115
8388
55401749 9435
Depth
(cm)
Schoe
nople
ctus
ac
utus /
S. tabe
rnaem
ontan
i
seed Naja
s flex
ilis
se
ed Menya
nthes
trif
oliata
seed Tri
gloch
in
mari
tima
seed Bide
ns
ce
rnua
seed Sag
ittaria
lat
ifolia
seed Com
arum
pa
lustre
seed Hyp
ericu
m s.l. s
p.
se
edNymph
aea
od
orata
ssp.
tubero
sa
se
ed Nymph
aea
od
orata
seedPota
mogeto
n sp.
se
edCarex r
ostra
ta
se
ed Carex s
pp. (g
roup 2
)
with
plan
ocon
vex s
eed
Carex s
pp. (g
roup 1
)
with
trigo
nous
seed
Eleoch
aris s
p.
se
ed Dulich
ium
aru
ndina
ceum
seed
Cenoc
occu
m
ge
ophil
um
scler
otium
Cristat
ella
muc
edo
statob
last
Erioph
orum sp
.
se
ed
Charco
al
pie
cePtyc
hosto
mum s.
l.
ste
mSph
agnu
m spp.
ste
mBetu
la
pa
pyrife
ra
seed Myri
ca
ga
le
leaf, s
eed
Abies
ba
lsamea
leaf La
rix
lar
icina
leaf, s
eed
Chamae
daph
ne
ca
lycula
ta
seed And
romed
a
po
lifolia
var. l
atifol
ia
leaf, s
eed
Vaccin
ium cf
.
m
acroc
arpon
leaf, s
eed
Ericac
eae s
pp.
lea
f
200
150
100
50
0
200
150
100
50
0
200
150
100
50
0
MOSSES TREES and SHRUBS ERICACEOUS SHRUBS MYCORHIZAE
CYPERACEAE FORBS
BRYOZOA CHARCOAL
0 2000
0 20 0 20 0 20 0 20 0 20 0 20 0 20 0 200 0 20 0 20 0 200 20 0 20 40 0 20 0 20 0 20 0 20
0 20 0 20 40 60 0 20
40 60
0 200 0 2000 4000 6000 0 20 0 200 400 600 800 0 20 0 200 400 0 20 0 2000 4000 0 20 0 2040 60
Amblyste
giace
ae or
stem
Callieg
onac
eae s
pp.
pseu
dotriq
uetru
m
99
200
150
100
50
0
200
150
100
50
0
200
150
100
50
0Macrof
ossil
zo
ne
II
I
Stratig
raphy
2421 15+-
Date (c
al. yr
BP)
8037 20+-
2762 15+-
II
I
2421 15+-
8037 20+-
2762 15+-
b
48
13 120
11 990
1171
1416 16 033
Depth
(cm)
Sphag
num sp
p.
ste
m Amblyste
giace
ae or
ste
m Ptycho
stomum
s.l.
pseu
dotriq
uetru
m
ste
m Betula
pa
pyrife
ra
seed La
rix
lar
icina
leaf, s
eed
Picea s
p.
lea
f, see
d
Myrica
gale
lea
f, see
d
Abies
ba
lsamea
leaf
200
150
100
50
0
200
150
100
50
0
MOSSES TREES and SHRUBS
0 2000 0 2000 0 2000 4000 60000 20 0 200 20 0 200 200 4004000 6000
Callier
gona
ceae
spp.
13 831
Rhodo
dend
ron
gro
enlan
dicum
leaf And
romed
a
po
lifolia
var.
la
tifolia
leaf, s
eed
Chamae
daph
ne
ca
lycula
ta
leaf, s
eed
Vaccin
ium cf
.
macroc
arpon
leaf, s
eed
Ericac
eae s
pp.
lea
f
200
150
100
50
0
ERICACEOUS SHRUBS
0 20 40 0 20 0 20 0 2000 0 2000
Cenoc
occu
m
ge
ophil
um
scler
otium
Cristat
ella
muc
edo
statob
last
Charco
al
pie
ce
200
150
100
50
0
MYCORHIZAE BRYOZOA CHARCOAL
0 20 0 20 40 0 20
Carex s
pp.
(gr
oup 1
)
with tri
gono
us se
ed
Carex r
ostra
ta
se
edEleo
chari
s sp.
se
ed Dulich
ium
aru
ndina
ceum
seed Care
x spp
.
(gr
oup 2
)
with pl
anoc
onve
x
s
eed
200
150
100
50
0
CYPERACEAE
0 20 0 20 0 20 40 60 80 0 200 200 20
Schoe
nople
ctus
ac
utus /
S. tabe
rnaem
ontan
i
seed
Najas
fle
xilis
seed Sag
ittaria
lat
ifolia
seed Men
yanth
es
trif
oliata
seed Bide
ns
ce
rnua
seed Call
a
pa
lustris
seedPoly
gonu
m sp.
se
edSpa
rganiu
m sp.
se
edCom
arum pa
lustre
se
edPota
mogeto
n sp.
se
edHyp
ericu
m s.l. s
p.
se
ed
200
150
100
50
0
FORBS
0 20 40 0 20 0 20 0 20 0 20 0 20 0 20 0 20 0 20 0 20
Date (c
al. yr
BP)
Stra
tigrap
hy
8027 20+-
3595 15+-
3513 15+-
3198 15+-
Macrof
ossil
zo
ne
II
I
8027 20+-
3595 15+-
3513 15+-
3198 15+-
II
I
Moderately decomposed peat with moss and sedge
Moderately decomposed peat with moss and wood remains
Gyttja
Sand and clay
a
Legend
Number of macrofossil pieces / 100 cm3 of peat
Macrofossil diagrams of the Bic-Saint-Fabien BSF peatland at (a) R1, (b) R2, (c) R3 and (d) N. The dotted lines mark the different vegetation zones.
26 552
16 820
12 608
26 552
13 028
76
35 868
40 852
19820
20350
76
556
Depth
(cm)
Pseud
ocall
iergo
n
trif
arium
stemCam
pyliu
m
ste
llatum
stem
Callier
gon s
.l. sp
.
ste
mAmblyste
giace
ae or
ste
mPtyc
hosto
mum s.
l.
ste
mDrep
anoc
ladus
s.l. s
p.
ste
m Mnium sp
.
ste
mPalu
della
sq
uarro
sa
stem
Abies
ba
lsamea
leaf
Betula
pa
yrifer
a
seedLa
rix
lar
icina
leaf, s
eed
Najas
fle
xilis
seed Sag
ittaria
lat
ifolia
seed Bide
ns sp
.
se
edMen
yanth
es
trif
oliata
seed Sarr
acen
ia
pu
rpurea
seed Pota
mogeto
n sp.
se
edSpa
rganiu
m cf.
mini
mum
seed Tri
ental
is
bo
realis
seed Ly
simac
hia
thy
rsiflo
ra
seed Tri
gloch
in
mari
tima
seed Poly
gonu
m sp.
se
ed
Carex s
pp. (g
roup 2
)
with
plan
ocon
vex s
eed
Eleoch
aris
ov
ata
seed Duli
chium
aru
ndina
ceum
seed Tri
chop
horum
alp
inum
seed Rhy
ncho
spora
alb
a
seed Care
x
ros
trata
seed Cen
ococ
cum
ge
ophil
um
scler
otium
Cristat
ella
muc
edo
statob
last
Charco
al
pie
ceEleo
chari
s cf.
tenu
is
seed
Myrica
ga
le
leaf, s
eed
Picea s
p.
lea
f, see
d
Thuja
oc
ciden
talis
leaf
And
romed
a
polifo
lia va
r.
la
tifolia
leaf,
seed
Chamae
daph
ne
ca
lycula
ta
seedEric
acea
e spp
.
lea
fKalm
ia
po
lifolia
leaf
Vaccin
ium cf
.
mac
rocarp
on
leaf, s
eed
Sphag
num sp
.
ste
m
350
300
250
200
150
100
50
0
MOSSES
TREES and SHRUBS ERICACEOUS SHRUBS
MYCORHIZAECYPERACEAE BRYOZOA CHARCOAL
FORBS
0 2000 4000 6000 8000 10000 12000
0 2000 4000 6000 8000 10000 0 2000 4000 6000 8000 10000 12000
0 2000 4000 6000 8000 10000 0 2000 0 20 0 204000 6000 0 2000 4000 6000 0 2000 4000 6000 0 2000 4000 6000 0 2000 4000 6000 80008000 10000
350
300
250
200
150
100
50
0
350
300
250
200
150
100
50
0
350
300
250
200
150
100
50
0
350
300
250
200
150
100
50
0
350
300
250
200
150
100
50
0
0 20 0 200 20 0 20 0 20 0 20 40 60 800 200 0 200 0 200400
0 200
0 200 0 20 0 20 0 20 40 0 20 0 20 0 20 0 20 0 20 40 0 200 200 400
0 200 0 20 0 20 0 20 40 0 20 0 20 0 20 0 200 2000 4000 0 2000 4000 6000 8000 10000
Callier
gona
ceae
spp.
pseu
dotriq
uetru
m
Schoe
nople
ctus
ac
utus /
S. ta
berna
emon
tani
seed
Stratig
raphy
Macrof
ossil
zo
ne
III
IV
II
I
Date (c
al. yr
BP)
4422 20+-
2665 15+-
719 15
6734 20+-
4870 20+-
+-
III
IV
II
I
4422 20+-
2665 15+-
719 15
6734 20+-
4870 20+-
+-
III
IV
II
I
4422 20+-
2665 15+-
719 15
6734 20+-
4870 20+-
+-
III
IV
II
I
4422 20+-
2665 15+-
719 15
6734 20+-
4870 20+-
+-
c
Well humified peat with wood remains
Moderately decomposed peat with moss and sedge
Gyttja
Sand and clay
Legend
Number of macrofossil pieces / 100 cm3 of peat
65 272 28 336
Carex c
f.
dian
dra
see
dEleo
chari
s cf.
obtus
a
see
d
Betula
pa
pyrife
ra
see
d
Mosse
s
(v
ariou
s
spe
cies)
stem Call
iergo
n s.l.
sp.
ste
mPtyc
hosto
mum s.
l.
pseudotriq
uetrum
ste
mStra
minergo
n cf.
stra
mineum
stem Drep
anoc
ladus
s.l. s
p.
ste
mPalu
della
sq
uarro
sa
stem Ambly
stegia
ceae
or
ste
mSph
agnu
m
se
ction A
cutifo
lia
stemSph
agnu
m
se
ction C
uspid
ata
stem Sph
agnu
m spp.
ste
mCall
iergo
n
gi
gante
um
stem Cam
pyliu
m
st
ellatu
m
stem Sco
rpidiu
m
co
sson
ii
s
tem
Abies
ba
lsamea
leaf La
rix
la
ricina
con
e, lea
f, see
d
Myrica
ga
le
leaf,
seed
Picea s
p.
lea
f, see
d
Alnus i
ncan
a
ss
p. rug
osa
see
dThu
ja
oc
ciden
talis
leaf,
seed
, stem
Vaccin
ium sp
.
se
edAnd
romed
a
po
lifolia
var.
la
tifolia
carp
el, le
af, se
ed
Ericac
eae s
pp.
lea
fCha
maeda
phne
ca
lycula
ta
carpe
l, see
d
Kalmia
an
gusti
folia
see
dVac
cinium
cf.
mac
rocarp
on
leaf Vac
cinium
ox
ycoc
cos
leaf,
seed
Cenoc
occu
m
ge
ophil
um
scle
rotium
Cristat
ella
m
uced
o
stat
oblas
t
Charco
al
pi
ece
Carex s
pp. (g
roup 2
)
with
plan
ocon
vex s
eed
Carex s
pp. (g
roup 1
)
with
trigo
nous
seed
Eleoch
aris c
f.
tenu
is
see
dTri
chop
horum
al
pinum
see
dRhy
ncho
spora
al
ba
see
dTri
chop
horum
ce
spito
sus
see
dEleo
chari
s sp.
se
edCare
x ros
trata
se
ed
0 200 0 200 400500
450
400
350
300
250
200
150
100
50
0
500
450
400
350
300
250
200
150
100
50
0
0 200 400
754
97
64
9276 12 828
12 431
101
0 20 0 20 0 20 0 20 40 0 20 0 20 0 20 0 20 0 200 20 40 60 0 20 40
MYCORHIZAECYPERACEAE
TREES and SHRUBS
MOSSES
FORBS
ERICACEOUS SHRUBS
BRYOZOA CHARCOAL
500
450
400
350
300
250
200
150
100
50
0
500
450
400
350
300
250
200
150
100
50
0
500
450
400
350
300
250
200
150
100
50
0
500
450
400
350
300
250
200
150
100
50
0
0 20
0 20 0 200 0 20 0 20
Dulich
ium
ar
undin
aceu
m
see
d
0 20
0 2000 0 02000 2000 4000 6000 8000 10000 12000 0 0 200 0 2002000 4000 6000 8000 0 2000 4000 6000 0 02000 2004000 6000 8000 1000010000 12000 14000400040
0 20 0 20 0 20 40 0 20 0 20 400 20 0 200 200 200 0 200 4000 0 2000 4000 6000 8000 0 2000 4000 6000 800020
88
0 20 40 60 80 0 20 0 20 40 0 200 200 0 20 0 20 0 20 0 20 0 20 0 20
Najas
fle
xilis
see
dPota
mogeto
n sp.
se
edSag
ittaria
la
tifolia
see
dMen
yanth
es
tri
foliat
a
see
dSarr
acen
ia
pu
rpurea
see
dBide
ns sp
.
se
edLy
simac
hia
th
yrsiflo
ra
see
dSpa
rganiu
m sp.
se
edTri
gloch
in
m
aritim
a
see
dPoly
goniu
m sp.
se
edCall
a palu
stris
se
ed
Depth
(cm)
Callierg
onac
eae s
pp.
Schoe
nople
ctus
ac
utus /
S. ta
berna
emon
tani
seed
Stratig
raphy
Macrof
ossil
zo
ne
IV
III
II
I
Date (c
al. yr
BP)
7090 20+-
5497 20+-
4944 15 +-
4182 15+-
1269 15 +-
IV
III
II
I
7090 20+-
5497 20+-
4944 15 +-
4182 15+-
1269 15 +-
IV
III
II
I
7090 20+-
5497 20+-
4944 15 +-
4182 15+-
1269 15 +-
IV
III
II
I
7090 20+-
5497 20+-
4944 15 +-
4182 15+-
1269 15 +- Well humified peat with wood remains
Moderately decomposed peat with moss and sedge
Moderately decomposed peat with moss and wood remains
Gyttja
Sand and clay
d
Legend
Number of macrofossil pieces / 100 cm3 of peat
