Siebold and Zuccarini’s Type Specimens and Original ...—279— Siebold and Zuccarini’s Type...

—279—

Siebold and Zuccarini’s Type Specimens and Original Materials from Japan, Part 5. Angiosperms. Dicotyledoneae 4

Shinobu Akiyamaa,*, Gerard Thijsseb,†, Hans-Joachim Esserc

and Hideaki Ohbad

aDepartment of Botany, National Museum of Nature and Science,4-1-1, Amakubo, Tsukuba, 305-0005 JAPAN;

bNationaal Herbarium Nederland, Naturalis Biodiversity Center,P. O. Box 9514, THE NETHERLANDS;

†Present address: Department of Botany, Naturalis Biodiversity Center, P. O. Box 9517, THE NETHERLANDS;cBotanische Staatssammlung München,

Menzinger Straße 67, D-80638 München, GERMANY;dDepartment of Botany, the University Museum, the University of Tokyo,

7-3-1, Hongo, Tokyo, 113-0033 JAPAN*Corresponding author: [email protected]

(Accepted on May 10, 2014)

This is part 5 of the list of type specimens and original materials of the taxa described by Siebold and Zuccarini from Japan. Part 5 concerns the taxa belonging to the families ranging from Saxifragaceae to Euphorbiaceae according to the 12th Engler system (Melchior 1964). The followings five new combination names are proposed. These are: Amygdalus persica L. f. pendula (Siebold) H. Ohba & S. Akiyama, stat. nov., Amygdalus persica L. f. stellata (Siebold & de Vriese) H. Ohba & S. Akiyama, stat. nov., Amygdalus persica L. f. versicolor (Siebold) H. Ohba & S. Akiyama, comb. et stat. nov., Armeniaca mume (Siebold & Zucc.) K. Koch f. pendula (Siebold) H. Ohba & S. Akiyama, comb. et stat. nov., Cerasus itosakura (Siebold) H. Ohba & S. Akiyama, comb. nov.

(Continued from J. Jpn. Bot. 89: 209–229, 2014)

Key words: Japanese flora, lectotypification, Siebold collections, Zuccarini.

J. Jpn. Bot. 89: 279–330 (2014)

SAxifRAGACEAECardiandra Siebold & Zucc., fl. Jap. 1: 119 (1835); in Abh. Math.-Phys. Cl. Königl. Bayer. Akad. Wiss. 4(2): 192 (1845).

Type: Cardiandra alternifolia (Siebold) Siebold & Zucc.

Present treatment: Cardiandra Siebold & Zucc.

Deutzia Thunb.184) Deutzia crenata Siebold & Zucc., fl.

Jap. 1: 19, t. 6 (1835); in Abh. Math.-Phys. Cl. Königl. Bayer. Akad. Wiss. 4(2): 192 (1845).

Present treatment: Deutzia crenata Siebold & Zucc.

Japanese name: Utsugi.Lectotype (designated here): Japonia. Bürger

(L0170287) [Category 5 (fl. Jap. 1: t. 6); the branch on the far right corresponds with the plant illustrated in Table 6 of Flora Japonica] (fig. 116).

Original material: Herbar. de Siebold

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coll. 1829 (L0170284, excl. central flowering branch) [Category 2]. Legit in Japonia Bürger. Communicavit v. Siebold, Herbarium Zuccarinii (M0120974) [Category 2]. Japonia. [Siebold s.n.] (MAK S0889) [Category 2].

The specimen designated here as the lectotype of Deutzia crenata Siebold & Zucc. was not annotated by Siebold and/or Zuccarini, but the branch on the far right corresponds particularly well with the plant illustrated in Table 6 of their Flora Japonica under D. crenata. Both the far right branch of the specimen and the branch in the illustration have widely to moderately ovate petiolate leaves with an acute apex and widely cuneate to truncate or sometimes shallowly cordate base and filaments with terminal teeth, and are within the range of D. crenata.

Siebold and Zuccarini distinguished Deutzia crenata from D. scabra Thunb. by 1) the shape of leaves, i.e. short petiolate, widely ovate with acute apex, rounded or subcordate base, lightly or finely “tenuiter” crenulate margins vs. subsessile or very short petiolate, ovate with acuminate apex, rounded or subcordate base, serrulate margins, 2) stellate hairs on upper surface with 4–6 vs. 3 or 4 arms, and 3) filaments with two terminal teeth vs. without teeth. Except for the presence of teeth on the filaments these two species are not clearly distinguished by the features given by them.

Usually D. crenata is distinguished by the petiolate leaves (usually oblong with acute or acuminate apex) and dentate filaments; D. scabra has sessile leaves (usually ovate with obtuse apex) on flowering branches and toothless filaments. In and around Nagasaki the range of variation of the leaves, especially in the shape and length of the petioles, and also teeth on the filaments, overlaps between the two species.

in D. scabra the leaves are usually ovate or orbicular-ovate or widely oblong-ovate, although in the plants around Nagasaki the leaves show a tendency toward being oblong to oblong-ovate, as illustrated in Table 7 of Flora Japonica. As pointed by Hara (1957), these plants belong to

a local variety, var. sieboldiana (Maxim.) H. Hara (D. sieboldii Koernicke) with different tendencies from var. scabra in leaf shape and teeth of filaments.

A specimen collected by Siebold in 1829, L0170284, consists of two species, D. crenata and D. scabra, i.e., a branch with flowers mounted in the central part of the sheet surrounded by an elongate branch of D. crenata corresponds well with the drawing in Table 7 of Flora Japonica under D. scabra, which is D. scabra var. sieboldiana.

One of the original specimens collected by Bürger, M0120974, consists of two branches. Both branches have narrowly oblong-ovate leaves (with acute or acuminate apex and truncate or widely rounded base), which are sessile or nearly so, except the uppermost ones on the flowering branches. The right hand branch has a paniculate inflorescence somewhat transitional to a pyramidal inflorescence with numerous flowers. The filaments have two rather distinct teeth at the apex. We consider these plants to be D. crenata, but doubt whether they might be Deutzia scabra var. sieboldiana still remains.

185) Deutzia gracilis Siebold & Zucc., fl. Jap. 1: 22, t. 8 (1835); in Abh. Math.-Phys. Cl. Königl. Bayer. Akad. Wiss. 4(2): 192 (1845).

Present treatment: Deutzia gracilis Siebold & Zucc.

Japanese name: Hime-utsugi.Lectotype (designated here): Japonia.

Herbar. de Siebold coll. 1829 (L0176009) [Category 2] (fig. 117).

Original material: Legit Bürger in Japonia. Communicavit v. Siebold, Herbarium Zuccarinii (M0120975) [Category 2]. Japonia. Herb. Zuccarinii (M0120976) [Category 2]. Japonia. [Siebold s.n.] (MAK S0890) [Category 2].

Hoteia C. Morren & Decne.186) Hoteia thunbergii Siebold & Zucc.

in Abh. Math.-Phys. Cl. Königl. Bayer. Akad. Wiss. 4(2): 191 (1845).

October 2014 TheJournal of Japanese Botany Vol. 89 No. 5 281

Present treatment: Astilbe thunbergii (Siebold & Zucc.) Miq., Ann. Mus. Bot. Lugd.-Bat. 3: 96 (1867).

Japanese name: Aka-shôma.Lectotype (designated here): Legit in

Japonia & Communicavit de Siebold anno 1842, Herbarium Zuccarinii (M0120971) [Category 2] (fig. 118).

Original material: in Japonia legit Bürger. Communicavit d. Siebold, Herb. Zuccarinii (M0120970) [Category 2b]. Japonia. [Siebold s.n.] (MAK S0408) [Category 2].

Siebold and Zuccarini published Hoteia thunbergii with a description based on their materials. Although they cited Spiraea aruncus of Thunberg’s Flora Japonica (1784, p. 211), there is no indication that they studied Thunberg’s specimens in UPS.

Hydrangea L. (see Ohba and Akiyama 2013a, 2013b, 2014).

187) Hydrangea acuminata Siebold & Zucc., fl. Jap. 1: 110, t. 56, 57 i (1839).

Present treatment: Hydrangea serrata (Thunb.) Ser. in DC., Prodr. 4: 15 (1830) (var. serrata).

Japanese name: Yama-ajisai. Lectotype (Ohba and Akiyama 2013b):

Japonia. Herb. Siebold (L0043373).Original material: Legit in Japonia et

Communicavit d. Siebold, Herbarium Zuccarinii (M0153946) [Category 2]. Legit Dr Siebold in Japonia. Communicavit Prof. Zuccarini anno 1843, Herbarium Zuccarinii (M0153945) [Category 2b]. in Japonia legit et Communicavit Siebold, Herb. Zuccarinii (M0153948) [Category 2b].

188) Hydrangea acuminata Siebold & Zucc. var. buergeri Siebold & Zucc., fl. Jap. 1: 111, t. 57 ii (1839), as “Bürgeri.”

Present treatment: Hydrangea serrata (Thunb.) Ser. in DC., Prodr. 4: 15 (1830) (var. serrata).

Japanese name: Yama-ajisai. Lectotype (Ohba and Akiyama 2013b):

Japonia. Herb. Burger (Communicavit Bürger, Hb. Siebold) (L0103049) [Category 2, 5 (left- hand branch, fl. Jap. 1: t. 57 ii)].

Ohba and Akiyama (2013b) recognized Hydrangea buergeri Siebold & Zucc. as a variety of H. acuminata because in the text Siebold wrote: Occurrit varietas foliis robustioribus longioribusque ...

189) Hydrangea alternifolia Siebold in Nova Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. Cur. 14(2): 692 (1828).

Present treatment: Cardiandra alternifolia (Siebold) Siebold & Zucc., fl. Jap. 1: t. 65 (1839); 121, t. 66 (1840, versim.).

Japanese name: Kusa-ajisai; Kusasimots’ke (Siebold 1828), Kusa-Kaku (Siebold).

Lectotype (designated here): Japonia. Hb. itoo Keiske ex Owari (L0420368) [Category 2, 3] (fig. 119).

Original material: Japonia. Communicavit Dr. Bürger, Hb. v Siebold (L0043362) [Category 2]. Legit in Japonia et Communicavit Siebold 1839, Herbarium Zuccarinii (M0120979) [Category 2]. Legit in Japonia & Communicavit de Siebold (M0120981) [Category 2b]. in Japonia legit Bürger. Communicavit d. Siebold (M0120980) [Category 2b].

Hydrangea alternifolia was originally described by Siebold in 1828 and transferred to Cardiandra by Siebold and Zuccarini in 1839. The specimen in M annotated “Cardiandra alternifolia S & Z” by Zuccarini, M0120979, is considered to have been given by Siebold and is part of the original material.

190) Hydrangea azisai Siebold in Nova Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. Cur. 14(2): 689 (1828). Siebold & Zucc., fl. Jap. 1: 104, t. 51 (1839).

Present treatment: Hydrangea macrophylla (Thunb.) Ser. in DC., Prodr. 4: 15 (1830) (f. macrophylla).

Japanese name: Ajisai; Azisai (Siebold 1828).

Lectotype (Ohba and Akiyama 2014):

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Japonia. Hb. von Siebold (L0043369) [Category 2, 5 (lower branch, fl. Jap. 1: t. 51)].

Original material: Legit in Japonia et Communicavit Siebold anno 1839, Herbarium Zuccarinii (M0153951) [Category 2]. in Japonia legit Bürger. Communicavit d. Siebold, Herb. Zuccarinii (M0153952) [Category 2b]. Japonia. [Siebold s.n.] (MAK S0433) [Category 3]. Japonia. [Siebold s.n.] (MAK S0837) [Category 2].

The specimen in M annotated “Hydrangea Azisai Sieb. & Zuccar.” by Zuccarini, M0153951, is considered to have been given by Siebold and is part of the original material.

191) Hydrangea azisai Siebold var. foliis aureo-variegatis Siebold & Zucc., fl. Jap. 1: 105 (1839), nom. illeg. [phrase name]

Present treatment: probably Hydrangea macrophylla (Thunb) Ser.

Original material: not found in L, M, or MAK.

192) Hydrangea belzonii Siebold & Zucc., fl. Jap. 1: 109, t. 55 (1939).

Present treatment: Hydrangea serrata (Thunb.) Ser. var. yesoensis (Koidz.) H. Ohba in iwatsuki et al. (eds.), fl. Jap. 2b: 90 (2001).

Japanese name: Ezo-ajisai.Lectotype (Ohba and Akiyama 2013b):

Japonia. Hb. von Siebold (L0329008) [Category 2, 5 (upper right-hand branch, fl. Jap. 1: t. 55)].

Original material: Hb. Siebold (L0043370) [Category 2]. Legit Bürger in Japonia. Communicavit d. Siebold, Herbarium Zuccarinii (M0153955) [Category 2]. in Japonia legit Bürger. Communicavit d. Siebold, Herb. Zuccarinii (M0153954) [Category 2b]. Japonia. [Siebold s.n.] (MAK S0851) [Category 2].

193) Hydrangea bracteata Siebold & Zucc., fl. Jap. 1: 176, t. 92 (1841).

Present treatment: Hydrangea petiolaris Siebold & Zucc.

Japanese name: Tsuru-ajisai.

Lectotype (Ohba and Akiyama 2014): Japonia. Herb. Siebold (L0043367) [Category 5, right hand branch (fl. Jap. 1: t. 92, upper) and left hand branch (fl. Jap. 1: t. 92, lower)].

194) Hydrangea cordifolia Siebold & Zucc., fl. Jap. 1: 113, t. 59. ii (1839).


Japanese name: Tsuru-ajisai.Lectotype (Ohba and Akiyama 2014):

Japonia. Communicavit Bürger, Hb. von Siebold (L0043365) [Category 2].

Original material: Legit in Japonia Bürger. Communicavit d. Siebold, Herbarium Zuccarinii (M0153957) [Category 2]. in Japonia legit Bürger. Communicavit d. Siebold, Herb. Zuccarinii (M0153958) [Category 2b]. in Japonia legit Bürger. Communicavit d. Siebold, Herb. Zuccarinii (M0153959) [Category 2b]. in Japonia legit Bürger. Communicavit d. Siebold, Herb. Zuccarinii (M0153960) [Category 2b]. Japonia. [Siebold s.n.] (MAK S0850) [Category 2].

195) Hydrangea hortensia Siebold in Nova Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. Cur. 14: 688 (1828), nom. superfl.

Present treatment: probably Hydrangea macrophylla (Thunb.) Ser. (var. macrophylla).


Hydrangea hortensia is regarded as a variant name for Hydrangea hortensis J. Sm.

196) Hydrangea involucrata Siebold in Nova Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. Cur. 14: 691 (1828). Siebold & Zucc., fl. Jap. 1: 118, t. 63, 64 (1839).

Present treatment: Hydrangea involucrata Siebold

Japanese name: Tama-ajisai; Ginbaisoo and Kinbaisoo (Siebold 1828)

Lectotype (Ohba and Akiyama 2014): Japonia. Hb. von Siebold (L0043366) [Category


2, 5 (upper branch, flowers with doubled petals, fl. Jap. 1: t. 64, left hand branch)].

Original material: Japonia. Hb. von Siebold, in July to Aug. (L0101479) [Category 2, 5 (lower branch, flowers with single petals, fl. Jap. 1: t. 64, right hand branch)]. Japonia. Hb. von Siebold (L0103032) [Category 2, 5 (fl. Jap. 1: t. 64, left hand branch)]. Legit in Japonia et Communicavit Siebold anno 1839, Herbarium Zuccarinii (M0153961) [Category 2].

The specimen in M annotated “Hydrangea involucrata S. & Z.” by Zuccarini, M0153961, is considered to have been given by Siebold and is part of the original material.

197) Hydrangea japonica Siebold in Nova Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. Cur. 14(2): 689 (1828). Siebold & Zucc., fl. Jap. 1: 106, t. 53 (1839).

Present treatment: Hydrangea serrata (Thunb.) Ser. var. japonica (Siebold) H. Ohba & S. Akiyama in Bull. Natl. Sci. Nat. Sci, B, 39: 179 (2013).

Japanese name: Beni-gaku; Kakusoo (Siebold 1829).

Lectotype (Ohba and Akiyama 2013a): Japonia. Hb. v Siebold, in July (L0043374) [Category 2].

Original material: Japonia. Hb. Siebold, in June (L0329010) [Category 2].

The original specimen, L0329010, is also the lectotype of H. japonica var. benkaku Siebold.

198) Hydrangea japonica Siebold var. benkaku Siebold in Nova Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. Cur. 14(2): 689 (1828), nom. superfl. Siebold & Zucc., Fl. Jap. 1: 106 (1839).

Present treatment: Hydrangea serrata (Thunb.) Ser. var. japonica (Siebold) H. Ohba & S. Akiyama.

Japanese name: Benkaku (Siebold 1828).Lectotype (Ohba and Akiyama 2013b):

Japonia. Hb. Siebold, in June (L0329010) [Category 2].

199) Hydrangea japonica Siebold var. konkaku Siebold in Nova Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. Cur. 14(2): 689 (1828). Siebold & Zucc, fl. Jap. 1: 106 (1839).

Present treatment: Hydrangea serrata Ser. var. japonica (Siebold) H. Ohba & S. Akiyama in Bull. Natl. Sci. Nat. Sci, B, 39: 179 (2013).

Japanese name: Konkaku (Siebold 1828).Lectotype (Ohba and Akiyama 2013b):

Japonia. Hb. von Siebold (L0103054).

200) Hydrangea otaksa Siebold & Zucc., fl. Jap. 1: 105, t. 52 (1839).

Present treatment: Hydrangea macrophylla (Thunb.) Ser. ‘Otaksa.’

Japanese name: Ajisai [Otaksa-ajisai as the horticulture name]; Otaksa (Siebold and Zuccarini 1839).

Lectotype (Ohba and Akiyama 2014): Japonia. Hb. Siebold (L0113146) [Category 2].

Original material: Japonia. Hb. Siebold, in July (L0328086) [Category 2]. Legit in Japonia Bürger. Communicavit d. Siebold, Herbarium Zuccarinii (M0153965) [Category 2]. in Japonia legit Bürger. Communicavit d. Siebold, Herb. Zuccarinii (M0153966) [Category 2b].

201) Hydrangea paniculata Siebold in Nova Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. Cur. 14(2): 691 (1828). Siebold & Zucc., fl. Jap. 1: 115, t. 61 (1839).

Present treatment: Hydrangea paniculata Siebold.

Japanese name: Nori-utsugi, Nori-no-ki.Lectotype (Ohba and Akiyama 2014):

Japonia. Hb. von Siebold (L0043375) [Category 2].

Original material: Legit Bürger in Japonia. Communicavit d. Siebold, Herbarium Zuccarinii (M0153968) [Category 2]. in Japonia legit Bürger. Communicavit d. Siebold, Herb. Zuccarinii (M0153969) [Category 2b]. Japonia. [Siebold s.n.] (MAK S0846) [Category 2].

The specimen in M annotated “Hydrangea paniculata S. & Z.” by Zuccarini, M0153968, is

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considered to have been given by Siebold and is part of the original material.

202) Hydrangea petiolaris Siebold & Zucc., fl. Jap. 1: 108, t. 54 (1835).


Japanese name: Tsuru-ajisai; Jama demari (Siebold and Zuccarini 1835).

Lectotype (Ohba and Akiyama 2014): Japonia. Hb. von Siebold, in July (L0043364) [Category 2, 3].

203) Hydrangea sitsitan Siebold in Nova Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. Cur. 14(2): 692 (1828).

Present treatment: Hydrangea serrata Ser. var. serrata f. prolifera (Regel) Nakai, Siebold Hydrang. Kaisetsu: 14 (1938).

Japanese name: Shichidanka.Lectotype (Ohba and Akiyama 2013b):

Japonia. Hb. von Siebold (L0043368) [Category 2].

Original material: Japonia. Hb. von Siebold (L0170300) [Category 2, 3]. Japonia. [Siebold s.n.] (MAK S0819) [Category 3].

The lectotype specimen, L0043368, is also the lectotype specimen of H. stellata Siebold & Zucc.

204) Hydrangea stellata Siebold & Zucc., fl. Jap. 1: 112, t. 59. I (1839), nom. superfl. for H. sitsitan Siebold.

Present treatment: Hydrangea serrata Ser. var. serrata f. prolifera (Regel) Nakai, Siebold Hydrang. Kaisetsu: 14 (1938).

Japanese name: Shichidanka.Lectotype (Ohba and Akiyama 2013b):

Japonia. Hb. von Siebold (L0043368) [Category 2] (fig. 133).

205) Hydrangea thunbergii Siebold in Nova Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. Cur. 14(2): 690 (1828), nom. superfl. for H. serrata (Thunb.) Ser. Siebold & Zucc., fl. Jap.

1: 11, t. 58 (1839). Present treatment: Hydrangea serrata

(Thunb.) Ser. (var. serrata).Japanese name: Yama-ajisai; ‘Ama-tsja’ in

sensu Siebold (1830).Type: Type of Hydrangea serrata Ser.Hydrangea thunbergii Siebold is a

superfluous name for H. serrata (Thunb.) Ser., but H. thunbergii [sensu Siebold, excl. type] is the plant called Amatya (Amats'ja in Siebold, 1828) in Japan, i.e., H. serrata var. thunbergii H. Ohba & S. Akiyama (Ohba and Akiyama 2013b). A specimen, “Japonia. Cultivated in Hortus Botanicus, Hb. von Siebold, in July,” (L0043372), is an authentic specimen of H. thunbergii Siebold [sensu Siebold] based on our category 2 criteria. Another specimen, “Japonia. [Bürger s.n.],” (L0103047, branch in upper left), is also an authentic specimen of H. thunbergii Siebold [sensu Siebold] based on our category 2 criteria.

Mitellopsis Meisn.206) Mitellopsis japonica Siebold & Zucc.

in Abh. Math.-Phys. Cl. Königl. Bayer. Akad. Wiss. 4(2): 190 (1845). nom. nud.

Present treatment: Mitella japonica Maxim. in Bull. Acad. imp. Sci. St. Pétersb. 11: 431 (1867).

Japanese name: Ô-charumeru-sô. Original material: Japonia. in montium locis

humidis. Siebold s.n. (L0176011) [Category 3].The Japanese name of Mitellopsis japonica,

Charumeru-sô, was not appeared in the protologue but appears on the original label of Siebold on sheet L0176011. Although there is no annotation by Siebold or Zuccarini, we regard this specimen to be original material.

Parnassia L.207) Parnassia mucronata Siebold & Zucc.


Present treatment: Parnassia palustris L. var. multiseta Ledeb., fl. Ross. 1: 263 (1842).


Japanese name: Umebachi-sô. Lectotype (designated here): Legit in

Japonia & Communicavit de Siebold anno 1842, Herbarium Zuccarinii (M0120972) (fig. 120).

Original material: Legit Dr Siebold in Japonia. Communicavit Prof. Zuccarini anno 1843, Herbarium Zuccarinii (M0120973) [Category 2b].

The lectotype specimen, M0120972, has no annotation by Siebold and/or Zuccarini, however Zuccarini retained the name Parnassia punctulata S. et Z. on the label. The epithet “mucronata” apparently derives from the description “... differt ... foliis omnibus mucronato-acutis ...” or “[cauline leaves] sessili e basi cordata suborbiculari mucronato.”

The epithet punctulata appears to be related to the “ferrygino-punctatis” feature of the leaves. it might be thought that mucronata and punctulata are candidates for the epithet.

Siebold s.n. in Leiden (L0170304) may be original material. Three plants on the right side of the sheet accompany Siebold’s annotation “Parnassia palustris an nova species” and “Parnassia Japonica, Sieb;” and a plant of the left side (no. 134) accompanies a sketch of a gland showing a mucronate (or bifurcate) apex. The epithet mucronata might possibly be derived from this characteristic of the gland.

Siebold s.n. in Leiden (L0170302), with his annotation “Parnassia palustris. Nov. lusus Higo.” is also attributed to this species, but it is not certain that it is original material.

Philadelphus L.208) Philadelphus satsumanus Siebold,

Catalogue: 5 (1856), nom. nud.Present treatment: Philadelphus satsumi

Siebold ex Lindl. & Paxt.Japanese name: Baika-utsugi.Original material: Japonia. Keiske s.n.

(L0170309) [Category 2].

208b) Philadelphus satsumanus Siebold [Catalogue: 5 (1856), nom. nud.] ex Miq., Ann.

Mus. Bot. Lugd.-Bat. 3: 99 (1867), nom. superfl.Present treatment: Philadelphus satsumi

Siebold ex Lindl. & Paxt.Japanese name: Baika-utsugi.Lectotype (Ohba et al. 2005): Japonia.

Siebold s.n. (L0329021).The original material of P. satsumanus

Siebold, Keiske s.n. (L0170309), is the only specimen with Siebold’s annotation “Philadelphus satsumanus S” but it was not cited by Miquel (1867).

Philadelphus satsumanus Siebold was reported by Siebold and Co. (1856) and validly published by Miquel in 1867, but it is a superfluous name for P. satsumi Siebold ex Lindl. & Paxt. Miquel cited the name, “P. satsumi Sieb. in hort.—Paxt. fl. Gard. ii. n. 375 [p. 102–103] icon. xylogr.” in synonymy, and four specimens collected by Siebold (L0329021, this is the lectotype of P. satsumanus Siebold ex Miq.), Keiske (L0329023), Mohnike (L0170308) and Oldham 258 (L 908, 24-200). These four specimens were used for the description of Philadelphus satsumanus by Miquel.

Hu (1955) distinguished P. satsumi Siebold ex Lindl. & Paxt. and P. satsumanus Siebold ex Miq. by the pubescence of the leaves, i.e. uniformly pubescent in P. satsumanus and glabrous, except pubescent primary nerves and axils on lower surface, in P. satsumi. Miquel’s original material (syntypes) used for the description of P. satsumanus, however, have nearly glabrous leaves except for the veins and the axils.

209) Philadelphus satsumi Siebold [in sched., nom. nud.] ex Lindl. & Paxt. in Paxton’s fl. Gard. 2: 102, fig. 188 (1852).

Present treatment: Philadelphus satsumi Siebold ex Lindl. & Paxt.

Japanese name: Baika-utsugi.Original material: not found in L, M, or

MAK.The name Philadelphus satsumi Siebold was

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published by Lindley and Paxton in 1852. Hu (1955) reported Siebold’s specimen in “Roy. Hort. Soc. Gard., Wisley” as being the type.

Platycrater Siebold & Zucc., fl. Jap. 1: 62 (1837 or 1838); in Abh. Math.-Phys. Cl. Königl. Bayer. Akad. Wiss. 4(2): 192 (1845).

Type: Platycrater arguta Siebold & Zucc. Present treatment: Platycrater Siebold &

Zucc.

210) Platycrater arguta Siebold & Zucc., fl. Jap. 1: 64, t. 27 (1835); in Abh. Math.-Phys. Cl. Königl. Bayer. Akad. Wiss. 4(2): 192 (1845).

Present treatment: Platycrater arguta Siebold & Zucc.

Japanese name: Baika-amacha; Bai kwa ama tsja (Siebold and Zuccarini 1835).

Lectotype (designated here): Japonia. Herbar. de Siebold, in 1829 (L0104524) [Category 2] (fig. 121).

Original material: Japonia. Siebold s.n. (L0104521) [Category 2]. Legit in Japonia & Communicavit de Siebold anno 1842, Herbarium Zuccarinii (M0120977) [Category 2]. in Japonia legit Bürger. Communicavit d. Siebold, Herb. Zuccarinii (M012978) [Category 2b].

Ribes L.211) Ribes fasciculatum Siebold & Zucc.


Present treatment: Ribes fasciculatum Siebold & Zucc.

Japanese name: Yabu-sannzashi.Lectotype (designated here): in Japonia

legit Bürger. Communicavit d. Siebold, Herb. Zuccarinii (M0153974) [Category 2b] (fig. 122).

Original material: in Japonia legit Bürger. Communicavit d. Siebold, Herb. Zuccarinii (M0153975) [Category 2b].

There are two specimens annotated “Ribes Jabusansasi” by Siebold in L, L0327091 and

L0327094. The epithet of the provisional name, Ribes Jabusansasi, is derived from the Japanese name of the plant. We cannot choose the Leiden specimens as original material because they have no indication of Siebold’s and/or Zuccarini’s description in 1845.

Saxifraga L.212) Saxifraga cortusifolia Siebold & Zucc.

in Abh. Math.-Phys. Cl. Königl. Bayer. Akad. Wiss. 4(2): 190 (1845), as “cortusaefolia.”

Present treatment: Saxifraga cortusifolia Siebold & Zucc.

Japanese name: Jinji-so.Lectotype (designated here): Legit in

Japonia et Communicavit d. Siebold, Herbarium Zuccarinii (M0120982) [Category 2] (fig. 123).

Schizophragma Siebold & Zucc., fl. Jap. 1: 58 (1837 or 1838); in Abh. Math.-Phys. Cl. Königl. Bayer. Akad. Wiss. 4(2): 192 (1845).

Type: Schizophragma hydrangeoides Siebold & Zucc.

Present treatment: Schizophragma Siebold & Zucc.

213) Schizophragma hydrangeoides Siebold & Zucc., fl. Jap. 1: 60, t. 26 (1837 or 1838), t. 100 i (1839); in Abh. Math.-Phys. Cl. Königl. Bayer. Akad. Wiss. 4(2): 192 (1845).

Present t reatment: Schizophragma hydrangeoides Siebold & Zucc.

Japanese name: iwagarami; Tsuru-demari (Siebold and Zuccarini 1837 or 1838).

Lectotype (designated here): Japonia. Herbar. de Siebold coll. 1829 (L0104501) (fig. 124).

Original material: Japonia. Hb. Siebold (L0104500) [Category 2]. Japonia. Nagasaki. Siebold s.n. (L0104499) [Category 3].

ROSACEAEAmygdalus L.

214) Amygdalus persica L. var. stellata Siebold & de Vriese in Ann. Hort. Bot. (fl.


Jardin) 2: 65, t. [6] (1859). Present treatment: Amygdalus persica L. f.

stellata (Siebold & de Vriese) H. Ohba & S. Akiyama, stat. nov. Basionym: Amygdalus persica L. var. stellata Siebold & de Vriese in Ann. Hort. Bot. (fl. Jardin) 2: 65 (1859) ≡ Prunus persica (L.) Batsch. var. β. vulgaris (DC.) Maxim. f. stellata (Siebold & de Vriese) Makino in Bot. Mag. (Tokyo) 22: 119 (1908).

Japanese name: Genpei-kiku-momo (nov.); Kiku-momo, Genji-guruma (Makino 1908).

Makino (1908) reported “Kiku-momo” and “Genji-guruma” as Japanese names, but this plant has both rose and white flowers. A new Japanese name, Genpei-kiku-momo [meaning plant with narrow petals and with both rose and white flowers], is presented here.

Lectotype (designated here): illustration t. [6] in Ann. Hort. Bot. (fl. Jardin) 2: t. [6, between pages 64 and 65] (1859).

We designate the illustration in the protologue as the lectotype. The plant illustrated in the table has double flowers with both pink and white narrow petals. in L, M and MAK there are some specimens of the cultivar of Amygdalus persica, but none of them represents var. stellata Siebold & de Vries. One of them in L, L0102983, is a specimen annotated “Amygdalus persica erecta, Genpei too” by Siebold. “Genpei too” means having both pink and white flowers. That specimen resembles var. stellata with bicolored double flowers, but the petals are wide. in M there is also a specimen, M0154003, named Persica vulgaris var. flore pleno albo with double petaled flowers, but appears to be only white.

Armeniaca L.215) Armeniaca mume Siebold, Syn. Pl.

Oeconom.: 69 (1830), nom. nud.Present treatment: Armeniaca mume (Siebold

& Zucc.) K. Koch, Hort. Dendrol.: 140 (1853).Original material: Japonia. Herbar. de Siebold

coll. 1829 [and 1827] [flowers: Mar.1829 and leaves: 亥六月 June 1827 by lunar calendar]

(L0329142, this is also original material of Prunus mume) [Category 2]. Japonia. [Bürger s.n.] (L0328748) [Category 2].

Armeniaca mume Siebold (1830) is an invalid name. Koch (1853) made a new combination, Armeniaca mume (Siebold & Zucc.) K. Koch based on Prunus mume Siebold & Zucc.

216) Armeniaca mume Siebold var. a. floribus simplicibus plenisque niveis Siebold, Syn. Pl. Oeconom.: 69 (1830), nom. illegit. [phrase name]

Present treatment: a cultivar of Armeniaca mume (Siebold & Zucc.) K. Koch.


217) Armeniaca mume Siebold var. b. floribus simplicibus plenisque roseis Siebold, Syn. Pl. Oeconom.: 69 (1830), nom. illegit. [phrase name]



218) Armeniaca mume (Siebold & Zucc.) K. Koch var. praecocissima rubiflora Siebold & de Vriese in de Vriese, Tuinb. fl. 1: 1, t. 1 (1855), nom. illegit. [phrase name]


Original material: illustration in de Vriese, Tuinb. fl. 1: 1, t. 1 (1855).

According to the original description and the illustration, praecocissima rubiflora appears to be an early flowering double flowered cultivar with rose colored petals like ‘Kankôbai.’

Aronia Mitch.219) Aronia asiatica Siebold & Zucc., fl.

Jap. 1: 87, t. 42 (1839); in Abh. Math.-Phys. Cl. Königl. Bayer. Akad. Wiss. 4(2): 131 (1845).

Present treatment: Amelanchier asiatica

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Endl. in Walp., Rep. 2: 53 (1843).Japanese name: Zaifuri-boku; Zaifuri,

Zaifuribok (Siebold and Zuccarini 1839).Lectotype (designated here): Herb. de

Siebold in 1829 (L0176017) [Category 2] (fig. 125).

Original material: Japonia. [April. Jedo]. Siebold s.n. (L0170335) [Category 3].

Ceraseidos Siebold & Zucc. in Abh. Math.-Phys. Cl. Königl. Bayer. Akad. Wiss. 3(3): 743 (1843); 4(2): 122 (1845).

Present treatment: Cerasus Mill.Type: Ceraseidos apetala Siebold & Zucc.

in Abh. Math.-Phys. Cl. Königl. Bayer. Akad. Wiss. 3(3): 744, t. V 2 (1843); 4(2): 122 (1845).

220) Ceraseidos apetala Siebold & Zucc. in Abh. Math.-Phys. Cl. Königl. Bayer. Akad. Wiss. 3(3): 744, t. V 2 (1843); 4(2): 122 (1845).

Present treatment: Cerasus apetala (Siebold & Zucc.) Ohle ex H. Ohba in J. Jpn. Bot. 67: 277 (1992) (var. apetala).

Japanese name: Miyama-chôji-zakura. Lectotype (designated here): Japonia.

Siebold s.n. (L0176026) [Category 2] (fig. 126).

Crataegus L.221) Crataegus alnifolia Siebold & Zucc.


Present treatment: Sorbus alnifolia (Siebold & Zucc.) K. Koch in Ann. Mus. Bot. Lugd.-Bat. 1: 249 (1864).

Japanese name: Azuki-nashi. Lectotype (designated here): Japonia. Legit

Bürger in Japonia, communicavit de Siebold, Herbarium Zuccarinii (M0153991) [Category 2] (fig. 127).

Original material: Herb. Bürger (L0329395) [Category 2].

Aldasoro et al. (2004) selected Bürger s.n. (L0245120) as the lectotype, but the specimen has no annotation or other information to indicate it to be original material. We designate

M0153991 collected by Bürger, and which was cited by Aldasoro et al. as an isolectotype, as the lectotype.

222) Crataegus cuneata Siebold & Zucc. in Abh. Math.-Phys. Cl. Königl. Bayer. Akad. Wiss. 4(2): 130 (1845).

Present treatment: Crataegus cuneata Siebold & Zucc.

Japanese name: Sanzashi. Lectotype (designated here): Legit Bürger in

Japonia. Communicavit de Siebold, Herbarium Zuccarinii (M0153734) [Category 2] (fig. 128).

Original material: in Japonia legt. Bürger comcvt. de Siebold, Herb. Zuccarinii (M0153733) [Category 2b].

The specimens in L, Siebold s.n. (L0170337 and L0170338), with accompanying slips with the Japanese name “Crataegus sansasi, Jap. and サンザシ ” by Siebold, are also representative. The specimen, Siebold (L0170336), with the Japanese name “Crataegus oosansasi, J[ap.]” by Siebold, also has no evidence of being original material.

Kerria DC.223) Kerria japonica (L.) DC. var. a

Floribus simplicibus Siebold, Oeconom.: 69 (1830), nom. illegit. [phrase name]

Present treatment: Kerria japonica (L.) DC. in Trans. Linn. Soc. London 12: 157 (1817) (f. japonica).

Japanese name: Hitoe-yamabuki.Original material: Japonia. [Siebold s.n.]

(MAK S0261a) [Category 2].

224) Kerria japonica (L.) DC. var. b. Floribus plenis Siebold, Oeconom.: 69 (1830), nom. illegit. [phrase name]

Present treatment: Kerria japonica (L.) DC. f. plena C. K. Schneid., ill. Hand. Laubholzk. 1: 501 (1905).

Japanese name: Yae-yamabuki.Original material: not found in L, M, or

MAK.


225) Kerria japonica (L.) DC. var. c Floribus simplicibus foliis argentatis Siebold, Oeconom.: 69 (1830), nom. illegit. [phrase name]

Present treatment: Kerria japonica (L.) DC. ‘Variegata.’

Japanese name: fuiri-yamabuki.Original material: not found in L, M, or

MAK.

226) Kerria japonica (L.) DC. var. α. sejo-jama-buki Siebold & Zucc., fl. Jap. 1: 183 (1841), as “Sejô jama buki.”

Present treatment: Kerria japonica (L.) DC. f. plena C. K. Schneid., ill. Hand. Laubholzk. 1: 501 (1905).

Japanese name: Yae-yamabuki; Sejô jama buki (Siebold and Zuccarini 1841).

Lectotype (designated here): Hb. von Siebold (L0328743) [Category 5 (fl. Jap. 1: t. 98 iii)] (fig. 129).

227) Kerria japonica (L.) DC. var. β. hitoje-jamabuki Siebold & Zucc., fl. Jap. 1: 183 (1841), as “Hitoje jamabuki.”

Present treatment: Kerria japonica (L.) DC. in Trans. Linn. Soc. London 12: 157 (1817) (f. japonica).

Japanese name: Hitoe-yamabuki; Hitoje jamabuki. (Siebold and Zuccarini 1841).

Lectotype (designated here): illustration in Siebold & Zucc., fl. Jap. 1: t. 98 i (1841).

228) Kerria japonica (L.) DC. var. γ. fuiri-jamabuki Siebold & Zucc., fl. Jap. 1: 183 (1841), as “Fuiri jamabuki.”

Present treatment: Kerria japonica (L.) DC. ‘Variagata.’

Japanese name: fuiri-yamabuki; fuiri jamabuki. (Siebold and Zuccarini 1841).

Lectotype (designated here): illustration in Siebold & Zucc., fl. Jap. 1: t. 99 ii (1841).

The specimen, Herb. Siebold (L0328745), with the note “Kerria japonica DC. Sieb. & Zucc., fl. jap. t. 98. variet. foliis maculatis,”

corresponds well with this variety but lacks Siebold and/or Zuccarini’s handwriting.

The illustration of Table 99 ii represents well the variegated form of Kerria japonica, although there is no explanation by Siebold and Zuccarini. Yamaguchi et al. (2003) reported that the illustration of Table 98 of Flora Japonica was based on this specimen.

229) Kerria tetrapetala Siebold, Syn. Pl. Oeconom.: 69 (1830), nom. nud.

Present treatment: Rhodotypos scandens (Thunb.) Makino in Bot. Mag. (Tokyo) 27: 126 (1913).

J apanese name: Sh i ro -yamabuk i ; Sirojamabuki (Siebold 1830).

Original material: Herb. von Siebold (L0329269) [Category 2]. Japonia. Keiske [Sukerok] s.n. (L0175824) [Category 3].

The specimen, L0329269, is also the lectotype of Rhodotypos kerrioides Siebold & Zucc. (see p. 293).

Malus L.230) Malus floribunda Siebold, Cat. Rais. 5

(1856), nom. nud. Present treatment: probably Malus halliana

Koehne, Gatt. Pomac.: 27 (1890).Japanese name: probably Kaidô. Original material: not found in L, M, or

MAK.

231) Malus [Jap.] Ringo Siebold, Cat. Rais. 5 (1856), nom. nud.

Present treatment: Malus domestica Borkh., Theor. Prakt. Handb. forstbot. 2: 1272 (1803).

Japanese name: Ringo. Original material: Japonia. Siebold

(L0329359) [Category 3].

232) Malus spectabilis Desf. var. [Jap.] Kaido Siebold, Cat. Rais. 5 (1856), nom. nud.

Present treatment: Malus halliana Koehne, Gatt. Pomac.: 27 (1890).

Japanese name: Kaidô.

290 植物研究雑誌　第 89巻　第 5号 2014年 10月

Original material: Japonia. Siebold s.n. (L0329350) [Category 3]. Japonia. Siebold s.n. (L0329358) [Category 3].

233) Malus toringo Siebold, Cat. Rais.: 5 (1856).

Present treatment: Malus toringo (Siebold) Siebold, Cat. Rais.: 5 (1856). De Vriese, Tuinb.-fl. 3: 368, t. [17, between pages 368 and 369], excl. fig. fl. (1856).

Japanese name: Zumi; Ko-ringo; Kimi-zumi (Siebold and de Vriese 1856).

Type: Type of Sorbus toringo Siebold in Jaarb. Kon. Ned. Maatsch. Aanm. Tuinb. 1848: 47 (1848).

When Siebold published Malus toringo Siebold (Cat. Rais.: 5, 1856) validly, he adopted the epithet of Sorbus toringo Siebold (1848) without referring to that name. According to ICN Art. 41.4 it is treated as a new combination, Malus toringo (Siebold) Siebold, not as the name of a new species. in the same year de Vriese wrote Malus toringo Siebold (Tuinb. fl. 3: 368, t. [17, between pages 368 and 369], 1856) with referring to Sorbus toringo Siebold and Siebold’s Catalogue Raisonné.

Persica Mill.234) Persica pendula Siebold, Cat. Rais.: 36

(1863).Present treatment: Amygdalus persica L. f.

pendula (Siebold) H. Ohba & S. Akiyama, stat. nov. Basionym: Persica pendula Siebold, Cat. Rais.: 36 (1863) ≡ Prunus persica (L.) Batsch var. pendula (Siebold) Koidz. in Bot. Mag. (Tokyo) 44: 95 (1930).

Japanese name: Beni-shidare-momo. Lectotype (designated here): Japonia.

Siebold s.n. (L0102984) [Category 5] (fig. 130).The lectotype specimen is not annotated as

‘Persica pendula’ but has Siebold’s annotation “Amygdalus persica pendula” and the Japanese name “Sdale momo” [it is now written ‘Sudare’ or ‘Shidare’ and means a pendulous form of Amygdalus persica in Japanese].

235) Persica stellata Siebold, Cat. Rais.: 36 (1863), nom. nud.

Present treatment: Amygdalus persica L. f. stellata (Siebold & de Vriese) H. Ohba & S. Akiyama.

Japanese name: Genpei-kiku-momo. Original material: not found in L, M, or

MAK.

236) Persica vulgaris Mill. var. versicolor Siebold, Cat. Rais.: 36 (1863).

Present treatment: Amygdalus persica L. f. versicolor (Siebold) H. Ohba & S. Akiyama, comb. et stat. nov. Basionym: Persica vulgaris Mill var. versicolor Siebold, Cat. Rais.: 36 (1863) ≡ Prunus persica (L.) Batsch f. versicolor (Siebold) Koidz. in Bot. Mag. (Tokyo) 44: 94 (1930).

Japanese name: Genpei-momo. Original material: not found in L, M, or

MAK.

Prunus L.237) Prunus domestica L. var. [a] zumomo

Siebold, Syn. Pl. Oeconom.: 68 (1830).Present treatment: probably a cultivar of

Prunus domestica L.Japanese name: Sumomo.Original material: not found in L, M, or

MAK.

238) Prunus domestica L. var. [b] awozumomo Siebold, Syn. Pl. Oeconom.: 68 (1830).

Present treatment: probably a cultivar of Prunus domestica L.

Japanese name: Ao-sumomo.Original material: not found in L, M, or

MAK.

239) Prunus domestica L. var. [c] sirozumomo Siebold, Syn. Pl. Oeconom.: 68 (1830).

Present treatment: probably a cultivar of Prunus domestica L.


Japanese name: Shiro-sumomo.Original material: not found in L, M, or

MAK.

240) Prunus donarium Siebold, Syn. Pl. Oeconom.: 68 (1830), nom. nud.

Present treatment: Cerasus serrulata (Lindl.) G. Don in Loudon, Hort. Brit.: 480 (1839).

Japanese name: Shiogama(-sakura); Sakura (Siebold 1830).

Original material: Japonia. Siebold s.n. (L0834824) [Category 2, 3]. Japonia in templis. Legit v. S. (MAK S0251, upper branch) [Category 2].

241) Prunus itosakura Siebold, Syn. Pl. Oeconom.: 68 (1830).

Present treatment: Cerasus itosakura (Siebold) H. Ohba & S. Akiyama, comb. nov. Basionym: Prunus itosakura Siebold, Syn. Pl. Oeconom.: 68 (1830).

Japanese name: Shidarezakura, itozakura; itosakura (Siebold 1830).

Lectotype (designated here): Japonia. Nagas.[aki]. Siebold s.n. (L0102980) [Category 2] (fig. 131).

Original material: Japonia. Siebold s.n. (L0102977) [Category 2]. Japonia. [Siebold s.n.] (L0102981) [Category 2].

Ohba mistakenly treated Prunus itosakura Siebold as an illegitimate name (nomen nudum) and adopted Cerasus spachiana Lavallée ex E. Otto (in Hamburger Garten-Blumenzeitung 35: 57, 1879) according to Sargent (1888) for it (Ohba 2001, 2012, Ohba et al. 2007). However, it is recognized as a legitimate name with a short diagnosis. Siebold wrote on this species: “Cerasus ramulis pendulis florentibus adornans idolorum sacra.” The “Cerasus ramulis pendulis” [Cerasus with pendulous branchlets], is considered sufficient for a diagnosis. in Siebold’s time, this was the only flowering cherry with pendulous branchlets.

Maximowicz (1883) published Prunus pendula Siebold ex Maxim. and cited Synopsis

Plantae Oeconomicarum (1830, No. 368[sic]) as the source of the Siebold name. Number 368 of Siebold’s Synopsis Plantae Oeconomicarum is neither Cerasus nor Prunus but a species of Gramineae, “CCxVii. GRAMiNEA, Makomo, Japon.”

242) Prunus jamasakura Siebold, Syn. Pl. Oeconom.: 68 (1830), nom. nud.

Present treatment: Cerasus jamasakura (Siebold ex Koidz.) H. Ohba in J. Jap. Bot. 67: 278 (1992).

Japanese name: Yamazakura.Original material: Japonia. [Siebold s.n.]

(L0834829) [Category 3]. Japonia. K[Keiske] s.n. (L0329119) [Category 2, 3]. Japonia. Legit v. S.[Siebold s.n.] (MAK S0247) [Category 2]. Japonia. [Siebold s.n.] (MAK S0249) [Category 2].

Prunus jamasakura, native to Japan, was validly published by Koidzumi (1911), but he did not designate a lectotype. The specimen, L0834829, consists of three branches with simple petaled flowers. The leaves on a post flowering branch are narrowly oblong-obovate or elliptic with glabrous surfaces and margins and single (partly double) serrations and have interpetiolar glands. These features characterize Cerasus jamasakura.

243) Prunus macrophylla Siebold & Zucc. in Abh. Math.-Phys. Cl. Königl. Bayer. Akad. Wiss. 4(2): 122 (1845), non Poir. (1816).

Present treatment: Laurocerasus zippeliana Miq., fl. Ned. ind., Eerste Bijv. (also as fl. ind. Batav.) 1(1): 367 (1855).

Japanese name: Bakuchi-no-ki. Lectotype (designated here): Japonia.

Siebold s.n. (L0329088) [Category 2] (fig. 132).Original material: Legit Bürger in Japonia.

Communicavit de Siebold, Herbarium Zuccarinii (M0154001) [Category 2]. in Japonia legt Bürger comct. de Siebold, Herb. Zuccarinii (M0154002) [Category 2b]. Japonia. Siebold s.n. (L0329091) with Siebold’s annotation

292 植物研究雑誌　第 89巻　第 5号 2014年 10月

“Laurocerasus japonica S[iebold]” and Japanese name “バクチノキ [Bakuchi-no-ki in Japanese].” Japonia. Siebold s.n. (L0176030) with Siebold’s annotation “Laurocerasus japonica Siebold.”

Two specimens, Siebold s.n. (L0329091 and L0176030), have only Siebold’s annotation “Laurocerasus japonica S[iebold]” and also the Japanese name “バクチノキ [Bakuchi-no-ki in Japanese]” on L0329091, and are considered to be derived from the same source as lectotype.

244) Prunus mume Siebold & Zucc., fl. Jap. 1: 29, t. 11 (1836); in Abh. Math.-Phys. Cl. Königl. Bayer. Akad. Wiss. 4(2): 122 (1845).

Present treatment: Armeniaca mume (Siebold & Zucc.) K. Koch, Hort. Dendrol.: 140 (1853).

Japanese name: Ume; Mume (Siebold and Zuccarini 1836).

Lectotype (designated here): Japonia. Siebold s.n. (L0329140) [Category 3, 5 (fl. Jap. t. 11 i)] (fig. 133).

Original material: Japonia. Legit v. S.[Siebold] (L0329135, MAK S0259) [Category 2]. Legit Bürger in Japonia Communicavit de Siebold, Herbarium Zuccarinii (M0153997) [Category 2]. in Japonia legt. Bürger comct. de Siebold, Herb. Zuccarinii (M0153998) [Category 2b]. in Japonia legt. Bürger comct. de Siebold, Herb. Zuccarinii (M0153999) [Category 2b]. Legit Dr Siebold in Japonia. Communicavit Prof. Zuccarini, Herbarium Zuccarinii (M0154000) [Category 2b]. Japonia. [Siebold s.n.] (MAK S0373) [Category 2]. Japonia. Planta culta introducta in H. Sieb. [Siebold s.n.] (MAK S0553) [Category 2].

245) Prunus mume Siebold & Zucc. var. floribus purpureis semiplenis adjecto ramulo foliifero Siebold & Zucc., fl. Jap. 1: 30, t. 11 iii (1836), nom. invalid. [phrase name]

Present treatment: probably a cultivar of Armeniaca mume (Siebold & Zucc.) K. Koch.


246) Prunus mume Siebold & Zucc. var. pendula Siebold in Jaarb. Kon. Ned. Maatsch. Tuinb. 1848: 47 (1848).

Present treatment: Armeniaca mume (Siebold & Zucc.) K. Koch f. pendula (Siebold) H. Ohba & S. Akiyama, comb. et stat. nov. Basionym: Prunus mume Siebold & Zucc. var. pendula Siebold in Jaarb. Kon. Ned. Maatsch. Tuinb. 1848: 47 (1848).

Japanese name: Shidare-ume.Lectotype (designated here): Japonia.

Siebold s.n. (MAK S0551, left) [Category 3] (fig.134).

Original material: Japonia. s.n. (MAK S0551, right) [Category 3]. Japonia.

247) Prunus spinulosa Siebold & Zucc. in Abh. Math.-Phys. Cl. Königl. Bayer. Akad. Wiss. 4(2): 122 (1845).

Present treatment: Laurocerasus spinulosa (Siebold & Zucc.) C. K. Schneid., ill. Handb. Laubh. 1: 649, f. 354 o–p, f. 355 f (1906).

Japanese name: Rin-boku. Lectotype (designated here): Legit in

Japonia et Communicavit de Siebold, Herbarium Zuccarinii (M0154004) [Category 2] (fig. 135).

Original material: in Japonia legt. v. Siebold, Herb. Zuccarinii (M0154005) [Category 2b].

Pyrus L.248) Pyrus gracilis Siebold & Zucc. in Abh.

Math.-Phys. Cl. Königl. Bayer. Akad. Wiss. 4(2): 131 (1845).

Present treatment: Sorbus gracilis (Siebold & Zucc.) K. Koch, Hort. Dendrolog.: 178 (1853).

Japanese name: Nankin-nanakamado. Lectotype (designated here): Legit. Bürger in

Japonia. Communicavit de Siebold (M0154368) [Category 2] (fig. 136).

Original material: Japonia. Siebold (L0329393) [Category 2].

A specimen, Herb. Bürger (L0043355), has an accompanying type slip, Pyrus gracilis Siebold & Zucc., in Hara’s hand, but the specimen is not regarded as original material


because there is no evidence of the original description.

Raphiolepis Lindl.249) Raphiolepis japonica Siebold & Zucc.,

fl. Jap. 1: 162, t. 85 (1841); in Abh. Math.-Phys. Cl. Königl. Bayer. Akad. Wiss. 4(2): 130 (1845).

Present treatment: Raphiolepis indica (L.) Lindl. var. umbellata (Thunb.) H. Ohashi f. umbellata (Thunb.) Hatus. in J. Geobot. (Kanazawa) 25: 126 (1978).

Japanese name: Sharin-bai; Hama mokkok’ (Siebold and Zuccarini 1841).

Lectotype (designated here): Japonia. Hb. von Siebold (L0176032) [Category 2, 3] (fig. 137).

Original material: Japonia. Hb. von Siebold (L0176031) [Category 3]. Legit Bürger in Japonia. Communicavit de Siebold, Herbarium Zuccarinii (M0153736) [Category 2]. in Japonia legt. Bürger comct. v. Siebold, Herb. Zuccarinii (M0153735) [Category 2b].

250) Raphiolepis mertensii Siebold & Zucc., fl. Jap. 1: 164 (1841); in Abh. Math.-Phys. Cl. Königl. Bayer. Akad. Wiss. 4(2): 130 (1845).

Present treatment: Raphiolepis indica (L.) Lindl. var. umbellata (Thunb.) H. Ohashi f. umbellata (Thunb.) Hatus. in J. Geobot. (Kanazawa) 25: 126 (1978).

Japanese name: Maruba-sharin-bai. Original material: not found in L, M, or

MAK.

Rhodotypos Siebold & Zucc., fl. Jap. 1: 185 (1841); in Abh. Math.-Phys. Cl. Königl. Bayer. Akad. Wiss. 4(2): 125 (1845).

Type: Rhodotypos kerrioides Siebold & Zucc.

Present treatment: Rhodotypos Siebold & Zucc.

251) Rhodotypos kerrioides Siebold & Zucc., fl. Jap. 1: 187, t. 99 (1841); in Abh. Math.-Phys. Cl. Königl. Bayer. Akad. Wiss.

4(2): 125 (1845). Present treatment: Rhodotypos scandens

(Thunb.) Makino in Bot. Mag. (Tokyo) 27: 126 (1913).

Japanese name: Shiro-yamabuki; Siro jamabuki.

Lectotype (designated here): Herb. von Siebold (L0329269) (fig. 138).

Original material: Japonia. Siebold s.n. (L0329268) [Category 3]. Japonia. Siebold s.n. (L0175824) [Category 3]. Legit in Japonia Bürger. Communicavit de Siebold anno 1842, Herbarium Zuccarinii (M0153995) [Category 2]. Legit in Japonia & Comunicavit de Siebold anno 1842, Herbarium Zuccarinii (M0153994) [Category 2].

The lectotype is also the original material of Kerria tetrapetala Siebold, nom. nud.

Rosa L.252) Rosa iwara Siebold in Cat. Raisonné

des Pl. du Japon, 1856: 6 (1856), nom. nud. Present treatment: Rosa ×iwara Siebold

ex Regel, index Sem. Hort. Petrop. 1861: 53 (1861).

Japanese name: Ko-hamanasu. Original material: Japonia. Legit Siebold,

Herb. Zuccarinii (M0154345) with Siebold’s annotation “Rosa Iwara Sieb.” [Category 2].

Rosa ×iwara is a natural hybrid between Rosa multiflora and R. rugosa, and in Japan was rarely cultivated in gardens as an ornamental during Siebold’s stay (Ohba and Akiyama 2013a). Rosa iwara Siebold was validly published by Regel [index Sem. Hort. Petrop. 1861: 53, 1861]. As Regel did not designate a lectotype, we designate the following specimen as the lectotype.

Lectotype (designated here): Japonia. Legit Siebold, Herb. Zuccarinii (M0154345) with Siebold’s annotation “Rosa Iwara Sieb.” and Crépin’s annotation “Rosa rugosa × R. multiflora” (fig. 139) [Category 2].

253) Rosa polyantha Siebold & Zucc. in

294 植物研究雑誌　第 89巻　第 5号 2014年 10月

Abh. Math.-Phys. Cl. Königl. Bayer. Akad. Wiss. 4(2): 128 (1845).

Present treatment: Rosa multiflora Thunb. in Murray, Syst. Veg. ed. 14: 474 (1784).

Japanese name: No-ibara. Lectotype (Ohba and Akiyama 2013a):

Japonia. Legit Bürger, communicavit Siebold, Herbarium Zuccarinii (M0154318).

Rubus L.254) Rubus ribifolius Siebold & Zucc. in

Abh. Math.-Phys. Cl. Königl. Bayer. Akad. Wiss. 4(2): 127 (1845).

Present treatment: Rubus ×ribifolius Siebold & Zucc.

Japanese name: Hama-kiichigo. Lectotype (designated here): Legit in

Japonia & Communicavit Zuccarini, Herbarium Zuccarinii (M0120864) [Category 2] (fig. 140).

255) Rubus thunbergii Siebold & Zucc. in Abh. Math.-Phys. Cl. Königl. Bayer. Akad. Wiss. 4(2): 126 (1845).

Present treatment: Rubus hirsutus Thunb., Dissert. Bot. Med. Rubo.: 7, 10 (1813).

Japanese name: Kusa-ichigo. Lectotype (designated here): Japan.

(Thunberg s.n., UPS-Herb. Thunberg no. 12266).

Siebold and Zuccarini cited Rubus idaeus [auct. non L.: Thunb., fl. Jap.: 216, 1784] as a synonym of Rubus thunbergii. Seven specimens in Thunberg’s collection (UPS, Herb. Thunberg nos. 12261–12267) are identified as Rubus idaeus. We have selected no. 12266 as the lectotype which appears to be the most representative among them.

Specimens used for the description of Rubus thunbergii by Siebold and Zuccarini are as follows:

Legit in Japonia & Communicavit de Siebold anno 1842, Herbarium Zuccarinii (M0154011) [Category 2]. in Japonia legit Bürger comcvt. v. Siebold, Herb. Zuccarinii (M0154012) [Category 2b]. in Japonia legit Bürger comcvt. v. Siebold,

Herb. Zuccarinii (M0154013) [Category 2b].

256) Rubus tokkura Japon ex Siebold, Syn. Pl. Oeconom.: 65 (1830), nom. nud.

Present treatment: Rubus coreanus Miq., Ann. Mus. Bot. Lugd.-Bat. 3: 34 (1867).

Japanese name: Tokkuri-ichigo; Tokkuri-itsigo (Siebold 1830).


Sieversia Willd.257) Sieversia dryadoides Siebold & Zucc.


Present treatment: Geum pentapetalum (L.) Makino in Bot. Mag. (Tokyo) 24: 32 (1910).

Japanese name: Chinguruma. Lectotype (designated here): Japonia.

[Sukerok s.n.] (L0175827) [Category 2] (fig. 141).

Original material: Legit v. Siebold in Japonia. Communicavit de Siebold, Herbarium Zuccarinii (M 0154382) [Category 2].

Sorbus L.258) Sorbus japonica Siebold, Syn. Pl.

Oeconom.: 67 (1830), nom. nud. Present treatment: Aria japonica Decne. in

Nouv. Arch. Mus. Hist. Nat. 10: 166 (1874).Japanese name: Urajiro-no-ki; Urasironoki

(Siebold 1830). Original material: Japonia. Siebold s.n.

(L0366764) [Category 2, 3].

259) Sorbus toringo Siebold in Jaarb. Kon. Ned. Maatsch. Aanm. Tuinb. 1848: 47 (1848).

Present treatment: Malus toringo (Siebold) Siebold, Cat. Rais.: 5 (1856). De Vriese, Tuinb.-fl. 3: 368, t. [17, between pages 368 and 369] excl. fig. fl. (1856).

Japanese name: Zumi; Ko-ringo, Kimi-zumi (Siebold & de Vriese 1856)

Lectotype (designated here): Japonia. Siebold s.n. (L0176022) [Category 2] (fig. 142).


Siebold effectively published Sorbus toringo with the description “Arbuste nain à feuilles florales elliptiques et raméales trilobées. Couvert au premier printemps de fleurs blanches et rosées moins grandes que celles de notre Pyrus spectabilis.”

Veldkamp selected the specimens L0176022 and L0329344 as the neotype of Sorbus toringo Siebold. L0329344 (Siebold s.n.) has neither annotation nor notes by Siebold himself, but L0176022 has Siebold’s annotation “Sorbus tôringo” and is regarded as original material.

Spiraea L.260) Spiraea japonica Siebold, Syn. Pl.

Oeconom.: 65 (1830), nom. nud. Present treatment: Spiraea cantoniensis

Lour., fl. Cochinch. 1: 322 (1790).Japanese name: Ko-demari; Kogomebana

(Siebold 1830).Original material: Japonia. Siebold

(L0176039) [Category 2].Honda (1938) considered this to be Spiraea

thunbergii Siebold & Zucc.

261) Spiraea prunifolia Siebold & Zucc., fl. Jap. 1: 131, t. 70 (1840); in Abh. Math.-Phys. Cl. Königl. Bayer. Akad. Wiss. 4(2): 125 (1845).

Present treatment: Spiraea prunifolia Siebold & Zucc.

Japanese name: Shijimi-bana; fage bana (Siebold and Zuccarini 1840).

Lectotype (designated here): Japonia. Hb. von Siebold (L0176040) [Category 2] (fig. 143).

Original material: Japonia. Siebold s.n. (L0329293) [Category 2]. Legit in Japonia. Bürger. Communicavit de Siebold anno 1842, Herbarium Zuccarinii (M0154017) [Category 2].

Although the specimen M0154017 was transferred to M from Siebold in 1842 (two years later than the publication of the name), we believe that it is original material.

262) Spiraea thunbergii Siebold, Syn. Pl. Oeconom.: 65 (1830), non Blume (1827).

Present treatment: Spiraea thunbergii Siebold ex Blume, Bijdr. 17: 1115 (1827).

Japanese name: Yuki-yanagi; Jukiyanagi (Siebold 1830).

Lectotype (designated here): Japan. Thunberg s.n. (L0144310, van Royen Herbarium) (fig. 144).

This is a new name for the Spiraea reported by Thunberg as S. crenata L. in his Flora Japonica (1784, p. 210). Siebold cited Spiraea crenata [misprinted as “erenata”][auct. non Pallas; Thunb., fl. Jap.: 210, 1784] as a synonym of Spiraea thunbergii. from Spiraea crenata in Thunberg’s collection the lectotype must be selected.

Thunberg’s collections were deposited mainly in UPS (Spiraea crenata: nos. 12070–12072) and partly in the van Royen Herbarium (now in L). Since Siebold’s annotation “Dolichos polystachyos Th.” is on a Thunberg specimen of Wisteria in the van Royen collection, we believe that Siebold had an opportunity to examine Thunberg’s specimens in the van Royen Herbarium and we have selected a lectotype of Spiraea thunbergii from the van Royen’s collection.

Spiraea thunbergii Siebold was published validly by Blume in 1827, earlier than by Siebold himself in 1830. So Spiraea thunbergii Siebold (1830) is a later homonym (see below no. 262b).

Specimens used for the preparation of Siebold’s Spiraea thunbergii are as follows:

Japonia. Siebold s.n. (L0176042) [Category 2]. Japonia. Hb. von Siebold (L0176041) [Category 2].

The specimen L0176041 is considered to be the one used for the description of Spiraea thunbergii Siebold & Zucc., and the basis for the illustration in Flora Japonica, Tab. 69. L0176042 is the lectotype of Spiraea thunbergii Siebold ex Blume.

296 植物研究雑誌　第 89巻　第 5号 2014年 10月

262b) Spiraea thunbergii Siebold ex Blume, Bijdr. 17: 1115 (1827).

Present treatment: Spiraea thunbergii Siebold ex Blume.

Japanese name: Yuki-yanagi.Lectotype (designated here): Japonia.

Siebold s.n. (L0176042, left) [Category 2] (fig. 145).

A specimen, L0176041, with Siebold’s annotation was collected in 1829 and is not original material.

262c) Spiraea thunbergii Siebold & Zucc., fl. Jap. 1: 130, t. 69 (1840), nec Blume (1827) non Siebold (1830); in Abh. Math.-Phys. Cl. Königl. Bayer. Akad. Wiss. 4(2): 124 (1845).

Present treatment: Spiraea thunbergii Siebold ex Blume, Bijdr. 17: 1115 (1827).

Japanese name: Yuki-yanagi; Juki janagi, iwa janagi (Siebold and Zuccarini 1840).

Lectotype (designated here): Japonia. Siebold s.n. (L0176041, left) [Category 2] (fig. 146).

Original material: Japonia. Siebold s.n. (L0176042) [Category 2] (fig. 145). Legit in Japonia Bürger. Communicavit de Siebold anno 1842, Herbarium Zuccarinii (M0154020) [Category 2]. Legit in Japonia. Bürger. Communicavit Siebold anno 1839, Herbarium Zuccarinii (M0154021) [Category 2]. Legit in Japonia. Bürger comcvt. d. Siebold, Herb. Zuccarinii (M0154022) [Category 2b]. Legit Dr Siebold in Japonia. Communicavit Prof. Zuccarini, anno 1843 (M0154369) [Category 2b].

Stephanandra Siebold & Zucc. in Abh. Math.-Phys. Cl. Königl. Bayer. Akad. Wiss. 3(3) 739 (1843); 4(2): 191 (1845).

Type: Stephanandra flexuosa Siebold & Zucc. (= S. incisa (Thunb.) Siebold & Zucc. ex Zabel).

Present treatment: Stephanandra Siebold & Zucc.

263) Stephanandra flexuosa Siebold &

Zucc. in Abh. Math.-Phys. Cl. Königl. Bayer. Akad. Wiss. 3(3): 740, t. iV 2 (1843); 4(2): 191 (1845).

Present treatment: Stephanandra incisa (Thunb.) Siebold & Zucc. ex Zabel in Gart.-Zeitung (Berlin) 4: 510 (1885).

Japanese name: Kogome-utsugi.Lectotype (designated here): Japonia.

Siebold s.n. (L0329272) [Category 2] (fig. 147).Original material: Legit in Japonia &

Communicavit de Siebold anno 1842, Herbarium Zuccarinii (M0154024) [Category 2].

Stranvaesia Lindl.264) Stranvaesia digyna Siebold & Zucc.

in Abh. Math.-Phys. Cl. Königl. Bayer. Akad. Wiss. 4(2): 129 (1845), as “Stranvaisia.”

Present treatment: Pourthiaea villosa (Thunb.) Decne. in Nouv. Arch. Mus. Hist. Nat. 10: 147 (1874) (var. villosa).

Japanese name: Ke-kamatsuka. Lectotype (designated here): Legit. Bürger

comcvt. de Siebold, Herbarium Zuccarinii (M0154027) [Category 2] (fig. 148).

Original material: Legit. Bürger in Japonia. Communicavit de Siebold, Herbarium Zuccarinii (M0154028) [Category 2b]. in Japonia legt. Bürger comvt. de Siebold, Herb. Zuccarinii (M0154026) [Category 2b].

LEGUMiNOSAECaesalpinia L.

265) Caesalpinia japonica Siebold & Zucc. in Abh. Math.-Phys. Cl. Königl. Bayer. Akad. Wiss. 4(2): 117 (1845).

Present treatment: Caesalpinia decapetala (Roth) Alston in Trimen, Handb. fl. Ceylon 6: 89 (1931).

Japanese name: Jaketsu-ibara. Lectotype (designated here): Japonia

in monte Wunzen. Siebold s.n. (L0325745) [Category 2] (fig. 149).

Original material: Legit in Japonia et Communicavit de Siebold anno 1839, Herbarium Zuccarinii (M0153878) [Category


2].Siebold and Zuccarini described Caesalpinia

japonica based on Siebold’s collection and cited Caesalpinia crista L. in Thunberg’s Flora Japonica (1784, p. 179). The lectotype specimen L0325745 is a representative specimen upon which the description of Caesalpinia japonica was prepared. Regarding Caesalpinia japonica to be a new species described by Siebold and Zuccarini, we select the lectotype from the collections with annotation by Siebold and Zuccarini.

Crotalaria L.266) Crotalaria eriantha Siebold & Zucc.


Present treatment: Crotalaria sessiliflora L., Sp. Pl. ed. 2: 1004 (1763).

Japanese name: Tanuki-mame. Lectotype (designated here): Japonia.

Siebold (L0423600) [Category 2] (fig. 150).Original material: Legit Dr Siebold in

Japonia. Communicavit Prof. Zuccarini anno 1843, Herbarium Zuccarinii (M0153881) [Category 2]. Legit in Japonia Bürger. Communicavit de Siebold, Herbarium Zuccarinii (M0153882) [Category 2b].

Dolichos L.267) Dolichos hassjoo Siebold, Syn. Pl.

Oeconom.: 55 (1830), nom. nud. Present treatment: Mucuna pruriens (L.) DC.

var. utilis (Wall. ex Wight) Burck in Ann. Jard. Bot. Buitenzorg 11: 187 (1893).

J a p a n e s e n a m e : H a s s h ô - m a m e ; Hassjoomame (Siebold 1830).

Original material: Japonia. [Siebold s.n.] (L0104303) [Category 2].

268) Dolichos incurvatus Thunb. var. [a] Floribus albis seminibus albis Siebold, Syn. Pl. Oeconom.: 55 (1830), nom. illegit. [phrase name]

Present treatment: probably a cultivar of Canavalia gladiata (Jacq.) DC., Prodr. 2: 404

(1825).Japanese name: Natamame (Siebold 1830). Original material: not found in L, M, or

MAK.

269) Dolichos icurvatus Thunb. var. [b] Floribus purpureis seminibus roses Siebold, Syn. Pl. Oeconom.: 55 (1830), nom. illegit. [phrase name]

Present treatment: probably a cultivar of Canavalia gladiata (Jacq.) DC., Prodr. 2: 404 (1825).

Japanese name: Natamame (Siebold 1830). Original material: not found in L, M, or

MAK.

270) Dolichos lablab L. var. [a] Floribus purpureis seminibus nigris Siebold, Syn. Pl. Oeconom.: 55 (1830), nom. illegit. [phrase name]

Present treatment: probably a cultivar of Lablab purpureus (L.) Sweet, Hort. Brit.: 481 (1823).

Japanese name: ingenmame (Siebold 1830). Original material: not found in L, M, or

MAK.

271) Dolichos lablab L. var. [b] Floribus albis seminibus albis Siebold, Syn. Pl. Oeconom.: 55 (1830), nom. illegit. [phrase name]

Present treatment: probably a cultivar of Lablab purpureus (L.) Sweet, Hort. Brit.: 481 (1823).

Japanese name: ingenmame (Siebold 1830). Original material: not found in L, M, or

MAK.

272) Dolichos umbellatus Thunb. var. [a] Seminibus roseis leguminibusque pendulis Siebold, Syn. Pl. Oeconom.: 55 (1830), nom. illegit. [phrase name]

Present treatment: a cultivar of Vigna unguiculata (L.) Walp., Repert. Bot. Syst. 1: 779 (1842).

Japanese name: Sasage (Siebold 1830).

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Original material: Japonia. Siebold s.n. (L0367305) [Category 2].

273) Dolichos umbellatus Thunb. var. [b] Seminibus nigris leguminibusque pendulis Siebold, Syn. Pl. Oeconom.: 55 (1830), nom. illegit. [phrase name]

Present treatment: a cultivar of Vigna unguiculata (L.) Walp. in Repert. Bot. Syst. 1: 779 (1842).

Japanese name: Kurosasage (Siebold 1830). Original material: Japonia. Siebold s.n.

(L0367300) [Category 2b].

274) Dolichos umbellatus Thunb. var. [c] Seminibus nigris leguminibusque erectis Siebold, Syn. Pl. Oeconom.: 55 (1830), nom. illegit. [phrase name]

Present treatment: a cultivar of Vigna unguiculata (L.) Walp. in Repert. Bot. Syst. 1: 779 (1842).


A specimen, Siebold s.n. (L0367301) with Siebold’s annotation “Dolichos kuronagasasake, S.” appears to be part of the original material.

Dumasia DC.275) Dumasia truncata Siebold & Zucc.


Present treatment: Dumasia truncata Siebold & Zucc.

Japanese name: No-sasage. Lectotype (designated here): Legit Bürger in

Japonia. Communicavit de Siebold, Herbarium Zuccarinii (M0153850) [Category 2] (fig. 151).

Original material: in Japonia legt v. Siebold, Herb. Zuccarinii (M0153851) [Category 2b].

H. E. Hazelhorst selected Pierot specimen (L0043333 and L0043334) as holo- or lectotype, and also Siebold s.n. (L0043336) and Bürger s.n. (L0043335) as syntypes in 1977. Although these specimens are fine and well representative, none has indication that they are original material.

Glycine Willd.276) Glycine soja Siebold & Zucc. in Abh.

Math.-Phys. Cl. Königl. Bayer. Akad. Wiss. 4(2): 119 (1845). Verdcourt in Kew Bull. 24: 256 (1970).

Present treatment: Glycine max (L.) Merr. subsp. soja (Siebold & Zucc.) H. Ohashi in J. Jap. Bot. 57: 30 (1982).

Japanese name: Tsuru-mame. Lectotype (designated here): Japonia.


Communicavit de Siebold, Herbarium Zuccarinii (M0153853) [Category 2]. in Japonia legt. Bürger. Comcvt. de Siebold, Herb. Zuccarinii (M0153854) [Category 2b].

One of the original materials, M0153853, has an accompanying label indicating lectotypification by Tepljacova in 1990, but we were unable to find a paper by Tepljacova on this matter.

Glycyrrhiza L.277) Glycyrrhiza kansoo Siebold, Syn. Pl.

Oeconom.: 58 (1830), nom. nud. Present treatment: probably Glycyrrhiza sp. Japanese name: Kanzô. Original material: not found in L, M, or

MAK.

Lespedeza Michx.278) Lespedeza argyraea Siebold & Zucc.


Present treatment: Lespedeza cuneata (Dum. Cours.) G. Don, Gen. Hist. 2: 307 (1832).

Japanese name: Medo-hagi. Lectotype (designated here): Legit in Japonia

Bürger. Communicavit de Siebold, Herbarium Zuccarinii (M 0153856) [Category 2] (fig. 153).

Original material: Japonia. Siebold s.n. (L0103929) [Category 2]. in Japonia legt. Bürger comcvt. de Siebold, Herb. Zuccarinii (M0153857) [Category 2b].


Pachyrhizus DC.279) Pachyrhizus thunbergianus Siebold

& Zucc. in Abh. Math.-Phys. Cl. Königl. Bayer. Akad. Wiss. 4(3): 237 (1846), as “Pachyrrhizus.”

Present treatment: Pueraria lobata (Willd.) Ohwi in Bull. Tokyo Sci. Mus. no. 18: 16 (1947).

Japanese name: Kuzu; Kats, Kudsu, Kudsu Kadsura, Kadsune (Siebold and Zuccarini 1846)

Type: Type of Dolichos hirsutus Thunb. in Trans. Linn. Soc. Lond. 2: 339 (1794).

This was a new name for Dolichos hirsutus Thunb. under Pachyrhizus.

Specimens used for the preparation of the treatment of Pachyrhizus thunbergianus by Siebold and Zuccarini are as follows:

Legit in Japonia. Siebold s.n. (L0327145). Japonia. Siebold s.n. (L0327149). Japonia. Siebold s.n. (L0327148). Legit Bürger in Japonia. Communicavit de Siebold, Herbarium Zuccarinii (M0153873). in Japonia legt. Bürger comcvt de Siebold, Herb. Zuccarinii (M0153874). in Japonia legt. Bürger comcvt de Siebold, Herb. Zuccarinii (M0153875).

Phaseolus L.280) Phaseolus atsuki Siebold, Syn. Pl.

Oeconom.: 57 (1830). nom. nud. Present treatment: Vigna angularis (Willd.)

Ohwi & H. Ohashi in J. Jap. Bot. 44: 29 (1969) (var. angularis).

Japanese name: Azuki. Original material: Japan. Siebold s.n.

(L0367315) [Category 2]. Japonia. Siebold s.n. (L0176058) [Category 2]. Japonia. Siebold s.n. (L0367322, lower left) [Category 3].

281) Phaseolus atsuki var. ooatsuki Siebold, Syn. Pl. Oeconom.: 57 (1830), nom. nud.

Present treatment: a cultivar of Vigna angularis (Willd.) Ohwi & H. Ohashi (var. angularis).

Syn.: Phaseolus chrysanthos Savi f. ooatsuki (Siebold) Honda, Pl. Siebold.: 555 (1935).

Japanese name: Ooatsuki (Siebold 1830).

Original material: Japonia. Siebold s.n. (L0176057) [Category 2]. Siebold s.n. (L0367322, middle right and left) [Category 2].

282) Phaseolus bundoo Siebold, Syn. Pl. Oeconom.: 57 (1830), nom. nud.

Present treatment: Vigna radiata (L.) R. Wilczek, flore du Congo Belge et du Ruanda-Urandi: 6 (1954).

Japanese name: Yae-nari; Bundoo or futanarimame (Siebold 1830).

Original material: Japonia. Siebold s.n. (L0367326) [Category 2].

283) Phaseolus vulgaris L. var. [a] Seminibus albis Siebold, Syn. Pl. Oeconom.: 57 (1830). nom. illegit. [phrase name]

Present treatment: probably a cultivar of Phaseolus vulgaris L., Sp. Pl. 723 (1753).

Japanese name: Sirotorukusun (Siebold 1830).


284) Phaseolus vulgaris L. var. [b] Seminibus rubis Siebold, Syn. Pl. Oeconom.: 57 (1830). nom. illegit. [phrase name]

Present treatment: probably a cultivar of Phaseolus vulgaris L., Sp. Pl.: 723 (1753).

Japanese name: Akasasage (Siebold 1830).Original material: not found in L, M, or

MAK.The specimen, Siebold s.n. (L0367309)

with Siebold’s annotation “Phaseolus aka nagasasake,” appears to be part of the original material because the Japanese word “aka” means red.

285) Phaseolus vulgaris L. var. [c] Seminibus atro virentibus Siebold, Syn. Pl. Oeconom.: 57 (1830). nom. illegit. [phrase name]


Japanese name: Aosasage (Siebold 1830).

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286) Phaseolus vulgaris L. var. [d] Seminibus nigris Siebold, Syn. Pl. Oeconom.: 57 (1830). nom. illegit. [phrase name]


Japanese name: Kurotorukusun (Siebold 1830).


Soja Moench.287) Soja japonica Siebold, Syn. Pl.

Oeconom.: 56 (1830). nom. nud., as “Sooja.” Present treatment: Glycine max (L.) Merr.,

interp. Rumph. Herb. Amboin.: 274 (1917).Japanese name: Daizu; Sooju (Siebold 1830),

Daisu (Siebold 1830).Original material: not found in L, M, or

MAK.

288) Soja japonica Siebold var. [a] Seminibus albis Siebold, Syn. Pl. Oeconom.: 56 (1830). nom. illegit. [phrase name]

Present treatment: probably a cultivar of Glycine max (L.) Merr.

Japanese name: Daisu (Siebold 1830).Original material: not found in L, M, or

MAK.

289) Soja japonica Siebold var. [b] Seminibus albo bruscis Siebold, Syn. Pl. Oeconom.: 56 (1830). nom. illegit. [phrase name]


Japanese name: Tobimame (Siebold 1830).Original material: not found in L, M, or

MAK.

290) Soja japonica Siebold var. [c] Seminibus fuscis Siebold, Syn. Pl. Oeconom.: 56 (1830). nom. illegit. [phrase name]


Japanese name: Sinsjumame (Siebold 1830).Original material: not found in L, M, or

MAK.

291) Soja japonica Siebold var. [d] Seminibus nigris globosis Siebold, Syn. Pl. Oeconom.: 56 (1830). nom. illegit. [phrase name]


Japanese name: Kuromame (Siebold 1830).Original material: not found in L, M, or

MAK.

292) Soja japonica Siebold var. [e] Seminibus nigris combressis Siebold, Syn. Pl. Oeconom.: 56 (1830). nom. illegit. [phrase name]


Japanese name: Kurotokorosun (Siebold 1830).


293) Soja nomame Siebold, Syn. Pl. Oeconom.: 56 (1830), nom. nud.

Present treatment: Chamaecrista nomame (Siebold) H. Ohashi. in J. Jap. Bot. 64: 215 (1989).

Japanese name:. Kawara-ketsumei; Nomame, Jawaraketsmai (Siebold 1830).

Original material: Japonia. H. B. [Hortus Botanicus in Dezima]. Siebold s.n. (L0325738) [Category 2].

The spec imen c i ted above has an accompanying slip with a description of flower, but the specimen has no flowers.

Sophora L.294) Sophora angustifolia Siebold & Zucc.



Present treatment: Sophora flavescens Sol. in Aiton, Hort. Kew. 2: 43 (1789).

Japanese name: Kurara.Lectotype (designated here): Japonia.


Communicavit de Siebold, Herbarium Zuccarinii (M0121586) [Category 2]. “Legit Dr Siebold in Japonia. Communicavit Prof. Zuccarinii anno 1845, Herbarium Zuccarinii” and “in Japonia legt. Bürger comcvt. de Siebold, Herb. Zuccarinii” [This specimen has two labels] (M0121585) [Category 2b].

Wisteria Nutt.295) Wisteria brachybotrys Siebold & Zucc.,

fl. Jap. 1: 92, t. 45 (1839); in Abh. Math.-Phys. Cl. Königl. Bayer. Akad. Wiss. 4(2): 118 (1845).

Present treatment: Wisteria brachybotrys Siebold & Zucc.

Japanese name: Yama-fuji; Jamma fudsi.Lectotype (designated by Compton and Lack

2012): Japonia. Herb. de Siebold (L0176059) [Category 2] (fig. 155).

Original material: Legit in Japonia de Siebold. Communicavit de Siebold, Herbarium Zuccarinii (M0153884) [Category 2].

296) Wisteria chinensis (Sims) DC. var. beni-fudsi Siebold & Zucc., fl. Jap. 1: 90 (1839), as “sinensis.”

Present treatment: Wisteria floribunda (Willd.) DC., Prodr. 2: 390 (1825).

Japanese name: fuji; Beni-fudsi (Siebold and Zuccarini 1839).

Lectotype (designated here): Japonia. Herb. de Siebold in 1829 (L0170538) [Category 5 (fl. Jap. t. 44)] (fig. 156).

297) Wisteria chinensis (Sims) DC. var. siro-fudsi Siebold & Zucc., fl. Jap. 1: 90 (1839) , as “sinensis.”

Present treatment: probably a cultivar of Wisteria floribunda (Willd.) DC.

Japanese name: Shiro-fudsi (Siebold and

Zuccarini 1839).Original material: not found in L, M, or

MAK.

298) Wisteria japonica Siebold & Zucc., fl. Jap. 1: 88, t. 43 (1839); in Abh. Math.-Phys. Cl. Königl. Bayer. Akad. Wiss. 4(2): 118 (1845).

Present treatment: Wisteria japonica Siebold & Zucc.

Japanese name: Natsu-fuji; Ko-fudsi (Siebold and Zuccarini 1839).

Lectotype (designated probably by S. T. Dunn, 1912): Japonia. Herb. de Siebold (in) 1829 (L0059625) [Category 2] (fig. 157).

Original material: Legit Bürger in Japonia. Communicavit de Siebold, Herbarium Zuccarinii (M0153889) [Category 2]. in Japonia legt Bürger comcvt de Siebold, Herb. Zuccarinii (M0153890) [Category 2b]. in Japonia legt Bürger comct de Siebold, Herb. Zuccarinii (M0153891) [Category 2b].

GERANiACEAEGeranium L.

299) Geranium thunbergii Siebold & Zucc. in Abh. Math.-Phys. Cl. Königl. Bayer. Akad. Wiss. 4(2): 136 (1845).

Present treatment: Geranium thunbergii Siebold & Zucc.

Japanese name: Gen-no-shôko. Lectotype (designated here): Japonia.

Nagasaki, Kosido. Sept. Oct. Herb. Thunberg s.n. (L0059752) (fig. 158).

Original material: Japonia. Thunberg s.n. (L0059753).

This is a new name for Geranium reported by Thunberg as G. palustris L. (UPS Thunberg collection nos. 15733 and 15734) in his Flora Japonica (1784, p. 268). Siebold & Zuccarinii (1845) cited Geranium palustris as a synonym of Geranium thunbergii. So from Geranium palustris in Thunberg’s collection the lectotype must be selected.

As mentioned under Spiraea thunbergii (p. 295) we believe that Siebold was able to

302 植物研究雑誌　第 89巻　第 5号 2014年 10月

examine Thunberg’s specimens in the van Royen Herbarium.

Thunberg wrote: “pedunculis bifloris longissimis declinatis, petalis integris, foliis quinquelobis incisis. ... Nagasaki, in Kosido, alibi...” This description well corresponds to Thunberg’s specimens, (L0059752) and (L0059753) in the van Royen collection and Siebold’s specimens (L0176062, L0176063) identified G. thunbergii by Siebold himself.

The following specimens were also used for the preparation of the treatment of Geranium thunbergii by Siebold & Zuccarini:

Japonia. Siebold s.n. (L0176062). Japonia. Siebold s.n. (L0176063). Japonia. Herb. Thunberg (L0059753).

According to ICN Art. 38, Siebold and Zuccarini’s Geranium thunbergii is regarded as a new species, although Yeo (1992) did not accept Geranium thunbergii Siebold & Zucc. as such.

299b) Geranium thunbergii Siebold ex Lindl. & Paxton in Paxton’s flow. Gard. 1: 186, f. 115 (1851).

Present treatment: Geranium thunbergii Siebold & Zucc.

Japanese name: Gen-no-shôko. Lectotype (designated here): illustration in

Paxton’s flow. Gard. 1: 186, f. 115 (1851). Though Yeo (1992) regarded Lindl. &

Paxton as the author of Geranium thunbergii Siebold, the name was legitimately published by Siebold and Zuccarini as mentioned above.

EUPHORBiACEAEAntidesma L.

300) Antidesma japonicum Siebold & Zucc. in Abh. Math.-Phys. Cl. Königl. Bayer. Akad. Wiss. 4(3): 212 (1846).

Present treatment: Antidesma japonicum Siebold & Zucc.

Japanese name: Yama-hihatsu. Lectotype (designated here): Japonia.

[Siebold s.n.] (MAK S1587) [Category 2] (fig. 159).

Two specimens, Siebold s.n. (L0015981) and Siebold s.n. (L0015982), collected in Japan in L bear no annotation by Siebold or Zuccarini.

Cicca L.301) Cicca flexuosa Siebold & Zucc. in Abh.

Math.-Phys. Cl. Königl. Bayer. Akad. Wiss. 4(2): 143 (1845).

Present treatment: Phyllanthus flexuosus (Siebold & Zucc.) Müll. Arg. in DC., Prodr. 15(2): 324 (1866).

Japanese name: Koban-no-ki. Lectotype (designated here): Legit in

Japonia Bürger comcvt. v. Siebold, Herbarium Zuccarinii (M0120849) [Category 2] (fig. 160).

Original material: Japonia. Siebold s.n. (L0159844) [Category 2]. Japonia. [Siebold s.n.] (MAK S1595) [Category 2]. Japonia. [Siebold s.n.] (MAK S1596) [Category 2].

Croton L.302) Croton siraki Siebold, Syn. Pl.

Oeconom.: 30 (1830), nom. nud. Present treatment: Neoshirakia japonica

(Siebold & Zucc.) Esser in Blumea 43: 129 (1998).

Japanese name: Shiraki. Original material: Japonia. Siebold s.n.

(L0022693) [Category 2]. Japonia. Siebold s.n. (L0022695) [Category 2]. Japonia. [Siebold s.n.] (MAK S1592) [Category 2, 3].

302b) Croton siraki Siebold & Zucc. in Abh. Math.-Phys. Cl. Königl. Bayer. Akad. Wiss. 4(2): 144 (1845). nom. nud.

Present treatment: Neoshirakia japonica (Siebold & Zucc.) Esser.

Japanese name: Shiraki. Original material: Legit v. Siebold in Japonia

et Communicavit, Herbarium Zuccarinii (M0120848) [Category 2b]. Japonia. [Siebold s.n.] (MAK S1592) [Category 2]. These are also the original material for Croton Siraki Siebold (see above).


Glochidion J. R. forst. & G. forst.303) Glochidion obovatum Siebold & Zucc.


Present treatment: Glochidion obovatum Siebold & Zucc.

Japanese name: Kanko-no-ki. Lectotype (designated here): Legit Bürger in

Japonia. Communicavit v. Siebold, Herbarium Zuccarinii (M0120852) [Category 2] (fig. 161).

Original material: Japonia. Siebold s.n. (L0030948) [Category 2]. in Japonia legt. Bürger comunic. v. Siebold, Herb. Zuccarinii (M0120853) [Category 2b]. Legit Dr Siebold in Japonia. Communicavit Prof. Zuccarini anno 1843 (M0120851) [Category 2b]. Japonia. Siebold s.n. (MAK S1534) [Category 2]. Japonia. Siebold s.n. (MAK S1535) [Category 2].

Mercurialis L.304) Mercurialis leiocarpa Siebold & Zucc.

in Abh. Math.-Phys. Cl. Königl. Bayer. Akad. Wiss. 4(2): 145 (1845), as “lejocarpa.”

Present treatment: Mercurialis leiocarpa Siebold & Zucc.

Japanese name: Yama-ai. Lectotype (designated here): Legit. in

Japonia & Communicavit de Siebold, Herbarium Zuccarinii (M0120841) [Category 2] (fig. 162).

Original material: Japonia. Siebold (L0080015) [Category 2]. in Japonia legt Bürger comunic. v Siebold, Herb. Zuccarinii (M0120842) [Category 2b]. Japonia. [Siebold s.n.] (MAK S1538) [Category 2]. Japonia. [Siebold s.n.] (MAK S1539) [Category 2].

Phyllanthus L.305) Phyllanthus lepidocarpus Siebold &

Zucc. in Abh. Math.-Phys. Cl. Königl. Bayer. Akad. Wiss. 4(2): 143 (1845).

Present treatment: Phyllanthus lepidocarpus Siebold & Zucc. (see T. Kurosawa in Acta Phytotax. Geobot. 52: 18, 2001)

Japanese name: Ko-mikan-sô.

Lectotype (designated here): Communicavit de Siebold anno 1842, Herbarium Zuccarinii (M0088455) (fig. 163).

Original material: Japonia. [Siebold s.n.] (MAK S1542) [Category 2].

Siebold and Zuccarini (1845) compared Phyllanthus lepidocarpus with “Ph. Niruri Thunb. fl. Jap. p. 56” with a question mark. in L and M there are no specimens determined as P. lepidocarpus by Siebold or Zuccarini. The lectotype designated here, M0088455, was annotated at first as “Phyllanthus Niruri L.” and then “P. echinatus S. & Z.” by Zuccarini. Phyllanthus echinatus Siebold & Zucc. was not published, but the epithet echinatus has a meaning similar to lepidocarpus. We consider that Siebold and Zuccarini described P. lepidocarpus based on this specimen.

306) Stillingia japonica Siebold & Zucc. in Abh. Math.-Phys. Cl. Königl. Bayer. Akad. Wiss. 4(2): 145 (1845).

Present treatment: Neoshirakia japonica (Siebold & Zucc.) Esser in Blumea 43(1): 129 (1998).

Japanese name: Shiraki. Lectotype (redesignated here): Legit. v.

Siebold in Japonia et Communicavit, Herbarium Zuccarinii (M0120845) [Category 2] (fig. 164).

Original material: in Japonia legt. Bürger comunic. v. Siebold, Herb. Zuccarinii (M0120843) [Category 2b]. in Japonia legt. Bürger comunic. v. Siebold, Herb. Zuccarinii (M0120844) [Category 2b].

Peter van Welzen designated a specimen “Japonia. Siebold s.n. (L0081028)” as the lectotype (Esser 1998) but this is not original material because it lacks an annotation by Siebold and/or Zuccarini.

ReferencesAkiyama S., Thijsse G., Esser H.-J. and Ohba H.

2014. Siebold and Zuccarini’s type specimens and original materials from Japan, part 4. Angiosperms. Dicotyledoneae 3. J. Jpn. Bot. 89: 209–229.

Aldasoro J. J., Aedo C., Garmendia f. M., de la Hoz f. P.

304 植物研究雑誌　第 89巻　第 5号 2014年 10月

and C. Navarro. 2004. Revision of Sorbus Subgenera Aria and Torminaria. Syst. Bot. Monogr. 69: 1–148.

Compton J. and Lack H. W. 2012. The discovery, naming and typification of Wisteria floribunda and W. brachybotrys (Fabaceae) with notes on associated names. Willdenowia 42: 219–240.

Esser H.-J. 1998. Neoshirakia, a new name for Shirakia Hurus. (Euphorbiaceae). Blumea 43: 129–130.

Hara H. 1957. Critical notes on some specimens of East-Asiatic plants in foreign herbaria (8). J. Jap. Bot. 32: 134–140.

Hara H. 1978. Comments on the East Asiatic plants (5). J. Jpn. Bot. 53: 232–238.

Honda M. 1938. Plantae Sieboldianae. In: Japanisch-Deutsche Gesellschaft, Tokyo (ed.), Siebold Kenkyuシーボルト研究 , pp. 517–596. iwanami shoten, Tokyo.

Hu S.-Y. 1955. A monograph of the genus Philadelphus (continued). J. Arnold Arbor. 36: 325–368.

Koidzumi G. 1911. Notes on Japanese Rosaceae. ii. Bot. Mag. (Tokyo) 25: 74–76.

Koidzumi G. 1911. Notes on Japanese Rosaceae. iii. Bot. Mag. (Tokyo) 25: 183–188.

Koidzumi G. 1913. Conspectus Rosacearum Japonicarum. J. Coll. Sci. imp. Univ. Tokyo 34(2): 1–312.

Makino T. 1908. Observations on the flora of Japan. Bot. Mag. (Tokyo) 22: 113–120.

Maximowicz C. P. 1883. Diagnoses plantarum novarum asiaticarum, V. Bull. Acad. imp. Sci. Saint-Pétersbourg 29: 51–228.

Ohba H. 2001. Prunoideae. In: iwatsuki K,. Boufford D. E. and Ohba H. (eds.), flora of Japan, 2b: 124–144. Kodansha Scientific Inc., Tokyo.

Ohba H. (ed.) 2012. Tetsuya Kawasaki’s illustrated Works of Japanese flowering Cherries. 222 pp. Aboc-sha, Kamakura.

Ohba H. and Akiyama S. 2013a. Rosa collected by Siebold

and his collaborators from Japan. J. Jpn. Bot. 88: 197–211.

Ohba H. and Akiyama S. 2013b. A revision of the species of Hydrangea (Hydrangeaceae) described by Siebold and Zuccarini part 1. Bull. Natn. Mus. Nat. Sci., Ser. B, 39: 173–194.

Ohba H. and Akiyama S. 2014. A revision of the species of Hydrangea (Hydrangeaceae) described by Siebold and Zuccarini part 2. Bull. Natn. Mus. Nat. Sci., Ser. B, 40: 29–45.

Ohba H., Akiyama S., and Thijsse G. 2005. Miquel’s new taxa of the vascular plants described from Japan in Prolusio florae japonicae and some other works. in: Ohba H. and Boufford D. E. (eds.), The Botanical Collections: Proceedings of the Symposium 'Siebold in 21st Century' Held at the University Museum, the University of Tokyo, in 2003. The University Museum, the University of Tokyo, Bulletin no. 41: 31–140.

Ohba H., Kawasaki T., and Tanaka H. 2007. flowering Cherries of Japan, new ed. 263 pp. Yama-kei Publishers Co. Ltd., Tokyo.

Sargent C. S. 1888. Plant notes. Prunus pendula. Garden and forest 1: 196.

Thunberg C. P. 1784. flora Japonica. J. G. Mülleriano, Leipzig.

Yamaguchi T., fujii S., Tamura M. N., Nagamasu H., Kato N., Wada H., and Yamamoto A. 2003. The specimens of Japanese plants collected by Ph. f. von Siebold, H. Bürger and von Siebold's collaborators, and now housed in the National Herbarium of the Netherlands and the Botanische Staatssammlung München. Calanus (Bull. Aitsu Mar. Stat. Kumamoto Univ. Jap.) Special No. 5, 191–576.

Yeo P. f. 1992. A revision of Geranium L., in south-west China. Edinb. J. Bot. 49: 123–211.

秋山　忍 a， G. Thijsseb， H.-J. Esserc，大場秀章 d：Sieboldと Zuccariniが日本から記載した分類群レクトタイプと原資料，第五部．被子植物．双子葉植物綱　四　Sieboldと Zuccariniは，主に Siebold自身および，日本人植物学者，Bürgerらの採集品からなるシーボルト・コレクションを研究し，日本から多数の新種や新変種を発表し，日本植物相の分類学的研究に多大な貢献を行った．しかしながら，それらの新植物の発表に際しては大半で標本の引用がなく，またタイプの指定がなされなかった．　Sieboldと Zuccariniが命名した学名の安定化などの目的で，ライデン，ミュンヘン，牧野植物標本館（首都大学東京），東京大学総合研究博物館に収蔵されるシー

ボルト・コレクションから Sieboldと Zuccariniが記載に用いた標本（原資料）を標本上に記された手記ならびに記載文などを参考に特定した．また，タイプが指定されていない分類群については，特定した原資料のなかから記載に最もよく合致する標本をレクトタイプとして選定した．　本稿はその第五部で，被子植物双子葉植物綱（ユキノシタ科からトウダイグサ科）を収載した．レクトタイプの選定に関連して，とくに Deutzia crenata（ウツギ），Parnassia mucronata（ウメバチソウ），Philadelphus


satsumanus （バイカウツギ），Sorbus toringo （ズミ）， Geranium thunbergii （ゲンノショウコ），Phyllanthus lepidocarpus （コミカンソウ）では選定について問題点などをコメントした．　Amygdalus persica f. pendula（ベニシダレモモ），Amygdalus persica f. stellata（ゲンペイキクモモ，新称），Amygdalus persica f. versicolor（ゲンペイモモ），Armeniaca mume f. pendula（シダレウメ），Cerasus itosakura（エドヒガン）の新学名を提唱した．　エドヒガン（シダレザクラ）に Sieboldが与えたPrunus itosakuraは，これまで、記載を伴わない裸名とみなされてきたが，解説文中に “Cerasus ramulis pendulis（下垂性の小枝をもつサクラ）”とあり，当時の日本産自生種中で下垂性サクラはエドヒガンしかなく，この種の特徴を記述したものと理解することができる．なお，Sieboldはサクラは属としては Prunusに分類しているので，この解説文中の Cerasusは属名としてではなく，サクラ類の意味で用いている．そこでPrunus itosakuraを有効名と断定し，上記の新学名を提唱した．

　ズミの学名について，原 (1978)はすでに小泉が詳述しているとし，Malus toringo (Siebold) Sieboldが正名であるとした．しかし，小泉 (1911, 1913) はMalus toringo Sieboldの出典を Sieboldの Catalogue Raisonné (1856) としたが，原は de Vriese としCatalogue Raisonné には触れなかった．Catalogue Raisonné (1856)でのMalus toringoの発表は短い記載を伴い新種としての要件を備えているが，同じ 1856年に de Vrieseによって Sorbus toringo Sieboldにもとづく Malus toringo (Siebold) Sieboldが Tuinbouw-Flora 3巻 368ページに発表された．国際藻類・菌類・植物命名規約第 41条 4項に従い，Malus toringo Sieboldの学名は Sorbus toringo Sieboldにもとづく新組合せとし，その出典を Sieboldの Catalogue Raisonné (1856) とするのが適切である．

（a国立科学博物館植物研究部，bオランダ・国立標本館

現所属：オランダ・国家生物多様性解析センター植物部門，cドイツ・ミュンヘン州立植物標本館，

d東京大学総合研究博物館）

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Rhod

otyp

os k

erri

oide

s Si

ebol

d &

Zuc

c.

(L03

2926

9).

fig.

139

. L

ecto

type

of R

osa

iwar

a Si

ebol

d ex

Reg

el (M

0154

345)

.


fig.

140

. L

ecto

type

of R

ubus

ribi

foliu

s Sie

bold

& Z

ucc.

(M01

2086

4).

fig.

141

. L

ecto

type

of

Siev

ersi

a dr

yado

ides

Sie

bold

& Z

ucc.

(L

0175

827)

.


fig.

142

. L

ecto

type

of S

orbu

s tor

ingo

Sie

bold

& Z

ucc.

(L01

7602

2).

fig.

143

. L

ecto

type

of S

pira

ea p

runi

folia

Sie

bold

& Z

ucc.

(L01

7604

0).


fig.

144

. L

ecto

type

of S

pira

ea th

unbe

rgii

Sieb

old

(L01

4431

0).

fig.

145

. L

ecto

type

of S

pira

ea th

unbe

rgii

Sieb

old

ex B

lum

e (L

0176

042,

le

ft).


fig.

146

. L

ecto

type

of S

pira

ea th

unbe

rgii

Sieb

old

& Z

ucc.

(L01

7604

1,

left)

.fi

g. 1

47.

Lec

toty

pe o

f St

epha

nadr

a fle

xuos

a Si

ebol

d &

Zuc

c.

(L03

2927

2).


fig.

148

. L

ecto

type

of S

tranv

aesia

dig

yna

Sieb

old

& Z

ucc.

(M01

5402

7).

fig.

149

. L

ecto

type

of C

aesa

lpin

ia ja

poni

ca S

iebo

ld &

Zuc

c. (L

0325

745)

.


fig.

150

. L

ecto

type

of C

rota

laria

eria

ntha

Sie

bold

& Z

ucc.

(L04

2360

0).

fig.

151

. L

ecto

type

of D

umas

ia tr

unca

ta S

iebo

ld &

Zuc

c. (M

0153

850)

.


fig.

152

. L

ecto

type

of G

lycin

e soj

a Si

ebol

d &

Zuc

c. (L

0103

661)

.fi

g. 1

53.

Lec

toty

pe o

f Les

pede

za a

rgyr

aea

Sieb

old

& Z

ucc.

(M01

5385

6).


fig.

154

. L

ecto

type

of

Soph

ora

angu

istif

olia

Sie

bold

& Z

ucc.

(L

0421

682)

.fi

g. 1

55.

Lec

toty

pe o

f W

iste

ria

brac

hybo

trys

Sie

bold

& Z

ucc.

(L

0176

059)

.


fig.

156

. L

ecto

type

of W

ister

ia c

hine

nsis

Sieb

old

& Z

ucc.

var.

beni

-fuds

i Si

ebol

d &

Zuc

c. (L

0170

538)

.fi

g. 1

57.

Lec

toty

pe o

f Wist

eria

japo

nica

Sie

bold

& Z

ucc.

(L00

5962

5).


fig.

158

. L

ecto

type

of G

eran

ium

thun

berg

ii Si

ebol

d &

Zuc

c. (L

0059

752)

.fi

g. 15

9. L

ecto

type

of A

ntid

esm

a jap

onicu

m S

iebol

d & Z

ucc.

(MA

K S

1587

).


fig.

160

. L

ecto

type

of C

icca

flexu

osa

Sieb

old

& Z

ucc.

(M01

2084

9).

fig.

161

. L

ecto

type

of G

loch

idio

n ob

ovat

um S

iebo

ld &

Zuc

c. (M

0120

852)

.


fig.

162

. L

ecto

type

of M

ercu

rialis

leio

carp

a Si

ebol

d &

Zuc

c. (M

0120

841)

.fi

g. 1

63.

Lec

toty

pe o

f Ph

ylla

nthu

s le

pido

carp

us S

iebo

ld &

Zuc

c.

(M00

8845

5).

330 植物研究雑誌　第 89巻　第 5号 2014年 10月

fig. 164. Lectotype of Stillingia japonica Siebold & Zucc. (M0120845).

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Siebold and Zuccarini’s Type Specimens and Original ...—279— Siebold and Zuccarini’s Type...

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