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NOAA Technical Memorandum NMFS
SEPTEMBER 1987
ICHTHYOPLANKTON AND STATION DATA FOR CALIFORNIA COOPERATIVE OCEANIC FISHERIES
INVESTIGATIONS SURVEY CRUISES IN 1958
Elaine M. Sandknop Richard L. Charter H. Geoffrey Moser
James D. Ryan
NOAA-TM-NMFS-SWFC- 86
F
U.S. DEPARTMENT OF COMMERCE National Oceanic and Atmospheric Administration National Marine Fisheries Service Southwest Fisheries Center
NOAA Technical Memorandum NMFS
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The National Oceanic and Atmospheric Administration (NOAA), organized in 1970, has evolved into an agency which establishes national policies and manages and conserves our oceanic, coastal, and atmospheric resources. An organizational element within NOAA, the Office of Fisheries is responsible for fisheries policy and the direction of the National Marine Fisheries Service (NMFS).
In addition to its formal publications, the NMFS uses the NOAA Technical Memorandum series to issue informal scientific and technical publications when complete formal review and editorial processing are not appropriate or feasible. Documents within this series, however, reflect sound professional work and may be referenced in the formal scientific and technical literature.
P
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NOAA Technical Memorandum NMFS
This TM series s used for documentation and timely communication of prelimtnary results. imterim reporb. or Special purpose inlormation. and have not received complete format review. edttorial control. or detarled editing
SEPTEMBER 1987 8 c
J' OF, ___ 4,
4 Q r ~ G t c@
ICHTHYOPLANKTON AND STATION DATA FOR CALIFORNIA COOPERATIVE OCEANIC FISHERIES
INVESTIGATIONS SURVEY CRUISES IN 1958
Elaine M. Sandknop Richard L. Charter H. Geoffrey Moser
James D. Ryan
Southwest Fisheries Center National Marine Fisheries Service
La Jolla, CA 92038
NOAA-TM-NMFS-SWFC-86
U.S. DEPARTMENT OF COMMERCE C. William Verity, Jr., Secretary National Oceanic and Atmospheric Administration Anthony J. Calio, Administrator
National Marine Fisheries Service William E. Evans, Assistant Administrator for Fisheries
CONTENTS
Page
List of Figures ............................................ iii
List of Tables ............................................. iv
Abstract ................................................... 1
Introduction ............................................... 1
Sampling Area and Pattern .................................. 2
Sampling Gear and Methods .................................. 3
Laboratory Procedures ...................................... 4
Identification ............................................. 5
Computer Entry and Editing ................................. 10
Species Summary ............................................ 11
Explanation of Tables ...................................... 11
Acknowledgments ............................................ 13
Literature Cited ........................................... 14
Figures .................................................... 17
Tables ..................................................... 31
Index ...................................................... 244
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LIST OF FIGURES
Page
Figure 1 .
Figure 2 .
Figure 3 . Figure 4 . Figure 5 . Figure 6 . Figure 7 . Figure 8 . Figure 9 . Figure 10 . Figure 11 . Figure 12 . Figure 13 . Figure 14 .
Composite arrangement of diagrammatic charts showing areas sampled on each CalCOFI cruise during 1958 .................... Station pattern for CalCOFI Cruise 5801 showing tracks for each vessel ................ Station pattern for CalCOFI Cruise 5802 ....... Station pattern for CalCOFI Cruise 5803 ....... Station pattern for CalCOFI Cruise 5804 ....... Station pattern for CalCOFI Cruise 5805 ....... Station pattern for CalCOFI Cruise 5806 ....... Station pattern for CalCOFI Cruise 5807 .......
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Station pattern for CalCOFI Cruise 5808 ....... Station pattern for CalCOFI Cruise 5809 ....... Station pattern for CalCOFI Cruise 5810 ....... Station pattern for CalCOFI Cruise 5811 ....... Station pattern for CalCOFI Cruise 5812 ....... The basic station plan for CalCOFI cruises from 1950 to the present ..............
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LIST OF TABLES
Page
Table 1. Station and plankton tow data for ........................ CalCOFI cruises in 1958 31
Table 2. Pooled occurrences of fish larvae taken ................. during CalCOFI cruises in 1958 77
Table 3. Pooled numbers of fish larvae taken ................. during CalCOFI cruises in 1958 81
Table 4. Numbers of fish larvae taken on stations occupied during CalCOFI cruises in 1958 ........ 85
Table 5. Summary of pooled occurrences of fish larvae taken on CalCOFI cruises from 1951-1960 ...................................... 239
Table 6. List of stations with multiple occu- pancies in one month during 1958 ............... 243
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ABSTRACT
This report provides ichthyoplankton and associated station and tow data from California Cooperative Oceanic Fisheries Investigations (CalCOFI) cruises conducted off California and Baja California in 1958. It is the eighth report in a series that presents these data for all biological-oceanographic CalCOFI surveys from 1951 to the present. A total of 1884 stations was occupied during 12 monthly multivessel cruises over the quarter- million square mile survey area which extends from the California-Oregon border to Cape San Lucas, Mexico and seaward to several hundred miles. The data are listed in a series of 6 tables: the background, methodology, and information necessary for interpretation and quantitative analysis of the data are presented in an accompanying text. All pertinent station and tow data, including volumes of water strained and standard haul factors are listed in the first table. Another key table lists, by station and month, standardized counts of each of the 151 larval fish categories identified from survey samples. This and previous and subsequent reports make the CalCOFI ichthyoplankton and station data available to all investigators and serve as guides to the newly developed computer data base.
INTRODUCTION
This report, the eighth of a series, provides ichthyoplankton and associated station and tow data from California Cooperative Oceanic Fisheries Investigations (CalCOFI) joint biological-oceanographic survey cruises conducted in 1958. This program was initiated in 1949, under the sponsorship of the Marine Research Committee of the State of California, to study the population fluctuations of the Pacific sardine (Sardinops sagax) and the environmental factors that may play a role in such fluctuations. CalCOFI, known as the California Cooperative Sardine Research Program from 1949 to 1953, was made up of representatives of the South Pacific Fisheries Investigations (SPFI) of the U.S. Fish and Wildlife Service [now the La Jolla Laboratory, National Marine Fisheries Service (NMFS)], the Scripps Institution of Oceanography (SIO), the California Department of Fish and Game (CDFG), the California Academy of Sciences (CAS) and the Hopkins Marine Station of Stanford University. The first three of these agencies supplied ships and personnel to conduct the sea surveys. NMFS processed the plankton samples and analyzed the ichthyoplankton from them. SI0 processed and analyzed the hydrographic samples and measurements and also analyzed invertebrate groups from the plankton samples.
The boundaries, station placement, and sampling frequency for the CalCOFI survey area were based on the results of joint biological and oceanographic cruises conducted by NMFS and SI0 during 1939-41. Those cruises were designed to collect sardine eggs and larvae and associated hydrographic data over the entire areal and seasonal spawning range of the species. On these
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survey cruises, a depth which encompassed the vertical distribution of sardine eggs and larvae. Wide-ranging joint biological and oceanographic survey cruises were resumed in 1949 with sardine as the focus: however, an increasing interest in other biological components resulted in the deepening of standard tows to 140 m in 1951. This marked the beginning of truly quantitative ichthyoplankton sampling on CalCOFI surveys.
Data resulting from CalCOFI surveys in 1958 have been published in a number of forms. Hydrographic data (Reid et al., 1965), zooplankton volumes (Thrailkill, 1961; Smith, 1971) and ichthyoplankton data for selected species (Kramer, 1971) were presented in standard formats. The latter lists counts for eggs and larvae of sardine and for larvae of northern anchovy ( E n g r a u l i s mordax) , jack mackerel (Trachurus s y m m e t r i c u s ) , Pacific mackerel (Scomber j a p o n i c u s ) , Pacific hake ( M e r 1 u c c i u s p r o d u c t u s ) , and rockfishes ( S e b a s t e s spp.) . Also, length frequencies are listed for sardine, anchovy, jack mackerel, and Pacific mackerel larvae. Distribution maps of larvae of 5 of these taxa taken on CalCOFI surveys during 1958 are presented in the CalCOFI Atlas series (Kramer and Ahlstrom, 1968; Ahlstrom, 1969; Kramer, 1970; Ahlstrom et al., 1978). Other atlases provided distribution maps of 6 mesopelagic fish larvae (Ahlstrom, 1972) and 8 flatfish taxa (Ahlstrom and Moser, 1975) taken during 1958.
A computer data base for eggs and larvae of sardine and anchovy and for larvae of hake, and the two mackerels was established in 1969. The development of a data base for other fish larvae is a complex undertaking because competency of identification has evolved steadily over the past 3 8 years. We began the task of producing a CalCOFI ichthyoplankton data base and associated data report series in 1983. All available original records for 1958 were subjected to an extensive verification and editing process to produce this report. This and previous (Ambrose et al., 1987a, b; Sandknop et al., 1987; Stevens et al., 1987a, b; Sumida et al., 1987a, b) and subsequent reports make the CalCOFI ichthyoplankton and station data available to all investigators and serve as guides to the computer data base. The data base will be modified when additional errors are discovered and when composite taxa from the earlier years are reidentified. These reports are the fundamental reference documents against which subsequent changes in the data base can be compared.
plankton tows were made to 70 m,
SAMPLING AREA AND PATTERN
In 1958, CalCOFI survey cruises were conducted at monthly intervals. A total of 1884 stations included in this data base was occupied on 12 cruises, with an average of 157 stations per cruise (range of 37-272). Coverage of the survey station pattern varied among cruises and the entire quarter-million square mile survey area was not covered on any single cruise
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(Figures 1-13; Table 1). The area off northern California (lines 40-57) was not routinely surveyed; this area was only partially covered on three cruises in April, June, and July. The area off central California (lines 60-77) was surveyed on seven cruises: January, March through July, and October. The area between Pt. Conception, California and Pt. San Juanico, Baja California (lines 80-137) was surveyed monthly. In September coverage north of line 110 was limited to two lines in the southern California Bight and in November and December there was no coverage south of line 100. The area off southern Baja California (lines 140-157) was surveyed in January, February, March, and October. Coverage extended seaward to station 145 (ca. 400-500 miles offshore) on lines 80-100 during April, but typically did not extend beyond station 90 (ca. 160-250 miles offshore). Offshore coverage was greatest betwe n January and July and generally diminished during later cruises. f
Six vessels were employed on these cruises: the Black Douglas of NMFS, and the Horizon, Orca , Paol ina T . , Spencer F . B a i r d , and the S t r a n g e r of SIO. One to four vessels participated on each cruise, with cruises in April, July, and October employing the maximum of four vessel. The Black Douglas was used on all cruises except February, November, and December. The S t r a n g e r participated in 8 cruises (January-July, October) and the Paol ina T participated in 7 cruises. The other 3 vessels were used on a total of 6 cruises (Thrailkill, 1961).
SAMPLING GEAR AND METHODS
The standard CalCOFI net sed from 1949 to 1969 had a l-m. diameter mouth opening (0.785 my area) and an overall length of about 5 m. The net was constructed of 3Oxxx gauze, a heavy duty grade of silk bolting cloth, with a mesh size of 0.55 mm after shrinkage. The last 40 cm of the cone and the cod end were constructed of 56xxx grit gauze which had a mesh size of 0.25 mm after shrinkage. On parts of 3 cruises during 1958 (5804, 5805, 5807) the standard net was replaced with one constructed of nylon. Construction of the nets was similar; however, the nylon
'CalCOFI lines (Figure 14) are arranged perpendicular to the coastline and extend from the Canadian border (line 10) to below Cape San Lucas, Baja California (line 157). Stations were established on the basis of a perpendicular to line 80 (off Pt. Conception) at a point designated as station 60. Stations were plotted seaward and shoreward from station 60 on each line. Cardinal CalCOFI lines (those ending in lrO1l) are 120 miles apart and usually bracket two ordinal lines (ending in rr311 or 11711), so that lines are 40 miles apart over most of the pattern. Cardinal stations are 40 miles apart and typically these are separated by a station number ending in I15l1 so that stations are 20 miles apart out to station 90 on most lines. Stations are placed at closer intervals near the coast and islands to accommodate these features (see Kramer et al., 1972 for further details).
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nets had mesh sizes of 0.471 mm for the net body and 0.280 mm for the end of the cone and the cod end (Smith, 1971). The net ring was fastened to a short 3-lead bridle connected to several meters of line which attached to the towing cable by a clamp. A current meter was suspended in the center of the net mouth to measure volume of water filtered (see Kramer et al., 1972, for further details).
The standard tow from 1951 through 1968 was an oblique haul to 140 m depth (to 15 m of the bottom in shallow areas) designed
filter a constant amount of water per depth interval (ca. 4g3/m of depth) over the vertical range of most ichthyoplankters. Hauls were made at a ship speed of 1.5-2.0 knots and initiated by clamping the net line to the towing cable with the 45 kg terminal weight about 10-15 m below the surface. The net was lowered to 140 m depth by paying out 200 m of wire over a 4 minute period (35 m of depth/min.). After fishing at depth for 3 0 seconds, the net was retrieved at 20 m/min. The angle of stray of the towing cable was recorded every 30 seconds and maintained at 4 5 O (+3O) by adjusting the ship speed and course. After reaching the surface, the net was washed down and the samples preserved in 5% formalin buffered with sodium borate. Flowmeter readings were made at the beginning and end of each tow. Detailed descriptions of gear and methods are given by Kramer et al. (1972) and Smith and Richardson (1977).
(14 m depth/min.).
LABORATORY PROCEDURES
Laboratory processing began with the determination of a displacement volume for each sample (methods described in Staff, SPFI, 1953 and Kramer et al., 1972). Zooplankton volumes (including ichthyoplankton) of samples collected in 1958 are listed in Thrailkill (1961) and presented graphically in Smith (1971) .
Sorting involved the removal of ichthyoplankton from the sample and identification and separation of eggs and larvae of selected species (see introduction). Usually, each sample was sorted completely; however, some of the samples were fractioned into aliquots using a Folsom plankton s p itter (McEwen et al., 1954) prior to sorting. Several criteria’ were used to determine wheth’er a sample was fractioned: samples containing an abundance of thaliacians and coelenterates and exceeding 150 ml in total plankton volume were fractioned (to 50%, 25%, 12.5%, or 6.25%) to approximate a reduced volume of 50 ml for sorting; samples with an excessive quantity of fish eggs and/or larvae were occasionally fractioned to expedite the sorting process in order to meet scheduled deadlines. If the identified fraction of an aliquot yielded rare or interesting species of fish larvae, the remaining fraction was frequently sorted and identified with the
2Personal communication, James R. Thrailkill, National Marine Fisheries Service, Southwest Fisheries Center, La Jolla, CA.
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intent of finding additional specimens. Aliquot percentages for fractioned samples from 1958 are listed in Table 1 under the 'IPercent Sorted" column.
A "standard haul factor" (SHF) was calculated for each tow to make them comparable and allow estimations of areal abundance. This factor ad'usts the number of eggs or larvae in a haul to the number in 10 m3 of water strained per meter of depth fished. If
2 the vertical distribution of the species has been encompassed then the adjusted value is equivalent to the number under 10 m of sea surface. The S H F is calculated for each haul by the
10 D V
formula :
S H F =
where D =
v =
v = where R =
a =
P =
Tow depth, volume of listed in Table 1
depth of haul = cosine of the average angle of stray of the towing cable multiplied by cable length (m)
total volume of water (m3) strained during the haul
R 0 a . p
total number of revolutions of the current meter during the haul
area (m2) of the mouth of the net
length of column of water (m) needed to produce one revolution of the current meter.
water strained, and standard haul factor are for each tow taken during 1958. Detailed
descriptions of factors involved in calculating these values are presented in Ahlstrom (1948), Kramer et al. (1972), and Smith and Richardson (1977).
IDENTIFICATION
Identification of ichthyoplankton species beyond those separated during the sorting process was carried out by a separate group of specialists. Ontogenetic stages of fishes are inherently difficult to identify and this is further complicated by the large number and diversity of species which contribute to the ichthyoplankton of the California Current region. Most identifications were accomplished by establishing ontogenetic series on the basis of morphology, meristics, and pigmentation and then identifying these series by relating them to known metamorphic, juvenile, or adult stages with overlapping features (Powles and Markle, 1984). A total of 149 taxa was identified for 1958, with 79 taken to species, 31to genus, 34 to family,
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and 5 to order. Some of the developmental series recognized originally could not be assigned scientific names, particularly in the Bathylagidae, Myctophidae, and Pleuronectiformes. These were given descriptive names, which later were changed to scientific names as they became known.
The task of producing a reliable and equitable ichthyoplankton data base required extensive procedures to verify, correct, and edit the original identifications. The primary data source was the original identification sheets (see Kramer et al., 1972, for examples); however, a critical resource used in all phases of this process was the CalCOFI ichthyoplankton collection in which the samples are archived. Throughout the course of CalCOFI ichthyoplankton studies, samples have been identified to the lowest taxon possible. In reviewing these identifications for the data base, our approach has been conservative and we have preserved those identifications and counts which we could confirm, while correcting as many of the errors as possible. During the coding of the identification sheets, the I1descriptive types" were assigned scientific names and reexamined, if necessary. After computer entry, taxonomic errors and inconsistencies in the data base were corrected and the most obvious identification errors were corrected. Our current knowledge of ichthyoplankton techniques coupled with a precise understanding of the development of identification competency in the program over the years allowed us to critically judge the historical records. Identifications were changed to different taxa, lumped to a higher taxonomic category, or given a more precise taxonomic name. In many cases, identifications of a taxon were inconsistent among cruises in a year, because of varying competency of identifiers. These records were made equitable by lumping to the higher taxonomic category to avoid biases that could result in quantitative misinterpretations.
Next, statistical, seasonal, and geographic outliers were identified, employing a series of graphic summaries and listings. Examination of geographic outliers proved to be especially effective because of our accumulated knowledge of species distributions. In the course of examining samples for these outliers, other identification errors were discovered and eventually all taxa were scrutinized to some extent. Lastly, certain taxa were reexamined in all samples for the entire CalCOFI time series. These taxa were selected because of their commercial, ecological, phylogenetic, or zoogeographic importance or because taxonomic confusion was at the ordinal level. The following is a list of the taxa for 1958 which received special attention, with explanations and caveats intended to aid in quantitative interpretations:
Anguilliformes - tentative and sporadic identifications to family or lower taxon lumped to order.
Sardinops sagax - all specimens south of line 120 checked for misidentification of Opisthonema spp.
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Engraulidae - includes nearshore taxa (mostly Anchoa spp.) large enough to separate from Engraul i s mordax . Some nearshore samples of small E. mordax may contain other anchovy genera, but could not be differentiated.
Nansenia spp. - all specimens checked and identified as N. candida or N. c r a s s a ; all specimens of these species near their range boundaries checked.
Bathylagus spp. - all specimens checked; residuals are small, poorly preserved specimens.
Sternoptychidae - tentative and sporadic identifications of
Bathophi lus spp. - all specimens checked. hatchetfishes to genus were lumped to family.
Tactostoma macropus - all specimens checked. Paralepididae - tentative and sporadic identifications to genus
lumped to family.
Scopelarchidae - tentative and sporadic identifications to genus lumped to family.
Lampanyctus spp. - tentative and sporadic identifications to species (mostly descriptive types) lumped to genus; identification of L . r e g a l i s and L. ritteri begun in 1954.
Lampanyctus r e g a l i s - underrepresented because of inability to differentiate small larvae (<5 mm) from those of other species of the genus; counts may include other species of the genus because of difficulty in identifying larvae of this large and complex genus.
Lampanyctus ritteri - comment for L. r e g a l i s applies to this species.
Diogenichthys a t l a n t i c u s - all specimens at margins of range checked.
Diogenichthys l a t e r n a t u s - all specimens at margins of range checked.
Elec trona rissoi - recognition of this species was inconsistent and others may be included in Protomyctophum crockeri or Myctophidae.
Hygophum spp. - all specimens reidentified to species; residuals are small, poorly preserved specimens.
Myctophum aurolaternatum - all specimens checked; originally identified as llAstronesthidaetl.
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Protomyctophum crockeri - some samples on northern lines may contain P . thompsoni , which was not identified at the time.
Symbolophorus c a l i f o r n i e n s i s - all specimens south of line 120 checked for confusion with Hygophum spp., stemming from descriptive names.
Bregmaceros spp. - all gadiform types (see Index), except
Ophidiiformes - this category did not exist originally and M e r l ucc i u s product u s and Macrouridae , reexamined.
ophidiiform larvae were included in Brosmophysis marginata , Carapidae, llOtophidiumll , llZoarcidaevl , and llblennyll ; identi- fications of B. marginata and Carapidae proved to be mostly correct and vlZoarcidaell to be a yet unidentified ophidiiform species; all llOtophidiumlf and llblennyll were reexamined and the former included Ophidion sc r ippsae , Chi lara t a y l o r i and other ophidiiform taxa (moved to order) ; I1blennyv1 contained 0. s c r i p p s a e , C. t a y l o r i , and other ophidiiform taxa in addition to true blennioids.
Ceratioidei - identifications of this group were inconsistent and additional specimens may be in the unidentified fish larva category.
Hemiramphidae - specimen checked. Trachipteridae - tentative and sporadic identifications to genus
were lumped to family.
Melamphaes spp. - all identifications ascribed to Melamphaidae were reexamined and assigned to genus (Melamphaes, Poromi t ra ) or species (Scope1 ogadus b i s p i n o s u s ) : larvae originally identified as Melamphaes spp. were not reexamined and this category may contain other melamphaid genera.
Cottidae - some samples may include specimens of Scorpaenichthys marmorat u s , hexagrammids (e. g . , Oxy1 ebi u s p i ct u s , Z a n i 01 e p i s spp. ) , and some blennioids (e. g. , Hypsobl enni u s spp. ) .
Oxy leb ius p i c t u s - all specimens checked. Z a n i o l e p i s spp. - all specimens checked. S e b a s t e s spp. - in addition to other scorpaenid genera,
category includes Pr iono tus spp., serranids, scombrids, and other spiny-headed shorefishes, particularly in samples south of line 120.
Sebas to lobus spp. - this category is underrepresented and additional specimens
Hypsoblennius spp. - some
- - -
may be in S e b a s t e s spp.
specimens remain in Cottidae.
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Clinidae - some specimens remain in Cottidae or unidentified fish larva category.
Labridae - tentative and sporadic identifications to genus were lumped to family.
Pomacentridae - specimens checked; now includes species other
C h r o m i s p u n c t i p i n n i s - records south of about line 120 may than C h r o m i s p u n c t i p i n n i s , primariy in the south.
include other pomacentrid taxa.
Apogonidae - all specimens checked. Carangidae - all specimens checked; tentative and sporadic
identifications to genus or species s y m m e t r i c u s and Seri ol a 1 a1 and1 )
S e r i o l a l a l a n d i - all specimens checked.
(except Trachurus were lumped to family .
Gerreidae - tentative and sporadic identifications to genus were lumped to family.
Haemulidae - tentative and sporadic identifications to genus were lumped to family.
G i r e l l a n i g r i c a n s - all specimens checked. Medi a1 una c a l i f orni ensi s - all specimens checked. C a u l o l a t i l u s p r i n c e p s - all specimens checked. Sciaenidae - tentative and sporadic identifications to genus
lumped to family.
Gempylidae - all specimens checked. Scombridae - all larvae identified to this family or constituent
taxa (except Scomber j a p o n i c u s ) were reexamined and reas- signed; underrepresentation or absence of these taxa may be attrip uni- dentn,,&d fish larva category.
cl. to misidentification or they may be in the
Nomeidae - tentative Identifications to genus lumped to family. Pleuronectiformes - all
(originally called ttflatfishll) were examined and reidenti- fied; residuals are small, poorly preserved specimens.
assigned to various paralichthyid genera or
available specimens of this category
Bothidae - all specimens examined and reassigned; most were to Bothus spp.
C i t h a r i c h t h y s spp. - all larvae identified to genus or to a species of the genus from 1954 to 1960 were rechecked and
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identified to species; residuals are small, poorly preserved specimens or those with variable taxonomic characters.
C i t h a r i chthys spp. ; present records result from complete reidentification of C i t h a r i c h t h y s spp.
Etropus spp. - larvae of this taxon were originally lumped with
Hippoglossina spp. - all specimens of this genus (originally called 'Ipigrnented bothidfl) were examined and assigned to H. s tomata .
P a r a l i c h t h y s spp. - all specimens of this genus were examined and most were assigned to P . c a l i f o r n i c u s or X y s t r e u r y s 1 i ol ep i s .
headed bothid") . S y a c i urn oval e - all specimens examined (originally called "spiny-
X y s t r e u r y s l i o l e p i s - originally misidentified as P a r a l i c h t h y s c a l i f o r n i c u s ; all specimens reidentified.
Glyptocephal u s z a c h i r u s - all specimens examined. Hypsopset ta g u t t u l a t a - specimens were originally identified as
P1 euroni ch thys spp . Microstomus p a c i f i c u s - all specimens examined. Pleuron ich thys spp. - all larvae of this genus and constituent
species were examined and assigned to species; residuals are small, poorly preserved specimens.
COMPUTER ENTRY AND EDITING
Each taxon on the original identification sheets was given a 3-digit code based on the list of codes in Haight et al. (1979). Taxon codes and counts from these sheets were keypunched by cruise and station, along with pertinent station and tow data and entered into the VAX 11/780 computer at the University of California, San Diego Computing Center. After entries were completed f o r an entire year, print-out listings of taxa and counts on each station were compared with the original sheets to eliminate keypunch errors. Next, data in the file were cross-checked with data on an existing file which contained: station and tow data; numbers of eggs of sardine, anchovy, and saury (Cololabis s a i r a ) ; numbers of larvae of sardine, anchovy, hake, jack mackerel, and Pacific mackerel; total number of fish eggs; and total number of fish larvae.
Discrepancies in ichthyoplankton data in these two files were corrected by inspecting original records from the sorting laboratory, the original ichthyoplankton identification sheets, and the samples themselves. Station and tow data discrepancies between the two files were corrected by reviewing ships1 logs and
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deck tow sheets, original records from the sorting laboratory, cruise announcements, publications, header information on the ichthyoplankton identification sheets, and station plots generated for each cruise. Eventually all station and tow data were checked by comparing these sources.
The corrected ichthyoplankton data base was then examined statistically and outliers were found and checked as above. Distributional plots were then prepared for each taxon and these were checked by reviewing the data sources mentioned above and by examining archived specimens. A listing of each taxon by station (Table 4) was produced, which became the primary document for subsequent checks. Misidentifications found in geographic outlier checks and other misidentifications and data problems discovered in the course of examining archived samples resulted in several iterations of Table 4. Finally, totals in Table 4 were checked against annual summaries of incidence and abundance (Tables 2 and 3 ) . Ecological analyses of the data (Moser et al., 1987) were conducted concurrently with editing procedures and provided cross-checks that allowed correction of errors.
SPECIES SUMMARY
Larvae of northern anchovy, Engraul i s mordax, represented 44.8% of all fish larvae taken on CalCOFI cruises during 1958 and numbered three and one-half times as many as V i n c i g u e r r i a l u c e t i a , a midwater gonostomatid, the next most abundant species with 12.8% (Tables 2 and 3 ) . These species ranked 2nd and 1st in occurrences, respectively. The third most abundant species, Pacific hake, Merlucc ius produc tus , constituted 12.7% of all larvae and ranked 5th in occurrence. Larvae of S e b a s t e s spp., a composite of about 70 species, ranked 4th in abundance and occurrence. Two midwater lanternfishes, Tr ipho turus mexicanus and Stenobrachi u s 1 eucopsarus , ranked 5th and 6th in abundance and 3rd and 9th in occurrence. Larvae of Pacific sardine, Sardinops sagax , ranked 7th in abundance but only 18th in occurrence, indicating relatively large sample sizes. A deepsea smelt, Bathylagus wesethi , ranked 8th in both abundance and occurrence. Another lanternfish, Diogenichthys l a t e r n a t u s , and jack mackerel, Trachurus symmetricus, ranked 9th and 10th in abundance and 6th and 12th in occurrence. These 10 top-ranking taxa contributed 88.7% of all larvae taken during 1958. The remaining 11.3% was represented by 151 taxa plus the unidentified and disintegrated categories. Of the 10 taxa, 5 were midwater species, 3 were coastal pelagic species, and 2 were coastal demersal species or generic groupings.
EXPLANATION OF TABLES
Table 1 - This table lists by cruise the pertinent station tow data for 1958, the volume of water filtered standard haul factor for each tow, the percent sample sorted, and the total numbers of fish eggs
11
and and of and ~
larvae. CalCOFI cruises are designated by four digits; the first two indicate the year and the second two the month. Within each cruise the data are listed in order of increasing line and station number (southerly and seaward directions); the order of station occupancy is shown on the station charts (Figures 2-13). Stations are designated by two groups of digits; the first set indicates the line and decimal fraction and the second set indicates the station on the line. Decimal fractions were not used in 1958. Time is listed as Pacific Standard Time at the start of each tow in 24- hour designation. Methods for determining tow depth, volume of water strained, standard haul factor, and percent sorted were described in the methods section. The values for total fish eggs and larvae represent raw counts (unadjusted for percent sorted or standard haul factor). Ship codes are as follows: BD, Black Douglas; HO, Horizon, OR, Orca; PT, Paol ina T ; SB, S p e n c e r F . Baird ; ST, S t r a n g e r .
Table 2 - This table lists pooled occurrences of all larval fish taxa taken during 1958 in ranked order.
Table 3 - This table lists pooled counts of all larval fish taxa taken during 1958 in ranked order. Numbers are adjusted for percent sorted and standard haul factors.
Table 4 - This table gives numbers of fish larvae for each taxon, listed by station and calendar month in which the tow was taken. Counts are adjusted for percent of sample sorted and standard haul factor. Average values are given for stations occupied more than once during a month. See Table 1 for station and tow data and Table 6 for listing of stations with multiple occupancies during a month. Multiple occupancies occurred when a station was occupied more than once in the same calendar month; in some cases, multiple occupancies resulted from separate cruises. The orders are listed in "phylogeneticf1 sequence modified from Nelson (1984). Subtaxa within each order are listed alphabetically. Page numbers for each taxon are given in the index at the end of the report.
Table 5 - This table is a summary of pooled occurrences of all larval fish taxa taken on CalCOFI surveys from 1951 to 1960. Taxa are listed in the same order as in Table 4.
Table 6 - List of stations with multiple occupancies in'one month during 1958.
12
ACKNOWLEDGMENTS
Lois Hunter was primarily responsible for the original identifications of larvae from CalCOFI cruises in 1958 (5801- 5809). David Kramer identified larvae from cruises 5810, 5811, 5812, and also a portion of 5804. Ronald Whyte coded each larval fish taxon or type and Rita Ford entered them into the computer. Debby Snow efficiently assisted in all aspects of data editing and retrieval. Cindy Meyer and Larry Zins provided programming assistance. Dorothy Roll designed the CalCOFI data acquisition system and provided data processing support. Ken Raymond, Roy Allen, the report. Lorraine Prescott and Diane Forsythe prepared the manuscript for printing. Paul Smith determined statistical outliers, provided assistance during geographical outlier checks and offered helpful suggestions throughout the project. Izadore Barrett, Director of the Southwest Fisheries Center and Reuben Lasker, Chief, Coastal Fisheries Resources Division, SWFC, provided the support critical to the completion of the project. James Thrailkill planned CalCOFI surveys and supervised cruises, data handling, and plankton sorting from 1949 to 1986 and is largely responsible for the high quality of these operations. Without the vision and direction of Elbert Ahlstrom and Elton Sette and the dedicated efforts of the many people who collected, processed, and analyzed the samples, this data base would not exist.
and Henry Orr helped with graphics and production of
13
LITERATURE CITED
Ahlstrom, E. H. 1948. A record of p lchard eggs anc larvae collected during surveys made in 1939 to 1941; U.S. Fish Wildl. Serv. SSRF 54, 82 p.
Ahlstrom, E. H. 1969. Distributional atlas of fish larvae in the California Current region: jack mackerel , Trachurus s y m m e t r i c u s , and Pacific hake, M e r l u c c i u s p r o d u c t u s , 1951 through 1966. CalCOFI Atlas No. 1l:xi + 187 p.
Ahlstrom, E. H. 1972. Distributional atlas of fish larvae in the California Current region: six common mesopelagic fishes - V i n c i g u e r r i a l u c e t i a T r i p h o t u r u s mex icanus , S t e n o b r a c h i u s 1 eucopsarus Leurogl ossus stilbi u s , Ba thy laqus wesethi , and Bathy lagus ochotensis, 1955 through 1960. CalCOFI Atlas N o . 17: xv + 306 p.
Ahlstrom, E. H. and H. G. Moser. 1975. Distributional atlas of fish larvae in the California Current: Flatfishes, 1955 through 1960. CalCOFI Atlas No. 23: xix + 207 p.
Ahlstrom, E. H., H. G. Moser, and E. M. Sandknop. 1978. Distributional atlas of fish larvae in the California Current region: Rockfishes, S e b a s t e s spp., 1950 through 1975. CalCOFI Atlas No. 26: xxi + 178 p.
Ambrose, D. A., R. L. Charter, H. G. Moser, and C. R. Santos Methot. 1987a. Ichthyoplankton and station data for California Cooperative Oceanic Fisheries Investigations survey cruises in 1951. U.S. Dep. Commer., NOAA Tech. Memo., NMFS, SWFC, No. 79, 196 p.
Ambrose, D. A., R. L. Charter, H. G. Moser, and C. R. Santos Methot. 198733. Ichthyoplankton and station data for California Cooperative Oceanic Fisheries Investigations survey cruises in 1955. U . S . Dep. Commer., NOAA Tech. Memo., NMFS, SWFC, No. 83, 185 p.
Haight, C. A., H. G. Moser, and P. E. Smith. 1979. Data entry programs: CalCOFI. 11. Fish eggs and larvae identification sheet. National Marine Fisheries Service, Southwest Fisheries Center, La Jolla, Admin. Rept. N o . U-79-25.
Kramer, D. 1970. Distributional atlas of fish eggs and larvae in the California current region: Pacific sardine, S a r d i n o p s c a e r u l e a (Girard) , 1951 through 1966. CalCOFI Atlas No. 12:vi + 277 p.
Kramer, D. 1971. Sardine eggs and larvae and other fish larvae of the Pacific coast, 1958 and 1959. U . S . Dep. Commer., NOAA, NMFS, Data Rep. 68, 132 p.
14
Kramer, D. and E. H. Ahlstrom. 1968. Distributional atlas of fish larvae in the California Current region: Northern anchovy, Engraulis mordax (Girard) , 1951 through 1965. CalCOFI Atlas No. 9: xi + 269 p.
Kramer, D., M. Kalin, E. G. Stevens, J. R. Thrailkill, and J. R. Zweifel. 1972. Collecting and processing data on fish eggs and larvae in the California Current Region. NOAA Tech. Rep. NMFS Circ. 370, 38 p.
McEwen, G. F., M. W. Johnson, and T. R. Folsom. 1954. A statistical analysis of the performance of the Folsom Plankton Sample Splitter, based on test observations. Arch. Meteor. Geophys. Bioklim. Ser. A, 7:502-527.
Moser, H. G., P. E. Smith, and L. E. Eber. 1987. Larval fish assemblages in the California Current region during 1954- 1960, a period of dynamic environmental change. CalCOFI Rep. 28:97-127.
Nelson, J. S. 1984. Fishes of the world. John Wiley and Sons, N.Y., 523 p.
Powles, H. and D. F. Markle. 1984. Identification of larvae, p. 31-33. In: Ontogeny and systematics of fishes. H. G. Moser, W. J. Richards, D. M. Cohen, M. P. Fahay, A. W. Kendall, Jr., and S. L. Richardson (eds.). Spec. Publ. No. 1. Amer. SOC. Ichthyol. Herpetol., 760 p.
Reid, J. L., Jr., R. S, Arthur, and E. B. Bennett, (eds.). 1965. Oceanic observations of the Pacific: 1958. Univ. Calif. Press, Berkeley, 707 p.
Sandknop, E. M., R. L. Charter, H. G. Moser, and J. D. Ryan. 1987. Ichthyoplankton and station data for California Cooperative Oceanic Fisheries Investigations survey cruises in 1952. U.S. Dep. Commer., NOAA Tech. Memo., NMFS, SWFC, No. 80, 207 p.
Smith, P. E. 1971. Distributional atlas of zooplankton volume in the California Current region, 1951 through 1966. CalCOFI Atlas No. 13: xvi + 144 p.
Smith, P. E. and S. L. Richardson. 1977. Standard techniques for pelagic fish egg and larva surveys. FA0 Fish. Tech. Pap. No. 175, 100 p.
Staff , South Pacific Fishery Investigations. 1953. Zooplankton volumes of the Pacific Coast, 1952. U.S. Fish Wildl. Serv. SSRF 100, 41 p.
Stevens, E. G., R. L. Charter, H. G. Moser, and M. S. Busby. 1987a. Ichthyoplankton and station data for California Cooperative
1
Oceanic Fisheries Investigations survey cruises
15
in 1953. U.S. Dep. Commer., NOAA Tech. Memo., NMFS, SWFC, No. 81, 186 p.
Stevens, E. G., R. L. Charter, H. G. Moser, and M. S. Busby. 1987b. Ichthyoplankton and station data for California Cooperative Oceanic Fisheries Investigations survey cruises in 1956. U.S. Dep. Commer., NOAA Tech. Memo., NMFS, SWFC, No. 84, 189 p.
Sumida, B. Y., R. L. Charter, H. G. Moser, and D. L. Snow. 1987a. Ichthyoplankton and station data for California Cooperative Oceanic Fisheries Investigations survey cruises in 1954. U.S. Dep. Commer., No. 82, 207 p.
Sumida, B. Y., R. L. Charter, 1987b. Ichthyoplankton and Cooperative Oceanic Fisheries in 1957. U.S. Dep. Commer., No. 85, 225 p.
NOAA Tech. Memo., NMFS, SWFC,
H. G. Moser, and D. L. Snow. station data for California Investigations survey cruises NOAA Tech. Memo., NMFS, SWFC,
Thrailkill, J. R. 1961. Zooplankton volumes off the Pacific coast, 1958. U.S. Fish Wildl. Serv. SSRF 374, 70 p.
I
16
)-----I 5806 1
Figure 1. Composite arrangement of diagrammatic charts showing areas sampled on each CalCOFI cruise during 1958.
17
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w57 50 I I I I I I I I 1 I I I I I 125O 1200 115' I IO"
Figure 2. Station pattern for CalCOFI Cruise 5801 showing tracks for each vessel. Stations with plankton tows only are indicated by a dot: those with plankton tows and hydrographic measurements are shown by a dot and circle. Modified from charts in Reid et al. (1965) to include only those stations listed in Table 1 of this report.
1
J
1
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19
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20
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22
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30'
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Figure 8. Station pattern for CalCOFI Cruise 5807. Symbols as in Figure 2.
24
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Figure 10. Station pattern f o r CalCOFI Cruise 5809. Symbols as in Figure 2.
26
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F i g u r e 11. S t a t i o n pa t t e rn f o r CalCOFI C r u i s e 5810. Symbols as i n F i g u r e 2 .
27
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TABLE 2. Pooled occurrences of fish larvae taken during CalCOFI cruises in 1958.
Rank Taxon Occurrences
mi-
l 2 3 4 5 6 6 8 9
1 0 11 1 2 1 3 1 4 1 5 16 1 7 18 1 9 20 2 1 2 2 23 2 4 25 25 27 28 29 30 3 1 3 1 3 3 3 4 35 36 37 38 39 40 4 1 42 43 4 4 45 46 47 48
V i n c i g u e r r i a l u c e t i a Engraul i s mordax Tr ipho turus mexicanus S e b a s t e s spp. Merluccius productus Lampanyctus r i t ter i D i o g e n i c h t h y s l a t e r n a t u s Bathylagus wesethi S tenobrachius leucopsarus Pro tomyc tophum crockeri Leuroglossus s t i l b i u s Trachurus symmetricus Cyclothone spp. Disintegrated fish larva Melamphaes spp. Bathylagus ochotensis Symbofophorus c a l i f o r n i e n s i s Sardinops sagax Stomias a t r i v e n t e r Unidentified fish larva Myctophidae Ceratoscopel u s townsendi Paralepididae C i t h a r i c h t h y s s t igmaeus D i o g e n i c h t h y s a t l a n t i c u s G o n i c h t h y s t e n u i c u l u s Lampanyctus spp. C i t h a r i ch th y s spp . C i t h a r i c h t h y s f r a g i l i s Hygophum atra tum Tar le tonbeania c r e n u l a r i s Diaphus spp. Sternoptychidae Gobiidae Labr idae Scomber japon icus C i t h a r i c h t h y s xanthostigma Ici ch th y s 1 ocking t oni Sciaenidae Chauliodus macouni C i t h a r i chth y s s o r d i d u s Parophrys v e t u l u s Ophidiiformes Chiasmodontidae Myctophum n i t i d u l u m Synodus spp. Hypsoblennius spp. Lyopse t ta ex i l i s
882 7 8 5 7 6 8 665 5 4 1 416 416 377 3 6 1 360 350 328 317 258 238 237 216 1 9 3 182 1 8 1 1 7 4 1 5 9 1 4 5 1 3 6 1 2 6 1 2 6 1 2 1 1 1 8 1 0 1
9 6 9 0 90 86 8 4 82 8 1 80 7 9 7 6 7 5 69 62 6 1 59 56 53 5 1 50
77
a
TABLE 2. (cont.)
Rank
48 50 5 1 52 53 54 55 5 6 57 5 8 59 6 0 6 0 6 2 6 2 6 2 65 6 5 65 68 69 7 0 7 1 7 1 7 1 7 4 7 4 7 4 7 7 7 7 7 9 7 9 7 9 7 9 8 3 8 3 8 3 8 6 8 7 8 7 8 7 8 7 9 1 9 2 9 2 9 2 9 2 96 9 6
Taxon
Scopelarchidae Nansenia crassa Paralichthys californicus HY90Phum SPP. Etrumeus acuminatus Lampadena urophaos Symphurus spp. Ichthyococcus spp. Anguilliformes Diogenichthys spp. Microstoma microstoma Hippoglossina stomata Idiacanthus antrostomus Trachipteridae Argentina sialis Serranidae Lampanyctus regalis Diplophos taenia Cololabis saira Scopelogadus bispinosus Peprilus simillimus Prionotus spp. Trichiuridae Bathylagus pacificus Notoscopelus resplendens Cottidae Clinidae Microstomus pacificus Tetragonurus cuvieri Nansenia candida Chromis punctipinnis Ceratioidei Brosmophycis marginata Etropus spp. Ophidion scrippsae Sphyraena argentea Carangidae G1 yptocephalus zachirus Pleuronichthys spp. Myctophum aurolaternatum Coryphaena hippurus Bathylagus spp. Oxylebius pictus Aristostomias scintillans Pleuronectiformes Agonidae Bregmaceros spp. Stomiiformes Scorpaena spp.
78
Occurrences
50 49 48 47 45 4 4 40 37 36 35 34 33 33 3 1 3 1 3 1 28 28 28 27 26 25 24 24 2 4 20 20 20 1 7 1 7 1 6 1 6 1 6 1 6 1 5 1 5 1 5 1 4 1 3 1 3 1 3 1 3 1 2 11 11 11 11
9 9
TABLE 2. (cont.)
Rank
96 99 99 99 99 103 103 103 106 106 106 106 106 111 111 111 111 111 116 116 116 116 116 116 116 116 124 124 124 124 124 124 124 131 131 131 131 131 131 131 131 131 131 131 142 142 142 142 142
Taxon
Pomacentridae Bothus spp. Loweina rara Syacium ovale Thunnus albacares Seriola lalandi Macrouridae P1 euronichth ys vertical is Gempylidae Haemulidae Hygophum reinhardtii Zaniolepis spp. Scorpaenichthys marmoratus Psettichthys melanostictus Pleuronichthys coenosus Brama spp. Ger reidae Physiculus spp. Poromitra spp. Cyclopteridae Scombridae Opisthonema spp. Girella nigricans Carapidae Pleuronichthys decurrens Bathophilus spp. Hypsopsetta guttulata Apogonidae Ophiodon elongatus Auxis spp. Scopelosaurus spp. Scomberomorus spp. Pleuronichthys ritteri Hexagrammidae Sarda chiliensis Xystreurys liolepis Tactostoma macropus Syngnathus spp. Nome idae Engraulidae Sebastolobus spp. Medialuna californiensis Caulolatilus princeps Atherinidae Exocoetidae Balistidae Hemiramphidae Bathylagus miller1 Paralichthys spp.
Occurrences
9 8 8 8 8 7 7 7 6 6 6 6 6 5 5 5 5 5 4 4 4 4 4 4 4 4 3 3 3 3 3 3 3 2 2 2 2 2 2 2 2 2 2 2 1 1 1 1 1
me 79
TABLE 2. (cont.)
Rank
142 142 142 142 142
Taxon
Uranoscopidae Gobiesocidae I s o p s e t t a i s o l e p i s Fistulariidae Notol ychnus v a l d i v i a e
80
Occurrences
1 1 1 1 1
t
I
TABLE 3. Pooled numbers of fish larvae taken during CalCOFI cruises in 1958. Counts are adjusted for percent of sample sorted and standard haul factor (see text).
Rank
1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 29 30 31 32 33 34 35 36 37 38 39 40 41 42 43 44 45 46 47
Taxon Count
Engraul i s mordax V i n c i guerr i a 1 u c e t i a Merluccius productus Sebas te s spp. Tr ipho turus mexicanus S tenobrachius leucopsarus Sardinops sagax Bathylagus wesethi D i o g e n i c h t h y s l a t e r n a t u s Trachurus symmetricus Leuroglossus s t i l b i u s Lampanyctus ritteri C i t h a r i c h t h y s f r a g i l i s C y c l o t h o n e spp. Protomyctophum crockeri Sciaenidae Bathylagus ochotensis C i t h a r i c h t h y s spp . Ceratoscopelus townsendi Scomber japon icus Pr ionotus spp. Symbolophorus c a l i f o r n i e n s i s Synodus spp. Stomias a t r i v e n t e r Melamphaes spp. Disintegrated fish larva Unidentified fish larva Myctophidae Lanlpanyctus spp. C i t h a r i c h t h y s s t i gmaeus Diaphus spp. C i t h a r i c h t h y s xanthostigma G o n i c h t h y s t e n u i c u l u s Hygophum atra tum Paralepididae D i o g e n i c h t h y s a t l a n t i c u s Etrumeus acuminatus Ophidiiformes Labridae Tarletonbeania c r e n u l a r i s Icichthys lockingtoni Para l i ch thys c a l i f o r n i c u s Gobiidae Serranidae Lampadena urophaos C i t h a r i c h t h y s sord idus Sternoptychidae
81
204848 58461 58261 23645 16679 11829 11393 7011 7010 6443 4844 3129 3046 2925 1824 1772 1569 1536 1446 1341 1307 1299 1225 1206 1155 1118 1075 899 865 864 816 763 733 692 686 641 574 564 526 525 425 406 404 382 373 341 327
TABLE 3. (cont.)
Rank
48 49 49 51 52 53 54 55 56 57 58 59 60 61 62 63 64 65 66 67 68 69 70 71 72 73 74 75 76 76 78 79 80 80 82 83 84 85 86 87 88 89 90 91 91 93 94 95
Taxon
Parophrys vetulus Chauliodus macouni Symphurus spp. Chiasmodontidae Argentina sialis Lyopsetta exilis Hygophum SPP* Hypsoblennius spp. Anguilliformes Myctophum nitidulum Opisthonema spp. Myctophum aurolaternatum Nansenia crassa Diplophos taenia Bregmaceros spp. Scopelarchidae Diogenichthys spp. Idiacanthus antrostomus Ichthyococcus spp. Clinidae Ophidion scrippsae Peprilus simillimus Bath yl agus paci f i cus Notoscopelus resplendens Hippoglossina stomata Scopelogadus bispinosus Trachipteridae Microstoma microstoma Trichiuridae Lampanyctus regalis Cololabis saira Chromis punctipinnis G1 yptocephalus zachirus Carangidae Cot tidae Microstomus pacificus Nansenia candida Sphyraena argentea Pleuronectiformes Etropus spp. Tetragonurus cuvieri Haemulidae Bathylagus spp. Pomacentridae Brosmophycis marginata Ceratioidei Pleuronichthys spp. Coryphaena hippurus
82
Count
323 288 288 280 273 272 262 251 245 230 227 225 224 221 218 175 151 150 139 138 119 115 114 113 112 111 107 101 97 97 96 91 89 89 88 87 83 78 77 74 66 64 62 59 59 50 49 47
6
FI
TABLE 3. (cont.)
Rank
96 97 98 99 100 101 102 103 104 105 106 107 108 108 110 111 112 113 114 114 114 117 117 119 120 120 120 120 120 120 120 127 128 128 130 131 132 132 132 132 136 136 138 138 140 140 142 142
Taxon
Seriola lalandi Psettichthys melanostictus Oxylebius pictus Thunnus albacares Stomiiformes Aristostomias scintillans Scorpaena spp. Agonidae Gerreidae Syacium ovale Scorpaenichthys marmoratus Bothus spp. P1 e uroni ch t h ys vert i ca 1 is Tactostoma macropus Scomberomorus spp. Loweina rara Macrouridae Gempylidae Scombridae Zaniolepis spp. Auxis spp. Exocoetidae Hygophum reinhardtii Cycloptesidae Physiculus spp. Poromi tra spp . Sarda chiliensis Pleuronichthys coenosus Bathophilus spp. Apogonidae Brama spp. Pleuronichthys decurrens Carapidae Engraulidae Hexagrammidae P1 euroni ch th ys ri t teri Nome i dae Girella nigricans Hypsopsetta guttulata Ophiodon elongatus Uranoscopidae Scopelosaurus spp. Medialuna californiensis Caulolatilus princeps Sebastolobus spp. Syngnathus spp. Xystreurys liolepis Gobiesocidae
83
Count
43 41 40 39 38 36 34 32 30 29 28 25 24 24 23 22 21 19 18 18 18 16 16 15 14 14 14 14 14 14 14 13 12 12 11 10 9
c 9 9 9 8 8 7 7 6 6 5 5
TABLE 3. (cont.)
Rank
142 145 145 145 145 149 149 149
Taxon
Paralichthys spp. Atherinidae Fistulariidae Balistidae Bathylagus milleri Notolychnus valdiviae Isopsetta isolepis Hemiramphidae
Total
84
Count
457093
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TABLE 6. List of stations with multiple occupancies in one month during 1958. Stations were occupied twice in one month except those indicated by an asterisk, which were occu- pied three times.
Station Month
73.0 55.0 90.0 80.0 90.0 90.0 97.0 30.0 97.0 32.0 97.0 40.0 97.0 50.0 90.0 28.0 * 90.0 30.0 * 90.0 37.0 * 90.0 45.0 * 90.0 50.0 * 90.0 55.0 * 90.0 60.0 * 90.0 65.0 90.0 70.0 93.0 27.0 * 93.0 30.0 * 93.0 35.0 * 93.0 40.0 * 93.0 45.0 * 93.0 50.0 * 93.0 55.0 * 93.0 60.0 * 93.0 65.0 93.0 70.0
1 2 2 2 2 2 2 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9 9
243
INDEX
This index lists taxa included in Table 4 with their page numbers.
Page
85 Anguilliformes ............................................ Clupeiformes
Clupeidae Etrumeus acuminatus ............................. Upisthonema spp. ................................ Sard inops sagax ................................. E n g r a u l i s mordax ................................ Engraulidae ..........................................
salmoniformes Argentinidae
A r g e n t i n a s i a l i s ................................ 93 Nicrostoma microstoma 94 Nansenia candida ................................ 95 Nansenia c r a s s a ................................. 95
Bathy lagus spp .................................. 96 Bathy lagus rnilleri .............................. 96 Bathy lagus ochotensis ........................... 96 B a t h y l agus p a c i f i c u s ............................ 99 Bathy lagus wesethi .............................. 100 Leurog lossus s t i l b i u s ........................... 103
Stomiiformes .............................................. 107
...........................
Bathylagidae
Gonostomatidae
85 86 86 88 88
Cyclothone spp .................................. 107 Diplophos t a e n i a ................................ 111 I c h t h y o c o c c u s spp. .............................. 111 v i n c i g u e r r i a 1 u c e t i a ............................ 112
Sternoptychidae ...................................... 118 Stomiatoidea Chauliodontidae
Idiacanthidae
Malacosteidae
Melanostomiidae
Chaul iodus macouni .............................. 12 0
I d i a c a n t h u s antros tomus ......................... 121
A r i stostomi a s scintill ans ....................... 12 2
B a t h o p h i l u s spp ................................. 122 T a c t o s t o m a macropus ............................. 122
S t o m i a s atriventer .............................. 123 Stomiidae
Myctophiformes Alepisauroidei
Chloropthalmoidei Paralepididae ........................................ 125
Scope losaurus spp. .............................. 128 Notosudidae
244
Scopelarchidae ....................................... Myctophidae .......................................... Lampanyctinae
Ceratoscopelus townsendi ........................ Diaphus spp ..................................... Lampadena urophaos .............................. Lampanyctus spp ................................. Lampanyctus r e g a l i s ............................. Lampanyctus ritteri ............................. Notoscopelus resp lendens ........................ Stenobrachius leucopsarus ....................... Tr ipho turus mexicanus ........................... Diogenichthys spp ...............................
Myctophoidei
Notolychnus v a l d i v i a e ...........................
Myctophinae
Diogenichthys a t l a n t i c u s ........................ D i ogeni chth ys 1 a t e rna t u s Gonichthys t e n u i c u l u s ...........................
Hygoph urn rei nhard ti i
Myct oph um ni ti d ul um Symbolophorus cal i forniensis .................... Tar1 etonbeania crenul a r i s .......................
........................ Hygophum spp .................................... Hygophum atratum ................................ Loweina rara .................................... Myctophum aurolaternatum ........................ Protomyctophum crockeri .........................
............................
.............................
Synodontoidei Synodontidae
Bregmacerotidae Bregmaceros spp .................................
Merlucciidae Merlucc ius productus ............................
Moridae Phys icu lus spp ..................................
Macrouridae .......................................... Ophidiiformes .............................................
Bythitidae
Carapidae ............................................ Ophidiidae
Ophidion sc r ippsae .............................. Lophiiformes Ceratioidei ............................................... Gobiesociformes
Gobiesocidae ......................................... Beloniformes
Exocoetidae .......................................... Hemiramphidae ........................................
Synodus spp ..................................... Gadiformes
Brosmophysis marginata ..........................
Page
128
129
131 134 136 137 139 139 144 144 145 148
153 154 156 159 161 162 164 164 164 164 166 170 172
174
175
175
180 180 180
181 182
182
182
183
183 . 183
245
Page
Scomberesocidae C o l o l a b i s s a i r a ................................. 183
Atherinidae .......................................... 184
Trachipteridae ....................................... 184
Atheriniformes
Lampri f ormes
Beryciformes Melamphaidae
Nelamphaes spp .................................. P o r o m i t r a spp ................................... Scope1 ogadus b i s p i n o s u s .........................
Fistulariidae ........................................ Syngnathidae
Syngnathus spp ..................................
Agonidae ............................................. Cottidae .............................................
S c o r p a e n i c h t h y s marmoratus ...................... Cyclopteridae ........................................ Hexagrammidae ........................................
Ophiodon e l o n g a t u s .............................. O x y l e b i u s p i c t u s ................................
Syngnathiformes
Scorpaeniformes Cottoidei
Z a n i o l e p i s spp .................................. Scorpaenoidei
Scorpaenidae Scorpaena spp ................................... S e b a s t e s spp .................................... S e b a s t o l o b u s spp ................................
Triglidae P r i o n o t u s spp ...................................
Perciformes Blennioidei
Blenniidae
Clinidae ............................................. Gobiidae ............................................. Labridae ............................................. Pomacentridae ........................................
C h r o m i s p u n c t i p i n n i s ............................ Apogonidae ............................................ Bramidae
Brama spp ....................................... Carangidae ...........................................
S e r i o l a l a l a n d i ................................. Trachurus symmetr icus ...........................
Hypsoblennius spp ............................... Gobioidei
Labroidei
Percoidei
Coryphaenidae Coryphaena h i p p u r u s .............................
Gerreidae ............................................
184 188 188
189
189
189 189 190 190 190 190 190 191
191 191 196
196
196 197
198
199 201 201
201
201 202 202 202
206 206
246
Page
Haemulidae ........................................... 2 0 7 Kyphosidae
G i re1 1 a ni g r i c a n s ............................... 2 0 7 Medi a1 una c a l i f orni ensi s ........................ 2 0 7
Malacanthidae C a u l o l a t i l u s p r i n c e p s ........................... 2 0 7
Sciaenidae ........................................... 2 0 7 Serranidae ........................................... 2 0 8
Gempylidae ........................................... 2 0 9 Scombridae ........................................... 2 0 9
A u x i s spp ....................................... 2 1 0 Sarda chiliensis ................................ 2 1 0 Scomber j a p o n i c u s ............................... 2 1 0 Scomberomorus spp ............................... 2 1 1 Thunnus a l b a c a r e s ............................... 2 1 1
Trichiuridae .......................................... 2 1 2
Scombroidei
Sphyraenoidei
Stromateoidei
Sphyraenidae
Centrolophidae Icichthys lockingtoni ........................... 2 1 3
Nomeidae ............................................. 2 1 4 Stromateidae
P e p r i l u s s i m i l l i m u s ............................. 2 1 4 Tetragonuridae
Sphyraena argen tea .............................. 2 1 2
Te t ragonurus c u v i e r i ............................ 2 1 5
Chiasmodontidae ...................................... 2 1 5 Uranoscopidae ........................................ 2 1 6
Pleuronectiformes ......................................... 2 1 7 Pleuronectoidei
Trachinoidei
Bothidae
Paralichthyidae Bothus spp ...................................... 2 1 7
C i t h a r i c h t h y s spp ............................... 2 1 7 C i t h a r i c h t h y s f r a g i l i s .......................... 2 1 9 C i t h a ri ch t h y s sordi d u s .......................... 2 2 0 C i t h a r i c h t h y s s t i gmaeus ......................... 2 2 1 C i t h a r i chthys xanthosti gma ...................... 2 2 3 E t r o p u s spp ..................................... 2 2 4 Hippog loss ina s tomata ........................... 2 2 4 P a r a l i c h t h y s spp ................................ 2 2 5 Para1 i ch th ys c a l i f orni cus ....................... 2 2 5 Syacium o v a l e ................................... 2 2 6 X y s t r e u r y s l i o l e p i s ............................. 2 2 6
Pleuronectidae G l y p t o c e p h a l u s z a c h i r u s ......................... 2 2 6 Hypsopse t ta g u t t u l a t a ........................... 2 2 7 I s o p s e t t a i s o l e p i s .............................. 2 2 7 L y o p s e t t a e x i l i s ................................ 2 2 7 Microstomus p a c i f i c u s ........................... 2 2 8
2 4 7
Page
Parophrys vetulus ............................... P1 euroni ch th y s spp . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . P1 euronichthys coenosus . . . . . . . . . . . . . . . . . . . . . . . . . P1 euronichthys decurrens . . . . . . . . . . . . . . . . . . . . . . . . Pleuronichthys r i t t e r i . . . . . . . . . . . . . . . . . . . . . . . . . . Pleuronichthys ver t ica l i s . . . . . . . . . . . . . . . . . . . . . . . Psettichthys melanostictus . . . . . . . . . . . . . . . . . . . . . .
Soleoidei Cynoglossidae
Synphurus spp ................................... Tetraodontiformes
Balistidae ........................................... Disintegrated fish larva .................................. Unidentified fish larva ...................................
2 4 8
229 229 230 230 230 230 231
231
231 232 235
RECENT TECHNICAL MEMORANDUMS
Copies of this and other NOAA Technical Memorandums are available from the National Technical Information Service, 5285 Port Royal Road, Springfield, VA 22167. Paper copies vary in price. Microfiche copies cost $4.50. Recent issues of NOAA Technical Memorandums from the NMFS Southwest Fisheries Center are listed below:
NOAA-TM-NMFS-SWFC- 76 Report of a marine mammal survey of the Eastern Tropical Pacific aboard the research vessel David Starr Jordan, July 29-December 5,1986.
. SEXTON
77 Report of a marine mammal survey of the Eastern Tropical Pacific aboard the research vessel McArthur, July 29-December 6,1986. R.S. HOLT and A. JACKSON (August 1987)
78 Results of the Bay Area Sportfish Economic Study (BASES) C.J. THOMSON and D.D. HUPPERT
79 lchthyoplankton and station data for California Cooperative Oceanic Fisheries Investigations survey cruises in 1951. D.A. AMBROSE, R.L. CHARTER, H.G. MOSER and C.R. SANTOS METHOT (September 1987)
80 lchthyoplankton and station data for California. Cooperative Oceanic Fisheries Investigations survey cruises in 1952. E.M. SANDKNOP, R.L. CHARTER, H.G. MOSER, and J.D. RYAN (September 1987)
81 lchthyoplankton and station data for California Cooperative Oceanic Fisheries Investigations survey cruises in 1953. E.G. STEVENS, R.L. CHARTER, H.G. MOSER, and M.S. BUSBY (September 1987)
Rn (August 1987)
82
83
lchthyoplankton and station data for California Cooperative Oceanic Fisheries Investigations survey cruises in 1954. B.Y. SUMIDA, R.L. CHARTER, H.G. MOSER, and D.L. SNOW (September 1987)
lchthyoplankton and station data for California Cooperative Oceanic Fisheries Investigations survey cruises in 1955. D.A. AMBROSE, R.L. CHARTER, H.G. MOSER, and C.R. SANTOS METHOT (September 1987)
84 lchthyoplankton and station data for California Cooperative Oceanic Fisheries Investigations survey cruises in 1956. E.G. STEVENS, R.L. CHARTER, H.G. MOSER, and M.S. BUSBY (September 1987)
85 lchthyoplankton and station data for California Cooperative Oceanic Fisheries Investigations survey cruises in 1957. B.Y. SUMIDA, R.L. CHARTER, H.G. MOSER, and D.L. SNOW (September 1987)