Page 1
Taxonomic Composition and Ecological
Structure of the Species-Rich Butterfly
Community at Pakitza, Parque Nacional
del Manu, Peril
ROBERT K. RoBBuqs'
GEltkm1o
Jj,j,2
OLAF H. H. MIEu&
DONAW J. HMvEY'
MlpJ.JA M. C,c&cow&
`Department of Entotnolog. NHB Stop 127. Nazional Museum
of Natural History, Snuthsonian Inuirugion, Washintton DC 20560
2Museo de Historia Natural,
U,iiversidad Nacio,ial Mayor tie San Marcos,
Apanado 14-0434, Limo-I 4, Pad
`Deparrmnenzo de Zoologia, Unioersidade Federal do Parand,
Caixa Postal 19020, 81531-970 Cmitiba, Parand, Brazil
ARsTRACr
1,300 butterfly species were sampled on five field-trips to Pakieza
and arc listcd with thcir first dates ofcapture and whether they have
been recorded at Tambopata, a reserve 235 km to the southeast.
Approximately one-third of Pakitza's fauna are Hesperiidae,
one-third Lycacnidac+Riodinidae, and one-third Nym
phalidae+Papilionidae+Pieridae, which we suggest may bc gener
ally true for Neotropical butterfly communities. Slightly more than
10% of the species appear to be taxonomically undescribed. Almost
2% of the species have larvae that feed on plants other than
angiosperms, and about 28% have larvae that eat monocotyledons.
About one-third of the Riodinidae belong to tribes with larvae that
arc myrmecophilous. Adult butterflies were attracted to many
substances, including wet sand, bird droppings, and flowers, but
only about 10% of the fauna was attracted by decaying fruits,
carrion, and excrement. Many of the most widespread, common
Neotropical species, which are typical of disturbed habitats, were
either unrecorded or rarely recorded at Pakitza. Butterflies that
were "fogged' from the canopy by insecticide were mostly species
of open areas and also were collected by other methods.
1-Robbins, R. K., G. Lamas, 0. H. H. Mielke, D. 3. Harvey, H.Casagrande. 1996. Taxonomic composition and ecological structureof the species-rich butterfly community at Paicitza, PargueNacional del Hanu, Peru. Pp. 217-252, in Manu: The biodiversity 0Southeastern Pen D. E. Wilson and A. Sandoval, eds..Smithsonian Institution Press, Washington DC. 679 pp.
-I
217
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R. Rohhins, C. Lamas, 0. Mielkc, D. Harvey and M. Casapande
INTRODUCTION
Localities with the richest butterfly communities in the world occur in the
lowland drainages of the Rio Solimoes Upper Amazon River and Rio Madeira
in Colombia, Ecuador, Peró, and Brazil Brown, 1984; Emmel & Austin, 1990;
Lamas et a!., 1991; Robbins & Opler, 1996. More species may be found at
3,000-5,000 hectare sites in the Upper Amazon Basin than occur in most, if not
all, African or Indo-Australian countries Robbins, 1993. These communities
are of great scientific interest because of their unusually high species richness, hut
they have not been been well-documented. The taxonomy of Amazonian
Hesperiidae, Lycaenidae, and Riodinidae is poorlyknown, so specific identification Fhas been difficult. Consequently, information on the taxonomic and ecological
composition of these communities is scarce hut see Ehert, 1969; Drurnmond,
1976a; Hutchings, 1991.
The purpose of this paper is to provide basic information on the composition
of the butterfly fauna of Pakitza, a biological station located in lowland rain forest
356 m elevation on the east bank of the Rio Manu in the Reserved Zone of
Parque Nacional Manu, Madre de Dios, Peru 11 °56'47"S, 7101700W. We list
1,300 identified species recorded on five field-trips, making Pakitza the richest
documented site in the world for butterflies, and overview the raxonomic and
ecological composition of Pakitza's fauna. We report on the diversity and
dynamics of this community elsewhere.
STUDY SITE AND METHODS
Erwin 1991 mapped the trail system and many major streams at Pakitza. We
sampled most of the forest types and other habitats that occur within 5 km of the
base camp Erwin, 1991, hut concentrated our efforts along the banks of the Rio
Manu and on the Tachigali, Castañal and Pacal trails. Consequently, the study
site for our project was roughly a semi-circle of radius 5 km with an area of
approximately 3,925 hectares. We usually collected within 10 in of trails in the
forest, so 3,925 hectares is an upper limit of the actual area sampled.
Our field-work was limited to those short time-periods when the camp was
open. Most field-work was done in September and October during the transition
between the dry and wet seasons when lycaenid, riodinid, and hesperiid butterflies
are usually most common and diverse. Consequently, our comments about
seasonality are based primarily on Tambopata, a protected reserve 235 km to the
southeast at similar elevation, where we collected at more different seasons than
Pakitza Lamas, 1981, 1983, 1955; Lamas etal., 1991.
Five field-trips were made. 8-23 September 1989, 2-21 October 1990, 27
October-16 November 1990. 19 April-14 May 199!, and 26 September-20
218
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Butterfly Community w Pakicza
October 1991. There were two field-workers on each trip except 8- 14 September
1989 three people and 26 Septemher-20 October 1991 four people. We
recorded data on 97 days, totalling 247 person-days, and 1,311 person-hours. No
data were taken on ten other days that were either too cold or rainy for butterfly
activity.
A variety of collecting methods was used, including standard insect nets with
pole extensions, baits, standard butterfly bait-traps, commercially obtained malaise
traps, and "imitation" bird droppings Austin eta!., 1993; Lamas et al., 1993.
Baits included rotting fish, decaying fruits, excrement, and withered Hehorropium
rndicum L, Boraginaceae, a source of pyrroliziditie alkaloids Beebe, 1955; Pliske,
1976. Besides the species that we collected as part of our project, we also recorded
a few sightings of species which could he identified unambiguously. Additionally,
other scientists at Pakitza gave us butterflies that they had trapped or collected.
It would he hard to overemphasize that specimen preparation and species
identification were the most time-consuming and costly parts of this study.
Responsibility for identification was apportioned as follows: Casagrande for
Brassolinae; Lamas for other Nymphalidae, Pieridae, and Papilionidae; Mielke for
Hesperiidae; Harvey for Riodinidae; and Robbins for Lycaenidae. Specimens were
identified as well as possible using our taxonomic expertise, and the collections,
type photographs, and literature of our respective institutions, where all specimens
are being deposited. Even though we believe that our species list is the most
authoritative one ever made for a South or Central American locality, some of the
identifications are still provisional, and some are based only on males.
TAXONOMIC COMPOSITION
We list the 1,300 identified butterfly species Appendix that were collected
at Pakitza or positively identified by sight in 7 cases. There are 448 Hesperiidae,
25 Papilronidae, 31 Pieridae, 181 Lycaenidae, 246 Riodinidae, and 369
Nymphalidae. For each species, we note the first day on which it was sampled,
the first day of capture during the fifth field-trip, and whether or not the species
has been found at Tamhopata, a protected reserve in Madre de Dios at a similar
elevation Lanias, 1981, 1983, 1985, 1994; Lamas et al., 1991, where 1,234
species have now been recorded. Although quantification of the sampling method
and analysis of the resulting data are being published elsewhere, we present our
data here so that they are available.
The taxonomic composition of the Pakitza sample Table 1 is very close to
one-third Hesperidac 34.5%, one-third Lycaenidae + Riodinidae 32.8%, and
one-third rapilionidae * Pieridae + Nymphalidae 32.7%. Similar partitions are
also found Table 1 at Tambopata Lamas, 1994, Panama Rohbins, 1982, and
ltatiaia, a park n Rio de Janeirostate ZikAn & Zikán, 1968. Although this 1:1:1
partition is not found in temperate North America Miller & Brown, 1981 or in
Serra do lap in southetn Brazil Brown, 1992 Table I , primarily because
219
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Table 1. For Pakitza, Tamhopata Lamas, 994, Panama
RoIbins and Small, 198!, ltatiai., ZiLin and ZiHn, 1968,
Serra do Jan Brown, 1992, and North America Miller and
Brown, 1981, the percentage of true butterfly species
Papilionoidea and of all butterflies Papilionoidea +
Hesperioidea that belong to families Pipilionidae, Pietidae,
and Nymphalidae.
%OFTRUE %OFALL
LOCALITY BUTTERFLIES BUTTERFLIES
Pakit:a 49.9% 32.7%
Tanibopai:i 48.7% 314%
Panama 49.6
ratiaia 51.3% 32.3%
Serra do lipi 60 9'X 38.0%
United States 65.6X 402%
tigators who do not have a particular interest in
N. Rohhins, C. Lanws, 0. Mielke, V. I4arvn and M. Casagwnde
riodinid diversity is low, we
suspect that it is a robust
`rule of thumb" for the taxo
nomic composition of lo
wland Neorropical butter
fly communities.
Although we doubt that
there is biological significance
behind this non-phylogenetic
partition of species richness, it
may prove useful for species
richness studies in the Neo
tropics. Given the taxono
mic difficulties that we had
_______ _________________
in handling and identifying
1,300 species, future inves
Lycaenidae, Riodinidae, and
Hesperiidae can locus on Papilionidae, Pieridae, and Nymphalidae and multiply
by 3 to get an estimate for the entire fauna. For example, at least 204 Papilionidae,
Pieridae, and Nympahlidae have been recorded from La Selva, a biological station
in Costa Rica DeVries, 1994. The other families have been largely unsampled,
hut we would estimate that at least 612 butterfly species occur at La Selva.
Of the 1,300 species in the Pakitza list, we could nor identify 144 species 11%
rhat we believe are undescribed, These species belong to the Riodinidae 44,
Hesperiidae 39, Lycaenidac 33. and to the nymphalid suhfamilies Satyrinae
25, Brassolinae 2, and Nymphalinae 1. This result is further reason for
limiting most diversity studies to Papilionidae, Picridae, and Nymphalidae. Also,
since the South American fauna is more poorly-known taxonomically than orhers,
it is probably lair to conclude that at least 90% of the world's butterfly fauna is
described.
COMMUNITY STRUCTURE
Although the purpose of this study was to assess the butterfly diversity of
Pakirra, we also recorded incidental data on behavior and ecology. We present
his information, even though it is incomplete, because we hel eve that it is the
first attempt to look at the community structure of all butterflies at a Neottopical
site.
Approximately 28% of the butterfly species at Pakitza feed as larvae on
monocoryledons while most of the test eat dicotyledons. The monocot feeders
include 101 Satyrinae excluding Eupnchia, 22 Brassolinae, 225 Hespetiinae, and
about 20 species in other suhfamilies. A noteworthy feature of the Pakitza fauna
220
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Butterfly Community at Pakitza
is the 30 species olspfendeuptychia and Caerueuptychia, nearly hail of which are
undescribed. Larvae of Splendeuptychia eat bamboo Kendall, 1978; D. Murray,
pers. comm., which is very common at Pakitza Erwin, 1991, while larvae of
Caeruleuptychia eat palms K. Brown, pets. comm.. Pakitza is one of the richest
known sites ior Ithomiinae 62 species, most of whose larvae feed on Solanaceae.
A small proportion of Pakitza's butterflies have larvae that do not eat
angiosperms. Two species of Euntaeus Lycaenidae presumably feed on cycads
the foodplant of their sister species Robbins, in preparation; some species of
Calycopis Lycaenidae and C/saris Riodinidae appear to be detritivores S.
Johnson, 1985; Harvey, unpuhl.; ten species of Sarota probably eat epiphylls
DeVries, 1988; and larvae of Euprychia Nymphalidae eat Selaginella or mosses
Singer et al., 1971, 1983, 1986. About 30 larvae and pupae of Mirnocasrnia
rochschuldi Riodinidae were found in an ant nest Cephalores acratus in a dead
branch. Larval feeding was not observed, but larvae may have been fed by ants
Harvey, in preparation. At least some species of Setabis Riodinidae have been
reported as predaceous on Homoptera Harvey, 1987.
Myrmecophily symbiotic relationships between butterfly larvae and ants is
restricted, with few exceptions, in the Riodinidae to tribes Eurybiini, Lemoniini,
and Nymphidiini Harvey, 1987. Thus, a third of the riodinids at Pakitza 82
species are expected to be myrmecophilous, which is a hit higher than most other
Neorropical mainland areas, but less than Trinidad Harvey, 1987. The
distribution of myrmecophily in Lycaenidae is too poorly-known to allow similar
estimates for them.
A conspicuous adult-feeding behavior at Pakitra was "puddling', in which
males that appear to he freshly eclosed sip moisture in sunny spots from the dirt
banks of the Rio Manu and some streams inside the forest. Most "puddlers" appear
to be species of open areas, such as tree-fall gaps, river edges, and the upper canopy,
and not, as a general rule, species restricted to undistuthed forest. Sodium in the
soil attracts males, prolongs their feeding, and is transferred, in part, to females
during mating Arms et al., 1974; Adlet & Peatson, 1982; Pivnick & McNeil,
1987; Lederhouse et al., 1990. Occasionally females also may "puddle", such as
Eunice Nymphalidae in October 1991. Also, both sexes ofMmisrr-vrnon zilda and
14. clean Lycaenidae regularly "puddle" in the late afternoon along the Rio
Manu.
Perhaps the easiest way to summari:e the extent of "puddling" behavior is to
list those taxa in which it was not observed. They are Parides P. sesosms puddles
at other Ama7onian sites, K. Brown, pers. comm. Papilionidae; Dismcnphia, but
not Enantia and Pseudopieris Pieridae; all Lycaenidae except some Ministnmon
and Ocaria ocrisia; most Riodinidae with Lvropteryx, Rherus, Ancylwis, Monethe,
Paycdlla, Lasaia, Baeutis, and Melanis being exceptions; lthomiinae; Saryrinac; and
Brassolinae. The phylogenetic incidence of `puddling" among Hesperiidae is not
evident except that it occurs in all subfamilies at Pakit:a.
Many Lycaenidae, particulatly Cakcopis ceraw and Ce!mro celmu.c, congregate
at drying stream beds at the end of the dry season. Individuals usually alight on
221
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R. Rohbins, C. Lamas, 0. Mielke, I. Harvey and M. Casagrande
vegetation, hut also may land on moist dirt banks, where they sip mmsture. In
retrospect, it was our impression that this behavior occurred most often during the
hottest part of the day and may have been a thermoregulatory behavior. It does
not appear to be the same behavior as "puddling"
Many butterflies in primary Neotropical forest are associated with army ants
Zikan, 1929; Drummond, 1976b; Ray & Andrews, 1980; Lamas, 1983; Austin et
at., 1993. These butterflies sip liquid from the ground or from bird droppings on
leaves. Although Arias Charaxinae, Satyrinae, Euselasia Riodinidae, and
others were associated with army ants at Pakitza, Hesperiidae seemed to he the
major butterfly participants, and "imitation" bird droppings attracted many more
Hesperiidae than other butterflies Lamas et al., 1993.
Few flowers at Pakitza attracted many butterflies, but we found adults feeding
on other substances. As mentioned, many Hesperiidae and others eat bird
`droppings", even when not associated with army ants Lamas et al., 1993. Aboui
10% of the species at Pakitza were attracted by decaying fruits, carrion, and
excrement. Withered Heliorropiurn indicum L. Boraginaceae attracted many
species of lthorniinae in October 1990, but was less successful in other years.
Pakitza and Tambopata have a marked dry season from April-May to
September-October. The abundance and diversity of most butterflies is highest
during the transition between dry and wet seasons and lowest during the wet
season. This pattern is particularly true for Lycaenidae and, to a lesser extent, for
Riodinidae and Hesperiidae. However, adults oI some species appear to fly only
during the wet season, such as Morpho mene!aus Nyrnphalidae, or are most
common at this time, such as Saliano Hesperiidae. Most Nymphalinae and
Limenitidinae Nymphalidae are conspicuous in the middle of the dry season,
such as Hamadryas and Eunica maja noerma, and are often very worn by September,
indicating that some of them may he in reproductive diapause.
Many of the most widespread, common, and weedy Neotropical butterflies,
which are common in the vicinity of Puerto Maldonado, are absent or rare at
Pakit:a. Danaus plexippus, D. gilippus Danainae, Ananiajatrophae Nymphalinae,
Rekoa pategon, Strymon mulucha, Lepcotes, and He,niargus Lycaenidae are
unrecorded at Pakitra, and Phoehis sennae Pieridae was collected only twice,
Consequently, sites in the Rio Madeira drainage that include a greater amount of
disturbed habitat than at Pakit:a would be expected to have more than 1,300
species. The biologically important question, though, is whether other sites have
as many species of undisturbed habitats. These species are less able to survive in
areas that have been modified by man and have more restricted distributions than
species of disturbed habitats Thomas, 1991; Spitzer et a!., 1993.
Because butterflies insomegroupsily primarily in one vertical stratum Niedmna
et al. , 1996, a major question is whether our sampling methods missed a set of
species restricted to the forest canopy. Fortunately, our colleagues, T. Erwin and
NI Pogiie, segregated butterflies that they "fogged" with insecticide from the forest
canopy Erwin 1983, 1990. Most oI these species were widesptead taxa of open
areas, such as Rekoa mecon Lycaenidae, not species restricted to the canopy.
222
J
Page 7
Bucteifly Community at Pakicza
Further, we collected each of the "fogged' canopy species by other means.
Although some species may live only in the upper strata of the forest, we have no
evidence that such a fauna, if it exists, is very large.
ACKNOWLEDGEMENTS
We thankjohn MacDonald, William Rowe, and Nancy Clarke for participating
in the project; field companions Mirian Medina, Christoph Hauser, Isabel
Bohorquez, Terry Erwin, Michael Pogue, Jerry Louton, and Nancy Adams for
providing specimens; Robert Lederhouse for providing references; Keith Brown
and John Brown for reading and commenting on the manuscript, the staff of
BIOLAT in Washington, Lima, and Pakitza, for arranging logistics and taking care
of us; and the officials of the Peruvian Ministry of Agriculture who sanctioned our
work at Pakitza. This paper is contribution number 92, Biological Diversity in
Latin America BIOLAT Project, Smithsonian Institution, and contribution
numbet 810 of the DepartrnentofZoology, Universidade Federal do Paraná, Brazil.
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Moss-feeding by a satyrine butterily. 3. Res.
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Spitzer, K., V. Novotny, IA. Tonner, and
3. Leps. 1993. Habitat preferences, distribu
tion and seasonality of the butterflies Lepi.
doptera, Papilionoidea in a montane tropi
cal rain forest, Vietnam. 3. Biogeogr. 20:
109-121
Thomas, C. D. 1991. Habitat use and
geographic ranges of butterflies from the wet
lowlands 0f Costa Rica. Biol. onserv. 55
269-28 1.
ZiUn, J. F. 1929. Myrmekophilie bei
Hesperiden? F.ntomol. Rundschau 46: 27-28.
Zikãn, 3. F. and W. ZiUn. 1968.
Inseto-fauna do ltatiai e da Mantiqueira. Ill,
Lepidoptera. Pesq. Agropec. Bras. 3:45-109.
APPENDIX
Taxonomic list of the Pakitza butterflies. Species names with an asterisk *
belong to groups in which only males were used for identification. Higher
taxonomic categories follow Evans 1951-1955 for Hespertidae, Miller 1987 for
Papilionidae, Klots 1933 for Pieridae, Eliot 1973 for Lycaenidae, Harvey
1987 for Riodinidae, and Harvey 1991 for Nymphalidae.
The column "1st Dayt' lists the first day of capture for that species. If it was
nor collected as part of our project, we note whether it was a sight record SR or
collected by another scientist XX. The column "5th Trip" Iisrs the first day of
capture during the filth field-trip or if it was not captured on this trip NC. The
column "Tambopata" notes whether or not the species was collected at Tambopata.
225
Page 10
NYMPHALIDAE: HELICONIINAE
1. Actinote pellenea hyalina Jordan, 1913
2. A. rhalia crassinia Hopifer, 1874
3. Philaerhria dido Linnaeus, 1763
4. Agraulis vanillae lucina C & K Felder, 1862
5. ljryas julia alcionea Cramer, 1779
6. Eucides aliphera aliphera Godart. 1819
7. E. Isabella hippolinus Butler, 1873
8. E. tybia lybia Fabricius, 1775
9. E. rates tabernuta Lamas, 1985
10. E. vibilia unifasciata Butler, 1873
11. Laparus doris doris Linnaeus, 1771
12. Neruda noSe manu Lamas, 1976
13. Heliconius burneyi koenigi Neukirchen, 1995
14. H. demeter tambopata Lamas, 1955
15. H. etevatus lapis Lamas, 1976
16. H. erato Iuscombei Lamas, 1976
17. H. hecale sisyphus Satvin, 1871
IS. H. leucadia Bares, 1862
29. F-I. melpomene schunkei Lamas, 1976
20. H. numara tyrcaeus Weymer. 1891
21. H. pardalinus macon Weymer, 1891
22. H. sara thamar Hubner, 1806
23. H. waltacei lavescens Weymer, 1891
24. H. xanthocles quindecim Lamas, 1976
NYMPHALIDAE NYMPHALINAE
25. Anarria amathea sticheli Fruhstorfer, 1907
26. Metamorpha elissa elissa HUbner, 1819
27. Siproeta stelenes meridionalis Frubstorfer, 1909
28. Junonia genoveva occidenratis C & K Felder, 1862
29. Castilia angusta Hewitson. 1868
30. C. perilla Hewirson. 1852
31. Eresia clara Clara Bates, 1864
32. E. eunice eunice Htibner, 1807
33. E. nauplius plagiat'a Raher, 1913
34. Eresia sp. n.
35. Ortilia gentina Higgins, 1981
36. Tegosa claudina Eschschottz, 1821
37. Telenassa burchelli Moulton, 1909
NYMPHAI.IDAE: LIMENITIDINAE
38. Historis acheronra acheronta Fabricius. 1775
39. H. odius dious Lunas. 1995
40. Bacotus `mazonicus Riley, 1919
1ST 5Th TAMBO
DAY TRIP PATA
I 82 YES
28 NC NO
16 76 YES
19 NC YES
I 77 YES
to 77 YES
19 NC YES
16 75 NO
4 NC YES
28 77 YES
12 77 YES
2 79 YES
I 83 YES
I 77 YES
2 80 YES
I 75 YES
4 75 YES
2 77 YES
2 77 YES
I 75 YES
16 NC YES
I 75 YES
76 76 YES
2 75 NO
40 NC YES
9 76 YES
23 NC YES
9 77 YES
13 91 YES
19 76 YES
22 78 YES
4 81 YES
4 85 YES
89 89 NC
16 80 YES
40 78 YES
8 76 YES
2 77 YES
19 77 YES
8 76 YES
41. B. deucalion C & K Felder, 1860
42. B. japetus Staudinger, 1885
43. Smyrna blomfildia blomflldia Fabricius, 1782
44. Colobura dirce dirce Linnaeus, 1758
45. Tigridia acesta tapajona Butler, 1873
46. Biblis hyperia laticlavia Thieme, 1904
47. Vila ateca azeca Doubleday. 1848
48. V. ernilia caecilia C & R Felder, 1862
49. Myscelia capenas ocromaculata Butler, 1873
50. Caronephele acontius acontius Linnaeus, 1771SI. C. antinoe Godart, 1824
52. C. nurnilia numilia Cramer. 177553. Nessaea hewicsonii botjvjensis Jenkins, 1989
54. N. obrina lesoudieri LeMoult, 1933
55. Eunica alcmena flora C& K Felder, 1862
56. F. alpais alpais Godart, 1824
57. F. amelia erroneata OberthUr, 1916
58. F. bechina bechina Hewitson. 1852
59. F. caelina atycia Frubstorfer, 1909
60. F. clytia Hewitson, 185261. F. concordla Hewirson, 1852
62. F. eurota eurota Cramer, 177563. E. maja noerina HaIl, 1935
64. E. malvina malvina Bates, 1864
65. F. margarita Godart, 1824
66. F. marsolia fasula Eruhstorfer, 1909
67. F. mygdonia mygdonia Godart, 2824
68. F. orphise Cramer, 1775
69. F. sophonisba agele Seth, 1915
70. F. sydonia sydonia Godart, 1824
7!. Hamadryasamphinome amphinome Linnaeus, 1767
72. H. arinome arinome Lucas, 1853
73. H. chloe chloe StolI, 1787
74. H. iphthime iphthime Bates, 1864
75. H. Iaodamja laodamia Cramer, 1777
76. Ectima iona Doubleday, 1848
77. E. lirides Staudinger, 1885
78. F. thecla peruviana Bryk, 1953
79. Panacea prola amazonica Fruhsrorfer, 1915
80. P. regina Bates, 1864
81. Batesia hypochiora hypoxantha Salvin & Oodman, 1868
82. Asterope marku hewitsoni &audinger, 188683. Pyrrhogyra crameri hagnodorus Frubstorfer, 1908
84. P. edocla cuparina Bates, 1865
85. P. neaerea amphiro Bates, 1865
86. P. otolais olivenca Fruhstorfer, 1908
87. Temenis laorhoe laothoe Cramer. 1777
88. T. pulchra pallidior Oberrhijr, 1901
89. Nica flavilla sylvestris Bates, 1864
90. Perks lamis Cramer, 1779
9!. Dynamine aerara aerata Butler. 1877
R. Robbing, C. Lamas, 0. Mielke, D. Harvey and M. Cacagran&
SPECIES
Butterfly Community at Pakitza
I
4 76
13 85
19 NC
2! 80
16 75
18 88
29 77
2 75
2 77
2 77
28 97
6 75
5 85
I 75
40 NC
19 79
II 76
Is 97
6 88
5 82
16 77
5 77
48 92
71 89
46 NC
I 76
2 76
10 91
53 75
36 81
Il 79
26 78
1 76
82 82
83 83
89 89
77 77
2! 93
I 77
3 76
I 76
46 NC
2 75
16 77
16 76
I 77
2 76
83 83
53 76
6 78
19 76
YES
YES
YES
YES
YES
YES
YES
YES
YES
YES
YES
YES
NO
YES
NO
YES
YES
NO
YES
YES
YES
YES
YES
YES
NO
YES
YES
YES
YES
YES
YES
NO
YES
YES
YES
YES
NO
YES
YES
YES
NO
YES
YES
YES
YES
YES
YES
YES
YES
YES
YES
226 227
Page 11
H. Hobbths, 0. Lavnas, 0. Miclke, D. Harvey and M. Casagrande
92. D. irternisia glauce Bates, 1865
93. D. athemon harreiroi Fern;Sndcz, 1928
94. D. chryseis Bates, 1865
95. D. coenus leucothea Bates, 1865
96. D. gisella Hewitson. 1857
97. D. intermedia Talboc, 1932
98. D. paulina paulina Bates. 1865
99. D. smerdis smerdis Tessmann, 1928
100. Elaematera pynme ssp. n.
101.. Catacore kolyma pasithea Hewirson, 1864
102. Diaethria clymena peruvians Guenee. 1872
103. Paulogramma pyracmon peristera Hewirson, 1853
104. Gillicore astarre stratiotes C & R FeIder, 1861
lOS. C. cynosura cynosura Doubleday, 1847
106. C. eunomia incarnata Rober, 1915
107. C. hesperis Ouérin, 1844
108. C. hystaspes zelpharna Hewitson, 1858
109. C. pygas cytlene Doubleday, 1847
110. C. texa maimuna Hewitson, 1858
III. Adelpha aethalia davisil Butler, 1877
112. A. attica C & K Felder, 1867
113. A. bocotia fulica Fruhatorfer, 1915
114. A. cocala urraca C & K Felder, 1862
115. A. cyrherea lanilla Fruhsrorler, 1913
116. A. eroria erotia Hewitson. 1847
117. A. iphiclus iphiclus Linnaeus, 1758
118. A, ixia pseudomcs.sana Fruhstorler, 1913
119. A. jordani Fruhstorfer, 1913
120. A. Irma Irma Hewitson, 1847
121. A. nesentina chanchn Sr.iiitlinger, 886
122. A. naxia naxia C & R Fekler, 1867
123. A. phylaca juruana Butler, 1877
124. A. plesaure phliassa Codart, 1824
125. A. rhesproria deiphicola Frubstorfer. 1909
126. A. uta Fruhstorfer, 1915
127. A. zunilaces 3 ssp. n.
128. Marpesia berania bernnia 1-lewirson, 1852
129. M. chiron marius Cramer, 1779
130. M. crethon Fabricius, 1776
131. M. egina Bates, 1865
132. M. furcula oechalia Westwnod, 1850
133. M. petreus petreus Cramer, 1776
134. M. themistecles norica Hewitson, 1852
NYMPHALIDAE: CHARAXINAE
135. Consul labius divisus Butler, 1874
136. Hypna clytemnestma negra C & K Felder, I 862
137. Polygraphs xenocrares xenocrates Westwood, 1850
138. Siderone galanthis rhctsais C & R FeWer, [862
139. S. syntyche mars Bates, 1860
76 76 YES
19 76 YES
2 76 NO
5 76 YES
72 NC NC
53 NC NO
2 NC NO
16 77 NO
32 NC YES
45 78 YES
18 76 YES
4 76 YES
84 84 YES
2 76 YES
28 76 YES
4 76 YES
30 80 YES
36 NC YES
18 76 NO
27 77 YES
5! 91 YES
79 79 NO
4 NC YES
41 NC YES
26 NC NO
2 76 YES
16 96 YES
16 79 YES
19 89 YES
2 75 YES
77 77 YES
2 76 YES
2 75 YES
19 78 YES
89 89 NO
84 84 YES
6 84 YES
2 76 YES
32 77 YES
96 96 YES
2 78 YES
16 81 YES
I 76 YES
16 77 YES
97 97 YES
22 77 NC
79 79 YES
90 90 NC
r Butterfly Community at Pakitza
140. Zaretis itys irys Cramer. 1777
141. Fountainea ryphea ryphea Cramer, 1775
142. Memphis basilia drucei Staudinger, 1887
143. M. cambyses Druce, 1877
144. M. glauce glauce C & K Felder, 1862
145. M. memphis memphis C & K Felder, 1867
146. M. phantes phantes Hopifer, 1874
147. M. monius morpheus Sraudinger, 1886
[48. M. philumena philumena Doubleday, 1849
149. M. pithyusa K. Felder, 1869
ISO. M. polycannes Ethricius, 1775
151. M. polyxo Druce, 1874
152. M. pmxias praxiag Hopifer, 1814
IS. M. xenocles xenoctes Hewirson, 1850
154. Archaeoprepona amphimachus symairhus Fruhstorfer, 1916
155. A. demophon muson Fruhstorfer, 1905156. A. demophoon andicola Fnihstorfer, 1904
157. A. licomedes Cramer, 1777
158. A. meander megabates Fnjhstorfer, 1916
159. Prepona dexamenus dexamenus Hopifer, 1874
160. P. lames demodice Godart, 1824161. P. pheridamas Cramer, 1777162. Agrias daudina sardanapalus Bares, 1860
NYMPHALIDAE APATURINAE
163. Doxocopa agathina agathina Cramer, 1777
164. D. tacit gñseldis C & K Felder, 1862
165. D. lavinia Butler, 1866
166. D. linda linda C& 8. Felder, 1862
[61. D. pavon pavon Latteille, 1809
168. 1. zunilda floris Fmhstorfer, 1907
NYMPHAL1DAE MOKPHINAE
169. Antirrhea bela C & 8. Felder, 1862
170. A. philoctetes avernus Hopifer, 1874
171. A. raygetina taygetina Butler, 1868
[72. Caerois chorinaeus protonce Fnihstorfer, 1912
173. Morpho achilles theodonjs Frubstorfer, 1907
174. M. deidamia grambergi Weber. 1944
115. M. eugenia ssp.
176. H. menelaus alexandrovna Dicer, 1874
177. M. reLemachus iphiclus C & R Felder, 1862
NYMPHALIDAE: BRASSOLINAE
178. Brassolis sophorae ardens Stichel, 1903
179. Narope cyllabarus Wesrwood, 1851
180. N. nesope Hewirson, 1869
181. N. panniculus Stichel. 1904
6 75 YES
10 77 NO
I 75 YES
50 75 YES
2 75 YES
1 76 YES
21 NC YES
92 92 YES
35 77 YES
19 88 NO
4 75 YES
79 79 YES
46 96 NO
2 76 YES
18 77 YES
I 76 YES
16 93 YES
30 81 YES
18 NC YES
79 79 YES
65 75 YES
3! 81 YES
4 77 YES
2 82 YES
37 NC YES
49 89 YES
18 83 YES
18 78 YES
16 NC YES
21 83 YES
3 78 YES
I 75 YES
20 76 YES
I 75 YES
54 8! YES
SR NC NO
SR NC YES
34 NC YES
23 83 YES
20 76 YES
91 91 YES
94 94 YES
228 229
Page 12
a
R. Robbins, 0. Lamas, 0. Mielke, 0. Harvey and M. Casngrande Butterfly Community at Pakitza
182. N. syllabus Staudinger, 1887
183. Narope sp. n.
184. Opsiphanes cassIae crameri C & R FeWer, 1862
185. 0. invirae amplificacus Srichel, 1904
186.0. quiteria quaestor Stichel, 1902
187. Opoptera aersa hilara Stichel, 1902
188. Opoptera sp. n.
189. Catoblepia berecynthia adjecta Scichel, 1906
190. C. soranus Wesrwood, 1851
191. C. xanthicles belisar Stichel, 1904
192. Selenophanes cassiope mapiriensis Bristow, 1982
193. Ei-yphanis automedon tristis Staudinger, 1887
194. Caligopsis seleucida seleucida Hewitson, 1877
195. Caligo euphorbus euphorbus C & R FeWer, 1862
196. C. eurilochus livius Staudinger, 1886
197. C. idomeneus idomenides Erubstorfer, 1903
298. C. placidianus Staudinger, 1887
199. C. reucer phorkys Fruhsrorfer, 1912
NYMPHALIDAE: SATYRINAE
200. Cithaerias pireta ssp. n.
201. Haetera piera ssp. n.
202. Pierella hortona alboIasciata Rosenberg & Talbot, 1914
203, P. lamia chalybaea Godman, 1905
204. P. lena brasiliensis C & K Felder, 1862
205. I3ia actorion rebeli Bryk. 1953
206. Manataria hercyna hyrnethia Fruhstorl'er, 1912
207. Hnrjesia blanda tvloschler, 1877
208. I-I. obscura Butler, 1867
209. I-I. oreba Butler, 2870
210. Harjesia U sp. n.
211. Pseudodebis griseola Weymer, 1911
212. P. marpessa Hewitson, 1862
213. P. valenrina Cramer, 1779
214. Taygetis celia Cramer, 1779
215. T. cleopatra C & R Felder, 1867
216. T. echo koepckei Forster, 1964
217. T. elegia Weymer, 1910
218. T. larua C & R Fekler, 1867
219. T. leuctra Butler, 1870
220. T. mermeria mermeria Cramer, 1776
221. T. sosis Hopffer. 2874
222. T. sylvia Bates, 866
223. T. thamyra Cramer, 1779
224. T. virgilia Cramer, 1776
225. Taygetis sp. n.
226. Caeruleuprychia aegrota Butler, 1867
227. C. brbcius Godart, 1824
228. C. cyanires Butler, 1871
229. C. glauca Weymer, 1911
20 76 YES
92 92 NO
91 91 YES
19 76 YES
44 78 YES
tO 79 YES
75 75 NO
I 76 YES
5 75 YES
I NC YES
2 77 YES
6 76 YES
26 76 YES
XX NC YES
20 78 YES
II 76 YES
21 NC YES
32 NC YES
4 75 YES
I 78 YES
I 75 YES3 76 YES
I 75 YES
I 80 YES
I? 8! YESI 75 YES24 78 YES
6 77 YES
I NC NO
2! 76 YES
78 78 YESI 75 YES
I 76 NO32 87 YES
83 83 YES
17 NC NOI 75 YES31 NC NO
4 76 YES6 87 YES
5 77 YES
2 87 YES
I 75 YES80 80 NOI 75 YESI 75 NO
20 NC YES
21 81 NO
230. C. helios Weymer, 1911
231. C. lobelia Butler, 1870
232. C. penicillata Godman. 1905
233. C. scopulata Godman, 1905
234. C. 2128 Butler, 1860
235. Caeruleuptychia sp. n. I
236. Caeruleuptychia sp. n. 2
237. Caeruleuptychia sp. n. 3
238. Caeruleuprychia sp. n. 4
239. Cepheuptychia cephus cephus Fabricius, 1775
240. Cepheuptychia sp. n.
24!. Chloreuptychia amaca Fabricius, 1776
242. C. catharina Staudinger, 1886
243. C. chlorimene Hubner, 1819
244. C. herseis Cedars, 1824
245. C. triarica Weymer, 1911
246. Chloreuptychia sp. n.
247. Cissia myncea Cramer, 1780
248. C. palladia Butler, 1867
249. C. proba Weymer, 1911
250. Erichthodes antonina C & R Felder, 1867
251. Euptychia enyo Butler 1867
252. Euptychia sp. n.
253. Hermeuptychia fallax C & R Felder, 1862
254. H. hermes Fabricius, 1775
255. Magneuptychia analis Godman 1905
256. M. iris C & R FeEder, 1867
257. M. `helle' Cramer, 1779 . homonym
258. M. lea philippa Butler. 1867
259. M. libye Linnaeus, 1767
260. M. moderata Weymer, 1911
26!. M. modesta Butler, 1867
262. M. ocypete Fabricius, 1776
263, M, segesta Weymer, 191!
264. Magneuptychia sp. n. I
265. Magneuptychia sp. n. 2
266. Magneuptychia sp. n. 3
267. Magneuprychia sp. n. 4
268. Magneuptychia sp. n. 5269. Magneuptychia sp. n. 6
270. Megeuprychia antonoe Cramer, 1775
27!. Pareuptychia binocuta binocula Butter, 1869
272. P. interjecta hesionides Forster, 1964
273. P. ocirthoe Fabricius, 1776
274. P. summandosa Gosse, 1880
275. Pareuptychia sp. n.
276. Paryphthimoides binalinea Butler, 1867
277. Postraygetis penelea penetea Cramer, 1777
278. Rareuptychia ctio Weymer, 191!
279. Splendeuptychia ashna Hewirson, 1869280. S. aurigera Weymer, 1911
19 81 YES
40 NC YES
20 75 NO1 75 YES
I 78 YES
I NC NO
63 80 NO
19 75 NO
80 80 NO
39 93 YES
6 76 YES2 76 YES
28 NC YES
I 78 NO
I 78 YES
I 95 NO
42 75 YES
2 85 YES
61 95 YES
4 83 YES
6 75 YES
2! 83 YES
89 89 NO
62 NC NO
I 75 YES
I 81 NO
I 87 YES18 77 YES
27 75 YES
2 NC YES
5 75 YESI 88 YES
29 85 YES
19 79 NO
Il 87 NO
18 87 YES
44 83 YES
2! 77 YES
6 77 NO20 83 NOI 76 YES
22 NC YES
I 79 YES
I 75 YES
22 NC YES
10 77 NO
16 76 YES
6 80 YES
3 76 YES
73 78 NO
34 NC NO
230 231
Page 13
R. Rabbins, 0. Larnas, 0. Mielke, V. Harvey and M. Casagrande
28!. 5, boliviensis Rxster, 1964
282. S. itonis Hewitson, 1862
283. S. purusana Aurivillius, 1929
284. S. quadrina Butler, 1869
285. S. rriangula Aurivillius, 1929
286. S. zischkai Forster, 1964
287. Splendeuptychia ap. n. I
288. Splendeuprychin sp. n. 2
289. Splendeuprychia sp. n. 3
290. Spkndeuptychia ap. it 4
29!. Splendeuprychia sp. n. 5
292. Splendeuptychia sp. n. 6
293. Splendeuptychia ap. n. 7
294. Spkndeuprychia sp. n. 8
295. Splendeuptychia sp. n. 9
296. Yphthimoides mytlna Weymer, 1911
297. Y. renata SmIl, 1780
298. Zischkaia amalda Weymer, 1911
299. 1 saundersii Butler, 1867
300. `Euprychia' ordinara Weytner. 1911
30!. Amphidecra calliomma C & R Felder, 1862
302. A. pigneraror pignemcor Butler. 1867
NYMPHALIDAE: DANAINAE
303. Lycorea ilione phenarete Doubleday, 1847
304. L halia pales C & R Felder, 1862
305. L pasinuntia concolor Scaudinger, 1885
306. Danaus eresimus sap. n.
NYMPHALIDAE: ITHOMIINAE
307. Athyrtis mechaniris salvini Srnka, 1884
308. Tirhorea harmonia brunnea Haensch, 1905
309. Melinaea machis lamasi Brown, 1977
310. M. marsacus Clara Rosenberg & Talbor, 1914
3!!. M. menophilus orestes Salvin, 1871
312. M. mnasias romualdo Fox, 1965
313. Paitiria neglecra Lamas, 1979
314. Thyridia psidii mo C & R FeLler, 1862
315. Forbestra olivencin aeneola Fox, 1967
316. Mechaniris lysimnth menecks Hewitson, 1860
3!?. M. mazaeus mazaeus Hewirson, 1860
318. NI. polymnia angusrifascia Talbor. 1928
319. Scada haresi baresi Haensch, 1903
320. 5. reckia labyrmntha Lamas, 1985
32!. Aeria eurimedia negrucola C & R FeLler, 1862
322. Merhona confusa psamathe Godman & Salvin, 1898
323. M. curvifascia Weymer, 1883
324. NI. grandmor asp. n.
13 75 YES
I 75 YES
8 NC YES
6 87 NO
3 76 YES
37 NC NO
2 88 NO
3 76 NO
10 75 YES
34 8! NO
8 76 YES
I 76 YES
74 76 NO
19 85 NO
89 89 NO
IS NC YES
23 89 NO
4 75 YES
28 84 NO
73 NC YES
34 8! YES
3! 79 YES
16 96 YES
3 80 YES
XX NC NO
IS NC YES
4 75 YES
I 76 YES
4 75 YES
I 87 YES
2 76 YES
24 NC NO
22 77 YES
3 78 YES
20 76 YES
9 76 YES
63 88 YES
22 NC YES
2 76 NO
I 76 YES
18 75 NO
I 75 YES
2 76 YES
I 78 NO
325. Rhodussa cantobrica pamina Haensch, 1905
326. Napeogenes aethra deucalion Haensch, 1905
327. N. machis patientia Lamas, 1985
328. N. pharo pharo C & R Felder, 1862
329. N. stella sap. n.
330. N. syiphis sylphis Ouérin, 1844
33!. Hypothyris euclea sap. n.
332. H. ninonia sap. n.
333. H. semifulva sap. n.
334. Hyposcada anchiala richardsi Fox, 194!
335. H. illinissa dobbe!!a Hewitson, 1876
336. H. zarepha sap. n.
337. Oleria alexina Hewitson, 1859
338. 0. didymaca didyn,aea Hewirson, 1876
339. 0. gunilla asp. n.
340. 0. onega lentira Lamas, 1985
341. C. ramona calarha Lamas, 1985
342. 0. vicrorine vjcrorine Cuerin, 1844
343. Ithomia agnosia agnosia Hewitson, 1855
344. I. arduinna arduinna d'Almeida. 1952
345. I. lagusa pemana Sa!vmn, 1869
346. I. Iichyi neivai d'Almeida, 1940
347. I. salapia ardea Hewirson, 1855
348. Callithomia alexirihoc thomax Bates, 1862
349. C. lenea zelie Guérin, 1844
350. Dircenna dero sap. n.
35!. D. loreta acreana dAlmeida, 1950
352. Cerarinia neso peruensis Haensch, 1905
353. C. tuna fuscens Haensch, 1905
354. Ceratiscada hymen hymen Haensch, 1905
355. Episcada suiphurea sulphurea Haensch, 1905
356. Episcada sp. n.
357. !`teronymia annisao gunnheri Lamas, 3985
358. P. orsreni Baumann, 1985
359. P. vestilla acaya Haensch, 1909
360. Godyris zavalena sap. n.
361. Hypoleria lavmnia cajona Haensch. L905
362. H. virginia viniosa Lamas, 1985
363. `Hypo!cria' aelia brevicula dAlmeida, 1951
364. `I-I.' orolmna arzalia Hewmnson, 1876
365. Mcclursgia cymo salonmna Hewitson, 3855
366. Pseudoscada rimna sap. n.
367. Heterosais nephele nephele flares, 1862
368. `Pseudoscada' florula sap. n.
Butterfly Community at Pakitza
5 75 YES
4 75 YES
5 76 YES
4 75 YES
2! 76 NO
2 75 NO
L8 76 YES
22 83 NO
17 75 NO
6 79 NO
XX NC NO
43 NC NO
23 NC NO
IS NC YES
XX NC NO
5 76 YES
I 77 YES
5 76 YES
35 NC YES
4 76 YES
22 NC NO
2 75 YES
4! 78 YES
54 NC YES
2 75 YES
19 NC YES
12 96 YES
II 75 YES
33 NC YES
IS 76 YES
19 76 NO
77 77 NO
II 76 YES
4 76 YES
6 76 NO
Ii 76 NO
II 76 NC
5 76 YES
30 90 NC
19 78 YES
7 76 NC
Ii 76 YES
7 76 YES
16 83 NC
233
I
232
NYMPHALIDAE: LIBYThEINAE
369. Libyrheana carinenra carmnenra Cramer, 1777 19 NC YES
Page 14
U. Robbins, 0. Lamas, 0. Mielke, D. Harvey and M. Casagrande Butterfly Community at Pakitza
234
RIODINIDAE: RIODININAE
404. Perophrhalma tullius rullius Fabricius. 1787
405. Mesophthalma idotea asp. nfl
406. Leucochimona maratha chionea Oodman &
407. L. matisca Hewitson, 1860
408. Semomesia croesus siccata Stichel. 1919
409. S. macaria Hewitson, 1859
410. S. tenella tenella Stichel, 1910
411. Mesosemia all. ephyne Cramer, 1776
412. Mesoscn,ia aff. merura I-Iewitson, 1873
413. Mesosemia all. gneris Wesrwoocl, 1851
414. Mesosemia sp. I
415. Mesosemia 5l,. 2
20 78 YES
I 76 YES
Salvin, 1885 5 75 YES
66 90 YES
4 75 YES
17 75 YES
89 89 YES
20 81 YES
18 83 YES
43 88 Nc
36 NC NO
46 77 Nc
L.
416. Mesosemia all. cyanira Stichel, 1909
417. M. cippus Hewitson, 1859
418. M. ibycus Hewitson, 1859
419. M. philocles thyesres Bruce, 1878
420. M. machaera asp.
42!. M. marerna Stichel, 1909
422. Mesosemia all. marerna Stichel, 1909
423. M. luperca Stichel, 1910
424. Mesosemia sp. 3 umbrosa ?
425. M. hedwigis Stichel, 1910
426. M. naiadella naiadella Stichel, 1909
427. M. sirenia sirenia Stichel, 1909
428. M. latissima Srichel. 1909
429. Mesosemia all. evias Stichel. 1923
430. M. menoeres paetula Stichet, 1915
43!. Mesosemia sp. 4 nr. atroculis
432. M. ulrica ulrica Cramer, 1777
433. M. eumene luria Stichel, [910
434. M. decolorata Lathy, 1932
435. M. macella Hewitson, 1859
436. M. giganrea Srichel, [915
437. Euryhia nicaea ssp.
438. E. caerulescens cacrulescens Bruce, 1904
439. B. dardus franciscana C & R Felder, 1862
440. B. promora Stichel, 1910 ?
441. E. halimede halimede HUbner, 1807
442. Alesa prema Godart. 1824
443. Alesa all. telephae Boisduval, 1836
444. A. amesis Cramer, 1777
445. A. hemiurga Bates, [867
446. Mimocastnia rorhachildi Seitz, 1916
447. Hyphilaria parthenis rigrinella Stichel. 1909
448. Cremna acroris meleagris Hopifer, 1874
449. C. rhasus subrurila Sticliel, 1910
450. Eunogyra satyrus Westwood, 1851
451. Lyropteryx apollonia apollonia Westwood, 1851
452. Cyrenia martin marcia Wesewood, 15l
453. Ancyluris meliboeus melibocus Fabricius, 1776
454. A. etias inelior Stichel, 1910
455. A. aulestes cryxo Saunders, [859
456. Rhetus arcius huanus Saunders, [859
457. R. periander laonome Morisse, 1838
458. Irhomeis lauronia Schaus, 1902
459. Isapis agyrtus sestus Stichel, 1909
460. Themone poecila Bates, 1868
46!. Norheme erora diadema Stichel, 1910
462. Monethe albertus alberrus C & It Felder, 1862
463. Meracharis lucius Fabricius, 1793
464. M. regalis regahis Butler. 1867
465. Cariomothis erythromelas lulvus Larhy, 1932 ?
466. Synnaria nyx Hubner, 1817
25 NC NO
20 79 YES
IC 83 YES
12 75 YES
2 77 YES9 77 YES
2 76 YES70 79 YES
2 81 NO
10 79 NO
6 NC YES
8 75 YES
14 NC NO
34 88 YES
8 77 YES
13 NC NO
27 82 YES
12 NC NO
I NC YES
95 95 NO22 8! NO
I 75 YES
8 81 YES
14 77 YES
43 NC YES
4 75 YES
91 9! NO
3 9! NOI 75 YES8 88 YES
IS NC NO3 NC YES3 78 YES
8 NC YES50 78 YES20 88 YES75 75 YESIS 77 YES21 79 YES6 77 YES
49 76 YES
3 76 YES
I NC YES3 75 YES14 NC NO16 81 YES
46 76 YES9 75 YES
I 75 YES
5 96 YES14 92 NO
I
RIODINIDAB EUSELAS!INAE
370. Euselasia euboea euboca Hewirson, 1853 27 77 YES
37!. E. pelor Hewitson, 1853* 16 77 NO
372. B. pellonia Stichel, [9j9* 16 75 YES
373. E. mirania Bates. l868 26 NC YES
374. E. roppini Sharpe. 1915 20 76 YES
375. E. curyonc euryone Hewirson, 1856 28 75 YES
376. E. violerta Bates, 1868 24 NC YES
377. E. arhas sap. 16 87 YES
378. B. euoras Hewitson, 1855 51 NC YES
379. B. eutychus Hewirson. 1856 19 85 YES
380. E. jugata Stichel, 1919 16 77 YES
38!. E. euodias euodias HewiNon, 1856 16 77 YES
382. E. orba spectralis Stichel, 1919 XX NC YES
383. E. euriteus euriteus Cramer, 1777 IS 76 YES
384. E. melaphaea condensa Scichel, 1927 19 77 YES
385. B. hygenius group, sp. 1 2! NC YES
386. B. hygenius group, sp. 2 26 NC YES
387. E. hygenius group, sp. 3 18 83 YES
388. B. hygenius group, sp. 4 19 75 YES
389. Euselasia afl. calusa flares, 1868 19 76 YES
390. E. alcmena Bruce. 1878 5 75 NO
391. B. crinon Stichel, 1919 16 85 YES
392. E. fervida hahneli Staudinger, [887 16 90 NO
393. E. gelanor erilis Stichel, 1919 21 84 YES
394. B. teleclus releclus Stoll, 1787 20 83 YES
395. Buselasia sp., midas group 19 91 YES
396. E. eugeon Hewirson, 1856 26 79 YES
397. E. brevicauda Lathy, 1926 2! 83 YES
398. E. uria angusrilascia Lathy, 1926 19 77 YES
399. B. eubotes eubotes Hewirson, [856 35 NC YES
400. B. lysiniachus Sraudinger, ¶888 5 83 NO
40!. B. angulara Bates, 1868 21 83 YES
402. E. utica euphacs Hewitson, 1855 16 84 YES
403. Merhone cecilia magnarca Seirz, 1913 26 85 NO
235
Page 15
R. Rohbins, 3. Lamas, 0. Mielke, Li Harvey and M. Casagrandc Butterfly Community at Pakirza
92 92 YES
23 96 NO
6 76 YES
76 76 NO
I 77 YES
26 NC NO
43 92 YES
28 81 NO
9! 9! NO
29 NC NO
6 NC NO
4 76 YES
20 75 NO
34 81 YES
4 77 NO
76 76 YES
82 82 YES
5 81 YES
2! NC NO
78 78 NO
29 NC NO
6 NC YES
25 NC YES
83 83 YES
15 79 YES
57 75 NO
42 NC NO
2 75 YES
2 75 YES
3 76 YES
19 79 YES
2 80 YES
16 84 YES
93 93 NO
74 NC YES
77 77 YES
97 97 NO
42 NC NO
3 NC NO
85 85 NO
92 92 NO
9 83 YES
2 76 YES
3 78 YES
14 76 YES
10 79 YES
13 84 YES
6 80 NO
92 92 NO
8 NC NO
S NC NO
467. Chamaelimnas tircis iaeris Bates, 1868 44
468-C. urbana Stichel, 1916 I 80
YES469. Parcella amarynthina C & R Felder, 1865 4
YES470. haris anius Cramer, 1776 2
YES47!. Charis sp. n. I
YES472. C. gynaca zama Bates. 1868 I
75 NO473. C- argyrea Bates. 1868
76 YES474. Chalodeta theadara rhccxlora C & R Felder. 1662
475.C. lypera Bates. [868 79 79
476. C. chaonitis flewitson. 1866
477. Carla mantlnea amazanica Bates, 1868
478. C. trochilus arete C & R Felder, 1861
3
IS
42
76
82
78
NC
YES
YES
NO479. C. `philema' Stichel, 1910
82 NO480. C. sponsa Staudinger, 1887
481. Crocozona coecias coecias Hewitson, 1866
482. Baeotls bacaenis bacaenita Schaus, 1902 U
483.13. euprepesorthotaenia Seirz, 1916
484. Lasais agestlas agesilas Litreille, 1809
46
9
2
3
77
78
76
78
YES
YES
NO
YES
YES485. L arMs Staudinger, 1887 4 89
486. L pseudomeris Clench, 1972 [9 90
YES487. Amarynthis meneria Cramer. 1776 8
488. Exoplisia cadmeis Hewitson. 1866 XX
489. Riodina lysippus lysias Stichel, 1910 29 78
490. Melanis xarifa qiiadripunctara Stichel, 1910
491. M. smirhiae Westwood, 1851
2!
18
79
76 NO
492. M. marathon stenotaenia RaIser, 1904 76 76
493. Mesene leuaçhrys Bates, 1868 I
YES494. M. nola eupteryx Bates, 1868 8
495. M. pyrrha Bates, 1868 19 77
NO496. Mescne sp. I 29
497. Mesene sp. 2 90 90 NO
498. Mesene monostigma Erlchson, 1848 U 83 83 YES
499. twlesene aft. silaris Codman & Salvin, 1878 77 77 NO
500. Symmachia mbina separata Lathy, 1932
501. 5. accusatrix Westwood, 1851
23
84
NC
84
NO
NO
502. Symmachia sp. I !cleonyma Hewitson, 1870 79 79
91 YES503. Synimachia sp. 2 !prabetor Stall, 1782 91
YES504. S. asclepia asclepia Hewirson, 1870 32
505. Phaenochitonia sophistes Bates. 1868
506. Sarota acantus Stall. 1781
507. Sarota sp. nr. acantus Stall, 1781
39
4
8
76
76
NC
YES
YES
NO508. Sarota sp. I 66
NO509. 5. flavicincta Lathy, 1932 92 92
510. Sarota aft. myrtea Godman & Salvin, 1886 14 NC YES
5!!. Sarota sp. 2 53 8! NO
512.Saratasp.3 22 75
513. Sarota Sr. 4 93 93 NO
514. 5. acanthoides spicata Staudinger, 1888 ? 20 NC NO
515. S. chrysus chrysus Stall, 1781
516. Anteros formosus formosus Cramer. 1777
20
6
83
79
YES
YES
518. A. renakius renaldus SroIl, 1790
519. Calydna caieta Hewitson, 1854
520. C- punctara C & R Felder, 1861
52!. C. thersander Stall, 1780 U522. C. maculosa Bates, 1868
523. C. hiria Godart, 1824
524. C. catana Hewitson. 1859
525. C. carneia Hewitson. 1859
526. C. cea Hewitson, 1859
527. C. calyce Hewitson, 1859
528. Emesis lucinda lucinda Cramer, 1775
529. E. castigata castigata Stichel, 1910
530. E. spreta Bates, 1868
53!. E. mandana mandana Cramer, 1780
532. E. diogenia Prirtwitz. 1865
533. E. fatimella fatimella Westwoocl, 1851
534. E. ocypore ocypore Oeyer, 1837
535. E. temesa emesina Staudinger, 1887
536. E. progne Godman, 1903
537. Emesis sp. ?heteroclita Stichel, 1929
538. Argyrogrammana stilbe Godart, 1824 U holosticta!
539. Argyrogrammana sp. I trochilia rameh?
540. Argyrogrammana sp. 2
541. Pachyrhone xarnhe Bates, 1868
542. Uraneis hyalina Butler. 1867
543, Thisbe irenea ssp.
544. Lemonias zygia ssp.
545. Juditha azan asp. fl.
5464. molpe molpe HUbner, 1808
547. Synargis orestessa HUbncr, 1819
548. 5. aharis Cramer, 1776
549.S. gela gela Hewirson, 1853
550.5. ochra ochra Bares, 1868
551.5. phillone Godart, 1824
552. Parnes nycteis Westwood. 1851
553. P. philotes Westwoocl, 185!
554. Menander caruscans Butler, 1867
555. M. pretus pretus Cramer, 1777
556. M. hebrus hebrus Cramer, 1775
557. Dysmarhia portia Bates, 1868
558. Dysmathia grosnyi Le Cerf, 1958
559. Calospila lucianus lucianus Fabricius, 1793
560. C. emylius emyliana Stichel, 1911
56!. C. rhodope amphis Hewitson, 1870
562. C. parthaun Dalman, 1823
563. C. zeanger pirene Oodman, 1903
564. Calospila sp. I rhesa ssp.?
565. C. rhara pulchra Lathy, 1904
566. C. apotheta Bates, 1868 U567. Calospila ,iff. heinileuca Bates, 1868
568. C. siaka siaka Hewirson, 1858
236237
Page 16
I?. Robbins, G. Lamas, 0. Mielke, 0. Harvey and M. Casagrande Butterfly Community at Pakicza
569. Calospila antonil Brévignon, 1995
570. Adelocypa annulilera Godman, 1903
57!. A. densemaculata Hewitson, 1870
572. A. amasis 1-lewitson, 1870
573. A. epixanthe Stichel, 191 I'
574. A. aminias aminias Hewitson, 1863'
575. Adelotypa sp. I'
576. A. leucocyana Oeyer, 1837'
577. A. huebneri pauxilla Stichel, 1911'
578. Adelotypa aristus.''
79. A. moths asemna Stichel, 1910
580. A. crinitatis ssp.
58!. `Adetotypa' tampTos Bates, 1868
582. Setabis epitus epiphanis Stichel, !910
s83 S. vetutina Butler, 1867
584 S pyçhsoides Butlet 867
55.S. crbentataButtdr lS6
Sg&S.flammutiBatesy!868
587. Setabis sp. I
588. Setabis sp. 2
589. Theope eudocia eudocia Westwood, 1851
590. T. hypoleuca Bates, 1868
591. Theope sp. in. hypoteuca Bates, 1868
592. T. lycaenina Bates, 1868
593. Theope sp.
594. T. pedias pedias Herrich-Schaffer, 1853
595. T. excelsa Bates, 1868
596. Theope all. mundula Stichd, 1926
597. Theope alt, theritas Hewirson, 1860
598. T. phaeo lotia Godman & Salvin, 1886
599. T. comosa Stichel, 1911
600. Theope afi. thootes Hewitson, 1860
60!. Theope aff. thestias Hewitson, 1860
602. Calociasma pulcherrima comparara Stichel, 1911
603. Nymphidium mann's Gamer. 1775
604. N. fulminans fuirninans Bates, 1868
605. N. baeotia Hewitson, 1853
606. N. minuta Druce, 1904
607. N. azanoides amazonensis Catlaghan, 1986
608. N. omois Hewitson, 1865
609. N. ascotia augea Druce, 1904
610. N. teucosia medusa Drtsce, 1904
611. N. acherois eryinanthus Ménétriès, 1855
612. N. caricae parthenium Stichel, 1924
613. N. tisimon lisimon Scott. 1790
614. Scalachtis calliope sap. n.
615. Setabis sp. n. 3
79 79
I 75
42 NC
6 NC
4 NC
13 76
4 75
1 75
I 76
I 75
I 75
16 76
4 NC
3 79
59 88Ii. 75
2 8I'
57
2 NC
NO
YES
NO
NO
YES
YES
NO
YES
YES
YES
YES
NO
YES
YES
YES
YES
YE&YES
NO
LYCAENIDAE: ThECLINAE
616. Eumaeus minijas Hubner, 1809
617. E. toxana Boisduval, 1870
618. Mithras nautes Cr,imer, 1779
619. `Thect? nr. orobia Hewitson, 1867
620. `mcd? coarnophila Tessn,ann, 1928
621. `Thect? macdata Lathy, 1936
622. Thestius meridionalis Draudt, 1920
623. "Thecta' ematheon Cramer, 1777
624. Event's gabrieta Cramer, 1775
625. E. batesii Hewitson, 1865
626. E. lioralia Druce, 1907
627. E. satyroides Hwitso.u 1865628. `Thecla' gibberbl4P,.itan, 1867
629,. "lThecIa' aterina Ijçwjtson. Q6* ``thcta myrtea fl-levcton jâei63V Thecta" mrcusa Hewirson, 1867
632. Attosmaitia strophius Godart, 1824
633. Arcas imperiatis Cramer, 1776
634. A. tuneta Hewitson, 1865
635. Theritas mayor, HUbner, 1818
636. Denivia acontius Goodson, 1945
637. 0. phegeus Hewitson, 1865
638. Denivia nr. viresco Druce, 1907
639. 0. viresco Druce, 1907
640.0. hemon Cramer, 1775
64!. 0. tisus StolI, 1790
642. Atlides polybe Linnacus, 1763
643. A. atys Cramer, 1779
644. Paiwairia tetemus Cramer, 1775
645. P. venulius Cramer, 1779
646. `Thecla' tigurina Hewitsor., 1874
647. `Thecla' ergina Hewitson, 1867
648. Thereus cotumbicota Strand, 1916
649. Arawacus separara Lathy, 1926
650. Rekoa meton Cramer, 1779
651. Ocaria ocrisia Hewitson, 1868
652. yanophrys amyntor Cramer, 1775
653. Panthiades bitias Cramer, [777
654. P. aeolus Fabricius, 1775
655. I'. phateros Linnacus, 1767
656. `Thecla' gemma Druce, 1907
657. `Thecla' minyia Hewitson, 1867
658. `Thecta' echelta Hewitson, 1867
659. Parrhasius polibetes Stall, 1781
660. P. orgia Hewitson, 1867
66!. Michaelus Ira Hewirson, 1867
662. M. vibidia Hewicson, 1869
663. M. thordesa Hewirson, 1867
664. M. jebus Oodarr, 1824
XX NC NO
3 83 YES
Il 77 NO
89 89 YES
4 85 NO
15 NC NO
2 75 YES
14 85 YES
80 80 NO
9 NC NO
14 90 NO
4 NC NO
10 85 YES
79 79 YES
I 75 NO
3 76 YES
3 75 YES
I 76 YES
2 75 YES
5 84 YES
22 NC YES
2 75 YES
I 75 YES
I 75 YES
1 75 YES
1 76 YES
ID 83 YES
80 80 NO
78 78
64 88
19 75
35 91
81 81
20 81
4 76
48 79
91 9!
9! 91
79 79
SR NC
10 97
4_ 75
46 81''
79 79
95 953 75
43 85
80 80
91 91
14 79
47 NC
ID 89
4 77
91 9!
92 92
93 93
26 8392 9296 96
48 92
4 NC
3 75
80 80
3 77
9 NC
I 75
79 79
2 78
4 84
93 93
14 NC
14 90
14 90
12 84
14 90
23 NC
89 89
NO
NO
YES
YES
YES
YES
YES
YES
YES
YES
YES
YES
NO
YES
YES
NO
YES
YES
YES
YES
YES
NO
YES
YES
NO
NO
NO
YES
NO
YES
YES
YES
YES
NO
YES
NO
YES
YES
YES
YES
YES
NO
YES
NO
YES
YES
YES
NO
2.38 239
Page 17
R, Robbins, 3. Lamas, 0. Mielke, 0. Harvey and M. Casagrande Butterfly Community at Pakitza
665. `Thecla' nr. gadira Ilewitson, 1867
666. "Thecla' norax Codman & Salvin, 1887
667. `Thecla" levis Druce, 1907
668. Olynthus obsoleta Lathy, 1926
669. Olynthus essus I-lerrich-Schiiffer, 1853
670.0. nitor Druce, 1907
671. Oenomaus ortygnus Cramer, 1779
672. Oenomaus nr. arena 1-lewitson, 1867
673. Srryn,on cesrri Reakirt, 1867
674. 5. ziba Hewitson, 1868
675. S. megarus Oodart, 1824
676. Lamprospilus orcidia 1-Tewirson, 1874
677 Lamprospilus nv picentia Hewitson, 1868
678. L. netesca Draudt, 1920
679. "Thecla' arm Hewirson, 1874
2
3
89
10
91
12
14
92
96
2
89
12
84
14
90
79
90
89
84
91
79
NC
92
96
76
89
79
84
79
90
YES
YES
NO
YES
NO
NO
NO
YES
YES
YES
NO
YES
NO
NO
NO
`SO. `Thecla' raminella Schaus, 1902
68!. `Thecla' aruma Hewitson, 1877
682. `Thecla' syllis Godman & Salvin, 1877
683. Kisnrans hesperitis Butler & Druce, 1872
684. `1 cclii' ceromia 1-lewitson, 1877
685. "Theda' vesper Druce. 1909
686. Electrostrymon ecbatana Hewitson, 1868
687. Symbiopsis "peruviana' Lathy, 1936 homonym
688. S. aprica Moschler, 1883
689. Calycopis calus Godart, 1824
690. C. buphonia Hewitson, 1868
69!. C. demonassa Hewitson. 1868
692. C. atnius Herrich-Schiffer, 1853'
693. Calycopis fly. atnius Herrich-Scltlfler, 1853'
694. C. devia Moschler, 1883'
695. C. cenroripa Hewirson, 1868
696. C. nicolayl Field, 1967 U
697. C. anfracra Druce, 1907'
698. C. anastasia Field, 1967'
699. C. vitruvia Hewitson, 1877
700. C. caesaries Druce, 1907
70!. C. cerata Hewirson, 1877
702. C. trebula Hewitson, 1868
703. C. anapa Field, 1967
704. C. orcilla Hewirson, t874'
705. C. naka Field, 1967 0'
706. Calycopis nr. vidulus Druce, 1907'
707. C. dfla Field, 1967'
708. Calycopis nr. dfla Field, 1967'
709. Calycopis nr. orcilla Hewitson, 1874'
710. Calycopis nr. pisis Godman & Salvin, 1887'
ill. C. harm Field, 1967 ?`
712. Tmolus echion Linnaeus, 1767
14
2
9
2
4
4
14
44
I
6
8
I
1
3
3
2
42
3
I
I
I
6
12
77
13
I
15
8!
3
IS
24
I
2
79
76
83
76
76
75
84
85
79
79
76
77
76
84
83
75
NC
79
77
77
83
76
76
77
77
78
NC
81
81
75
NC
76
75
YES
YES
YES
YES
YES
YES
NO
YES
NO
YES
YES
YES
YES
NO
YES
YES
YES
YES
YES
YES
YES
YES
YES
NO
NO
YES
YES
NO
YES
YES
NO
YES
YES
713. Tmolus nr. cydrara Hewitson, 1868
714. T. cydrata Hewirson, 1868
715. T. ufentina Hewitson, 1868
2
6
4
77
79
79
YES
YES
YES
716. Tmolus or. ufenrina Hewitson, 1868
717. T. mutina Hewirson, 1867
718. Tmolus nr. murina Hewitson, 1867
719. "Thecla' emessa Hewirson, 1867
720. `Thecla' fly. opatia Hewirson, 1868
721. `Thecla' or. cupa Druce, 1907
722. `Thecla' fabulla Hewirson, 1868
723. `Thecla' nr. purpuriticus Druce, 1907
724. "Thecla' tympania Hewirson, 1869'
725. `Thecta' nr. rympania Hewitson, 1869
726. `Thecla' nr. empusa Hewitson, 1867
727. `Thecla" hatciones Butler & Druce, 1872'
728. `Tficcla" tarena Hewirson. 1874
729. `Thecla" sospes Draudr, 1920
730. Siderus leucophaeus Hubner, 1813
731.S. parvinorus Kaye, 1904
732. Siderus nr. guapila Schaus, 1913
733. S. guayra Jorgensen, 1935 ?
734. 5. athymbra Hewirson, 1867
735, 5. metanira Hewirson, 1867
736. S. viola Draudt, 1920
737.S. caninius Druce, 1907
738. Siderus nr. panchaea Hewitson, 1869
739. `Thecla' splendor Johnson, 1991
740. Theclopsis lydus HUbner, 1819
74!. T. gargara Hewitson, 1868
742. `Thecla' tephraeus Oeycr, 1837
743. `Thecla' nr. tephraeus Ocyer, 1837
744. "T." sphinx Fabricius, 1775
745. "T." phoster Druce, 1907
746. `T.' pulchrirudo Dame, 1907
747. `T.' strephon Fabricius, 1775
748. `Thecla' nr. strephon Fabricius, 1775
749. `Thecla' perola Hewitson, 1867
750. "T." parvipuncra L.arhy, 1926
75!. `T." agrippa Fabricius, 1793
752. `T.' carteis Hewitson, 1870
753. `Thecla' or. carteia Hewirson, 1870
754. `T.' ryriam Druce, 1907
755. `Thecla' fly. ryriam Druce, 1907
756. `Thecla' nr. malvania Hewitson, 1867
757. "Thecla' nr. Ioyi Schaus, 1902
758. `Thecla' syedra Hewitson, 1867
759. "Thecla' or. syedra Hewitson, 1867
760. `T." adela Staudinger, 1888
76!. `T.' arnbrnx Wesrwood, 1852
762. Minisrrymon zilda Hewirson, 1873
763. M. cruenta Gosse, 1880
764. Ministrymon or. cruenta Gosse, 1880
765. M. cleon Fabricius, 1775
766. "Thecla' rerenria Hewirson, 1868
3 79 YES
2 75 YES
91 91 NO
10 84 YES
3 79 NO
76 76 NO
97 97 NO
IS 90 NO
6 75 YES
6 76 YES
12 75 NO
6 NC NO28 8! YES
26 NC NO
19 87 YES
96 96 YES
5 NC NO
I 82 NO
14 8! YES
6 75 YES
6 NC NO
84 84 NC
77 77 NO
13 83 YES
3 77 YES
3 75 YES
2 80 YES
20 NC YES
13 79 YES
3 75 YES
77 77 YES
16 77 YES
tO 79 YES
8 76 NO
IC 79 YES
84 84 YES
4 76 YES
29 NC YES
4 75 YES
20 90 YES
76 76 NO
IC NC NO
IS 9! YES
5 77 YES
8 77 Nc
85 85 NC
6 76 YES
5 85 YES
12 89 YES
2 80 Nc5 78 YES
240 241
Page 18
U. Robbins, C. Larnas, 0. Mielk,2, 0. Harvey and M. Casogrande Butterfly Community at Pakitza
767. "Thecla' lycabas Cramer, 1777
768. Auberaina ald:s Hewitson, 186$
769. janthecla rocena Hcwitsan, 867
770. Janthecla it. rocena Hewitsors. 1867
77l.j. malvina Hewitson, 1867
772. j. leea Venabks & Robbins, 1991*
773.. sista Hewirson, 1867'
774. Hypostrymon nsa Hewitson, 1873
775. laspis nr. talayra Hewirson, 1868
776. I. rhabena Hewirson, 1868
777. laspis nr. beera Hewitson, 1870
778. I. remesa Hewitson, 1868
779. `Thecla' picus Dnsce, 1907
780. Brangas teucria Hewitson, 1868
781. B. gerus Fabricius, 1787
782. `Thecla' thespia Hewitson. 1870
783. `Thecla' cupentus Stoll. 1781
784. `Thecln' nr. biston Moschler, 1877
785. Nesiostrymors celona Hewitson, 1874
786. Erora oleris Druce, 1907 U
787. E. phrosine Druce, 1909
788. E. Carla Schaus, 1902
789. Erora nr. opisena Druce, 1912
790. E. badera Hewitson, 1873
791. `Thecla' terms Hewitson, 1867
792. CaeroIethra carnica Hewirson, 1873
793. C. iambe Godman & Snlvin, 1887
794. Celmia celmus Cramer, 1775
795. `Tbecla' color Drssce, 1907
796. `Thecla' mecrida Hewitson, 1867
80
3
6
26
91
25
3
I
80
92
5
72
91
92
75
90
0
13
79
32
41
3
91
79
I
43
79
I
46
3
80
88
84
83
91
75
76
78
80
92
NC
NC
91
92
75
90
84
84
79
NC
NC
85
91
79
75
84
79
75
85
97
NO
YES
YES
NO
YES
YES
YES
YES
NO
NO
NC
YES
YES
NO
YES
NO
YES
NO
YES
NO
YES
NO
NO
NO
YES
YES
NC
YES
YES
YES
PIERIDAE: DISMORPHIINAE
797. Pseudopieris nehemia melanin Lamas, 1985
798. Dismorphin amphione ssp. n.
799. D. theucharila argochloe Bates, 1861
800. Enantia lina galanthis Bates, 1861
801. E. melite linealis PrufIer, 1922
802. Moschoneura pisirbous asp. n.
803. Patia orise denigrara Rosenberg & Talbor, 1914
PIERIDAE: PIERINAE
804. Anteos clorinde Godart, 1824
805. A. menippe Hubner, 1818
806. Aphrissa fluminensis d'Almeida, 1921
807. A. starira statira Cramer, 1777
808. Pl'ioehis arganre larra Fabricius, 1798
809. P. phile;. philea Linnacus, 1763
810. P. sennae inarcellina Cramer, 1777
811. Rliabdodryas trite trite Linsiacus, 1758
16 78 YES
26 78 NO
6 88 YES
II 76 YES
19 77 YES
I 8! NO
12 NC NO
25 NC NO
2 77 YES
4 78 YES
2 77 YES
2 NC YES
2 NC YES
34 88 YES
25 78 YES
812. Eurema agave agave Cramer, 1775
813. 8. album espinosne Fernandez, 1928
814. E. arbela arbela Geyer, 1831
815.8. irma Bates, 1861
816. E. paulina Bates, 1861
817. Pyrisitia leuce flavilla Bates, 1861
818. P. nise ssp. n.
819. Cunizza hirlanda ninguida FruhstorIcr, 907
820. Glurophrissa drusilla drusilla Cramer. 1777
821. Ascin montrste automate Btirmcisrer, 1878
822. Ganyra phaloe suhijoenta Schaus, 1902
823. Itahallin demophile lucania Frubstorfer. 1907
824. I. pandosia pisonis Hewitson, 1861
825. Pierihallia viardi ruhecula Fruhsrorfcr, 1907
826. Ivleleme lycimnia peruviana Lucas, 1852
827. Perrhyhris pamela mazuka Limas, 1981
PAPILIONIDAE: PAPILIONINAF
828. Prorographmum agtsilaus autosilaus Bates, 1861
829. Eurytides dolicaon deileon C & R Felder. 1865
830. Protesilaus glaucolaus leucas Rorhsch. & Jord., 1906
831. P. telesilaus relesilaus C & Ii Felder, 1864
832. Mimoides ariar.srhes gayi Lucas, 1852
833. M. pausanias pausanias Hewitson, 1852
834. M. xynias xynias Hewirson, 1875
835. llartus velus varus KoIlar, 1850
836. 13. crassus crassus Cramer, 1777
837. 13. polydamas polydamas Linnnetis, 1758
838. Parides aeneas lamasi Racheli, 1988
839. P. anchises drucei Butler, 1874
840. P. echemon empisrocles KUppers, 1975
84!. P. neophilus olivencius Bates, 1861
842. P. pizarro kuhlntinni May, 1925
843. P. sesostris sesostris Cramer, 1779
844. P. verrurnnus asrorius Zikan, 1940
845. Prerourus zagreus zagreus Doubleday, 1847
846. Heraclides anchisiades anchisiades Esper, 1788
847. I-I. androgeus androgeus Cramer, 1775
848. H. astyal us phanias Rothschild & Jordan, 1906
849. H. chiarssiajes chiansmades Wesrwocxl. 872
850. H. garleppi inrerruptus Staudinger, 1892
851. H. thoas cinyras Menétriès, 1857
852. H. rorquarus torquatus Cramer, 1777
853, Pyrrhopygc pusca Evans, 1951
854. P. proculus draudri Re!!, 1931
855. P. rubricollis ssp. n.
856. P. comeres ssp. ii.
83 83 YES
I 75 YES
19 NC NO
22 NC NO
3 NC YES
27 84 YES
2 75 YES
95 95 NO
2 76 YES
25 NC YES
8 78 YES
2 75 YES
I 75 YES
76 76 YES
2 77 YES
16 80 YES
2 78 YES
25 76 YES
2 82 YES
18 76 YES
22 95 YES
2 78 YES
83 83 YES
78 78 YES
SR NC YES
41 NC YES
20 76 YES
13 77 YES
XX NC NO
20 NC YES
19 76 YES
10 75 YES
17 91 YES
88 88 NO
26 80 YES
95 95 YES
2 77 NO
48 84 YES
40 78 NO
78 78 YES
3 77 YES
23 NC NO
77 77 YES
90 90 NO
84 84 YES
HESPER!IDAE PYRRHOPYGINAE
242 243
Page 19
U. Robhin.c, C. Lama.s, 0. Mielke, 0. Harvey and M. Casagrande Buuctjly Community at Pakitza
HESPERIIDAE: PYRGINAE
244245
857. Elbella intersecta intersecta Herrich-Schaffer.
858. E. merops Bell, 934
859. E. theseus Bell. 1934
860. E. parrahas ringo Mielke, 1995
861. E. blanda Evans, 1951
962. E. azeta azeta Hewitson, 1866
863. Elbella madeira Miellce, 1995
864. E. erna Evans, 1951
865. Prorelhella alborna Mahille, 1891
866. Parelbella ahira ahir' Hewirson, 18SS
867. Nosphisria ronara Hewitson, 1866
868. Jemadia hospita hospira Butler, 1877
869. J. hewitsonii hewirsonii Mahille, 1878
870. J. gnetus Fabricius, 1782
871. Mysoria sejanus tsp. a.
872. Croniades pieria pieria Hewirson, 1857
873. Myscelus nobilis Cramer, 1777
874. M. amystis mysus Evans, 1951
875. M. epimachia epimachia Herrich-Schaffer,
876. M. assaricus mapirica Strand, 1921
877. Passova passova styx Moschler, 1879
878. Aspirha agennria sanics l3ruce. 1908
1869
1869
27
84
77
88
76
76
89
3
88
92
37
77
2
iS
22
16
97
88
76
I
4
23
75
84
77
88
76
76
89
76
88
92
97
77
85
NC
95
96
97
88
76
NC
NC
NC
YES
YES
NO
YES
YES
YES
YES
NO
NO
NO
YES
YES
YES
YES
YES
NO
YES
YES
NO
NO
YES
YES
905. E. spina spina Evans, 1952
906. E. clavicomis clavicomis 1-!errich.Schalfer, 1869
907. Polygonus manueli manueli Bell & Comsrock, 1948
908. Aguna sp. a.
909. A. aurunce Hewitson, 1867 ?
910. A. coelus Stall, 1782 U91!. A. memphis Latreille, 1824
912. Aguna sp. n. I
913. Aguna sp. n. 2
914. Agunaclina Evans, 1952
915 Agunasp.n.3
916 Polythrix octomaculata octomaculara Sepp, 1844
917 P. minvanes Williams, 1926
918 P. auginus Hewitson, 1867 1
919. P. metallescens Mabille, 1888
920. l-leronia labriaris Butler, 1877
921. Chrysoplectrum pervivax HUbner, 1819
922. . pcmiciosus perniciosus Herrich-Schaffer, 1869
923. Codatracrus sp. n.
924. Urbanus proteus proteus Liunaeus, 1758
925. U. pronta Evans, 1952
926. U. esmeraldus Butler, 1877
927. U. esma Evans, 1952
928. li. velinus Plotz, 1880acawoios Williams, 1926; n. syn.
929. U. releus Hubner, 1821
930. U. mona Evans, 1952
93!. U. simplicius Stoll, 1790
932. U. reductus Riley, 1919
933. U. doryssus doryssus Swainson, 1831
934. U. virescens Mabille, 1877
935. U. chalco HUbner, 1823
936. Cephise cephise Herrich-Schuffer, 1869 U937. Asrraptes talus Cramer, 1777
938. A. fulgeracor fuigeraror Walch, 1775
939. A. aulus Plotz. 1881
940. A. enorrus Stall, 1782
941. A. janeira Schaus, 1902
942. A. alector hopfferi PIoct, 1881
943. A. crctarus cretatus Hayward, 1939
944. A. creteus creteus Cramer, 1780
945. Narcosius hercules Bell, 1956
946. N. narcosius narcosius Stoll, 1790
947. N. Samson Evans, 1952
948. N. parisi parisi Williams, 1927
949. N. nazaraeus Sreinhauser, 1986
950. CaIliades zeutus Moschler. 1879
95!. Aurochron neis Geyer, 1832
952. A. longipennis Plorz, 1882
953. A. zarex HUbner, 1818
954. Bungalotis erythus Cramer, 1775
955. B. asrylos Cramer, 1780
879. Phocides metrodorus metrodorus Bell, 1932
880. P. novalis Evans. 1952
88!. P. padrona Evans, 1952
882. P. pigmalion hewirsonius Mabille, 1883
883. Tarsoctenus coryrus corba Evans, 1952
884. T. praccia plutia Hewitson, 1857
885. Phanus virreus Stall, 1781
886. P. ecit000rum Austin, 1993
887. P. obscurior prestnni Miller, 1965
888. P. marshalli Kirby, 1880
889. ljdranomia kikkawai Weeks, 1906
890. Drephalys arinas Mabille, 1888
891. D. eous Hevitson, 1867
892. D. hypargus Mahille. 189!
893. Drephalys sp. n.
894. Augiades crinisus Cramer, 1780
895, Hyalothyrus leucomelis Geyer, 1832
896. H. neleus neleus Linnaeus, 1758
897. Phareas coeleste Wesrwoocl, 1852
898. Entheus eumelus ninyas Druce, 1912
899. Entheus sp., gentius group
900. Entheus sp., priasstis gossip
90!. Cahirus procas junta Evans, 1952
902. Proteides mercurius usercurius Ribriciris, 1787
903. Epargyreus socus sinus Evans, 1952
904. E. exadeus exadeus Cramer, 1779
25
90
91
3
92
81
3
3
75
37
80
88
2
34
80
2
42
4
5!
16
8
I
18
16
5
78
NC
90
91
86
92
8!
75
NC
75
81
80
88
NC
84
80
76
75
93
NC
90
79
79
76
87
78
78
YES
YES
YES
YES
YES
YES
YES
YES
Nc
YES
YES
NO
NO
YES
NO
YES
YES
NO
YES
YES
YES
YES
NO
YES
YES
YES
89 89 NO
iS 78 YES
2 76 YES
4 83 YES
10 84 YES
IS 97 YES
5 NC NO
76 76 NO
84 84 NO
92 92 NO
8! 8! NO
56 95 YES
75 75 YES
5 75 YES
92 92 YES
76 76 YES
21 9! YES
90 90 YES
SR NC YES
30 NC NO
44 78 YES
16 NC YES
28 87 NO
3 92 YES
20 77 YES
78 78 NO
82 82 YES
32 NC YES
I 75 YES
82 82 YES
I 95 YES
80 80 YES
80 80 YES
I 79 YES
28 83 YES
75 75 YES
46 79 YES
9 75 YES
3 79 YES
6 89 YES
96 96 NO
82 82 YES
92 92 NO
95 95 NO
26 NC NO
SI NC YES
2 81 YES
37 88 YES
I 76 YES
XX NC NO
37 NC YES
I.
Page 20
I?. kobhins, G. Lames, 0. Mid/cc, 0. Harvey and M. Ca.sagrandc Butterfly Community at Pakitza
956. Dyscophelltus nicephorus 1-lewirson. 1876
957. D. marion Evans, 1952
25
36
NC
85
YES
NO
958. 0. euribates euribates Stall, 1782 37 80 YES
959. D. porcius porcitus C & R Felder, 1862
960. 0. sebaldus Stall, 1781
14
XX
80
NC
NO
YES
961. Nascus phocus Cramer, 1777
962. N. paulliniae Sepp. 1842
963. Porphyrogenes passalus passalsis Herrich.Schiiffer, 1869
25
23
79
NC
95
79
NO
YES
NO
964. P. dcspecta despecta Butler, 870
965. Oikides atines Hewirson. 1867
74
54
NC
96
NO
YES
966. cIacnorrli ntis shenia she ml Hew son, I 877 56 76 YES
967. C. disjuncrus Bell, 1940 27 NC YES
968. Celaenorrhinus sp. similis group
969. C. syllius C & K Felder. 1862
85
8
85
91
YES
YES
970. C. jao Mabilk, 1889
971. Spathilepia clonius Cramer, 1775
972. Telensiades delalande Lacreille. 1824
11
43
I
80
NC
77
YES
NO
YES
973. T. nicomedes nicomedes Moschler. 1879 91 91 NO
974. T. epicalus Hubner, 1819 9 84 YES
975. T. penidas Hewitson, 1867 92 92 YES
976. T. anriope tosca Evans, 1953 68 87 YES
977. T. amphion misitheus Mahille. 1888
978. Pyrdaluscorhislo corbulo Stall, 1781
I
8!
79
81
YES
NO
979. Eracon clinlas Mahille, 1878 26 77 Nc
980. E. paulinus StolI, 1781 23 NC YES
98!. Spioniades Iibethra Hewitson. 1868 83 83 YES
982. Mictris crispus Herrich'SchI1fer, 1870 78 78 YES
983. Iliana purpurascens Mabille & Ilouller, 1912
984. Polyctor polycror polyctor Pritrsvitz, 1868
68
35
81
77
NC
YES
985. Nisoniades laca Steinhauser, 1989 78 78 NC
986. N. mimas Cramer, 1775 63 78 YES
987. N. ephora l-IerrichSch&iffcr, 1870 16 NC NO
988. N. evansi Steinhauser, 1989 78 78 YES
989. N. brunneata Williams & Bell, 1939 96 96 YES
990. N. macssrius Herrich-SchilIfer, 1870 8 8! YES
991. Pachyneuria I. lineatopuncrata Mob. & Boull., 1917 70 83 YES
992. P. herophile Hayward, 1940 3 78 YES
993. Pellicia klugi Williams & l3ell, 1939 14 78 YES
994. P. cosnrnacula cosrimacula Herricl's.SchiIIcr, 1870 76 76 YES
995. P. rrax Evans, 1953 95 95 YES
996. P. dimidiara dinsidiara HerricIs.Sclsf1cr, 1870 61 NC NC
997. Pellicia sp. nJ 97 97 NC
998. Morvina morvus cyclopa Evans, 1953 32 75 NC
999. M. fissilnacula ren';s Evans, 1953 2 92 YES
1000, M. buses balia Evans, 1953 34 79 NO
1001. Myrinia binoculus Mosclsler, `877 25 NC NO
1002. M. myris Msshille, 1898 36 78 NC
1003. M. santa monks Evans, 1953 82 82 NC
1004. Xispia qtlatlrata Mahille. 1889 78 78 NC
1005. Cyclosemia earina Hewirson, 1878 31 NC YES
1006. Gorgopas trochilus Hopffcr, 1874 41 78 YES
1007. BoIls nsancoi Lindsey, 1925
1008. B. cupreiceps Mabille, 1891
1009. B. morons morons Bell, 1940
1010. B. zorilla Plorz, 1886
lOll. Staphylus eblora Evans, 1953
1012.5. purumayo Bell, 1937
1013. S. lizeri uteri Hayward, 1938
1014.5, corunsba Williams & Bell, 1940
1015.5. oera Plorz, 1884
1016.5. astra Williams & Bell, 1940
1017. S. minor minor Schaus, 1902
1018. Plumbago plumbago Plorz, 1884
1019. Gorgyrhion begga pyralina Moschler, 18771020.0. beggina escalophoides Evans, 1953
1021. Ouleus juxra juxta Bell, 1934
1022.0. barinitta Plan. 1884
1023. 0. accerlens noctis Lindsey. 1925
1024. Zen tens difficilis Weeks, 1901
1025. Z. rermstigma rerrastigma Sepp, 1847
1026. Quadrus cerialis Sroll, 1782
1027. Q. conruhernalis conrubernalis Mabille, 1883
1028. Q. deyrollci porra Evans, 1953
1029. Pythonides jovianus Iabricii Kirby, 1871
1030. P. lerina Hewitson, 1868
103!. P. gmndis assecla Mabille, 1883
1032. P. herearsius herennius Geyer, 18381033. P. eminus eniinus Bell, 1934
1034. Pyrhonides maraca ssp. n.
1035. Sostrata esriva Erichson, 1848
1036. S. pusilla pusilla Oodrnan & Salvin, 1895
1037. Paches rribasciatus Lindsey. 1925
1038. P. exosa Butler, 1877
1039. Haemactis sanguinalis Wesrwood, 1852
1040. Milanion hemes ssp.
1041. H. pilumnus pilumnus Mabillc & BoWler, 19171042. Mylon ander nuder Evans, 1953
1043. H. menippus Fabricius, 1776
1044. H. pelopidas Fabricius, 1793
1045. M. jason Ehrmann, 1907
1046. Carrhenes fuscescens conia Evans, 1953
1047. C. canescens leads Butler, 1870
1048. C. sanres Bell, 1940
1049. Clito cliro Fabricius, 1787
1050. C. zelores l-lesvitson, 1873
1051. Xenophanes tryxus Stoll, 1780
1052. Antigonus nearchus Larreille, 1817
1053. A. erosus HUbner, 1812
1054. A. decens Butler. 1874
1055. Anisochoria pedaliodina pedaliodin:s Butler, 1870
1056. Aethilla echina ecluna llcwirsoi,, 18701057. Achlyodes husirus hems Ehrniann, 1909
65 NC YES
32 95 YES
32 97 YES
16 NC NO
66 NC YES
96 96 YES
8 NC YES
20 NC YES
80 80 YES
32 NC YES
29 NC NO
28 75 YES
tO 77 YES
2 NC YES
4 79 YES
92 92 NO
6 NC YES
78 78 NO
79 79 NO
6 77 YES
13 75 YES
I NC YES
I 81 YES
2 76 YES
25 91 YES
66 75 YES
72 NC NO
16 76 YES
27 76 YES
2 84 YES
4 76 YES
7! NC NO
SR NC YES
40 NC YES
6 75 YES
89 89 YES
3 77 YES
45 NC YES
3 77 YES
2 75 YES
16 NC YES
XX NC YES
77 77 YES
82 82 NO
40 77 YES
3 79 YES
22 75 YES
22 78 YES
49 NC YES
3 76 YES
14 76 YES
246 247
Page 21
R. Robbins, 3. Lamas, 0. Mielke, D. Harvey and M. Casagrande Butterfly Community at Pakitza
1058. A. rnithridates thraso HUbner, 1807
1059. Grais stigmaticus stigmaticus N4abille, 1883
1060. Anastrus sempiternus simplicior MOschkr, 1877
1061 A. rolimus rohigus Plorz. 1884
1062. A. petius perius Mosclilcr, 1877
1063. A. meliboca bactra Evans, 1955
1064. A. obscurus narva Evans, 1955
1065. Ebrietas infanda Butler, 1877
1066. F. anacreon anacreon Staudinger, 1876
1067. F. evanidus Mahille, 1898
1068. Cycloglypha thrasibulus thrasihulus Pabricius, 1793
1069. C. risias Godman & Salvin, 1896
1070. C. enega M6SChICT, 1877
1071. Helias phalaenoides phalaenoides HUbner. 1812
1072. Camptopleura therarnenes Mabille, 1877
1073. C. auxo MOschler, 1879
1074. PyTgus oileus orcus Stoll, 1780
1075. Heliopetes alana Reakirt. 1868
HESPERIIDAE: HESPERIINAE
1076. Synapte silius Latreille, 1824
1077. Lenro sp. n. 1
1078. L. ferrago Plan, 1884
1079. L. imerius PIotz, 1884
1080. Lenro sp. n. 2
1081. Anthoptus epictetus Fabricius, 1793
1082. A. insigrsis Ptprz, 1882
1083. Cornicea cornicca Plan, 1882
1084. Cantha calva Evans, 1955
1085. Vinius sagitta Mahille, 1889
1086. V. tryhana cryhana Kaye, 1914
1087. Pheraeus fastus Hayward. 1939
1088. P. maria Steinhauser, 1991
1089. Pheraeus sp. n. I
1090. Pheraeus sp. n. 2
1091. Misius misius Mahille, 189!
1092. Mob mango Ouenee, 1865
1093. M. calcarea ssp.
1094. Racra apella raza Evans, 1955
1095. Apausrus gracilis smarri Evans, 1955
1096. Callimormus radiola radiola Mabille, 1878
1097. Eurocus matildae vinda Evans, 1955
1098. E. quichua Lindsey, 1921
1099. Ludens ludens Mabille, 1891
1100. L. silvanicus Hayward, 1940, nom, rev.
1101. Methionopsis na Ptarz, 1882
1102. M. dolor Evans, 1955
1103. Artines sp.n.nr. aepirus Oeyer, 1832
1104. A. focus Evans, 1955
1105. A. trogon Evans, 1955
29 78 YES
41 NC NO
2 76 YES
76 76 YES
26 77 YES
29 NC YES
32 77 YES
2 75 YES
2 76 YES
6 75 YES
37 76 YES
8 96 YES
8 76 NO
10 NC YES
85 85 YES
2 76 YES
2 76 YES
76 76 YES
1 78 YES
25 75 YES
39 75 YES
27 81 YES
46 75 YES
81 81 YES
12 NC YES
78 78 YES
16 75 YES
6 96 YES
I 77 YES
68 NC NO
19 75 YES
IS 75 YES
9 77 YES
5 NC YES
7 75 YES
2 81 YES
32 76 NO
ID NC NO
.34 NC YES
29 87 YES
I 95 YES
6 78 NO
87 87 YES
4 77 YES
56 77 YES
23 78 YES
34 95 YES
17 75 YES
1106. Flaccilla aecas StoIl. 1781 91 91 YES1107. Mnaseas hicolor inca Bell, 1930
1108. GalIlo sp. n.
1109. Thargella caura caura Plan, 1882
1110. Venas evans I3sutler, 1877
34
78
4
8
77
78
75
85
YES
YES
YES
YESIll I. V. caerulans Mabille, 1878 52 76 YES1112. Phanes aletes Oeyer. 1832
1113. Phanes sp. n.
1114. Vidius nappa Evans, 1955
83
81
30
83
81
NC
YES
NO
NOll15.Vidiussp.n.
1116. Cymaenes hazarma Hewitson, 1877
7
17
84
82
YES
YES1117. C. cavalla Evans, 1955 II NC YES1118. C. laureolus loxa Evans, 1955 18 NC NO1119. C. uruba taheri Weeks, 1901
1120. Vehilius srictomenes sticromenes l3trrler, 1877
13
33
NC
75
YES
YES1121. V. seriatirs scriatus Mahille, 1891 55 79 NO1122. V. danius ssp. n.
1123. V. putus Bell, 1941
1124. V. madius ssp. n.
88
23
I
88
90
77
NO
YES
YESIlls. Mnasilus allubira Butler, 18771126. Mnasirheus chrysophrys Mabille, 1891
37
89
NC
89
YES
YES1127. M. geniignanii Hayward, 1940
1128. M. simplicissima Herrich-Schiiffer, 1870
25
76
NC
76
NC
NO1129. Mnasir}reus sp. n. 80 80 NO1130. Remella remus Fabricius, 1798 2 79 YES1131. Moeris submerallescens Hayward, 1940 22 78 YES1132. Parphorus storax storax Mabille, l891 52 75 YES1133. P. decora Herrich'Schiiffer, 1869 16 82 YES1134. P. prosper Evans, 1955 84 84 YES1I35. Parphonrs sp. n. I 31 83 NO1136. Parphorus sp. n. 2 3 95 NO1137. Parphorussp. n. 3 81 81 YES1138. Parphorus sp. n. 4
1139. Papias phainis Godman, l900
I
5
75
75
YES
YES1140. P. sukostulata sukostulata Herrich-Schaffer, 1870 3 NC NC1141. Propapias proximus Bell, 1934
1142. Gbalopsis nero Herrich'Schaffer, l869
I
4
75
75
YES
YES1143. Anna arita Schaus, 1902 23 75 YES1144. Morys geisa geisa Maschler, 1879 16 81 YESl145. Morys sp. n. 79 79 NO1146. Psoralis cIricrara ssp. n. 8 NC NO1147. Psorrrlis sp. n. I 16 77 YES1148. Psoralis sp. n. 2 65 76 NO1149. Tigasis fusca Hayward, 1940 SI NC NO1150. Tigasis sp. n. I 77 77 YESIlSI. Tigasis sp. n. 2 27 86 NO1152. Verrius richardi Weeks, 1906 2 79 YES1153. V. monaclra Plan, 1882 5 76 YES1154. V. phyllus phyllus Cramer, 1777 10 75 YES1155. V. marcus marcus Fabricius, 1787 6 75 YES1156. V. arrona Hewirson, 1868 20 84 YES
248 249
Page 22
1157.V. arva Evans, 1955
1158. V. fuldai Bell, 1930
I 75
75 75
65 8!
4 93
9 79
13 NC
8 79
85 85
32 79
22 78
72 95
46 76
5 75
16 83
4 79
SI 83
5! NC
13 78
80 80
96 96
23 80
42 77
1 75
57 NC
16 77
3! 84
27 76
42 87
21 89
80 80
27 89
80 80
19 NC
84 84
90 90
76 76
23 80
80 80
86 86
13 NC
4 NC
69 81
97 97
5 79
93 93
89 89
2 79
22 77
25 NC
I 77
25 80
NO
YES
YES
YES
YES
YES
YES
YES
YES
YES
YES
YES
NO
YES
YES
YES
NO
YES
YES
YES
NO
NO
YES
YES
YES
YES
YES
YES
YES
YES
YES
NO
YES
YES
NO
NO
YES
NO
YES
YES
YES
YES
YES
YES
NO
YES
YES
YES
YES
YES
YES
C. sicania orbius Godman, 1901
C. rnaroma Maschler, 1877
C. cathaea Hewirson, 1866
Perichares philetes philetes Omelin, 1791
P. lotus Butler. 1870
Orses cynisca Swainson, 1821
Alera haworrhiana Swainson, 1821
Alera sp. n
Lycas godart boisduvalii Ehrmann, 1909
L. argentea Hewitson, 1866
Satumus saturnus saturnus Fabricius, 1787
1219. S. meronidia Schaus, 1902
1220.5. rericulata meron MabjIle, 1891
1221. Phlebodes pertinax ScoT!. 1781
1222. P. campo sifax Evans, 1955
1223. P. notex Evans, 1955
1224. P. virgo Evans, 1955 ?
1225. P. torax Evans. 1955
1226. P. creocla Pk,tz, 1882
1227. P. xanthobasis Hayward, 1939
1228. Phlebodes sp. n. nfl. torax Evans, 1955
1229. Joanna boxi Evans, 1955
1230.
1231.
1232.
1233.
1234.
1235.
1236.
1237.
1238.
1239.
124c
1241.
1242. Decinea sp. n.
1243. 1. dama I-lerrich'Schiilfer, 1869
1244. Cyclosma altama Schaus. 1902
1245. Ortlios orthos orthos Godman, 1900
1246.0. trinka Evans, 1955
1247.0. potesta Bell, 1941 ?
1248. Orthos sp. n.
1249. Hylephila phyleus phyleus Drury, 1773
1250. Pompeius posnpeius Lacreilte, 1824
1251. Quasimellana angra Evans, 1955
1252. Quasimellana pandora Hayward, 1940
1253. I-lansa devergens devergens Draudc, 1923
1254. H. hyboma Plan, 1886
1255. Metron leucogascer ambrosci Weeks, 1906
1256. M. schroctkyi hypochlorn Draudt. 1923
1257. Pnpcrrius propertius Fabricius, 1793
1258. Phemiades pohli ddra Evans, 1955
14 80 YES
30 79 NO
57 93 YES
20 76 YES
25 86 YES
27 NC YES
23 76 NO
1 79 YES
26 75 YES
80 80 YES
36 75 YES
I 78 YES
8 NC YES
6 76 YES
13 79 YES
3 NC YES
80 80 YES
24 NC YES
6 78 NO
26 NC NO
56 NC NO
3 85 YES
16 75 YES
29 NC YES
3 NC YES
2 79 YES
80 80 NO
89 89 NO
31 NC YES
83 83 YES
2 82 YES
6 8l YES
19 75 YES
95 95 YES
2 NC NO
28 77 NC
30 77 NO
2 77 YES
21 79 YES
95 95 YES
96 96 NO
16 NC YES
48 NC YES
76 76 NO
50 90 YES
I 78 YES
27 92 NO
19 90 YES
19 76 NO
47 76 YES
91 91 YES
I?. Robbins, C. Lamas, 0. Mic!ke, 0. Harvey and M. Casagrande Butterfly Community at Pakitza
1208.
l20
1210.
1211.
1212.
l213.
1214.
1215.
1216.
1217.
1218.
1159. Paracarysrus hypargyra Herrich'Schalfer. l869
1160. P. rnenesrries rona Hewitson, 1866
lot. Turesis complanula Herrich.SchaIIer, 1869, nom. rev.
1162. T. basta Evans, 1955
1163. Thoon canra Evans, 1955
1164. T. modius Mabille, 1889'
1165. T. duhia Bell, 1932
1166. T. taxes Godman, !900
1167. T. ponka Evans, 1955
1168. T. ranka Evans, 1955
1169. Thoon sp. n. I
1170. l'hoon sp. n. 2 nr. yesEs Evans, 1955
1171. Justinia phacrusa phaerusa Hewirson, 1866
1172. J. justinianus dappa Evans, 1955
I173.J. maculata Bell, 1930
1174. Eutychide complana Herrich-Schiiffer, 1869
1175. E. subcordara sukordaca Herrich-Sch6fer, 1869
1176. Onophas columbaria flossires Butler, 1874
1177. Onophas sp. n.
1178. Sryriodes quadrinorata Mabille. 1889
1179.5. badius Bell, 1930
1180. S. quaka Evans, 1955
118!. Sryriodes sp. n.
1182. Enosis pruinosa pruinosa Plötz. 1882
1183. E. iccitss Evans, 1955
1184. E. blnrra Evans, 1955
1185. E. immacrilara denson Evans, 1955
1186. Verrica verricalis ssp. n.
1187. Ehusus ebusus ebusus Cramer. 1780
1188. Evansiella cordela Plan, 1882
1189. Talides sinois sinois ElUbner, 1819
1190. Tromba tromba Evans, 1955
1191. Nycrus crinirus Mabille, 1891
1192. Carysrus periphas periphas Mabille, 1891
1193. Tisias quadrara quadrata Herrich-SchaIfer, 1869
1194. T. rinds, Evans, 1955
1195. T. lesucur canna Evans, 1955
1196. Macros moeros Msschler, 1877
l197. ohalus virbius virbius Cramer, 1777
1198. C. calvina Hewirson, 1866
1199. Duhiella liscella fiscella Hewitson, 1877
1200. I. duhius Stall, 1781
1201. Carysrina lysireles Mabille, 189!
1202. Tellona variegata Hewitson, 1870
1203. D'amas clavus HerrichScbaffer, 1869
1204. Orphe vatinius Godman, 1901
1205. C. gerasa Hewirson. 1867
1206. Carysroides basoches Larreille, 1824
1207. C. noseda Hewitson, 1866
I
Quinn, cannae HerrichSchiiffer, 1869
ynea iquica Bell, 1941
C. corisana Moschler, 1883
C. popla Evans, 1955
C. megalops Godman, 1900
C. robba robba Evans, 1955
C. bisrrigula HerrichSchaffer, 1869
C. dilura Herrich-Schtlifer, 1869
Penicula bryanri Weeks, 1906
P. advena advena Draudt, 1923
P. crista Evans, 1955
Decinea decinea derisor Mabille, l891
250 25!
Page 23
N. Rohbins, C. Lanias, 0. Mielke, D. Harvey and M. Casagrande
1259. P. niilvius milor Evans, 1955
1260. P. pseudophincus de Jong, 1983
1261. Calpodes ethlius StoIl, 1782
1262. Panoquina lucas Fabricius, 1793
sylvicola Herrich-SchMIer, 1865; syn. n., comb. nov.
1263. P. fusina fusina Hewitson, 1868
1264. P. evadnes StoIl, 1781
1265. Panoquina sp. n.
1266. Zenis jebus ssp. n.
1267. Tirynthia conflua l-IerrichSchaffer, 1869
1268. Thespieus dalman Latreille. 1814
1269. Lindra simulius Deuce, 1876
1270. L. vsinewrighti Mielke, 1978
L. boliviana Mielke, 1993
Oxynthes corusca Herrich-Schffer, 1869
Niconiades xanthaphes Hubner, 1821
N. linga Evans, 1955
1275. N. nabona Evans, 1955
1276. N. centralis Mielke, 1967
1277. Aides duma argyrina Cowan, 1970
1278. A. brino StolI, 1781
1279. A. aegita Hewitson, 1866
Cravera laureata Draudt, 1923
Saliana triangularis Kaye, 1914
S. fusra Evans, 1955
S. fischer Latreille, 1824
S. nigel Evans, 1955
S. esperi Evans, 1955
S. longirostris Sepp, 1840
S. morsa Evans, 1955
S. salius Cramer, 1775
S. saladin culta Evans, 1955
Thracides cleanthes relmela Hewitson, 1866
T. thrasea Hewitson, 1866
1292. Neoxeniades braesia braesia Hewitson, 1867
1293. N. bajula ssp. n.
Aroma aroma Hewirson, 1867
hloeria psittacina C & R Felder, 1867
Pyrrhopygopsis socrates orasus Druce, 1876
Unidenrilied I Tigasis ?
Unidentified 2 Eprius ?
Unidentified 3 genus? nr. Psoralis
Unidentified 4 genus?
1294.
1295.
1296.
1297.
1298.
1299.
1300.
1271.
1272.
1273.
1274.
1280.
1281,
1282.
1283.
1284.
1285.
1286.
1287.
1288.
1289.
1290.
1291.
9! 91 NO
3 93 NO
10 83 YES
3 84 NO
3 76 YES
80 80 NO
XX NC NO
22 91 YES
9 75 NO
32 NC NO
89 89 NO
9! 91 NO
82 82 YES
56 84 YES
2 84 YES
3 79 YES
87 87 NO
84 84 NO
6 80 YES
91 91 YES
22 80 YES
91 91 NO
19 80 YES
92 92 YES
59 79 YES
75 75 NO
96 96 YES
75 75 YES
22 NC YES
61 77 YES
2 NC YES
76 76 YES
86 86 YES
6 79 YES
26 NC NO
10 NC YES
SR NC YES
76 76 YES
16 NC NO
26 NC NO
6 81 YES
78 78 NO
252
![Untitled-1 [] · taxonomic characters. Zoological Nomenclature, origin of code, ICZN. UNIT V : Taxonomic Records And Publications Taxonomic keys, Taxonomic characters description,](https://static.documents.pub/doc/80x56/5e60819b810ee55ab507dd7d/untitled-1-taxonomic-characters-zoological-nomenclature-origin-of-code-iczn.jpg)
