala amblyopsis monochroa Bates, 1888 (synonym)

Cyclocephala ampliata Bates 1888

Cyclocephala atripes Bates, 1888

Cyclocephala batesi Delgado and Castañeda, 1994

Cyclocephala brevis Höhne, 1923

Cyclocephala pubescens Burmeister, 1847 (primary junior homonym)

Cyclocephala carbonaria Arrow, 1911

Mononidia trachypyga Prell, 1934 (synonym)

Mononidia punctulata Prell, 1934 (synonym)

Cyclocephala howdeni Endrödi, 1967 (synonym)

Cyclocephala complanata Burmeister, 1847

Cyclocephala obliquata Casey, 1915 (synonym)

Cyclocephala emacerata Casey, 1915 (synonym)

Cyclocephala concolor Burmeister, 1847

Cyclocephala confusa Endrödi, 1966

Cyclocephala conspicua Sharp, 1877

Cyclocephala conspicua gregaroides Dechambre, 1992 (synonym)

Cyclocephala conspicua fusca Dechambre, 1992 (synonym)

Cyclocephala curta Bates, 1888

Cyclocephala fusciventris Arrow, 1902 (synonym)

Cyclocephala deceptor Casey, 1915

Cyclocephala discolor (Herbst, 1792)

Melolontha unciata Schönherr, 1817 (synonym)

Cyclocephala aurantiaca Prell, 1937 (synonym)

Cyclocephala discolor andina Bréthes, 1904 (synonym)

Cyclocephala erotylina Arrow, 1914

Cyclocephala freudi Endrödi, 1963

Cyclocephala fulgurata Burmeister, 1847

Cyclocephala gravis Bates, 1888

Cyclocephala guttata Bates, 1888

Cyclocephala kaszabi Endrödi, 1964

Cyclocephala larssoni Endrödi, 1964

Cyclocephala ligyrina Bates, 1888

Cyclocephala lunulata Burmeister, 1847

Cyclocephala nubeculosa Burmeister, 1847 (synonym)

Graphalia oblita Casey, 1915 (synonym)

Cyclocephala maculiventris Höhne, 1923

Cyclocephala mafaffa Burmeister, 1847

Cyclocephala grandis Burmeister, 1847 (synonym)

Stigmalia cuernavacana Casey, 1915 (synonym)

Stigmalia fallaciosa Casey, 1915 (synonym)

Stigmalia deficiens Casey, 1915 (synonym)

Stigmalia mafaffa histrionica Casey, 1915 (synonym)

Cyclocephala melanocephala (Fabricius, 1775)

Melolontha leucophthalma Fischer von Waldheim, 1823 (synonym)

Melolontha ventralis Erichson, 1847 (synonym)

Cyclocephala dimidiata Burmeister, 1847 (synonym)

Cyclocephala elegans Horn, 1871 (synonym)

Dichromina ocularis Casey, 1915 (synonym)

Cyclocephala melolonthida Ratcliffe and Cave, 2002

Cyclocephala multiplex Casey, 1915

Cyclocephala mutata Harold, 1869

Cyclocephala laevicauda Arrow, 1902 (synonym)

Cyclocephala pseudisabellina Endrödi, 1980 (synonym)

Cyclocephala vitracelis Dechambre, 1999 (synonym)

Cyclocephala ovulum Bates, 1888

Cyclocephala picopijola Ratcliffe and Cave, 2006

Cyclocephala porioni Dechambre, 1979

Cyclocephala prolongata Arrow, 1902

Cyclocephala sanguinicollis Burmeister, 1847

Cyclocephala divisa Casey, 1915 (synonym)

Cycocephala politicauda Casey, 1915 (synonym)

Cyclocephala sexpunctata Laporte, 1840

Cyclocephala pubescens Erichson, 1847 (synonym)

Cyclocephala lucida Burmeister, 1847 (synonym)

Cyclocephala sexpunctata spermophila Ohaus, 1910 (synonym)

Stigmalia triangulifer Casey, 1915 (synonym)

Stigmalia discoidalis Casey, 1915 (synonym)

Stigmalia costaricana Casey, 1915 (synonym)

Stigmalia circulifer Casey, 1915 (synonym)

Cyclocephala pubescens nigripes Höhne 1923 (synonym)

Cyclocephala sororia Bates, 1888

Cyclocephala flava Dechambre, 1999 (synonym)

Cyclocephala sparsa Arrow, 1902

Cyclocephala landini Endrödi, 1964 (synonym)

Cyclocephala virkii Howden and Endrödi, 1966 (synonym)

Cyclocephala stictica Burmeister, 1847

Cyclocephala sexnotata Burmeister, 1847 (synonym)

Cyclocephala microspila Bates, 1888 (synonym)

Cyclocephala stictica bilineata Höhne, 1923 (synonym)

Cyclocephala tutilina Burmeister, 1847

Cyclocephala weidneri Endrödi, 1964

Dyscinetus dubius (Olivier, 1789)

Melolontha geminatus Fabricius, 1801 (synonym)

Geotrupes lugubris Quensel in Schönherr, 1806 (synonym)

Dyscinetus frater Bates, 1888 (synonym)

Dyscinetus obtusus Casey, 1915 (synonym)

Dyscinetus laevipunctatus Bates, 1888

Palechus histrio Casey, 1915 (synonym)

Erioscelis columbica Endrodi, 1966 Mimeoma acuta Arrow, 1902

Stenocrates bicarinatus Robinson, 1947

Stenocrates difficilis Endrödi, 1966 (synonym)

Stenocrates canuli Delgado, 1991

Stenocrates hardyi Dechambre, 1985

Stenocrates laevicollis Kirsch, 1871

Tribe Pentodontini

Bothynus complanus (Burmeister, 1847)

Bothynus simplicitarsis Bates, 1888 (synonym)

Bothynus monstrosus Bates, 1888 (synonym)

Bothynus quadridens (Taschenberg, 1870)

Euetheola bidentata (Burmeister, 1847)

Cyclocephala brevis Perty, 1830 (synonym? Type unknown)

Heteronychus globosus Burmeister, 1847 (synonym? Type unknown)

Euetheola humilis (Burmeister, 1847)

Ligyrus rugiceps LeConte, 1856 (synonym)

Dyscinetus parvus Casey, 1915 (synonym)

Dyscinetus hondurana Casey, 1915 (synonym)

Orizabus botox Ratcliffe and Cave,2006

Orizabus clunalis (LeConte, 1856)

Orizabus hondurensis Ratcliffe and Cave, 2006

Orizabus puchicus Ratcliffe and Cave, 2006

Orizabus teamscaraborum Ratcliffe and Cave,2006

Parapucaya amazonica Prell, 1934

Tomarus bituberculatus (Palisot de Beauvois, 1811)

Ligyrus latifovea Bates, 1888 (synonym)

Ligyrus maximus Arrow, 1913 (synonym)

Ligyrus latus Arrow, 1914 (synonym)

Tomarus cicatricosus (Prell, 1937)

Tomarus ebenus (Degeer, 1774)

Scarabaeus cordatus Fabricius, 1792 (synonym)

Cyclocephala scarabaeinus Perty, 1830 (synonym)

Tomarus fossor (Latreille, 1813)

Ligyrus castaneipennis Apolinar, 1927 (synonym)

Tomarus gyas Erichson, 1848

Ligyrus amazonicus Arrow, 1914 (synonym)

Tomarus laevicollis (Bates, 1888)

Ligyrus bryanti Rivers, 1891 (synonym)

Tomarus maternus (Prell, 1937)

Tomarus nasutus (Burmeister, 1847)

Ligyrus pygidialis Bates, 1888 (synonym)

Tomarus sallaei (Bates, 1888)

Ligyrodes propinquus Casey, 1915 (synonym)

Ligyrodes aztecus Casey, 1915 (synonym)

Tribe Oryctini

Coelosis biloba (Linnaeus, 1767)

Coelosis biloba lepesmei Bourgin, 1944 (invalid name)

Coelosis biloba incana Bourgin, 1944 (invalid name)

Coelosis biloba cacica Bourgin, 1944 (invalid name)

Coelosis biloba tibialis Bourgin, 1944 (synonym)

Coelosis biloba tibialis pauliani Bourgin, 1944 (invalid name)

Enema endymion Chevrolat, 1843

Enema lupercus Burmeister, 1847 (synonym)

Enema paniscus Burmeister, 1847 (synonym)

Enema gibbicollis Sternberg, 1908 (synonym)

Enema pan (Fabricius, 1775)

Scarabaeus chorinaeus Fabricius, 1775 (synonym)

Scarabaeus quadrispinosus Fabricius, 1781 (synonym)

Scarabaeus enema Fabricius, 1787 (synonym)

Scarabaeus aeneas Kirby, 1818 (synonym)

Scarabaeus titornus Perty, 1830 (synonym)

Enema lupercus Chevrolat, 1843 (synonym)

Enema infundibulum Burmeister, 1847 (synonym)

Heterogomphus chevrolati Burmeister, 1847

Heterogomphus chevrolati eurytus Bates, 1888 (synonym)

Heterogomphus chevrolati punctatissimus Prell, 1912 (synonym)

Heterogomphus chevrolati insignis Prell, 1912 (synonym).

Heterogomphus mniszechi (Thomson, 1859)

Daemonoplus pehlkei Kolbe 1906a: 273 (synonym)

Daemonoplus kolbei Prell 1912b: 155 (synonym)

Irazua dilicra Ratcliffe, 2003

Megaceras morpheus Burmeister, 1847

Megaceras ixyon Reiche, 1859 (synonym)

Megaceras septentrionis Bates, 1888

Megaceras septentrionis crassicornis Dechambre, 1975 (synonym)

Podischnus agenor (Olivier, 1789)

Scarabaeus barbicornis Latreille, 1812 (synonym)

Podischnus propinquus Prell, 1911 (synonym)

Strategus aloeus (Linnaeus, 1758).

Geotrupes semiramis Fabricius, 1801 (synonym)

Scarabaeus aesalus Laporte, 1840 (synonym)

Strategus julianus Burmeister, 1847 (synonym)

Strategus piosomus Kolbe, 1906 (synonym)

Strategus roosevelti Casey, 1915 (synonym)

Strategus frontalis Casey, 1915 (synonym)

Strategus tarsalis Casey, 1915 (synonym)

Strategus gaillardi Casey, 1915 (synonym)

Strategus hipposiderus Ratcliffe, 1976

Strategus jugurtha Burmeister, 1847

Strategus longichomperus Ratcliffe, 1976

Xyloryctes ensifer Bates, 1888

Xyloryctes lobicollis Bates, 1888

Xyloryctes splendidus Prell, 1914

Xyloryctes teuthras Bates, 1888

Tribe Phileurini

Amblyodus taurus Westwood, 1878

Amblyoproctus chalumeaui Endrödi, 1977

Archophileurus simplex (Bates, 1888) Goniophileurus femoratus (Burmeister, 1847)

Hemiphileurus cavei Ratcliffe, 2003

Hemiphileurus cylindroides (Bates, 1888)

Hemiphileurus costaricensis Endrödi, 1978 (synonym)

Hemiphileurus jamesonae Ratcliffe, 1988 (synonym)

Hemiphileurus dejeani (Bates, 1888)

Hemiphileurus euniceae Ratcliffe and Cave, 2006

Hemiphileurus laevicauda (Bates, 1888)

Hemiphileurus simplex (Prell, 1914)

Hemiphileurus fraternus Arrow, 1937 (synonym)

Hemiphileurus variolosus (Burmeister, 1847)

Epiphileurus irregularis Prell, 1914 (synonym)

Hemiphileurus variolosus striatus Endrödi, 1978 (synonym)

Homophileurus luedeckei Kolbe, 1910

Homophileurus quadrituberculatus (Palisot de Beauvois, 1806)

Scarabaeus bajulus Perty, 1830 (synonym)

Phileurus cephalotes Laporte, 1840 (synonym)

Homophileurus muticus Prell, 1914 (synonym)

Homophileurus tricuspis Prell, 1914

Phileurus carinatus Prell, 1914

Phileurus carinatus declivis Prell, 1914 (synonym)

Phileurus didymus (Linnaeus, 1758)

Phileurus affinis Reiche, 1859 (synonym)

Phileurus truncatus Palisot de Beauvois, 1807

Phileurus recurvatus Casey, 1915 (synonym)

Phileurus valgus (Olivier, 1789)

Scarabaeus castaneus Haldeman, 1843 (synonym)

Phileurus capra Bates, 1888 (synonym)

Phileurus valgus septentrionis Kolbe, 1910 (synonym)

Phileurus meridionalis Kolbe, 1910 (synonym)

Phileurus valgus antillarum Prell, 1912 (subspecies)

Phileurus texensis Casey, 1915 (synonym)

Phileurus carolinae Casey, 1915 (synonym)

Phileurus sulcifer Casey, 1915 (synonym)

Phileurus floridanus Casey, 1915 (synonym)

Phileurus clathratus Casey, 1915 (synonym)

Phileurus voirinae Endrödi, 1985

Phileurus youngi Ratcliffe, 1988

Tribe Agaocephalini

Spodistes beltianus (Bates, 1888)

Lycomedes reichii Burmeister, 1847 (synonym)

Spodistes mniszechi (Thomson, 1860)

Spodistes monzoni Warner, 1992

Tribe Dynastini

Dynastes hercules (Linnaeus, 1758)

Scarabaeus scaber Linnaeus, 1764 (synonym)

Scarabaeus hercules oculatus Scopoli, 1772 (synonym)

Scarabaeus alcides Fabricius, 1781 (synonym)

Scarabaeus iphiclus Panzer, 1782 (synonym)

Scarabaeus perseus Olivier, 1789 (synonym)

Dynastes lagaii Verrill, 1906 (synonym)

Dynastes vulcan Verrill, 1906 (synonym)

Dynastes argentatus Verrill, 1907 (synonym)

Dynastes hercules ecuatorianus Ohaus, 1913 (synonym)

Dynastes hercules niger Endrödi, 1947 (synonym)

Dynastes hercules baudrii Pinchon, 1976 (synonym)

Dynastes hercules reidi Chalumeau, 1977 (synonym)

Dynastes hercules septentrionalis Lachaume, 1985 (synonym)

Dynastes hercules occidentalis Lachaume, 1985 (synonym)

Dynastes hercules lichyi Lachaume, 1985 (synonym)

Dynastes hercules paschoali Grossi and Arnaud, 1993 (synonym)

Dynastes hercules tuxtlaensis Morón, 1993 (synonym)

Dynastes hercules bleuzeni Silvestre and Dechambre, 1995 (synonym)

Dynastes hercules trinidadensis Chalumeau and Reid, 1995 (synonym)

Dynastes hercules morishimai Nagai, 2002 (synonym)

Dynastes hercules takakuwai Nagai, 2002 (synonym)

Dynastes maya Hardy, 2003

Golofa pizarro Hope, 1837

Golofa hastata Burmeister, 1847 (synonym)

Golofa imperialis Thomson, 1858 (synonym)

Golofa sallei Thomson, 1860 (synonym)

Golofa clavicornis Thomson, 1860 (synonym)

Golofa tersander Burmeister, 1847

Mixigenus leander Thomson, 1859 (synonym)

Mixigenus barbicornis Fairmaire, 1878 (synonym)

Golofa dohrni Nonfried 1890, (synonym)

Megasoma elephas (Fabricius, 1775)

Megasoma mexicanum Fischer, 1968 (synonym)

Megasoma elephas iijimai Nagai, 2003 (synonym)

Sitios de Colecta

Portada del Libro Los Dynastinae de

Honduras, Nicaragua y El Salvador

The Dynastine Scarab Beetles of Honduras, Nicaragua, and El Salvador

(Coleoptera: Scarabaeidae: Dynastinae)

by Brett C. Ratcliffe

and Ronald D. Cave 2006

Bulletin of the University of Nebraska State Museum 21: 1-424, 6 color plates, 3

color maps, 645 illustrations

Resumen.

Se realizó una revisión comprensiva de las 116 especies de escarabajos dinástinos que

ocurren en Honduras, Nicaragua, y El Salvador. Se proveen claves, descripciones,

distribuciones y notas sobre la biología de todas las especies, también se presentan

ilustraciones y mapas. Además, se incluyen sinópsis de los taxones de niveles

superiores, un glosario, un listado de las localidades de colecta y un listado de las

especies.

Se describen las siguientes especies nuevas: Cyclocephala picopijola, Orizabus botox,

O. hondurensis, O. puchicus, O. teamscaraborum, y Hemiphileurus euniceae.

Se reconocen los siguientes sinónimos: Cyclocephala flava Dechambre (= C. sororia

Bates); Golofa imperialis Thomson (= G. pizarro Hope); y Megasoma elephas iijimai

Nagai (= M. elephas [Fabricius]).

Los siguientes NUEVOS REGISTROS DE PAIS de Cyclocephalini se reportan por

primera vez en el área de estudio:

El Salvador: Cyclocephala confusa Endrödi, Stenocrates bicarinatus Robinson, y S.

canuli Delgado.

Honduras: Ancognatha sellata Arrow, Cyclocephala alexi Ratcliffe y Delgado, C.

atripes Bates, C. batesi Delgado y Castañeda, C. confusa Endrödi, C. curta Bates, C.

erotylina Arrow, C. kaszabi Endrödi, C. ligyrina Bates, C. maculiventris Höhne, C.

mutata Harold, Stenocrates canuli Delgado, y S. laevicollis Kirsch.

Nicaragua: Ancognatha sellata Arrow, Cyclocephala confusa Endrödi, C. erotylina

Arrow, C. guttata Bates, C. kaszabi Endrödi, C. larssoni Endrödi, C. maculiventris

Höhne, C. mutata Harold, C. tutilina Burmeister, Erioscelis columbica Endrödi,

Stenocrates bicarinatus Robinson, S. canuli Delgado, S. hardyi Dechambre, y S.

laevicollis Kirsch.

Las siguientes especies han sido reportadas en el área de estudio, pero no han sido

recolectadas desde entonces, o se reportan tan alejadas de sus rangos definitivos de

distribución que se las consideran registros erróneos o de algún modo sospechosos:

Cyclocephala amazona (L.), C. laminata Burmeister, C. obesa Burmeister, y C.

paraguayensis Arrow.

Los siguientes NUEVOS REGISTROS DE PAIS de Pentodontini se reportan por

primera vez en el área de estudio:

El Salvador: Euetheola bidentata (Burmeister), Orizabus clunalis (LeConte), Tomarus

cicatricosus (Prell), T. fossor (Latreille), y T. laevicollis (Bates).

Honduras: Euetheola bidentata (Burmeister), Orizabus clunalis (LeConte), Tomarus

cicatricosus (Prell), y T. maternus (Prell).

Nicaragua: Euetheola bidentata Burmeister, Tomarus cicatricosus (Prell), T. laevicollis

(Bates), y T. maternus (Prell).

Los siguientes NUEVOS REGISTROS DE PAIS de Oryctini se reportan por primera

vez en el área de estudio:

El Salvador: Xyloryctes ensifer Bates.

Honduras: Enema pan (Fabr.), Megaceras morpheus Burmeister, Xyloryctes ensifer

Bates, X. splendidus Prell, y X. teuthras Bates.

Nicaragua: Megaceras septentrionis Bates.

Los siguientes NUEVOS REGISTROS DE PAIS de Phileurini se reportan por primera

vez en el área de estudio:

El Salvador: Goniophileurus femoratus (Burmeister), Hemiphileurus cylindroides

(Bates), Homophileurus luedeckei Kolbe, y Homophileurus tricuspis Prell.

Honduras: Goniophileurus femoratus (Burmeister), Hemiphileurus dejeani (Bates),

Hemiphileurus laevicauda (Bates), Hemiphileurus variolosus (Burmeister),

Homophileurus tricuspis Prell, Phileurus carinatus Prell, P. truncatus Palisot de

Beauvois, P. voirinae Endrödi, y P. youngi Ratcliffe.

Nicaragua: Amblyoproctus chalumeaui Endrödi, Hemiphileurus cylindroides (Bates),

Hemiphileurus dejeani (Bates), Hemiphileurus laevicauda (Bates), Hemiphileurus

simplex (Prell), Hemiphileurus variolosus (Burmeister), y Phileurus youngi Ratcliffe.

Las siguientes NUEVOS REGISTROS DE PAIS de Dynastini se reportan por primera

vez en el área de estudio:

El Salvador: Golofa tersander Burmeister.

Honduras: Dynastes maya Hardy.

Literatura consultada

Ratcliffe, B. C. and R. D. Cave. 2006. The Dynastine Scarab Beetles of

Honduras, Nicaragua, and El Salvador. Bulletin Univ. Nebraska State Museum 21:

1-500.