by'

J~lanta Koszteyn

Not to. bc ci.tcil,:-"itl1oJIt prior: refq,rcnce"to thc author

o.M. 1983/L: 11Biological OceanographyOomittee "

t . . .

International Couneil forthc Exploration ot thc Sea

Sea F~scheri~s Institute, 'Al~ Zjednoczenia 1

81-345 GdYn1a, Poland

Differcnces in the species composition of mesozoopl~onic

communities in the southern Balt1c in July, 1981

'Abstract,

", During a research oruise of the "Profesor Siedlecki" in' ,July, 1981

over '100 mesozooplankton1c 'samples 'wo're taken in the' Polish' flsheries. ,', .' . . .

. zone. After 'qualitative and quantitative analysis ofthe collected'mr\te~iai,'fiv~ areas~: d1fferingwith respect to the species composi­tion'of mes~zooPlanktoniC communitiesliving il:1'the waters of the'southern Baltic were differentiated.

:-' ' , !

. ','" Bet\<reen 'July 9" ~'d29,' 1981-~ du.rlng a' eruise by R.V. i'Profesor,

Sic(~lecki"~.:co~piex"investigations of the environment and the liv1ng" r'esource~;''.~ft'h~,:Pol1sh 'fisheries zone inthe 'Baltic as weIl aD of

water" poifuti~n wE!re ,~onducted. Hesozooplanktonic samples from' over.',1:00 stations'were'al'so 'taken; their l~cation i8 presented in Fig. 1.

" I', .," ", '~,.' - •• •

" Thc ',preserit ,-Paper' discusses the re~mlts'of thc first, stage of in-vt~~tig~tlO~S into the horizontal distrib~tlon of mcsozoo'plankton,with sp~cial.empha~is'on: '

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, J , /,.

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,,; '- • .' •' " .' '.' ,:., " . '. " - : ": ... ',. ' ... '. .. ~ ,,' I:" '. ~ , " , ,; :,'. ':,;. '... , ; . \ " ,

, 1/: the. differenoes in, the quantitative distribution of 0.11': mesozoo;';'·:pi~kto~ :a~d '1ts sele'oted repres~nta:t1ve~~'·~:.:;"'. \:,::.,,:.,~::.<~<: .i,:";,.:, .. , '

2/:·ble,diifere~ce~, inth~ 'sp~o'ie~ compositi~n of'~eri~~o~~la:ni~to~ic': "., ~.c~~uriitie·s:li~ing in:~'the" ~aters' :~Of: the"'aouth~tn, BaitId.' :",;:.' ,'.. ':.:,." ' ;, -,,"

, , \' .'~,"',' ~" .' .. . . . ' : .,;" \ :.·::, ,,;.!,., ':'~. ,~. . . .... . :- -' ;:- . : , : ~ ., ... ~ .':' .. . .,.' ..'" '.-,' -, '. >

Matkriai~~ci:m~th6d.··:;", ,"," ,: .. ',:.. .,,' ". ",',\', "

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.' ','. "'l<: ,.".... ·l.~. .' "," '.. ", ,.,,' '~I_;.. ; '.,":' ":.,' . ~./,' " ".' '. - ;_ .' :" ~ .. ~. " ::~':' .

'... :;;.Z~?pl.c.mk1;0:nio:samples' were' tuken' bet''1een July. ~ 10 :'and '25,. .~ 1981 ':,', ': :',W:ith,: 0:'., ~o1s'eri net·"/~:'202::unil·· in"singl~', haulS:.fro~··t'he .bottomt6: the···,'::. '~ur'f~ee~\ 'T'he ~6:11~cted :tnat~riai'was' p'r~s~i-ved <lli: 4 %'-fornialiI~. -< ': ;,, ::':.' '::<:ci~aii t:~{i~~': ~d' q~~nt it'ati've: b.na.lY·Si~-'~6~~'red'~ai~iy: mci'sozciopi~­. ," ,t6i(~hich~,~6c~rdi~~ 'tci',the':~;~,e'co~ertdati'o~f/ ,,"n" the, lri~th6do'för: marine·_, 'tfi~i6'gic;ai::atudiea '. in 'the:Baltic 'Sea" ,)1'976/ inolud~s"org~isms'with ', • ) :; ~ +. ,_ .' ' • , ." \ , • ., ,'. " ~: l : .' •• , , > .' • • ..' • • ~ • • ,

':.' dimensions ·'trom 200)lIIl' to .1' II'JD.. ,:" .. " ".,' :, \,'", '",",:, ", :. ., .'

':':" ),:'::Me"sriioopl'hrt~6'n', c6rri:pö~e~t's'\'1efe:, in'mos't 'oa~es '.d~te~niined. t6, ~p~­,'o.i:eo:~:or,' ge'rt~'s·~.::"{ith':.th~:'exceptibn,'of;na~plüil""~tage~, 'of "Cop~poda,o.n:d ", :Cyr~ipe~Ua';:a:nd:' l~~~~l :' s'tage~ ;~f, :Poiychae't~" ~d,Biov~i~i~~" .,.,:~ ~" ': ',' ~ '.

'.!. .,;,•.•.".,'.~' ',' .\:~,.' ,:.,'.~,"... ",. ':,~' ',.' ....'~'.~ • ',' ' 1. "', .,' ",' '.",'.;.,.,',:' "'\':' ...

: "<The':,rosults ·.at· th.'e·se·analyses, were ,to serveto 'croate:a, ;sylithetic,',\ ~i6tci:~'·"'of::t1ie·:qh~t;it~ti~e",~listr'ibut.i~n' of' o.ll~'~es~z·O()Pl~kt~n~nd:,'

f •• ,.".':: ':.',' '":,,,.,,', ',- ••.• ,~' .... ' <' ,"" ...... '-' .~.:.-! .... ~ l .' ,i :'. " ", '. ',:' '. "_ ,'" ~ -.••••• ," , '.~ ,_ " ,.", •.\; ',},', ~•• ',',.:',',"

, :;its selected' speoies. 'Therefore~" it was' neoessary, to, find, a relative-,.•_....,1.'~·"._ f<.•_~":~~~_:, •. -";···.,~··::,, ,,', ' ''''' .. ,'' ,,' ",.~:.•••• :,' "'~ " ••••• : ••••.' •••.•• ".' .•• : •.••••.,:•• ':,'.' • " ....

ly, objective'method' of estimation of ,the,·size of differenoeo'in the' ,,ab~~d~~e,'o'i" org~i'smsamong 'l~div.iduai, samplirig' stat'io~s';~~ .~'ori,;,ert '

<. :.... '. i .,::. : ~. .I, : i .. '. '. ; . ',' '" '.: "',: ,. . ' : ' ' . '..,' " " . ','. I.. .' '. : ~. , •

,a ·set,of"singular' abundance.'values. into a frequenoy distr1bution~" :,-'::>:' ',':~ To'·.'~6iii6~(:':th1;~,.':,'~li'ciai a', ~erc': 'converte'd .. lxit0 'no~m~liz~d" ~ie~ents .. '

./~hi~h' ,'aet~~in.~,.;'bY,':"hb~',nianY.~t~d~~d>~deviation. ':,/a: c~ricrät'e: eie~ents .• .. ~ I,', ':<,', ': ," '." /., :".~ "'~'': . ;', ,'; ;,:',',.' ,..1 " _' , .•.. ~., " t"" ...... " '''',•.'_ ',_ ,.'.,._ .', .,,'; .:':. ~" ~, : '.'

differs,;from~'the":'I:lean,ofthe ,setof',observations to wh!ohit·belongs .....The'-, ~·aicui~ti·~ri·~' :~er~' rdad'~:'~~·c:o;ding to th~, f611~~irig, fo:bn~ia.· iBi;gu:;.~ ~~i','.. ', /"'~'.',:"" ',. :. ,.. , ,",' "', ,e"", ".': ,,-.':' '., : -6_cki .1979 .". " ., .... ',.', : .,"'." .' ' ':" , : , . ,,' . '". .' ' •

':':","?;;""':,i,,':'::'.,;~ ••.~,._"~;~~-"" ", ." .....•'.,•.•. ,'.•' •,"', .~ .' ,.,where: '·z Q.',normalized:element,>" i ,"",., "",;" ,'.. . ...., :,.," ..

},:".' :'.:. <~,·:'x;~;"~umbe;.~f"or~anisms,at'a· Eitation,"" . '.': ':'.:'.':>." ",'::: ','::....:' ...: ~,:,;:'i:~::',~~an '~h.~b~~~oi org'Oni~k~':i~:the:'inveatigated area,' ,"

.. :,:':,,{f;jtrilid~d ,d~~iQr\o~\, "." ...•'. " . ..", .•~ ', '.'. :•"'. ' .'";',:' ,'.': The ,frequericy .:distribut fon' of ,'norma11zcd. elementa .made' i t ~ (lusier ,.'th.'di~tih~{likh·::~~'~~a' 'O{':vl~ter""'hich"diff~~ dlstiri~tly .,with.·r~spect to'

,:~th~~ '.~~bcr,;·oi:·6~g'alÜSriis·tI1~an'riucihe~' '01: inlUvldua.ls tind:the" startd~rd:':'·ri~ror>:~~f.,'th"e:'~~~".~~~e '·'o~lcUi~ted:'.iorthe's·e areaa. \:',:' , ,',"'" ".,' " " ','

. ". .. .. , "; "', ...: '. .. ..... .:", ..... -. '~ .. ' '. , , . . .' ,;' . . ',' ' ., :.. '

" '. ~ .. . .• .' _.::'! '. :'~: .1:' '..: .~ ,',~ .• '.' .. , '. ,: : . '. \ :,' ',: . ,": ,.:. '\ .. - ,j..... . ',' ~. ~ • . ,,< " • . ...

Results :,,> :,~':.:: < "'. '.";..;.,;;,;;;.;;;;.;;..;;.,;;;;" "'. " .' .,,' "

• ". ' .' .•••,t " '7'." '.' '", • • , • . •

In the, mcsozooplanktonio oamples investigated 23 ,speoie,s. or"highertuxono~io' unit~:ware .id~riti:f::l.e,d' /T~bie' 1/ ~ , .

,

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Thc reoults 'of tbc'quantitativc nnnlyoio'of,tho collectcd material,are preoente'd on thc' mnpo of th~:quantitative distrib~tlon'of0.11 .meoozoopi~nkton, nnd' ito oelectcd' :rcprc~cntativco,/Figo .2-9/ • '

. \'lhat 'ean bc ri~en' nt· first sight.:1sthe, exccptionul' rlchncoG of thc'~oa~tai.''1aters.'~whose boundo.r~'isa~prox1mateiy.dcsignated' by thc 30 misobath~ Esp~(dallY' large l1Umb~rs,:of mecioz~oplanktonwere .~bse~vednear. thc 'c~tuary: of the S~uPlaIÜver /aimoot 46,000 ind'./m3/ the lJebaRive~ /~bo:ut, 33;600 1nd./m3, ,theRega" Ri~erlo.bout '28,000 -ind./m3/., :'

: und the Viotula:/over ,22,500 ,1nd./m3/ aS\'/cll as'in the' coaotal'\'laters, oi th~ '. P~rn'orsk·a. Bay '/~bout "3,bOO fnd~/m3/. Tho, ~ituation ob~crved ..w~st 'OfKoa:obrz~gwas'untypicalfor 'these \'Ia'ter~-;' only .3; 600 lnd./m',

.w~re' :f6~d th~r:~",' ..' :.'. ".'" .' ..,,',.. . .., ': .. The lowest,values· of the abundance' of mesozooplankton \-lore noted.. in the' wat"~'rs "'~i ,'the ~orth-eastern part' of th~ invest1gated .'area ;"..' :from ab~ut .2,400 .1nd./m'. over the' slopes o:f the Götl~d Deep to

" :.' 6',100', irld./k3 '.in'the "S~pskFu~row. In the south";eastern und 'ooU:th­we~t'er~':part;s"'oi'th~"inve~tlgated ~ea the'abundance v~lues ·ranged

:'.i'ro~ 6~500.·'.t'~"1"~OOO i.nd·./in3~ ',"-' .".', 'Th!s I,'gen~rai '~ictu~e',1s ,supplemented by simlla'riy prepared maps

0:( the"qu~tltativedi~trlbutionof oeleeted :masozooplanktonie ope-.Ci~sjFlgs'·3~91.:·' ' :..., - ' .

, ',:Pseudocaf~nus 'elongatus /Fig., 3/ was ,most abUndant in the' central'. ~d,north~we'~ter~ part. cf the investig~ted area l:f~om about 2,000 ":in:d./ni3, tri' ·o~e~".1'O, 500·'ind./m3/ ~ J.large .];lUmb~rsoi 'tlil0 "ßpecies wer~ .,also :.oboerved,'lIl th~"'centrai part of thc Gulf' cf Gdansk Ifrom' 4~100

: '. t'o' 5~obo·'lrid./m3 i~·"'In. the coastal'water~, p~elongntu~ was les~ ',: .. ,:, ab~da.nt, }iibolit::"ob<irid./ni3 , ~~.·the aver~ge/ or "was~n~t ,obsQ~ed at' all•

· " ., ,.; ". • :. j .• ; I .•.~; : ". ,',.' • ; ". " • .'. . : • • '.' , • . • . •

· .... .In·.,the,icaoe. of·Acartia"sPP.·, ,thesituation was reversed': /Ffg. '4/..,"". Th~-:highest·· abuhd~e~' :values' \'ler'e 'noted i~ the coas't~' zon~ lup to'-

'; ;3~,O~O,,~~,~.~)in~(~"',~~tJl'~t~e eX:cePt1c?n-~r:>.~~)~~~.waterS"'ie's~ of-Ko3:o,brz'eg~. " In dee':peI.<wäters~:·'Aeartlaspp';" were\r~:Lati"ely 'scarse' /about 900 ", ..· :' :i~'d~/mj /."ahd.'~t':'~e~erai': statio~s they:'~~r~"~ot obse'rved at ~ll.' ".' Amo~·!6th~'~.r·~·p~es~~tativ~~' of Copepoda,' 'tll~"(Üot;ibution'~:fTemora I

" lo~gico~1~,·J.'~'~·6~·tii~6ting/Fig. 5/ •. The 'relatlvely low ~bund~ce 'of• ~. ",: ,.1 " ',.. • .• .' _ . ':' .•" ': ~ . • .' ,...' • • . . •

·thio species,in·the.waters·of almost the whole lnveatigated'area wao\, ," , , ,', " '. '.:', .' • ,: ., • , .' • • " . '. r '

': IJtriking~·. The exoeptlon was the south~\'Ieatern·.part of the area, 'espc-'ciali.y th~' GUlf' 6f Gqanok, where' T.1ongi~or~'l1S formed con~enti:atlonsnunlberin'g~b~ut 'S,000).nd./m3 arid even ov~r :11,000 lnd./m'.· .. " .':, ··It· l's ·,.aloo,' worth~entionin'g Eurytemora . op'~, which \'las most abundantin .shalio~·,water·s·'/Fig. 6/. Especially large numbers of thc. rcpreocn­tati.;/eo o~this' ~pecies \'/~re .'observed ·in the 'Gulf of', ~dan3k ,- ovcr

f

, I

~'"

4 -

32,000 1nd./m'.Oladocora ~ere ropreaented bY,Evadne n~rdmanni,Boomina co~oaoni,

maritima ,and Podon 'sPP. Thc, first of these specieo was 'most abundant/up ,to 4~OOO ind./m"3/, esp'eciallY in the central part, of the lnveoti­gated aren. tEig", 7/. The" r'emaining twospecies occurr'ed' in' thecoastal zone~, B~slllina, core'goni ~aritima was ID,?st abundant, /up to2~900 ind~/m3/in ,thc waters"ofthe central part.of:th~ const andatthe orlginof the-Hel Peninsula /Fig. 8/.,Thel~gest numbcra'of PodonsPP./up to3,'OOOlnd./m3/ were, observed off the' western coastand inthe:shaliow~wa.terz6ne'oftheGulf of··Gdansk /Fig. ,9/.,

,I'

Discusaion

'Aso. result' of the qualitative ~d quantitative analysis of meso­zooplankton 1t co.uld be seen that:

, , , " " -, \ , '

1/ the investigated waters of the southern Baltic differed with'reo-'-,. , :' , ' 'r

',' peet to, thc abundance of all' mesozooplankton, , "'2/ 'the,differences in ,the quantitative distribution of ,the sp'eciea '

and'genera.' diaoussed above distinctly showeddifferences in,thespe~i~s'compositionofmesozooplanktonic oomm~itiesiiving inthis aren.',.Thissecond' observation is illustrated by, Figs 3-9 and t\'10 maps

!Figs '1 Oand '11/ .' on whlch 'th~ areas of' the most abundant oocurrenoeI oi'Cope~oda an'dCladocera are marked. As c'an be, seen fromFig. 10,''the abundance aIld species 'composition of mesozooplankton in the oen­tral~ 'n(;r~hern,andnorth-\...estern' part of the inv~stigated ares. were'

, "domlIlated mostly by Pseudooalanus elongatu's, in the, coastal part _.:by' Aoart'ia'spp.'; 'in, the south ~ast, T~mora longicornis was also a .-quantitativelyimportant element of mesozooplankton. , '

'" ' ' . The infiu'~nce, of' Cladooeraon the abundance of mesozo~planktonwas.',, ,~mailer "but,'no:','dotibt \hOY' shaped the species, composition of 1ts' commu­":'riit"les. 'Ito~',.b~':seen from 'Fig. 11 that 'Podon spp. and Bosminacore~

, , '. gon!' mro:.ltl~n: 60noentrat~dmainlY in the ,coastal zone while 'Eva~ne:no~dm~i:"p~ef"'erredd~operwaters.· .• ~ .'

, " " ;", ,"., ' " 't .'.. ," " .",. ,- • ..' ..

':'.c,Thus, the,arialysis'of the distribution ,of individual speoies and" the' ,qu.mtit~tive"rel~tions among them allowed to dif'ferentiate' five,areaa differing "with re~pocit tothe specles composition C?f mesozoo~

':,plankt:onlc "c~m.'1lunities:;':F'ig. 12; Table II/.,. , , ' Area ,'A was'cha~a~:teriz~dbY aclearlY visible dominance ofAcartiaspp. ~whi~h constituted over 64 % of all organi~ms. As may be oasJ.lyseen, thlsarea.ovcrlapped 'more or less with the area:ofthe most~bundant occ~rence of thisspecies /Figs 4, 10,12/. Thls is under­

<:stand'ablo if.: we t'ake into account ecological requi~emonts of Acartia

I, ,I

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.;·spp·", 'eopeclallY A.·b1f11osa. It is 0. ~ surface, eurythermal' species, .,with an. oPtiin~ temperature of over1 o' Oe 1Ackofors' and Herl1r~th' '...

'. ":'1975';' Chojnäcki:'1973a; Drzycimski: et 0.1. 1'972'; Nikolaev- '1975;, '. ",'>.... :~ ·:·~~dr~w~c.197.9~ '1$8~; Siudzirtski:'19T7; .imij~wsk~ .1974/., .' :." :, : : ; '. ".'.:.; ." .. ' Ijt:'t(iow q(.'theoo facts, ;'one ~an explain ,...ith· high probäbiiity·· t'he' ::

,. 1 j I '/'. .,' /. .: '.•._' I. '\ ~ . \ .' , ' . " : . . . ' • '. .' . ,'. . , : '.",

/' '...;low;:89?~d~~.e.~:r. :A~(lrt.1o. spp•. in the w?-ters we.~t .. O:f.'.·.K~.~o~r~eg'·'·''1p.ich ',.' .:.. ;.. ciQ.used',o.~breo.k::in .. thEfcontiIlu1ty ofArea A.: k'phcnomenon:,,'of 'coo.stal' './' ,., •• , , ' ,.,... . " • '~.. . I... '. ':'. .' ,": • f !'. • " . . ',., ' I. I" ~ ...' ~.~.: ~ " . .

. . upwelling;'1P1ech~~a; ;,1n .'.Px.'intl was.·..obserVed··llereL·its 'J;Tlost visible ", ," reo'uii'; ~as·:,:~:·,·drop'·:'iri·::te~pel:atui:·~: oi'sUrta~~: wat~~~ .dö~~·t~ ~ 3;~1'4. 00'. I • .'.

,".' .. /1-. ~ ~'..'i>~.' ~bci~t', :'4" ·oe.. ''1~:::r.elo.t lori.~ q'; ~'utface .:'~ ~~p~~ätiire.j.n." t'~'~icodsta:l"" ...... 'z6~oi ',and ~:t·hb·:"~~pe~~n~e·,~f.':'e6~8·id~~~b·le· ~~ber~t ;~f::P~~~d~~al~~u~ '.. '

'. I••• ,I\}" ~I'''' ~,!;.I'."I •.. t'~\·"l;.:·:, ," . "t" ~, • ",., •• ::\', "'.,"'" "'/",.". ,.' ',' " • "'" I.',.' ..,'~·';.elongat.us·,.-:'~·:species,.typiclll:iforde.eper·and:colder..:waters.·. t .... · . ,''', ," '., , '. I t •• , ' , .!' I, ',- ,. ,!. . ~'. I, \, • • I • ',,' • ,'. '~ ~' .' ., f • 1'.. .' ,'. ~ \ I' " ".. \'. t • '.' • . ' ,.

... ;. ,'Another characteristic.'.feature of' th1s .area was.'·a.1::10. a conslderable "I '!' ',.' • \, , " . •. " I \ " , .' • • •• I .'. • 1" • • 1 I I • • \ • .... J~' .'

·.·sho.re:'of .BefSm1no... c'oregoni::·rnaritima·:'and.,·.Podon'. "spp~ ',)0. t·öt'a.i."of,'ove·r '} %/.· .' -', I' .' • "... ' I " , '~. .' I " I .', "1' " " ." ~ " ~.' /', " ".~ " •• ' .:.:', ". ..",.' ... , ,'I ' " '.,: I .,' \'..... . ,',..

· . This. 1s .. c·a.uscd;· by ,the::.faot'.tho.t ···.these ,:speoies. have' s'im1lar ecologioal·," . ..' • L~ "~ j. .' , ,. • • I .' t ""',. .'.'. .' •• ' I, " , '.. -- ",' , I .." I ." • .1

· -:~'i'equircments ,:to~'''A''oartia: spp.:.·although:·thelr, .thermal' ;,optiInUI~>is" ~ven:. I ,:1' "I:·.':·~·:f~:,:':'.·, .,1,'.· .. · :,.,.:.~., :,", I" ., .,' "".' ..:;~"'.:. ," ," '1,',. ,.',: ....•.•.•\ ., I "',.,

h1gher, 'cspecially,· .. in ,the', oase of B.coregoni: maritima" .whichiso. .'.' ' ".. , '1" '.• ,I. , ' '. ',~,-, • ," ;"" \ "I I: t' ", . . • .' , "." . '", '•• : .' • ,,1 " ~.. .' '.: l " • I ... ~ ...' .' • ,'; , , -.' \' •

. \-larm stenother~ ::repres<mtritive '. of: summer '.fauna jCho'jnacki. 197.3'0;"" . ,~ :.-• ., 1. . '. ..' .' .' ~ ", .' I • , " ',:, ' • ' • 'I'" '.' • _,.,' "" .; .' ' • _ ;' " " •• ' .",. .,' '. '. '. " , l. I I. ." " , .;

· Kos~richkimi~ Llne" Sidr.ewic·,. ;Vit1nia' 1980;: Sidrewic '1979, "1980;' I •• :., .'

, , , .\. : {/J ,. ".,' '., .... • • : / .. "~" ' ..,. ' ", :.,', .'. "':', .:.",'" • 't t , I' :...., ':~, ..." I., I' ..,... i ~

.Siudzl.nsk1 '1977; .. \'liktor 1961/., :: " .... :.... :.' .. ' ,. :""',""::: i ,. ',' ." ',' , :

.;.... :Th'~' ~ecorid:~~~stal:'~~a "";n /Fig~ '1:2(': :'~as .. ch~~~ct·~·;ii~a.'bY::~"· ~i~·..'. . \, ,:' ".' ;' • """ '. " \ • , ',' • t . " .', • " " . • .. • '. ~ " - • '.' I '.: ". .. I' ~.. , '. . '; :.. , '.' \ '

.milar species,.compo~d·l::~(mto tha.t".of ·o'rea.·A.Aoo.rtia spp~ 'W'as predomi-'. ·"n~nt~.:cons·1;itut·ing·~lmost:58\.··~~.'~f ~ii'~rgimi~~s )Table II! .:··The· ,:share.

• • ,.' ". ~ '" I .. l ,', I " .. .,. • • • , .' ,. .' " . • " '. '--- " .'"~ .' , •

. .. ~"r Dosm1rui'·cc)re'gbIl:C'm·aritinib::.'Md:·podon· spp~: ,.\ia:~·· even .higli~i</pvei-·'g"I ••" '. "" \ l 'i # .• l . ,'" ,".' " ,'1'," .• , " "'. ", "'" " '1 " 1 • " •.-",' ,""',' •

'. This·\{ri.s·...:probabiy.:·cbrtnec·t·~d·,"wit·h .. the~ fact :·.that" th~.' shelte:te·d···coasto.l.:,'." \. i""'""I·'~:''':I,.,.. ·;I;.: ".~':~I.··.'·' .,:.; ',.:." ..-"..... '.,'1:. ,,'.,',:"',"~~' • ",' ,,', ":".,,, J','" " . I

., wäters .:ofthe 'Gulf ·.·cf "Puck ·o.ndthe 'Gulf:·:·o!::Gdansk· o.re·.ca1ID.er and .::. '..f I 01 "." .. '~ " I '.,:,.' _. t'., ,1-:' :~; • ',' " ': I :... ".\", • '. _': J "', I. '\". •. :". f l '. I:,"': .,.. " ..'\ . • " I· '.', :1' .-,\ ' • '" l..• '·'-' ',;'.. ''t, I' '

.• _',: ,p~?v:i~e .~~?~?:::~p"~:~~~.~:":~+.t~,.?,?~:~,~~, :~:~n.~~~~~.~.s ',~~,r~.::~~y~~~~~.~:~~~ ~:":~.~~."""" .. . treatrnent\',o;f; ·th1a ·'.part "ot: .the· ooastalzone as·, aseparate ..area .was ,," ..

• ., ·'·.·..1·. : " 11\ Jl ~ ~ ~: ,. '~',' ,....... ,.' ,. • . ' I, .:, \' " "" ',', • ' >" • ,:: .~ •..• i ,':,' ; ~ " .. :' ,: '" I'. • '; t" '.::':. 1 ' "1 • .l I I I, •

. ..', caused. bY.a.·.:' cionäid(n~able.:· oliare··\ ·of.:·~ Tc·morri.·· .longfo.örriis·, lov~r.·:·2.1'.-5 C/J/ •. :," : ......-.. ,, . . .'~ ':',,': ,.? ... :·":,\'\~"I,'\:,I:. ~'\:h',.I'·:"I"I·:I •• ,.:'.: ~.~,ll··,r .... i' : ...\:\".\ ... ;.., ....... " .. 1,""':' ...... ".':!,.:.' ~.J. '\. I ~"l'~" .;.'1, :<'

.,' .As.: f.a.r ·a~.,:~,t,s.' .~ool.ogic.al: Ir~qui:r;.~~el?-ts \ a:r;e. c.oI?-~erne.d, ,~he ..o.uthC?r? ,...,.~J:. '. :.' qtioted ~bd·~·~~:~·g~~·~.;t·hät:':it·\;fo'.an ·,e·u·ryh~Ü.ne',,· etirYth~rmai.iopt·ÜIiWn '.of:· :'.· 6~'14': .oei ;.:~~~:~:ie~:~;! ~re·t~~~i·rig::·~'tiir~~:~;.':~at·hX'~:~::'·~:lie'~~··~ ~~~ ;,qii:C~~~e'nc~S:·'Of.'' •. ':.,,'" ,,';,~ .,~ ·,.\·"-" .. ,,l'\~.\., ~~ . ,':'l"\':~'l' :,'.1,: ',".' .. :' I,": : \.'/t ,,"",' 'f.,.,. ' .. '1_" .

'. opinr~n:·:whethe~ ..,;th~s:· ~p(3ciQO '.J>*,~f~rs,:;·.coao.t~l· .. :wa.te~s:·,?;-~',öpel';\, seil. :w~t~rs~• • ' • -' ., , t' I • ,'. 'I. ~ .'" ", : '. , I ' '..., '," • , , • ".", 1 . _'..., . ." ,"l " • 'l" I . . •

· 'Ob.se~va.t10n:S.\:.m~4~~·:"1.~",J1,.i.1Y;·::1.9.8r~·,~}lggecit·,;tha·t:·.:it'; 10 'o.s.,:a?u:n'dan~·"·i~ ~he ::• .... ' .,i'~'~~I.\I';I,II~,., .. ,~I\ ", ,'.:,;' " ',' ''l'''~\,,' I'~ 'h••• ~,' \ ,I ;,'. ; ' '.' ,"'1 .I~

- 'coliotal,lzone,l·o.s·farther"from.tha ·coast : ; ; , :,' .. -', '",:.:: ,... .,- I I. ',' .,\ •• ""'-"'I""U t",'I' '. I .' •• '~"' •• ".' 'l.·· "','" •• ' t \ 1 ",1 ~

.. "'~, s~~g .,:~?,:1.~'~':'\,,~i'~~~~.~.i,?ri,··.9f<~l1:e ,·*,i~·S:t ... ~\'1~ '..~~~a~ ,it· ·,~~.~.u~~ '. b~:. :'" ".stre'ssed::tho.t:,tii'EP'ov:erlapping':'of their 'range 'the are·as. of ·:the m~st·: .

" " ", i .....,' 1" • P~' ~ .. : ~. M' I '.' ; ~,., '\ 'I .• '. • :. , I. '. • ,I \ '.' • ", ' '. • ;. '.. \' I' • ." , . .'. ". " '. "

·a.bundant·"oc·currence::'o·f: meso'zooplankton ;/Figs.' 2. ~nd.12l',is·'not ,o.oci- :' .', '. .••. .'. ", '_ ;" .. ' ,I, • ; , • " ' • • . " " ' ' • ". .' ,. •. f

· dentäi·; ..,On..t'iie:,·op:e· hand~··bot~:. dOI:1inan~" ripecies, found:·. th.ere .,:fo.vourablehYd:roiogio'di':"ootl:~Ut~on"s,.and,' :o~: thQ.'. ot~e~,· .. en·c~~t.e~edabU?~an~ anÄ \" ::varled föoci ..··components. Th1S 18 supported by the results 01' investi-····· ,.... ';', .., ".:' ".;:: .'. " , , " .

I. '.

I

I '

" , .. ' ...- 6 - . /

III·I

"grita.on~' of .·the biomass and distribution of phytoplankton IBorys1ak .and·:~Ü..nge~,..ir{ ptintl and :O.f d~ssolved;~organic matter' 1r-1Q~YkO\'l~ki, .. '

in'. pririt/~ '. ',' " , . . : . ,,' ,.. ' ..'. The' ~tr~ctüre (,f 'dominance was quite dlffcr~nt'in area.·' ci ~lhich' .:dcc.~pies ä:.~id.e, 'belt'alo~g .the.no,:t:t~ern ~d .~orth-\'1est,e:r;n bound~ry'

.. ··6ithe. in~es'tigated'area~' The .dominant species~here ~/as' 'Pseudocala-, .,'. .'. ." .' '.' , ", ,," ''-...' .' .. . ..

:' nus··.;'e·lorlgattis~:"cb'n:stitutung oyer. 58 '~'ofall:organis~s~ AB can·be. " .. " ,.~ ...... r' ;:._ ~. . . \ "" ", .-' , ~; . .,". • ,'.. .• :.' .' I" '. ", • •

, seen' froIn-; Fig~3~' ;it was most' abundant in'. this: area, which is':clo":'I '" ,,' ',' "," ; .. I",. " \ • ",' _', .." ..__ . ,' .• I '.,': ~ • ." ," ,', , ,~.. ." '_. .' '. " ' i' • .,': ".' '.' •

. ', sely related:to lts acologlöal:.requiremcnts. P.elongatus'is. a eury-"', hYlin~':"~p~6i~'B'::) optl~~: bi 7.5~ ,"18 96/, l:1.ving i~ o'penwaters> Dus'':, ,to 1ta<'cQld,'stenoth

ierm' natui:e /optimum' of 3~5 - 6,',oel;it, mo~e~" ,', .,', int'~', dee~~r"'~atcr',layer~ in 'suinmer~" forming. theiarg~st oonc'entra-

- .'" , \ .',,' ',' • , ' " .' '1'" ' • " '. ,..... '.' . , .' '. .' • . • .\ ,,' ., "

"·:'tions ',at '-0.. depth: o'fabout>-:SOm IAckefors and' Hern'roth 1975;.Chojna":",'6kl "1·973a., "19'75;~'Dr'~Y~lm~ki' et ',al. "1'972; .::DrzY~'im8ki:~ and Sludzinski':',1'977:~'.;~e'~rirot'h ~d';,Ackef~~s''1979; Koatrichkina 'etal. 1980;' Niko~'"'" .\' . ' ,':' . .'. ',. "" '. , " ':: , """ ~ . :. . .' . ;. . . .., . .' . . . "

': laev ',1957;' Sldrewic,1979,'1980;.Siudzinskl 1977; 'Zmljowska 19741. . '.:':.'±n,lh~ ,a;ea: under ;,di~'cuasion,p.elcmgatuo ':f'ound fo.;ourablc livlrig' ,:, ."';:'c~ridiiii'6ris: '·1~','w'~t~'r~',,·belo\.i a depth of. 40 ni.As 'c~'be ;~een'f;O~, ",'./":',', Fi€f~ ':30> )pi~chura.', :in pr'i~tl~ '. the .o.rea,\'1ith· wo.te~swith a t~Iripera.tu~: ,>re '''.o:f'''·4 , oe>:' occrirringat~ :,depth of40 to 80' m,' 'cor'resPQnd~: to '~'" ',:':'',iarg~':'deg'~e~ ~ith' the areas of the higheat ~buncianc~' OfP.e:lorig~tua'~, .

·::: .... "Fa~',behiiid ·P~elongatus~ although quite abundant,'w~~ ':Acartla: spp~:I ~ i: .,:. ,", . '.' " ..,' ,. . . ,'. ' .' ". ..

, .. ":1 over' 18.?~ ofall., organisms/,. represented mainlY: by A~ longiremis.' ",' ,, , ..' • '.' 1,' : . • ;. " ., , , • ' , ~ • t. , " . "

· Thls','spe'cies',:,unlike .. 'mEmt1oned above"A~b'ilifo8a, prefers deeper and",';"'~~lder" w~t~~s 'I:pa~e~s,'quoted above/.· :' . ''','< '.: '. , ".. ,,' '.'i':' ,'.. ~, :' ".,: (Ji~docri~~ were' dorii!nated' bY Evadne' nordmanni,constituting almost,,'11 %J of', all·~e~~z'6~piD.nkto~.,It '1~ .~ speoleswith"'relativelY, hi'gh , '::t~ierance·,~~·"r~ga;ds'.t'einp·e;~ture·.arid:'saÜ.nltY but-:bcc~B 'in .greate~'t,',a~Widan'c'e:"in:'~tirf'~6:e:''i~yerslocated:above desper ~aters/pa~e'~,'qU~:"":":,.t~d' '~~6~~(~'::~:'\,:" ::'!:\,,:./::.:;::, >' .. ,",' " .\"::':":, .''-" ','/. . ...:.',',:,,;, ""', " ::. ':\ ,,"':':,'The,: spe'oies,:.comp'6aition lno..the" remaining two" areaa,' D'rind' E, '\-ias, ,· , .. , I ~'l' " • ',l,; ";, ",.,': .,.• , ...• ,", I \1 .:. -",.., .' (. . .: , ' .,.., .' ," •.• • " '.'., '.. .' •.•• ;.' '

.,' :as :,~t, ..were,,;a.,'res:ultant,öf·,the ,situation' observed ,:in,.mesozooP1.an~o-.~ • 'I. , ." , ,'" \ I' " ; '\ \ \ ... ,~ " \ ~.; : " " ' '. \' ': . " :. ,.'. • ' • " ..'..... •

• nic,:'communities 'o'f'the' adjecent'areas, ·l.e., on,the one hand, 'areas"':'A:~~d',.':B;':',·'arid~:,:~l1':'.t'h~'';cit'h~r~. '~re~ 9~:Thi~,was vi~ibie in 'ar~a'D ··1~·'t~e.;-,:,'äi~'o~t,<eq~~rsh~~(,oi:'A~arti~:'spp~and'P~e~docalanus elongatu's ''/3,3,%,

• ' ..;: .' '., , ~... ~ •. ,,\, -'::" • • '\. ,. .. ..:', ' '. . " ". .' " ~ '. ,t " ',". ' ,"

,and:'29::%',:.respectiveli/.ln areo. E,thelist of.' dominant specierJ. was· ·:~~{il~:pl~'Ihrint'e\i,~Y<T6~~~r~ 'l~rigico'rn:la which I similary to are~ BI formed'.1a.rge\6'~~h·~~tratio~~,'here. In ~ll,' AciLrtla spp., Pseudocalanus' elon­': gatu:'s~,"'a.rid :'T'ä~o~ä:. 'io'nglcorni~ had a ~iml1ar perccntage sha.re~" i~o ~

23'·%, :33'%~:' 31 '9~';' ail ma~ozooplankton" respe~tivelY. .,,:, .. '; ,." , . , .' .

, '

.)

,.

•

, --

-e

),

t

~

· 1. Th~:'waters,of:the:', inv~stigated area 'cf the southern Baltie' exhi-.' .'bit~d,substanti~l" differeneeri as regard~:,th~ ab~danee,'of all."

.' nießo·z'o~plalikiori:. arid.·it~· individuai' repres'ehtati~es:':':,:-." ~;." .:äl·~~f/er~·: of·the~:coaetal· z~ne:do~·'. ~o a dept·b.of. 30"m'w~rcl ':the ..,:' ':;,.~f6·hest ;,th~"'iowest ~.quant·it:ites.: ci!' m~~ozoopianktori:\'1ere··obse·r~

"';>":)'::~'ed~:'1ri :th~' ~~~th~ea~te;ri·p~~t· bf :th~ irivestig~ted afea,':.,.··· .'.::::."br'A'~~~tTa' spp~:;,'.EU1:Yteniora ·sp. "':Bo~mi~a ~o~egoni:.ma±itirna.. · and :. '.

:.: +:,:',",'" :Pod.oh·~pp. '"were:most:.;~b~dant··:-in: thc 'coa~'bil'zoIie;':Ps'e~docala~':',',~·",>.. ·::·.'~ti~,,~~longatus·· ';~~f: Evadne ~~rd~ni.:~ 'iii' the 'con~ra:iartd'n6rth-:'. ,'.' , .' ·,'.~~est.ern·p'a~t of .th~' investigat'ed are~ while:·Tem·ora·' -iongicor':'

, '.: '.' ':. nis·· '~""in' the., 'Southl.w~ste~ part ·end· in th~ 'Gulf of Gdansk•.:.,' .. ::,~·.:i~}·,,,,,.':,,'<,'.-:',\.,,·,~,-·~,.. ,:.·'~\.:j·::, ..:'.. ..-:..... ~ .:::.:<~.,.,'. ,.:~,,~.:.~:;,...,'.'::;",\ ";'~' " :".,." .:.... : .. <, ...":' .~",.":', ",:, .:,' 2/~The".invest1gated;area::of',tho:southern Baltic 'also: differedwith':·::'::,>....~~s~~'ct .. 'tb:::~he' :"s;e·~ie~:: c;~po'~i t'toIl" ·of ..Iriesozo6~1~kt~ni~. eouimimi~

·'::··.:tie's,·:·~hi~h'.·a.iio,'1ed<tO. dist'i~gui~Jifive' a~~~~s~ ,: diif~.titig in the'• .' • I • • • '. i' I '. ~. • " ~ '- ," , • t.' ., " , •

':'(. oompösi~iOI~:'of·,d6~ina.n.t,specles:· '. o' ': . ' '.,;::-'~.".'."""'" '.~:: :,:;:>~/:a~.ea.:·i... ':j+>ne. 6'~ast~l' :'~one"'of 't}H:~ .open>s~aL ,.fas: dord~nated' by· ./':)~.'::\.:/A~~~·~~ .:..~~~~.r.:, ...', ..~.<..... ':' .:' .:.' :.:) .~.:,'::' ': :;,,'; . '.: ..,'" ,... ;,,:.:::, . ,".'.::: :~;::'-::' :,.:'.' .... '::,

'··.:b/-·araa,B· /the·.. eoastal· z'one of the·Gulf 'cf Gdansk/'was' dominated' .:,;' '/: ~;~ bY Ac~~ti~' '~':p;. and T'e~ora' l~'ngie~r~i~', '. .:''': c.":· ...:;"

.', .. ' . ,. ., .. ." ." .... '" ' '.: . ,'. '. \ ',' .. ':' ,

,'e/area .C:,/n6rthern and north-western part of the ilwest.igated;·..... ::'~ .... " ··are~/;~~~~, dominated by: Pseudocaianus··elo~g~tl.1S,;:· ~"'."'." :.- ':,.,:. ':~ .:'\'dl: ·:itli·~~~:SX ..D;.:)o~ntr~l.·~d'·horth~~~sterh p;'~tI 'arid 'El~outh~', ""··...;·;)}:.:: ..~·~~~ti1h'·:·p·a~t ':'bf "the .investi~ated ·a~ea/,"::the· ..·ihfluen..c~ o:r"th~: ..

, '.:,',' -;". " ",: "'- : .... ~.'",; ~ .... , "\" ,•••• ':'.. ). "", '.•• "" I ,', ''':;' ..••.• ,:", ,", :. :. ", " •.• ,.,. "', , ." .... .,..... '.,,~""".' .. ' .~,. ',' •

. • L ,:. adjacent':areaf( was··visible.Thus~"Aeartla. ~Pi>. ·and Pseudöeala";' .~~':·i.·.:·\'.~·,:.l~·~.,:.:'.~I(I,\··'.'\,\~:,.·:.:,.· ...._...~,., ".,~"""; .:. ·, ..... u.;••,., •.••. '•. ," .", ",. ',,',' , •.• " .•• ,'." .. , •. , •• ':"':,' '., .:,.,,'

" ."':-: nus;were,~dominarit: ,in, area D,':' ahd .Temora'longioornis was . domi:",'.,/..,·'; ..'~~t·.:·i~ ·,·~ea ';E'.':". ,,: <'" ,", ' '" ::.:., .; <'~'''''' .' '. ':':"'~".""" .,.,~'.

, . '. ',' . "

:<:,;:::.:'~,.:~;.::n:,:F:~<:;.. /":;· ", :'.:..',.:'.:."~':':.'~ :'........>.. ,.:.: '.. c.::,.':":· J ' •.•. ':'. '~' ; ...•.•. '. ,

...,'

". :'.... . ,

~ .. '..

Coneltisions

•

••••

I' •

... '

.', '

,8' -

References

,Aokefors H., Her.nro~h L.,,1975; The distr1bution and'biomas'öf

, " zooplankto~1n the Baltio proper 1n1972. Medd.

.',

.Havsf1skelab. Lysekil No•.184 ,

~.,..~ .....'\~... /

,

•

'.',/

' ..". '

, ,,-197Jb;', W10alarki !Cladooerai,- Bosmina'oorogoni

':,;~' ',' ,maritima /P.E'.Ml\ll~r/i'EVadne,' no~dmSnni L~ven' na'"" '. ,\ , . . .;

':tle zooplanktonu baltyokiegow 1971 r. Z.Nauk. AR, ' '.' . .

,- ,

, t "; I

'; .'

Boguoki Z'~ 1979 ; Elemonty ,st'atyatyki Cua b1ologow. Porman -.. ,"" " ~ ~,' """ " .,". '. .

BorysiakM. ~" Ringer, Z., '/w 'druku/ ;Sk;}:ad gatunkol'Y 'oraz rozmioszoze~I;. /.', •. ' :",., ' " 1 . . ' • .• ' :.' , . , , . .,f,',: ,,« .. :-~ie :konoontraoji biomaBY f1toplanktonu' w BSd:tYkil po;}:ud....·

.' ' '," ;:,', 'n1o\~mw 11pou 1981 r. Praoo ~1IR, Gd3nia. ,', , . , .

Chojnaoki"J~',:"1973a; Zooplankton Ba;}:tyku po;}:udniowego w 1960 r.. ... >, .',' ." :" ~. '.,,:'. ..'. .\ . , " ,

.. ""',,Z~I1auk. AR'w Szozooinle~'nr 40

,'," '

, :. l~' 'f ....

, " \",',

, .",' "'" w S'zoz,ocinio, Nr 40, ' \

,",' ",',': .,": ',"- 1975 ; Zooplankton .po;}:udniowego Ba;}:tykuw, 1972 roku.. 'i.:;" ': ", ',,' , i " ' ,,' ,

';: ; ,,: ;', . " Z.Nauk. AR' w' Szczecin1e t Hr 51 ,.. : ", ',' , . '..

:.Dn:Y~1~ki 'I. ~ Chojna~ki J., Rad~1un M. t 1972; Ekolog1a zooplankto-, ~, .• f' '.,"': ' ": • ,,' ',' ",' ': . ,I." .. : ,.". • ..... • ,,: • ~~ .;" , . . '. '

" ;' ~::: ';,' ,!,' ,'nupo;l:udniowegoBaltyku~ Ekoeyatemy Morekie T.II." .!. :. '. " ,'I,' . <[, ~ .' • • • '. .' • " • •

DrzYo1maki'·I.;, Siudzineki K. t 1977; Zooplankton i 1ohtioplankton,,' '. .' , I,: "'.' ~ .:', : .: l ~ . " , . . .' " '. :, :.' ". .' , •

,::"".,,"i>,:,:;;";:,\',:., pOd:udniowego Ba;}:tyku w latach 1971-1972. Stud1a i Ma~

. " ',: ("'1::' ,'! 'teria~ MIR, A. t Nr 17'I ,'I', •• ' , .'.! ,1 •

, ' J I ' ' ,", I ':.. ; ~.' ; ,.'. , , '

" \ Hemorth: L. ,:Aokefors' H•• 19791'The zooplankton' cf the Baltic pro-.. \',,' ,\".'~,j~,:·'}.\"'"~;:'.,',':1,,..,:."~".'.:. :~. ..' '. " , '.

"',' ",,', :::por. F1ehery Board of Sweden, Rep. Uo. 2 ' :'. .. , ' . :~' :' I :,.," ;.'.' .", .

Koetrtc~k1na'E.M.'."'Lino R.i., Sidrewio L.L., Wit1nja II.T •• 1980;

, ':::','::' :,',\, \;"::,:.~k'OJ:Og~CZOak1j~ komplekey zo'oplanktona BaJ:tija~ogo,,' " ':',';'" ',',~~ria~_E~O~ietemy Baltiki, Gdynia

. ',. ',' " ' ' . ./

'" MO'zYkowek1' T., :/w druku/; Zawartos6 wybranych ek;l:adn1k6w rozpuez-, ,

'" czonej mater1i, organicznoj w wodz1e morek1ej, Praoe .

MIR, Gdynia'

-' " 9

.NIkol:ajew J.J., 19.57; Blol:oglczeskije sezony Bal:tIjskogo morl~•.

Tr. Lotw. Otd. WNIRO II,RIsa. .

. " '

.Piechura J., /w druku/;Temperatura i zasolenie w6d w polskiej,

, strefie VIYd:qcznego rybol:owstwa w lipcu 1981 r. Prace

, ,,' MIR, , Gdynia

·.S"ldreWIo L.L., 1979; Niekotoryje zakonomiemost1 wertlkalnogo

,_ I rozpredlelenja zooplanktons w oentralIioj Bal:t Ikie•., ~ " ~ . .

. Rya. Iaal. ,Bal:t. Morl~, Bal:tNIRCh,14, Riga

i Ma-

- 1980; Iaeledowanije ekol:ogiczeakich oharakteristik

osnownych widom zoopl:anktona W oentralnoj Bal:tikie•.

srodowlska na zmiennosc popula~", '

" " oji Bosmina ooregon1, Daphnia hyalina i Daphnia. oucu~

, ." "

'. ' .

" Ryb •. Isal. Baa:t. Moria, Bal:tNIROh ~ 15, Rlga'

Siudzinsk1K., 1977; ZooplanktonZatokl Gdansklej. Studia{.' . .

, terlal:;v Nm, A, 18

Wlktor K~, ,1961 i 'Wpl:yw warunk6w

•,"

" .llata.'Ekol. Pol. A, IX, 5

• I:'

, . , .' w 1969 roku. Z.Nauk. Wydz. BiNoZ UG, Nr'2

•

~~jewska M~I., 1974; Zm!a.ny. aezonowe sk;l:adu gatunkowego mikro- .

',: .. ' ' .. ' .: 'zooPlankton':! centralnego i pol:udniowego Bal:tyku\ :," \;,

,.. ""

Table 1.j. '

List of species and higher taxonomie units identlfledin:, th:e', inve~tigated mesozooplanktonic sampIes.

•

•

.Table 11.

,~er~entage sha.re of individuai mesozooplartkton componcmts". in thedifferentiated areas 'of the southern Baltlo.

"

, area..13 . C. D E·

9.50 0.04'

" .'

,0.18 ,,1l,14

0.30 '0.382.22" '.0.181l~.05··-

. 3.15 .1.89' '0.-03 ',0.72' 0.770~02'0.70. 0.03 ..

..- " ,0. 13 'Ö. 14 " o. 091~55 .. 3.37 . 0.43,0.84 .: 0.68

..' .1.56 . 6.07. ,0.38- 0.55·.0.793.23 0:9710.94" 8~'357"542 • 6..1 1 • 70, .58. 25' ,29. 07. :5 3 .42' .

. . , ..". . ,,'64.4757.97' 18~4733.07'22.83 .

• ,'., :.'. ','j' '>

'3.20 :,21 ~52 '. '6~20' "S.'37, )0~89'3.173.89'· 0.392~46' 0'.~6

0.28' 0~22 '0.17 0.43.'0.14. 0.01 . - 1.07':3.61' .

, . .

0.27', 0.5{ . ,0.48"

0~02 ····0.60 , O~O:3 .·~.4810.01f.28,:

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.Fig~2;Q'~~tit~t~';;~d1~tjib~t'i~~' ot'.mesozOO~lankton' 1Ii: .the··P~liS,h ·se.ct~r O:r t~e southern··Baltic· Ix ~~ of specimes' in, l' m., m ~ standard, error, ot' the me~/.:.· , ' . , .':' , . ' '.

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Fig. 3.Quantitative', distribution, of Pseudooalanus' elongatus 1il the Polish' sector . of the 'soutliern Baltio. /designations as in Fig.2./., . '., " '.' " .. ' . _ " .-

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Plg.5.Quantitatlve· distribit.1on 'of, Temora: longiaornis in,the Polish: 'seator' of, the. southern Baltic' /desfgnatio'nG.' as in Fig.2~/ .' , ." .. ~.', . \ ' -. I ," " ;.... . '. : ... ', '.', ...

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Fig~'i';Qua~titati~e -'dis:'t;i~~~i~n ~Of~'~~dne :n'~;dmanniin,the);O~i'sh-se-~tor~:'ot~the';southerxi 'B~iti~"-id~~r~~tionsin Fig.2./. . " ,", , - -

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Flg.8.Q~tit-itat.ive'::ciisttibuti~n-ÖT:Bo;m~;~~~i~g~~-~~iti~~-.tiie'- P011Sh: s'ect~r o:f the _southernBalt.1c /des.1gna-," ·.t1ans as in.F.1g.2./.·, .-, .. _. -, " -.e

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