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Residence Time of Coarse Woody Debris on the Forest Floor in Southeastern Indiana Forests
M. Ross Alexander and Darrin L. RubinoBiology Department, Hanover College, Hanover, IN
What is CWD?
• Downed branches and boles – greater than 10 cm
diameter – longer than 1 m in length
• Divided into classes based on degree of rot (punkiness)
• Why?
Decay Classes
• Common classes defined by Pyle and Brown (1998)
• 5 classes– Class I
Freshly fallen logs
– Class II Bark still present, solid
– Class IIIBark typically absent,
wood: Spongy, flaky
– Class IV• No longer solid• Crushes when struck (no
flaking)
– Class V• Predominantly powder
wood• Typically flattened shape
Goals
• Determine death dates of logs within decay classes
• Determine how long different species reside on forest floor
• Are there a differences in the decay rates of different hardwood species?
Methods
• Several forests in Southeastern Indiana were sampled.
• Purposefully chose samples that had either bark or wane.• Wane = outermost growth ring
• “Cookie” samples were taken using a chain saw
Surface Preparation
• Sanding– Finer grits of sandpaper
– 50, 80, 120, 180, 220, 320, 400, 600
– Table sanders– Palm (orbital) sanders– Hand sanding
1
Dating
• Place arbitrary years on sanded sample• Skeleton plots
Dating
• Measure sample
• Allows for “Crossdating”
19901980
1971 – 2003
Crossdating Process
19901980
? – ?
1971 – 2003
19901980
1971 – 2003
? – ?
1990
1960 1970
1980
1971 – 2003
1953 – 1979
Results
• 88 samples from 8 species (LITU, FRAX, SAAL, FAGR, ROPS, Red and White Oak, PRSE, ) were analyzed
• ANOVA analysis was performed where appropriate
• Analysis was not performed on PRSE, SAAL, or the Oaks
Class n
I 29
II 13
III 39
IV 7
I II III0
1
2
3
4
5
Oaks
Decay Class
Num
ber o
f sam
ples
I III0
2
4
6
LITU
Decay ClassNum
ber o
f Sam
ples
I II III05
101520
ROPS
Decay Class
Num
ber o
f Sam
ples
I II III IV05
101520
FAGR
Decay Class
Num
ber o
f Sam
ples
I II III0123456789
FRAX
Decay Class
Num
ber o
f Sam
ples
Beech
I II III IV0
2
4
6
8
10
12
Decay Class
Mea
n Ye
ars
dead
a
c c
b
F = 19.07P < 0.001n = 40
Decay Class III
FAGR FRAX LITU ROPS0
5
10
15
20
25
Species
Mea
n Ye
ars
Dea
d
ab
a
b
F = 5.70P = 0.03n = 35
a
Conclusions
• Decay rates are different from species to species
• More sampling is needed to increase depth
• Future work: Chemical analysis to determine nutrient release among different species.
Acknowledgements
• Biology Dept. at Hanover College• Patricia Walne Student Research Fund• Cassie Lothery• Abby Simpson