F O L I A E N T O M O L O G I C A H U N G A R I C A R O V A R T A N I K Ö Z L E M É N Y E K

X U X 1988 p. 123-151

The Scientific Results of Hungarian Soil Zoological Expeditions in New Guinea. Coleoptera, Tenebrionidae:

Lagruni

By

O. M E R K L (Received November 1, 1987)

A b s t r a c t : A to ta l of 37 species col lected by Hungarian zoologists in Papua New Guinea is l i s t ed . Three new genera (Acu togr i a , K a i n d i l a g r i a . T o m o g r i a ) and 21 new species (Acutogr ia falcata, B o t h r i c h a r a i n e r s , B . i n t r i c a t a , B . i r i - descens, B . wau, Casnonidea ap ica l i s , C. ba loghi , C. caudata, C. demet r ida , C. dobodura. C. f lavipes , C. h y s t r i x , C. l o k s a i , C. pallens, C. punct i thorax, C. setosa. K a i n d i l a g r i a fo rc ipa ta , L a g r i a amethyst ina , L . l i gu la t a , L . sapphi- r i n a . T o m o g r i a per la ta) are descr ibed . W i t h 49 f igures .

Hungarian zoologists have v i s i t e d New Guinea seve ra l t imes . F i r s t , S. F E N I C H E L made f i e ldwork there , but i t ended by his t r ag ic death (1893-95), w i th an emphasis on b i r d s ; he col lec ted also many insects but no l ag r i ine specimens. Fo l l owing F E N I C H E L ' s i n t e r r u p t ­ed w o r k , L . BIRÓ made extensive and meticulous co l lec t ings (1895- 1901) i n German New Guinea ("Kaiser Wi lhe lms land" ) of that t i m e . In his most valuable m a t e r i a l , some speci­mens of L a g r i i n i were also found - these have been w r i t t e n up by PIC (1956).

Between 1965 and 1980, D r . J . B A L O G H pe r fo rmed s ix expedit ions to Papua New Gui-j nea ( f o r m e r l y T e r r i t o r y of Papua and New Guinea). D r . I . LOKSA and D r . L . MÓCZÁR joint­ed h i m in one case each. The late D r . J . J . H . S Z E N T - I V A N Y (Adelaide, South A u s t r a l i a ) hac) an impor t an t share in the organiza t ion of these t r i p s and the co l lec t ing of m a t e r i a l . The Wau Ecology Ins t i tu te ( f o r m e r l y Rishop Museum F i e l d Station) was most ly the centre and s t a r t i n g place of these expedi t ions .

The present con t r ibu t ion submits the resul t s concerning the t enebr ion id t r ibe L a g r i i n i . | The m a t e r i a l compr i ses 106 specimens belonging to 37 species; of them, 21 are descr ibed hereunder as new and 3 of them represent new genera. The two O r e o g r i a species have recent­ly been descr ibed in a generic r e v i s i o n ( M E R K L 1988), whi le two fur ther species r e m a i n un-, iden t i f i ed or undescr ibed for cer ta in reasons (see L i s t of species).

Since impor t an t ma te r i a l s of va r ious col lec t ings were available to me dur ing the de­s c r i p t i o n s , the specimens of col lec t ings other than those of J . B A L O G H were also taken into! account. These are indicated by f o u r - l e t t e r abbreviat ions in the text; the name of reposi tory; i s fol lowed by the name of cu ra to r or p r iva te co l l ec to r in the subsequent l i s t :

A N I C A u s t r a l i a n Na t iona l Insect Co l l ec t i on , CSIRO, Canber ra , A C T , A u s t r a l i a . D r . J . F . L A W R E N C E . (

B M N H B r i t i s h Museum (Natura l H i s t o r y ) , London, Uni ted Kingdom. D r . R . J . W . A L D R I D G E . B P B M Bern ice Pauahi Bishop Museum, Honolulu , H a w a i ' i , U . S . A . D r . G . A . SAMUELSON.I CASC C a l i f o r n i a Academy of Sciences, Department of Entomology, San F ranc i sco , C A L ,

U . S . A . D r . D . H . K A V A N A U G H .

# -7th con t r ibu t ion to the knowledge of L a g r i i n i .

CIMCI Canadian Nat ional C o l l e c t i o n of Insect , Ottawa, O N T , Canada. D r . A . S M E T A N A . H N H M Hungarian N a t u r a l H i s t o r y Museum, Budapest, Hungary. HPCW Hornabrook ' s pr ivate c o l l e c t i o n . We l l i ng ton , New Zealand. D r . R . W . HORNABROOK

and D r . R . G . ORDISH. I F P E Ins t i tu t für Pflanzenschutzforschung, Ebe r swa lde -F inow, German Democra t i c Re­

publ ic . D r . L . D I E C K M A N N . MCZC Museum of Compara t ive Zoology, H a r v a r d U n i v e r s i t y , Cambr idge , MASS, U . S . A .

D r . A . F . N E W T O N , J r . _ . _ MHNG Muséum d ' H i s t o i r e N a t u r e l l e , Geneva, Swi tzer land . D r . C I . B E S ' U C H E T . SAMA South A u s t r a l i a n Museum, Adela ide , SA, A u s t r a l i a . D r . E . G . M A T T H E W S . SPCB Sedlacek's pr ivate co l l e c t i on , B r o o k f i e l d , Q L D , A u s t r a l i a . M r . J . S E D L A C E K . UQIC U n i v e r s i t y of Queensland, Insect Co l l ec t ion , Br i sbane , Q L D , A u s t r a l i a . M s . M . A .

SCHNEIDER. USNM United States Nat iona l Museum, Smithsonian Ins t i tu t ion , Washington, DC, U . S . A .

D r . G. F . H E V E L . Z M H U Zoologisches Museum, Humbold t Universität, B e r l i n , G e r m a n Democra t i c Republ ic .

D r . F . HLEKE.

In the case of species, at which the Hungar ian m a t e r i a l consis ts of females or s l igh t ly ; damaged males only , I endeavoured to designate holotypes f r o m the m a t e r i a l of the Be rn i ce I Pauahi Bishop Museum, as fa r as i t was possible .

In the desc r ip t ion of species , the abbrev ia t ion E I is applied to the r a t io of eye d iameter and in t e rocu la r distance in d o r s a l v iew. These measurements and the r e l a t ive lengths of an­tennái segments are values measured d i r e c t l y by the ocular m i c r o m e t e r .

The labe l data of holotypes are fu l ly c i ted . The text of one l abe l is in quotation marks and the di f ferent labels are separated by semico lon . Holotype labels are r ed wi th the fo l low­ing text: Holotypus , sex m a r k , generic and species name, M e r k l , 1988.

Data of paratypes are g iven i n the sequence of l o c a l i t y , a l t i tude ( i f poss ib l e ) , t i m e of co l l ec t ing , co l l ec t ing method ( i f possible) , name of co l l ec to r ( s ) . The number of specimens according to sexes and the abbrev ia t ion of r epos i t o ry are in parentheses. The paratype speci­mens are grouped in accordance of provinces of Papua New Guinea. Specimens co l lec ted in the western par t of New Guinea mainland, ( I r i an Jaya) are neglected in this paper and never designated as paratypes. Paratype labels are y e l l o w , w i t h the text as i n the holotype labels (of course, w i t h Paratypus instead of Holotypus) .

To save space, the fo l lowing fu r ther abbreviat ions and condensations seemed to be reasonable. On the one hand, an ove rwhe lming par t of the specimens was col lec ted by the S E D L A C E K ' s - the le t te rs JS, MS and JHS mean J . . M . and J . H . S E D L A C E K . On the other hand, by far the most specimens or ig ina te f r o m Wau (Morobe Prov ince ) and i t i s fel t super­fluous to r ec i t e a l l - often qui te nearby - al t i tudes and days of co l l e c t i ng . T h e r e f o r e , the Wau specimens of the respect ive species are s u m m a r i z e d , wi th an i nd i ca t ion of lowest and h igh­est a l t i tudines , the f i r s t and last dates of co l lec t ing and a l i s t of c o l l e c t o r s .

Acknowledgements . I am grea t ly indebted to a l l persons who provided me h igh ly va lu ­able l ag r i ine m a t e r i a l co l lec ted i n New Guinea and i t i s a pleasure to supply the i r col lec t ions w i t h type specimens. T h e i r names are indicated i n the l i s t of o r i g i n a l r epos i to r ies (see above).. My special thanks are due to M i s s Ágnes BARTQS for p repar ing the excellent habitus d raw- 1

ings .

L I S T O F SPECIES

Codes between square brackets r e f e r to D r . J . B A L O G H ' s f i e ld notes s t i l l i n manu­s c r i p t ; numera l s in parentheses indicate number of specimens. Abbrev ia t ions : JB = J . B A ­LOGH ;.JJSZ = J . J . S Z E N T - I V A N Y .

L a g r i i n a

1. L a g r i a heurni Bo rchmann , 1924 - One specimen: A n g o r a m , 13-16. V I I I . 1969 [ N G A -U . 11. ] , J B .

2. L a g r i a p u p i l l i c o l l i s B o r c h m a n n , 1936 - One specimen: Haus Copper, Wau, M t . M i s -s i m , 22-24. I V . 1965, JB et JJSZ.

3. L a g r i a l i g u l a t a s p . n . - H o l o t y p e a n d 16 p a r a t y p e s : s e e d e s c r i p t i o n . 4. L a g r i a a m e t h y s t i n a s p . n. - O n e p a r a t y p e : s e e d e s c r i p t i o n . 5. L a g r i a s a p p h i r i n a s p . n . - H o l o t y p e and one p a r a t y p e : s e e d e s c r i p t i o n . 6. L a g r i a s p . n . - O n e s p e c i m e n s : M u s g r a v e r i v . v a l l . , c a . 55 k m N N E of P o r t M o ­

r e s b y . 31 . I I I . - 5 . I V . 1965 , J B et J J S Z . It i s a f e m a l e s p e c i m e n without any c h a r a c t e r i s t i c f e a t u r e . I n the l a c k of m a l e , i t w o u l d be an e x a g g e r a t i o n to d e s c r i b e i t .

7. O r e o g r i a l a r v a t a M e r k l , 1988 - 5 s p e c i m e n s : W a u , Mt . K a i n d i , 24 . DC. 1969 [ N G W -M . 2 6 . ] , J B (ho lo type , S and a o p a r a t y p e ) ; i d . [ N G W - U . 3 1 . ] (2 g p a r a t y p e s ) ; M t . K a i n d i , 2400 m , 1 5 - 1 6 . I V . 1965 , J D et J J S Z {Q_ p a r a t y p e ) .

8. O r e o g r i a p l i c a t a M e r k l , 1988 - O n e s p e c i m e n : H a u s C o p p e r , W a u . M t . M i s s i m , 2 2 - 2 4 . I V . 1965 , J B et J J S Z (ho lotype , o j . ,

8. B o t h r i c h a r a m o d e s t a B o r c h m a n n , 1 9 1 5 - O n e s p e c i m e n : A n g o r a m , 1 3 - 1 6 . V I I I . 1 9 6 9 [ N G A - U . 1 1 . ) , J B .

9 . B o t h r i c h a r a p u l c h e l l a ( G u é r i n - M é n e v i l l e , 1830) - 8 s p e c i m e n s : V a n a p a R i v . , 32 m i l e s N of P o r t M o r e s b y , 2 . I V . 1965 , J B et J J S Z (1); B r o w n R i v . , 40 k m N of P o r t M o r e s b y , 6 - 8 . I V . 1965 , J B et J J S Z (1); W a u , M c A d a m P a r k , 1 8 - 2 1 . I V . 1965 , J B et J J S Z (1) ; H a u s C o p p e r . W a u , M t . M i s s i m , 2 2 - 2 4 . I V . 1965 , J B et J J S Z (1); W a u , bea ten f r o m l e a v e s , 8 - 9 . I X . 1968 [ N G W - C . 1 7 . ] , J B (1) ; W a u , K i l o l o C r e e k , 26 . V I I I . 1968 [ N G W - C . 14} , J B (1); B a i y e r R i v e r , S a n c t u a r y , 1 -5 . DC. 1969 [ N G B - U . 2 1 . ] , J B (1); i d . [ N G B - U . 2 3 . ] (1) .

10. B o t h r i c h a r a p a l l i a t a ( M a c l e a y , 1886) - O n e s p e c i m e n : B a i y e r R i v e r , S a n c t u a r y , 1 - 5 . DC. 1969 ( N G B . - U . 2 1 . 3 , J B .

11. B o t h r i c h a r a f u s c o p u b e n s B o r c h m a n n , 1936 - 3 s p e c i m e n s : W a u , B i s h o p M u s e u m F i e l d S t a t i o n , 1 5 - 2 5 . I V . 1965 , J B et J J S Z (1); W a u , M c A d a m P a r k , 1 8 - 2 1 . I V . 1 9 6 5 . J B et J J S Z (1); W a u , beaten f r o m l e a v e s , 8 - 9 . DC. 1968 [ N G W - C . 1 7 ] , J B (1) .

12. B o t h r i c h a r a i n s i g n i s B o r c h m a n n , 1936 - O n e s p e c i m e n : W a u , E d d y C r e e k , 20 50 m , 30 . V I I I . 1968 [ N G W - C . 1 3 . ] , J B .

13 . B o t h r i c h a r a i r i d e s c e n s s p . n . - H o l o t y p e : s e e d e s c r i p t i o n . 14. B o t h r i c h a r a i n e r s s p . n . - H o l o t y p e : s e e d e s c r i p t i o n . 15. B o t h r i c h a r a i n t r i c a t a s p . n . - O n e p a r a t y p e : s e e d e s c r i p t i o n . 16. B o t h r i c h a r a w a u s p . n . - 2 p a r a t y p e s : s e e d e s c r i p t i o n . 17. B o t h r i c h a r a s p . n . - 6 s p e c i m e n s : H a u s C o p p e r , W a u , M t . M i s s i m , 22-24 . I V . 1965 .

J B et J J S Z . T h e s e s i x f e m a l e s c e r t a i n l y r e p r e s e n t a n e w s p e c i e s . M a l e s f r o m o t h e r c o l l e c ­t i o n s m a y be a s s o c i a t e d wi th t h e s e f e m a l e s , t h e s e a r e g r o u p e d in to two s y m p a t r i c s p e c i e s s h a r p l y d i f f e r i n g f r o m e a c h o t h e r . F e m a l e s , h o w e v e r , s e e m to be e n t i r e l y u n i f o r m a n d , at the m o m e n t , I c a n n o t p l a c e t h e m in e i t h e r one of the g r o u p s of m a l e s .

18. T o m o g r i a p e r l a t a gen. et s p . n . - 7 p a r a t y p e s : s e e d e s c r i p t i o n . 19. A c u t o g r i a f a l c a t a gen. et s p . n . - H o l o t y p e and 3 p a r a t y p e s : s e e d e s c r i p t i o n . 20 . K a i n d i l a g r i a f o r c i p a t a gen . et sp. 1 n . - O n e p a r a t y p e : s e e d e s c r i p t i o n .

S t a t i r i n a

2 1 . S o r a b i r o i P i c , 1956 - O n e s p e c i m e n : B u s s u R i v . , L a e , 1 3 . I V . 1965 , J B et J J S Z . 2 2 . C a s n o n i d e a s e m i s u t u r a t a F a i r m a i r e , 1896 - 9 s p e c i m e n s : H a u s C o p p e r , W a u , Mt;

M i s s i m , 2 2 - 2 4 . I V . 1965 , J B et J J S Z (7); L a l o k i , 15 k m N of P o r t M o r e s b y , 30 . I I I . 1965 , J B et J J S Z (1); P o r t M o r e s b y , 2 - 3 . I V . 1965 , J B et J J S Z (1) .

2 3 . C a s n o n i d e a q u a d r i l i n e a t a ( B o r c h m a n n , 1936) - 4 s p e c i m e n s : W a u , 8. DC. 1968 ! N G W -R . 1 9 ! , I . L o k s a (2); i d . , 2 2 - 3 0 . DC. 1969 [ N G W - U . 3 0 . ] , J B (1); W a u , M t . K a i n d i , 1 9 - 2 4 . V I I I . 1969 [ N G W - U . 1 6 ] , J B (1) .

24 . C a s n o n i d e a v a r i a b i l i s ( M a c l e a y , 1 8 8 7 ) - O n e s p e c i m e n : S u n s h i n e P l a n t . , B u l o l o v a i l . , 20 . I V . 1965 , J B et J J S Z .

2 5 . C a s n o n i d e a d i m i d i a f a F a i r m a i r e , ! 1887 - O n e s p e c i m e n : K i u n g a . 2 3 . V I I - 2 . V I I I . 1969 [ N G K - U ( + M ) . 2 . ] . J B .

2 6 . C a s n o n i d e a s e m i v i o l a c e a ( P i c , 1912) T O n e s p e c i m e n : W a u , B i s h o p M u s e u m F i e l d S t a t i o n , 1 5 - 2 5 . I V . 1965 , J B et J J S Z .

2 7 . C a n o n i d e a h y s t r i x s p . n . - H o l o t y p e a n d 7 p a r a t y p e s : s e e d e s c r i p t i o n . 2 8 . C a s n o n i d e a p u n c t i t h o r a x s p . n . - 2 p a r a t y p e s : s e e d e s c r i p t i o n . 2 9 . C a s n o n i d e a a p i c a l i s s p . n . - H o l p t y p e : s e e d e s c r i p t i o n . 30. C a s n o n i d e a f l a v i p e s s p . n . - H o l o t y p e : s e e d e s c r i p t i o n .

31 . Casnonidea baloghi sp. n. - One para type: see desc r ip t ion . 32. Casnonidea loksa i sp. n . - One paratype: see desc r ip t i on . 33. Casnonidea dobodura sp. n . - One paratype: see desc r ip t i on . 34. Casnonidea caudata sp. n . - Ho lo type : see desc r ip t i on . 35. Casnonidea pallens sp. n . - 2 para types: see desc r ip t ion . 36. Casnonidea demet r ida sp. n . - Holotype and 3 paratypes: see desc r ip t ion . 37. Casnonidea setosa sp. n . - Holo type : see desc r ip t i on .

Notes on and r e d e s c r i p t i o n of L a g r i a p u p i l l i c o l l i s B o r c h m a n n . 1936

BORCHMANN (1936:43) descr ibed this species f r o m "Holländisch-Neu-Guinea" on the basis of an unrecorded number of specimens and without reference to the type depos i tory . I t i s v e r y l i k e l y that the types 'were included i n B O R C H M A N N ' s co l lec t ion and were des t roy­ed dur ing W o r l d War I I . I t is c lear f r o m the desc r ip t ion that he knew both sexes and some cha rac t e r i s t i c features leave l i t t l e doubt what species he had. The desc r ip t ion was based on specimens wi th b lu i sh e l y t r a . The e l y t r a l pa t t e rn , however , is ac tual ly much more v a r y i n g , and the un ico loured dark blue e l y t r a represen t m e r e l y an (although frequent) ex t reme of the spec t rum of v a r i a b i l i t y . An in te rmed ia te and c h a r a c t e r i s t i c pat tern is the e l y t r a having a

F igs 1-2. 1= L a g r i a l igu la ta sp. n . , | o \ - j 2= L . p u p i l l i c o l l i s Borchmann , <?

pale spot at the midd la ( F i g . 2) . The spots,! on the one hand, may extend to the overwhelming par t of the e l y t r a and only the base and apex r e m a i n b lu i sh . On the other hand, the spots may be da rke r and obsolete, and this t r end leads to the un ico loured dark e l y t r a .

T w o of the new species descr ibed subsequently ( L a g r i a amethyst ina and L . sapphir ina) appear to be c lose ly a l l i e d to L a g r i a pup i l l i co l l i s 'Borchmann , but the l a t t e r is eas i ly d i s t i n ­guished at the f i r s t glance by the dark green or blue p rono tum as w e l l as dark legs and an­tennae.

Redesc r ip t ion : Head, p rono tum and s c u t e l l u m dark olivaceous green to blue wi th strong me ta l l i c sheen; e l y t r a pu rp l i sh red to blue; w i t h a l a rge , ovoid ochreous spot at the middle ; th is spot leaving su tu ra i m a r g i n dark , n a r r o w e s t at inner side, g radual ly widening toward e l y t r a l m a r g i n , an te r io r border nea r ly h o r i z o n t a l , pos te r io r bo rde r oblique; spot may often be expanded or reduced to disappear; v e n t r a l surface and legs black; antennae dark reddish brown to black. Length: 6 .9 -10 .0 m m .

S. Head l a rge ly impunctate between eyes and on ve r t ex ; w i t h s m a l l pi ts between eyesji ve r tex w i t h 2 blunt , smooth protuberance standing near each other ; temples coarse ly punc-j täte; surface wi th moderate ly long , e rec t h a i r s ; EI= 12:8; length r a t ios of antennái seg­ments : 9: 4: 8: 9: 7: 8: 7: 7: 6;5: 5: 24; no t r a n s v e r s a l or modif ied segments; segment I about as long as ha l f the distance between antennái sockets . P rono tum a l i t t l e t r ansve r se ; widest j u s t a n t e r i o r to middle ; sides weakly arched; punctures coarse , v e r y close, somewhat con­fluent p o s t e r i o r l y ; wi th 2 i n d i s t i n c t , t r a n s v e r s a l impress ions before base; pubescence mod­era te ly long and sparse, e rec t . E l y t r a elongate b road ly , ovoid , convex; punctation moder­ately coarse , near ly u n i f o r m , in terspaces not p l i ca te ; pubescence ra ther shor t , r a the r dense, e rec t . Sterna without modi f i ca t ions . Legs n o r m a l , sparse ly pubescent; t ibiae un­a rmed . Aedeagus: F i g . 7. Habi tus : F i g . ; 2 .

Ç. L a r g e r on average; EI= 13: 13; length r a t i o s of antennái segments V I I I to X I : 7:5:7» 6: 16; smooth area of head less extended; p rono tum more t r ansve r se .

F igs 3-4. 3- L a g r i a amethyst ina sp. n T , 6. - 4= L . sapphi r ina sp. n . , #

Desc r ip t i on of new taxa

L a g r i a l igu la ta sp. n. (Figs 1, 5, 6)

Head, p rono tum and scu t e l l um da rk , shiny, m e t a l l i c green; e l y t r a b rownish w i t h s t rong greenish , pu rp l i sh or v io le t t inge; v e n t r a l surface redd i sh , meso- and metas ternum da rke r ; legs black; antennae l a rge ly y e l l o w i s h , ap ica l 4-5 segments g radua l ly da rke r . Length: 6 .5-10 _1 m m .

8, Head v e r y weakly i m p r e s s e d between eyes; coarse ly and sparse ly punctate; w i t h long , scat tered h a i r s ; eyes conspicuously l a rge , EI= 12: 5;ülength ra t ios of antenna! seg­ments: 7 :4 . 5 : 9 : 8 : 7 . 5: 1 0 : 9 : 9 : 7 . 5:23; segment I about as long as ha l f the distance between antenna! sockets; segment X I somewhat widened, v e r y weakly tw i s t ed , inner surface con­cave (F ig . 5). P rono tum as long as or a l i t t l e longer than broad; widest at the middle , s ides, v e r y weakly arched; punctures coarse , separated by interspaces as broad as puncture d ia­meter ; surface wi th 4 shallow i m p r e s s i o n s ; pubescence modera te ly long, erect . E l y t r a e lon­gate, feebly broadening, weakly convex; punctation coarse , modera te ly dense; interspaces s l i gh t ly ra i sed ; w i t h r a the r long, sparse pubescence cons is t ing of erec t and semierec t h a i r s . Sterna without modi f ica t ions . Legs t y p i c a l ; w i t h sparse , modera te ly long, semierec t h a i r s ; t ib iae una rmed . Aedeagus: F i g . 6. Habi tus : F i g . 1.

g. Considerably l a r g e r and broader on the average; EI= 1 5 : 1 1 ; length ra t ios of anten­nái segments V I I I to X I : 1 0 : 9 : 8 : 19; las t antennái segment s imp le ; p rono tum of same length and width or s l i g h t l y t r ansve r se ; e l y t r a much broader .

Holotype, 6, label led as fo l lows : "NEW GUINEA (NE) B a i y e r R i v e r Sanctuary 1-5. X I . 1969"; " (No. N G B - U . 24. ) leg . D r . J . Ba logh" . I t is deposited i n H N H M .

Paratypes (total 124 specimens) . Wes te rn Highlands P rov ince : label led as holotype | (3 68, 2 go., H N H M ) ; i d . [ N G B - U . 2 2 ^ (1 8, H N H M ) ; i d . [ N G B - U . 2 1 ] (1 8, HNHM); i d . £NGB-R . 8 . ] (1 o, H N H M ) ; i d . , 4000 ' , 6 -11 . V I I . 1974, H . Howden (1 8, 1 o, CNCI) ; J i m m i R i v e r , 4700 f t . . Ï6 . VI I -21 .DC. 1961, W . W . Brand t (1 6, 1 £. A N I C ) ; W u m , Upper J i m i V . , 840 m , 18. V I I . 1955, J . L . G r e s s i t t (3 og, B P B M ) . - Eas te rn Highlands P rov ince : K a r i m u i , 1080 m 8-10. V I I . 1963, JS (1 o, B P B M ) ; Okapa, Agakamatasa, 17.1.1965, R. Hornabrook (1 g, HPCW); Okapa, Okasa, 15. V . 1965, R. Hornabrook (1 g, HPCW); Okapa, Purosa, 7000 f t . , 23. V I . 1964, R. Hornabrook (1 o, HPCW). - Morobe P r o v i n c e : B u l o l o , 680 m , 4. V I . 1969, JS (2 go, B P B M ) ; i d . , 700 m , 18. V I . 1969, JS (1 8, B P B M ) ; i d . , 6. X I . 1969, JS (1 g, B P B M ) ; i d . , 26. X I . 1969, JS, MS (2 66, 3 oo, B P B M ; 1 0, H N H M ex B P B M ) ; 5 k m N Bulo lo , 630 m , 6. V I I I . 1964, JS, MS (1 6, B P B M ) ; Bu lo lo R. , 1Ï30 m , 17. I X . 1969, A . B . M i r z a , JS, MS (2 Z 8 , 1 o, B P B M ) ; F insch Haven, L . Wagner (3 oo, S A M A ) ; Gara ina , 830 m , 13-15.1.1968* JS. MS (2 oo, B P B M ) ; Huon Peninsula , P ind iu , 5ÓÓ-600 m , 19. I V . 1963, JS (1 6, B P B M ) ; i d . , 870-1300 m , 21-22. I V . 1963, JS (1 g. B P B M ) ; Umg. K a i a p i t , 30. I X . 1979, W . G . U l l r i c h (1 g, MHNG); Lae , 4-6 . D i . 1968 [ N G L - C . 1 6 . ] . J . Balogh (1 6, H N H M ) ; Watut R. , 900-1900 m , X . 1969, A . B . M i r z a (2 68, 1 g, B P B M ) ; Wau. 1100- 300 m . 16. X . 1961-3. V I I . 1974, R. Hon-nabrook. H . F . Howden, JS, JHS, MS (9 56, 6 oo, B P B M ; 5 55, C N C I ; 1 6, HPCW; 3 39, 2 Q 2 , H N H M ex B P B M , 1 Ó, H N H M ex CNCI) ; i d . , 22-30. I X . 1969 I N G W - U . 33. !, J . Balogh (3 oo, H N H M ) ; i d . , 2 5 . V I I I . 1968, beaten f r o m decayed leaves [ N G W - C . 10.3 , J . Balogh (1 oi H N H M ) ; Wau, B i g Wau Ck. , 1300 m , X I . 1965, P. Shanahan (1 6, B P B M ) ; Wau, Bu lo lo R. , ' 800-900 m , 3 1 . V I I I . 1965, JS (2 56, B P B M ) ; i d . , 850-950 m , 4. I I . 1966, JS ( l g , B P B M ) ; Wau, Hosp i ta l Ck. , 1150- 1250 m , 9.1.1966,; JS . iMS (1 6, B P B M ) ; Wau, Kaisenik , 1000 m , 1 9 . I V . 1965, JS, MS (1 o, B P B M ) ; Wau, Ka r inga Ck. , 950-1150 m , 3 0 . I X . 1 9 6 4 , MS (2 58. B P B M ; 1 S, H N H M ex B P B M ) ; Wau, KilokV Creek , 26. V I I I . 1968 [ N G W - C . 14.3, J . Balogh I (1 5, 1 o_, H N H M ) ; Wau, M t . K a i n d i , 24-25^ V I I I ; 1968 [ N G W - C . 9 . ] J . Balogh (1 g, H N H M ) ; ' Wau, M t . M i s s i m , 950-1300 m , 8-9.1.1966;, JS, : MS (1 8, 1 g, B P B M ) ; Wau, Wau Ck. , 1200-j 1500 m , 6 . V . 1963, JS (1 8, B P B M ) . - N o r t h e r n P rov ince : Dobodura, I I I - V I I . 1944, D a r l i n g ­ton (2 88, M C Z C ) ; Kokoda, 1200 f t . , V I I I . 1933, L . E . Cheesmann (1 g, B M N H ) ; i d . , DC.1933J (1 Q, B M N H ) ; i d . , 1300 f t . , V I I I . 1933, l ower r a i n f o r e s t , L . E . Cheesmann (2 oo. B M N H ) ; i d ] , 400 m , 1 4 - 1 6 . X I . 1965. JS, MS (1 8, 1 o, B P B M ) ; i d . , 1 8 . X I . 1966, JS (2 88, B P B M ) ; Mana-galase Plateau, V I I . 1972, R. Hornabrook (2, 88,'1 g, HPCW; 1 Ä, 1 p. H N H M ex HPCW); Mt . i Laming ton , 1300-1500 f t . . C . T . M c N a m a r a (6 88, 3 gg, SAMA; 2 88, 1 g, H N H M ex SAMA); ;

Popondetta, 29.1.1965, R. Hornabrook (1 g | HPCW); i d . , 27-28.11.1966, G. Monte i th (2 8 8 } 2 gg, UQIC) ; i d . , 60 m . 1-2. I X . 1963, JS ( l j g , B P B M ) . j

R e m a r k s : Males of L a g r i a l igu la ta may eas i ly be recognized by the s l igh t ly spoon-shap-

ed antennái segment X I . Females Have no d i s t inc t features that would d i s t ingu ish them f r o m the genera l ized females of other (par t ly undescribed) species belonging to L a g r i a and other genera of L a g r i i n a , though the lus t rous green forebody may help in the recogn i t ion .

L a g r i a amethyst ina sp. n . (Figs 3, 8)

Head, p rono tum, scu te l lum, legs and antennae y e l l o w i s h red ; e l y t r a pu rp l i sh v i o l e t ; v e n t r a l surface redd i sh , w i th metas te rnum and abdominal sterna more or less da rke r . Length: 6. 7-8. 2 m m .

3. Head impres sed between eyes; th is i m p r e s s i o n and 2 blunt, i nd i s t inc t p ro tuber ­ances near ly smooth; o therwise , head sparsely punctate w i th long erec t h a i r s ; EI= 12: 10; length ra t ios of antennái segments: 9 : 4 : 9 : 1 2 : 9 : 9 : 9 : 8 : 8 : 7 . 5 : 3 2 ; no t r a n s v e r s a l o r m o d i f i ­ed segments; segment I shor te r than half of distance between antennái sockets . P rono tum t r ansve r se ; widest jus t an te r io r to the midd le ; sides nea r ly p a r a l l e l ; punctures coarse and close, somewhat rugose before base; w i t h 2 to ta l ly i nd i s t i nc t t r ansve r sa l impress ions be­fore base; pubescence ra ther long and sparse , e r ec t . E l y t r a broadly ova l , modera te ly con­vex; punctat ion less coarse , u n i f o r m l y sca t tered , in terspaces not p l ica te ; pubescence shor t , dense, e rec t . Sterna without modi f ica t ions . Legs t y p i c a l , w i th sparse, long pubescence; 1

t ib iae una rmed . Aedeagus: F i g . 8. Habi tus : F i g . 3. o. L a r g e r on average; EI= 12: 15; length r a t ios of antenna! segments V I I I to X I : 8:8:

7: 19; i m p r e s s i o n between eyes weaker: pronotum more t ransverse . Holo type . 6*, label led as fo l lows : " N . Guinea: N E Bulo lo , 700 m 26. X I . 1969"; " J . et

M . Sedlacek Co l l ec to r s BISHOP M U S E U M " . I t is deposited i n B P B M .

F igs 5-12. 5= Las t s ix antennái segments pf L a g r i a l igu la t a sp. n . , S. - 6= Aedeagus of L . | l igu la ta sp. n . - 7 = Aedeagus of L . p u p i l l i c o l l i s Borchmann . - 8= Aedeagus of L . amethyst ina s p . n . - 9= Aedeagus of L , sapphir ina sp. n . ! - 10= Aedeagus of Bo th r i cha ra i r idescens s p . n [• 11= Aedeagus of B . ine r s sp. n . - 12= Aedeagus of B . i n t r i c a t a sp. n.

Paratypes (total 35 specimens). Morobe P rov ince : label led as holotype (2 <j><j>, B P B M , 1 o, H N H M ex B P B M ) ; i d . , JS ( 3 o o , B P B M ; 1 H N H M ex B P B M ) ; i d . , 6. X I . 1969, JS (3 oo, B P B M ; 2 H N H M ex B P B M ) ; k i d . , 690 m , 27. I I I . 1968, P. Colman (2 çç, B P B M ) i d . , 23. I V . 1969, JS (1 o. B P B M ) ; i d . , 4. V I . 1969, JS (1 ff, B P B M ) ; Bu lo lo Gorge , M c A d a m Park , 900 m , 28. X I I . 1977, u . v . l i g h t , R. Hamsen, P. Heber t et J. Ha rmsen (1 S, CNCI ) ; Bulo lo R i v . , 680 m , 23. I V . 1969, JS (1 SPCB); F i n s c h Haven d i s t r . , L . Wagner (2 çç.

SAMA) ; Wau, 1050- 1100 m , 9. D i - 1 1 . X I . 1961, JS, JHS (2 88, B P B M ; 2 So*. 3 gg,- SPCB; 1 g, H N H M ex SPCB); Wau, K i l o l o Creek, 26. V I I I . 1968 [ N G W - C 1 4 . ] , J . Balogh ( l g , HNHM) :

Wau, Mt . M i s s i m , 950- 1300 m , 8-9.1.1966, JS, MS (1 6, B P B M ) ; Zenag-Lae Road. 200 m , 17.1. 1965, JS (1 g, B P B M ) . - Madang Prov ince : Manumbo (1 g, SAMA) . - East Sepik P r o ­v ince : Hauptlager b. M a l u , 31.1.1913, Ka i se r in -Augus ta f luss Expedi t ion , S.G. Bürgers ( l g , Z M H U ) ; Lager a. Topfe r f lu s s , 1 8 . I V . 1913, Ka i se r in -Augus ta f lu s s Expedi t ion , S .G.Bürgers (1 g, Z M H U ) ; M a p r i k , Sepik, 16.11.1965, R. Hornabrook (1 g, HPCW).

R e m a r k s : L a g r i a amethyst ina is s i m i l a r to L . sapphi r ina but the e l y t r a are more broadly oval (s lenderer in sapphi r ina) , always p u r p l i s h v io l e t (bluish or greenish in sapphi­ r i na ) and pronotum is never d a r k l i n g (often so i n sapphi r ina) .

L a g r i a sapphir ina sp. n . (Figs 4, 9)

Head, p rono tum, s c u t e l l u m , legs and antennae pale reddish brown; p rono tum most of­ten (and head somet imes) da rke r , w i t h greenish t inge; last antennái segment somewhat dark­e r ; e l y t r a greenish blue (reddish brown in i m m a t u r e specimens) ; v e n t r a l surface da rke r r e d ­dish b rown. Length : 7 .2 -9 .0 m m .

8. Head impressed between eyes; w i th 2 blunt protuberances behind i m p r e s s i o n ; p ro tuberances and i m p r e s s i o n near ly smooth; o the rwise , head sparse ly punctate; w i t h some long , e rec t h a i r s ; EI= 13: 8; length ra t ios of antennái segments: 9: 4: 10: 10: 6: 8: 8: 7: 7: 6: 30; no t r ansve r sa l or modif ied segments; segment I s h o r t e r than half of distance between anten­nái sockets. P rono tum near ly as long as broad or s l i gh t l y t ransverse ; widest j u s t an t e r io r to the middle ; sides subpara l l e l to weakly arched; punctation coarse and dense, somewhat rugose p o s t e r i o r l y ; surface w i t h 2 or 4 weak i m p r e s s i o n s ; pubescence ra the r long , sparse , e rec t . E l y t r a r a t h e r elongate ova l , moderate ly convex; sides near ly s t ra igh t f r o m humerus to broadest point; punctation coarse , dense; interspaces a l i t t l e p l icate ; pubescence r a the r shor t , dense, e rec t to suberect . Sterna without modi f ica t ions . Legs typ ica l , w i t h sparse, r a t h e r shor t pubescence; t ibiae unarmed. Aedeagus: F i g . 9. Habi tus: F i g . 4.

g. L a r g e r on the average; EI= 14: 12; length ra t ios of antennái segments V I I I to X I : 8 : 8 : 7 : 2 2 ; i m p r e s s i o n between eyes weaker , but protuberances somewhat more p rominen t ; pronotum d i s t i nc t l y t r ansverse .

Holotype, 8, label led as fo l lows : "NEW G U I N E A : Wau, Bishop Museum, F i e l d Station, 15-25. I V . 1965"; " C o l l . D r . J . B A L O G H et D r . J . J . S Z E N T - I V A N Y " . I t i s deposited in H N H M .

Paratypes (total 60 specimens) . Morobe P rov ince : B u l o l o , 26. X I . 1969, JS, MS (1 g, B P B M ) ; Bu lo lo R i v . , 1180 m , 1 7 . I X . 1969, A . B . M i r z a (1 g, B P B M ; 1 g, H N H M ex B P B M ) ; Gara ina , 20. X I - 1 7 . X I I . 1969, A . B . M i r z a (1 g, B P B M ) ; i d . , 28. V I I . 1971, R. Hornabrook (1 8, 1 g, HPCW; 1 g, H N H M ex HPCW); U m g . Gurako r , 1 6 . D i . 1979, W . G . U l l r i c h (1 g, MHNG); M u m i n g , 600 m , 9-10. I I I . 1962, JS (1 g, B P B M ) ; Sat telberg, v . Bennigsen (1 8, I F P E ) ; Simbang, Huon Golf , 8. I I . 1899, L . Biró (1 g, H N H M ) ; Wareo, L . Wagner (1 g, S A M A ) ; Wau, 1050-1300 m , 20. V I I I . 1961-XI . 1978, Deharveng, R. Hornabrook , H . F . How­den, R . E . P a r r o t t , JS, MS, R. Straatman,; (4 88, 6 gg, B P B M ; 3 gg, CNCI ; 1 g HPCW; 1 MHNG; 2 88, 2 gg, SPCB; 1 8, H N H M ex B P B M ; 3 gg, H N H M . 1 each ex C N C I , HPCW, SPCB); Wau, 8-14. X I I . 1976, m e r c u r y vapor l i g h t , G . F . H e v e l e t R . E . Dietz (1 8, 7 gg, USNM; 1 8, 1 g, H N H M ex USNM); Umg. Wau, X I I . 1978, W . G . U l l r i c h (1 g, MHNG); Wau, B i g Wau Creek, 1300 m , X I . 1965, P. Shanahan (1 8, B P B M ) ; Wau, Hospi ta l Creek , 1200 m , 11. 1965, JS, MS (1 g, . B P B M ) ; Wau, M t . M i s s i m , 980 m , 20. V I I . 1969, J . L . G r e s s i t t et V. H i r a s h i m a (1 o, B P B M ) . - Madang Prov ince : F i n i s t e r r e Range, Saidor, Funyende, 1200 m , 24-30. I X . 1958, W . W . Brandt (1 g, B P B M ) ; U m g . Madang, 29. V I . 1979, W . G . U l l r i c h (1 g, MHNG). - Wes te rn Highlands Prov ince : B a i y e r R i v e r , Sanctuary, 1 -5 . IX. 1969 [ N G B - U . 2 3 . ] , J . Balogh (1 g, H N H M ) ; M t . Hagen iVal ley , K e l t i g a , 5600 f t . , 28. I X - 2 5 . X . 1961, W . W . B r a n d t (1 8, A N I C ) . - West Sepik P rov ince : T o r r i c e l l i M t s , Mohai V i l i . , 750 m , 8-15. X I I . 1958, W . W . Brandt (1 6, B P B M ) . - Western P roy ince : D a r u , mouth of F l y R i v e r , V I I I . 1941, R. G; Wind (1 g, CASC); Moorhead, 18 m , 14. V I I . 1964, H . C l i s s o l d (1 g, B P B M ) ; Morehead , 10. X I . 1974, R. Hornabrook (1 8, HPCW). - N o r t h e r n P rov ince : Mt . Lamington , 1300-1500 f t . , i C . T . McNamara (1 g, S A M A ) ; Dobodura, I I I - V I I . 1944, Da r l i ng ton (1 g, M C Z C ) .

R e m a r k s : L a g r i a sapphir ina is c losely a l l i e d to L . amethyst ina but e l y t r a are more elongate (more ovoid in amethyst ina) , g reen i sh -b lu i sh (purp l i sh v io le t i n amethyst ina ), the pronotum and last antenna! segment most ly d a r k l i n g (reddish in amethyst ina) .

B o t h r i c h a r a i r idescens sp. n . (Figs 10, 13)

Head, pronotum and scu te l lum black, densely covered by fine decumbent h a i r s ; e l y t r a l a rge ly p u r p l i s h - b l u i s h , i r idescen t , at places wi th greenish l u s t r e ; pubescence decumbent, incomple te ; pubescent places a l te rna t ing w i t h extensive glabrous par t s ; v e n t r a l surface black; legs dark blackish b rown; antennae reddish brown, a l i t t l e d a r k l i n g toward apex. Pub­escence s i l v e r y whi te . Length: 7. 5-7. 6 m m .

6. Head finely and densely punctate; eyes ra ther convex, t r ansve r se ; EI= 13:8; length r a t ios of antennái segments: 8: 5: 13: 15: 13: 12: 12: 12: 11 : 11: 22; no modif ied or t r a n s v e r s a l segments; segment I about as long as half the distance between antennái sockets. Pronotum very densely and finely punctate; punctuation i nv i s ib l e because of dense pubescence; of near­ly same length and wid th ; widest at base, s l i gh t l y cons t r ic ted ju s t pos t e r io r to the middle-

disc w i th flat impres s ions . E l y t r a elongate, moderate ly convex, s l i gh t l y widening in s t ra igh t l ine , widest in apical 1/4; punctures-coarse, more or less evenly scat tered, w e l l separated, interspaces a lmost f la t , not p l ica te ; basal 1/6 densely pubescent, then fol lowed by a t r ans ­verse glabrous s t r i p of same wid th ; r e s t of e l y t r a wi th l a rge , pa r t ly confluent, round,

F ig s 13-14. 13= B o t h r i c h a r a i r idescens sp. n . , - 14= B . iners sp. n . , <?.

glabrous spots surrounded by pubescent areas; glabrous spots somewhat depressed. Legs ra ther long and slender; f emora wi th scarce , decumbent pubescence; t ibiae wi th pubescence also decumbent, inner side w i t h shor t , s emie rec t h a i r s as w e l l ; fore t ibiae near ly s t r a igh t , unarmed; middle and hind t ib iae s l igh t ly but d i s t i nc t l y arched, inner ma rg in w i t h few s m a l l denticles placed far f r o m each other . Sterna without modi f ica t ions : last abdominal s te rn i te broadly rounded. Aedeagus: F i g . 10. Habi tus : F i g . 13.

o" Unknown. Holotype, 6\ l abe l led as fo l lows : "NEW GUINEA (NE) B a i y e r R i v e r Sanctuary 1-5. D i .

1969."; " (No. N G B - U . 22. ) l eg . D r . J . Balogh" . I t is deposited i n H N H M . Paratype. Western Highlands P rov ince : J i m i V a l l e y , Tabibuga, 23-24. I V . 1970, R.

Hornabrook (1 Í . HPCW). R e m a r k s : B o t h r i c h a r a i r idescens is somewhat s i m i l a r to B . pulchel la (Guérin-Méne­

v i l l e , 1830) but d i s t inc t ly d i f fers f rom i t in the f o l l o w i n g :

ground colour of e l y t r a

t r a n s v e r s a l glabrous s t r i p behind pubescent e l y t r a l base

glabrous spots on e l y t r a

apex of aedeagus

i r idescens sp. n .

b lu i sh - p u r p l i s h , i r idescen t , sh ining

ho r i zon ta l

d i s t i nc t , w e l l depressed

an acute process

pulchel la (Guérin-Méne­v i l l e )

b lu i sh , not i r idescen t , alutaceous

oblique

ind i s t inc t , poor ly depressed

t r i a n g u l a r

B o t h r i c h a r a iners sp. n . (Figs 11 , 14)

Body b lackish b rown; e l y t r a somewhat redd i sh ; su tu ra i ma rg in black. D o r s a l and ven­t r a l surface covered wi th shor t , decumbent, g r ey i sh white pubescence. Length: 9. 1 m m .

<?. Head finely and sparse ly punctate, sparse ly pubescent; eyes ra ther convex, t r ans ­verse ; EI= 13. 5: 10; antennae ra ther stouth; segment length r a t i o s : 9 :4 . 5: 13: 14: 13: 12: 11: 1 0 : 1 1 : 1 1 : 2 1 ; no modif ied o r t r a n s v e r s a l segments; segment I sho r t e r than half the distance between antennái sockets. Pronotum a l i t t l e broader than long; widest jus t an te r io r to the middle , r a the r cons t r i c ted an t e r io r to base; disc w i t h 4 d i s t inc t but flat impress ions ; su r -facely densely and modera te ly coarsely punctate; pubescence r a the r sparse, punctation w e l l v i s i b l e among h a i r s . E l y t r a modera te ly convex, elongate, widest in pos te r io r 1/3: sides arcuate; surface wi th coarse punctures; interspaces not p l i ca te , as broad as or w ide r than puncture d iamete r ; pubescence moderately dense, leaving a few s m a l l spots g labrous; these spots v e r y weakly impres sed . Legs ra ther long and slender, sca rce ly pubescent; t ibiae near­ly s t r a igh t , unarmed. Sterna without modi f ica t ions ; las t abdominal s t e rn i t a rounded. Aedea­gus: F i g . 11 . Habi tus: F i g . 14.

g. Unknown. , Holotype , S, l abe l led as fo l lows : "NEW GUINEA (NE) Wau, M t . K a i n d i , 19-24. V I I I .

1969"; " (No . N G W - U . 12.) l eg . D r . J . Balogh. I t i s deposited i n H N H M . R e m a r k s : Th i s species is somewhat s i m i l a r to B o t h r i c h a r a h e l l e r i Borchmann, 1915,

but the e l y t r a are less elongate, the e l y t r a l punctat ion is coarse r , the antennae are shor t e r and t h i c k e r .

B o t h r i c h a r a i n t r i c a t a sp. n . (Figs 12, 15)

Head, pronotum and scu te l lum black, densely covered by f ine, decumbent h a i r s ; e l y t r a dark redd ish brown wi th r e s t r i c t e d , decumbent pubescence; surface l a rge ly glabrous; ven­t r a l surface, legs and antennae dark reddish brown to black, basal par t of antennae some­what pa le r . Pubescence golden y e l l o w i s h . Length: 9 . 5 - 1 1 . 5 m m .

S. Head f inely and densely punctate; eyes r a the r convex, t r ansve r se ; EI= 12: 13; length ra t ios of antenna! segments: 10: 5: 15: 13: 13: 11: 11 : 11:12: 10: 2 1 ; no modif ied or t r a n s v e r s a l

S e g m e n t s ; segment I about as long as ha l f of distance between antennái sockets. P rono tum ve ry densely and f inely punctate, punctuation almost i n v i s i b l e because of dense pubescence; about as long as broad; widest at base but width measured an te r io r to cons t r i c t ion a lmost equal to basal wid th ; disc wi th 4 i nd i s t i nc t i m p r e s s i o n s . E l y t r a s t rong ly convex and broad, widest in apica l 1/3; surface e x t r e m e l y roughly scu lp tu red ; w i th coarse , sparse punctures and h ighly elevated, i r r e g u l a r , pl icate in t e rva l s making the surface creased; w i t h some barely d i sce rn ib l e , shallow i m p r e s s i o n s ; pubescence sho r t , decumbent, inconspicuous, somewhat denser in basal 1/6 and along su tu ra i m a r g i n , elsewhere sparse r ; impressed spots g labrous . Legs moderate ly long and s lender , w i t h scanty pubescence; fore t ibiae near­ly s t r a igh t , unarmed; middle and hind t ib iae very weakly arched, w i t h a row of s m a l l den­t i c les along inner m a r g i n . Sterna wi thout modi f ica t ions ; last abdominal s te rn i te broadly rounded. Aedeagus: F i g . 12. Habi tus : F i g . 15.

F igs 15-16. 15= B o t h r i c h a r a i n t r i c a t a sp. n . , Ô*. - 16= B . wau sp. n . , <?

p. L a r g e r and broader on the average; eyes s m a l l e r ; EI= 14: 18; length r a t i o s of an­tennái" segments V I I I to X I : 12 :12 :11 :22 ; p rono tum d i s t i nc t l y t r ansverse ; a l l t i b i ae unarmed; e l y t r a s l i gh t ly caudate.

Holotype , 8, l abe l led as fo l lows : " N E W G U I N E A : NE Wau, Morobe D i s t r . 1100 m, 31 . I . 1963"; " J . Sedlacek Co l l ec to r BISHOP". I t is deposited in B P B M .

Paratypes (total 31 specimens) . Morobe P rov ince : Gara ina , 1200 m , 15 .1 . 1968, JS, MS (1 g, B P B M ) ; M t . K a i n d i , 2350 m , 1. I V . 1966, J . L . et M . G r e s s i t t (1 g, B P B M ) ; i d . , 19-24. V I I . 1969, J . L . Gres s i t t (1 g, B P B M ) ; Wau, 1100- 1800 m , P. Colman, R. Hornabrook, R . E . P a r r o t t , JS, MS, P. Shanahan (2 83, 6 gg, B P B M ; 1 g, C N C I ; 1 g, HPCW; 3 gg, SPCB; 1 8, H N H M ) ; Wau, Edie Creek , 2050-2300 m , 17-18. V I I I . 1965, JS (1 g, B P B M ) ; i d . , 2000-2400 m , 12. V I I I . 1963, P. Shanahan. (1 g, B P B M ) ; i d . , 7000 f t . , 2 1 . I X . 1964, M . E . Bacchus (1 g, B M N H ) ; Wau, Haus Copper, M t . M i s s i m , 22-24. I V . 1965, J. Balogh et J . J . Szent-Ivány (1 o, H N H M ) ; M t . M i s s i m , 1600-2000 m , 21-24. I X . 1965, MS (1 o, B P B M ) ; i d . , 1500-1800 m , 6.1.1970, JS, MS (1 g, B P B M ) ; Wau, K u n a i C k . , 1580 m , 22. V . 1965, JS ( l g . S P C B ) ; Wau, K u j e r u , 1500 m , 27. I X . 1969, A . B . M i r z a (1 8, 1 g, B P B M ; 1 6, H N H M ex B P B M ) ; Wau, Wau Creek , 1200- 1500 m , 16-18. I X . 1964, MS (1 8, 1 g, B P B M ; 1 g, H N H M ex BPBM); Watut R. , 900- 1900 m , X . 1969, A . B . M i r z a (1 8, B P B M ) .

R e m a r k s : Here tofore , i t is the only descr ibed species of Bo th r i cha ra having very rough­ly p l ica ted e l y t r a .

B o t h r i c h a r a wau sp. n. (Figs 16-18)

Body black; head, p rono tum, s cu t e l l um and v e n t r a l surface densely covered wi th ve ry shor t , fine decumbent h a i r s ; e l y t r a l ve s t i t u r e uneven: densely pubescent spots a l te rna t ing w i t h places of sparser pubescence. Pubescence s l i v e r y whi te . Length: 9. 1-12. 5 m m .

3. Head f inely and densely punctate; eyes moderate ly convex, t r ansve r se , E I : 16: 11; length ra t ios of antennái segments: 11:6: 15: 15:12: 12: 11: 11: 10: 9: 2 1 ; no modif ied or t rans­v e r s a l segments; segment I about as long as half the distance between antennái sockets. P rono tum densely and finely punctate; punctuation w e l l v i s i b l e , not en t i re ly covered by fine pubescence; about as long as broad; widest jus t a n t e r i o r to the midd le , but base near ly as broad; w i t h 4 ind i s t inc t impre s s ions . E l y t r a modera te ly vconvex, elongate, wides t i n ante­r i o r 1/4; sides widening i n a s t ra igh t l ine to widest point ; punctation coarse and moderately dense; at places, interspaces somewhat broader and s l i gh t l y ra i sed ; pat tern of pubescence c h a r a c t e r i s t i c : denser pubescence cove r ing basal 1/6 (this surface obliquely bordered pos­t e r i o r l y ) , f o rming a p reap ica l spot (reaching side of e l y t r a but not reaching su tu ra i marg in and apex); r emain ing par t of e ly t r a wi th a number of s m a l l i r r e g u l a r spots a r ranged in loose longi tudina l rows ; space amongst densely pubescent spots w i th much sparser v e s t i t u r e ; no def in i te ly glabrous and impressed places. Legs long and s lender , wi th sparse , decumbent pubescence; fore and middle t ibiae nea r ly s t r a igh t , s imp le ; hind t ibiae ve ry weakly arched, w i th a weak but d i s t inc t preapica l swe l l i ng (F ig . 17). Sterna without modi f i ca t ion ; last ab­dominal s te rn i te rounded. Aedeagus: F i g . 18. Habi tus : F i g . 16.

g. Much l a r g e r on the average; eyes s m a l l e r ; EI= 15:20; length ra t ios of antennái seg­ments V I I I to X I : 12 :11 :10 :20 ; pronotum d i s t i n c t l y t r ansverse , impres sed mesa l ly ; e l y t r a l ve s t i t u r e usual ly much more conspicuous; hind t ib iae s i m p l e .

Holotype , 3, l abe l led as fo l lows : "NEW GUINEA (NE) Wau, Morobe D i s t r . 1200 m , 5. V . 1963"; " J . Sedlacek BISHOP". I t is deposited in B P B M .

Paratypes (total 81 specimens) . Morobe P rov ince : K a i n d i - N a m i , 1700 m , 15. V I . 1965, JS (1 g, SPCB); Wau, 1050- 1800 m , 1. I I . 1961-22. I I I . 1969, G. Monte i th . JS, JHS, MS (12 88, 18 gg, B P B M ; 1 8, 4 gg, SPCB; 3 88, 6 gg, H N H M ex B P B M ) ; Wau, 1200 m , 8-14. X I I . 1976, m e r c u r y vapor l igh t , G. F . Hevel et H . E. Die tz (1 g, USNM); Wau, B i g Wau Ck. , 1200-1500 m , I X . 1965, JS, MS (1 o, B P B M ) ; Wau, Haus Copper , M t . M i s s i m , 22-24. I V . 1965, J . Ba ­logh et J . J . Szent-Ivány (2 gg, H N H M ) ; Wau, M t . M i s s i m , 1500-1800 m . 6.1.1970, JS, MS (1 3, 1 g, B P B M ) ; i d . , 1600-2000 m , 21-24. I X . 1964, MS (2 gg, B P B M ) ; W a u . Ku je ru , 1500 m , 27. V . 1969, A . B . M i r z a (1 8, B P B M ) ; Wau, M c A d a m Park , 1200- 1300 m , 7 . V . 1965, JS (1 g, B P B M ) ; Wau, M t . K a i n d i , 1900-2200 m . 19. V I I I . 1965, JS, MS (1 o. B P B M ) ; Wau, Wau. Ck. , 18. IX. 1964, JS (1 g, SPCB); i d . , 1200- 1500 m , 28. I I I . 1963, JS (1 8, B P B M ) ; i d . , 1200-1500 m , 16-18. I X . 1964, MS (1 g, B P B M ) ; 6 k m W of Wau, N a m i Creek, 1700 m , 15. VI.1962 (1 3, 2 gg, B P B M ; 1 g, H N H M ex B P B M ) . - Madang P rov ince : F i n i s t e r r e M t s . , Budenu, cca

4000 f t . , 15-24. X . 1964, M . E. Bacchus (2 88, 2 go, B M N H ) ; F i n i s t e r r e M t s . , M o r o , cca 5550 f t . , 3 0 . X . - 1 5 . X I . 1964, M . E . Bacchus (1 g, B M N H ) . - Eas te rn Highlands P rov ince : Asakamotasa , 6000 f t . , 27.11.1964, R. Hornabrook (2 go, HPCW; 1 g, H N H M ex HPCW); Umg. Kainantu , Onerunka, I V . 1979, W . G . U l l r i c h (1 g, MHNG); i d . , 5 . X . 1979 (1 g, MHNG), i d . , 13 .X . 1979 (1 g, H N H M ex MHNG); Okapa, 23. V . 1965, R. Hornabrook (1 g, H P C W ) ; i d . , cca 5000 f t . , 4-15. I I . 1965, M . E. Bacchus (2 gg, B M N H ) ; Purosa Camp, Okapa area , 15 mis SSE Okapa P a t r o l Post , 1950 m . Camp 10, 2 1 . I X . 1959, Sixth A r c h b o l d Expedi t ion , | L . J . B r a s s (1 g, A M N H ) ; Waisa, N Okapa, c. 5000 f t . , 15.11. 1965, M . E . Bacchus (1 g, B M N H ) . - Western Highlands Prov ince : B a i y e r - R o h i n a , I V . 1979, W . G . U l l r i c h (1 g, MHNG); Mt. Hägen Va l l ey , K e l t i g a , 5600 f t . , 28. I X . - 25. X . 1961, W . W . Brandt (1 g, A N I C ) .

17 18 19 20 21 22

Figs 17-22. 17= Hind t i b i a of Bo th r i cha ra wau sp. n . , S. - 18= Aedeagus of B . wau sp. n . -19-20= Aedeagus of T o m o g r i a per la ta gen. et sp. n . , v e n t r a l view (19) and l a t e r a l view (20). r

21-22= Aedeagus of A c u t o l a g r i a falcata gen, et sp. n . , v e n t r a l view (21) and l a t e r a l v iew (22)'

R e m a r k s : The new species is somewhat s im i l a r to B o t h r i c h a r a kuntzeni Borchmann , 1936 on account of i t s e l y t r a l pubescent spots ar ranged in r o w s , but a large preapical spot is present ( lacking in kuntzeni ) , the e l y t r a l punctation is coarse (fine in kuntzeni) , the i n t e r spaces s l i g h t l y pl icate (not so in kuntzeni) and ftje Hind t ib iae of males are swollen before apex (simple i n kuntzeni) .

T o m o g r i a gen. n.

Diagnos is : Bather b road and convex,- med ium-s i zed ; colour dark; pubescence shor t , semierec t , inconspicuous; head, pronutum and e l y t r a w i t h d i s t inc t punctation.

S. Head wi th i n t e r o c u l a r distance much n a r r o w e r than eye d iameter ; antennae wi thout modified o r t r ansve r sa l segments; segmenf: X I longer than 4 preceding segments combined s imple . P rono tum of same length and wid th , w i th d i s t inc t impre s s ions . E l y t r a ra ther broad, punctural interspaces f o r m i n g i r r e g u l a r , smooth, blunt protuberances; apex s imply round­ed epipleura reaching apex. Legs t yp i ca l ; I middle and hind t ib iae f inely s e r r a t e along inner edge. Sterna s i m p l e ; last abdominal segment s l igh t ly t runca te .

Ç. Much l a r g e r on the average; antennái segment X I shor te r than 3 preceding combin ­ed; eyes s m a l l e r , i n t e rocu la r distance broader ; p rono tum t r ansve r se ; a l l t ib iae unarmed .

Type species: T o m o g r i a pe r la ta sp. n . , by present designation. Remarks : On account of the p e a r l - l i k e tuberc les of the e l y t r a , this genus sharp ly dif­

fers f rom L a g r i a or any other New Guineán genera of L a g r i i n a . The pol ished, rounded pro­tuberances are much more evident when seen by naked eye.

T o m o g r i a per la ta sp. n. (Figs 19, 20, 23)

Head, p rono tum, s cu t e l l um, v e n t r a l surface and legs dark b lackish brown w i t h green­i sh -b ronzy t inge ; e l y t r a dark n u t - b r o w n tinged w i t h bronze; su tu ra i marg in black; antennae da rk brown. Leng th : 7. 1-10. 5 m m .

F igs 23-24. 23= T o m o g r i a pe r l a t a gen. et s p . n . , S*. - 24= A c u t o g r i a falcata gen. et sp .n . , 6*

S. Head coarse ly and densely punctate; w i t h sparse, moderate ly long h a i r s ; EI= 15:6; length ra t ios of antennái segments: 8: 5: 10: 9: 7: 8: 8: 8: 7. 5: 7: 34. P rono tum as long as broad o r ve ry weakly elongate; wides t at base, weakly n a r r o w i n g a n t e r i o r l y ; w i th 2 d i s t inc t , t r ans ­v e r s a l impress ions before base and behind a n t e r i o r m a r g i n ; ouflctation coarse and sparse,

disc w i t h smooth areas at the midd le ; pubescence modera te ly long, sparse , e rec t . E l y t r a wides t in apical 1/4; punctation coarse , unevenly sca t tered; interspaces f o r m i n g pol ished, s t rong ly r a i sed , rounded protuberances s t rewn i r r e g u l a r l y ; pubescence r a the r reduced, cons is t ing of fine s emie rec t to e rec t , scat tered ha i r s . Legs t y p i c a l ; t ibiae s t ra igh t ; middle and hind tibiae wi th a row of s m a l l denticles in apical half . Aedeagus: F igs 19-20. Habi tus: F i g . 23.

g. D i f f e r i ng as stated in the generic descr ip t ion ; EI= 15: 12; length r a t ios of antenna! segments V I I I to X I : 10: 10: 9: 23: 5.

Holotype, 8, l abe l led as fo l lows : "NEW GUINEA: (NE) Wau, Morobe D i s t r . Wau Ck. 1200- 1500 m , 28. I I I . 1963"; " J . Sedlacek Col lec to r BISHOP". I t is deposited in B P B M .

Paratypes (total 101 specimens) . Morobe P rov ince : labe l led as holotype (2 SS, 6 oo, B P B M ; 1 S, 1 a, H N H M ) ; i d . , 3 . I V . 1963, JS (2 gg, B P B M ) ; i d . , 16-18. I X . 1964, MS (To 1 , 2 gg, B P B M ; I S , 1 g, H N H M ex B P B M ) ; Bulo lo , 700 m , 26. X I . 1969, JS, MS (1 S, B P B M ) ; . B u l o l o R i v . , 1180 m , 17. D i . 1969, A . B . M i r z a (2 SS, B P B M ) ; Gara ina , R. Hornabrook (l8, H P C W ) ; U . Watut , SW, 1300-1600 m . I . V . 1963, J . L . G r e s s i t t (1 g, B P B M ) ; Watut R. , X ; 1969, 900-1900 m , A . B . M i r z a (1 g, B P B M ) ; Wau, 1150- 1700 m , 17. V I . 1961-8. V I I I . 1968, J . L . Gre s s i t t , H . F . Howdeh, P. Shanahan, JS, MS (5 SS, 15 oo, B P B M ; 1 S, CNCI ; 1 g, SPCB; 2 SS, 5 gg, H N H M ex B P B M ) ; Wau, B i g Wau Ck . , 1200 m , X I I . 1965, P. Shanahan (1 g, B P B M ) ; i d . , 1200-1500 m , D i . 1965, JS, MS (2 gg, B P B M ) ; Wau, Hosp i t a l Ck. , 1150-1250 m , 9.1.1966, JS, MS (1 g, B P B M ) ; Wau, K u j e r u , 1500 m , 27. D i . 1969, A . B . M i r z a (1 8, 7 og, B P B M ; 1 8, 2 gg, H N H M ex B P B M ) ; Wau, M c A d a m Park , 1200-1300 m . 7. V . 1965, JS (1 g, B P B M ) ; Wau, M t . K a i n d i , • 1500-2200 m , 19. V I I I . 1965. JS, MS (1 g, B P B M ) ; i d . , 1700, 8. V I I I . 1968. JS, MS (1 g, B P B M ) ; i d . , 2000 m , 27. V I I I . 1968 [ N G W - R . l l j , I . Loksa (1 o. H N H M ) ; Wau, M t . K a i n d i , N a m i Creek, 22. V I I I . 1968 [ N G W - C . 4.J , J . Balogh (1 g. H N H M ) ; K a i n d i - N a m i , 22. V I I I . 1968, JS (4 SS, 2 gg, B P B M ; 2 SS, H N H M ex B P B M ) ; Wau, N a m i Creek, 1600-1650 m , 24.11.1963, JS (1 g, B P B M ) ; i d . , 1700 m , 12. V I . 1962, JS (2 SS, 1 g, B P B M ) ; i d . , 1700 m , 15. V I . 1962, JS (2 SS, 2 gg, B P B M ; 1 g, H N H M ex B P B M ) ; i d . , 1700 m , 5. I I . 1963. JS (1 S, 1 g, SPCB); i d . , 1700 m , 22. V I I I . 1963, JS (1 g, B P B M ) ; - Wau, M t . M i s s i m , 22-24. I V . 1965, J . . Balogh et J . J . Szent-Ivány (1 S, 4 g g , H N H M ) ; i d . , 1500-1800 m , 6. 1.1970. JS, MS (2 gg, B P B M ) ; i d . , 1700 m , 7 . I I I . 1963, JS (1 g, B P B M ) ; 24-32 k m SE of Wau, 1500-1900 m , 20. I I I . 1962, JS (1 g, B P B M ) .

A c u t o g r i a gen. n .

Diagnosis : Elongate, modera te ly convex, med ium-s i zed ; co lour da rk ; pubescence s h o r t , semie rec t , sparse; head, p rono tum and e ly t r a w i t h d i s t inc t punctat ion.

8, Head wi th i n t e r o c u l a r distance n a r r o w e r than eye d iameter ; antennái segments I I to X s i m p l e , not t r ansve r se ; segment X I as long as 5 preceeding segments combined, curved , ab rup t ly attenuate in apical 1/3, apex acute. Pronotum s l igh t ly longer than broad, w i th v e r y shal low impress ions . E l y t r a modera te ly widening p o s t e r i o r l y ; surface roughly sculp tured; apex s imp le ; inner m a r g i n of epip leura reaching apex. Legs ra ther s t rong , moderately long; fore femora d i s t i nc t ly longer and s touter than middle and hind ones; fore t ibiae curved and twis t ed i n apica l 1/3, inner m a r g i n w i t h a l a rge , broad, t oo th - l i ke widening jus t before the midd l e ; inner m a r g i n of middle and hind t ibiae finely s e r r a t e subapical ly . Sterna s imp le ; last abdominal s te rn i te rounded at apex.

g. L a r g e r on the average; .antennái segment X I s i m p l e , a l i t t l e longer than 2 preceed-i ing segments combined; i n t e r o c u l a r distance broader; p ronotum t r ansve r se ; fore femora not d i s t i n c t l y longer and s touter than o thers ; fore t ib iae not modi f ied .

Type species: A c u t o g r i a falcata sp. n . , by present designation. R e m a r k s : The most sa l ient cha rac t e r i s t i c s of this genus are the b i l l - shaped , acute

last antennái segment and the uniquely fo rmed fore t ib iae of the males .

A c u t o g r i a falcata sp.; n . (Figs 2 1 , 22, 24)

Head, pronotum and scu te l lum dark b lackish green w i t h me ta l l i c sheen; e l y t r a shiny, da rk b lack ish brown; a l a r g e , common spot jus t behind middle of e l y t r a d a r k e r black (more evident on i m m a t u r e specimens) ; v e n t r a l surface, legs and antennae dark redd ish brown. Length : 8. 0-10. 5 m m .

S. Head coarse ly and densely punctate; w i th sparse, modera te ly long h a i r s ; eyes mod­era te ly convex; EI= 14: 6; antennái segment length r a t i o s : 11 : 5: 10: 8: 7: 7: 7: 7: 7: 6: 34; seg­ment X I , in p r o x i m a l 2 / 3 , nea r ly s t r a igh t , ' evenly wide , subquadrate i n cross sec t ion , then curved and na r rowed , w i th pointed apex. Pronotum s l igh t ly longer than broad, widest just an t e r io r to the middle ; sides evenly arcuate; surface less convex, w i t h 4 v e r y shallow i m ­press ions; punctation ve ry dense and coarse , punctures subcontiguous or even coalescing to t r a n s v e r s a l w r i n k l e s ; pubescence consis t ing of longer , spa r se r , e rec t and sho r t e r , denser decumbent ha i r s . E l y t r a widest i n apical 1/3; punctation coarse , punctures unevenly clus­t e red , leaving i r r e g u l a r interspaces as r a i sed pl icae; pubescence ra ther shor t , sparse, semie rec t , mixed wi th severa l e rec t h a i r s . Legs rather s t rong ; fore femora conspicuously stout; fore t ibiae nea r ly s t r a igh t in p r o x i m a l 2 /3 , then curved and twis ted ; inner ma rg in w i t h a l a rge , a s y m m e t r i c a l , t o o t h - l i k e widening just pos t e r io r to the midd le ; edge of this widening f inely crenula te ; middle t ib iae s l i gh t l y curved , d i s t a l ha l f of inner m a r g i n finely se r ra te ; hind t ib iae nea r ly s t r a igh t , d i s t a l 2/5 of inner m a r g i n w i t h fine dent icula t ion; a l l legs wi th sparse, modera te ly long, s emie rec t h a i r s . Aedeagus: F igs 21-22. Habi tus : F ig .24 .

g. L a r g e r on the average; eyes s m a l l e r , EI= 14: 12; r e l a t i ve lengths of antennái seg­ments V I I I to X I : 9 : 9 : 8 : 2 1 ; segment X I v e r y weakly cu rved , f u s i f o r m , p rono tum s l igh t ly t r ansverse , w i t h a mesal spot much f inely punctate (often obsolete): legs wi thout any modi ­f icat ions , i

Holotype, S, l abe l led as fo l lows : "NEW G U I N E A ; Wau, M c A d a m park, 18 -21 . I V . 1965"; " C o l l . D r . J . B A L O G H et D r . J . J . S Z E N T - I V Á N Y " I t is deposited in H N H M .

Paratypes (total 142 specimens) . Morobe Prov ince : l abe l led as holotype (2 g g . H N H M ) ; i d . , 1200- 1250 m , 20. V I I . 1965, JS (1 g, B P B M ) ; Bu lo lo , 1700 m , 26. X I . 1969, JS, MS (2 6*3 1 Q, B P B M ) ; Gara ina , 20. X I - 1 7 . X I I . 1969, A . B . M i r z a (1 o, B P B M ) ; i d . , R. Hornabrook (1 S, HPCW); Komba, L . Wagner (1 S, 1 g, SAMA) ; Lae, 4-6 . I X . 1968 [ N G L - C . 1 6 ] , J . Ba­logh (1 g , H N H M ) ; Sat telberg (1 p , Z M H U ) ; Watut R. , 900- 1900 m , X . 1969, A . B . M i r z a (2 SS, 3 gg, B P B M ; 1 6, 1 g, H N H M ex B P B M ) ; Wau, 1050- 1800 m , 29. V I I I . 1961-9. 11.1968, G . A . Samuelson, JS, JHS, MS, P. Shanahan (29 SS, 25 gg, B P B M ; 1 g, SPCB; 7 SS, 7 gg, H N H M ex B P B M ) ; Wau, Bu lo lo R. , 850-950 m , 4. I I . 1966, JS (1 Ó, B P B M ) ; i d . , 1130 m , 17. I X . 1969, A . B . M i r z a (3 SS, 3 gg, B P B M ; " 1 S, H N H M ex B P B M ) ; Wau, Hosp i t a l Ck. , 1150- 1250 m , 9. I . 1966, JS, MS (2 SS, B P B M ; 1 S, H N H M ex B P B M ) ; i d . , 1200 m , I . 1965, JS (2 SS, B P B M ) ; Wau, K u j e r u , 1500 m , 27. I X , 1969, A . B . M i r z a (3 SS, 4 gg, B P B M ; 1 S,[ 2 gg, H N H M ex B P B M ) ; Wau, M t . M i s s i m , 950- 1300 m , 8-9.1 .1966, JS, MS (1 g, B P B M ) ; ' i d . , 1120 m , 13.11.1963, JS (1 g, B P B M ) ; i d . , 1500- 1900 m , 6.1.1970, JS, MS (1 g, BPBM); Wau, Wau Ck. , 1200- 1500 m , 28. I I I . 1963, JS (1 g, B P B M ) ; i d . , 1200-1500 m , 3. I V . 1963, JS (1 S, B P B M ) ; i d . , 1200- 1500 m , 16-18. DC. 1964, MS (1 S, 2 go, B P B M ; 1 g . H N H M ex B P B M ) ; 6 k m W of Wau, N a m i Creek , 1700 m , 12. V I . 1962, JS (1 S, B P B M ) . - Madang P r o ­v ince : F i n i s t e r r e Range, Saidor , K i a m b a v i v i l l . , 1-28. V I I I . 1958, W . W . Brand t (1 S, B PB M l -Western Highlands P r o v i n c e : J i m i V a l l e y , Tabibuga, 23-24. I V . 1970, R. Hornabrook (1 <?, HPCW). - Chimbu P rov ince : Chimbu Va l l ey , B i s m a r c k Rge, 5-7500 f t . , X . 1944, Dar l ing ton (1 S, M C Z C ) . - Eas te rn Highlands P rov ince : Asa roka , 20 k m W Goroka , 20. I I I . 1973, K . W . , Stroeder (1 S, CNCI) ; Goroka , 1550 m , 19. V I . 1955, J . L . G r e s s i t t (1 g, B P B M ) ; i d . , 26. V I . 1955, J . L . G r e s s i t t (2 SS, B P B M ) ; i d . , 5400 f t . , 16.1.1956, J . H . B a r r e t t (1 S, B M N H ) ; K a r i m u i , 1080 m , 13. V I I . 1963, JS (2 SS, B P B M ) ; Kassam, 1350 m , 48 k m E of Kainantu, 7. I X . 1959, T . C . Maa (1 S, B P B M ) ; Lapegu, 15. V I . 1977, ex Pinus Patula, H . Roberts (1 ő7, B M N H ) ; Okapa, 3500 f t . , 6. LX_1965, R. Hornabrook (1 S, HPCW); Okapa, Okasa, 1400-1600 m , 17 .1 . 1966, JS (1 S, B P B M ) ; Purosa , 1700 m , 17-25. V . 1966, Gres s i t t et T a w i (2 SS, B P B M ; 1 S, H N H M ex B P B M ) : i d . , 1700-2000 m , 18.1.1966, JS (1 g, B P B M ) . - N o r t h e r n P rov ince : Iongai , 1700- 1900 m . 9. X I . 1965, JS (1 S, B P B M ) ; Kokoda, 400 m , 14-16. X I . 1965, JS (1 S, B P B M ) ; i d . , 18. X I . 1966, JS (1 S, B P B M ) .

i .ündilagria gen. n .

Diagnosis : F a i r l y elongate, i v e r y weakly convex, med ium-s i zed ; colour b rownish ; dor­sal surface wi th duplex pubescence; head, p ronotum and e l y t r a w i th d i s t inc t punctation.

S. Head wi th i n t e r o c u l a r distance much n a r r o w e r than eye d iameter ; eyes convex; an­tennae long and slender; no modif ied or t r ansve r sa l segments; segment X I cu rved , c y l i n d r i ­ca l , somewhat longer than 2 preceding segments combined. P rono tum n a r r o w e r than head.

not t r ansverse . E l y t r a elongate, p r a c t i c a l l y not widening p o s t e r i o r l y ; surface w i t h d i spers ­ed, low protuberances; apex s imp le ; inner m a r g i n of epipleura reaching apex. Legs long and s lender; inner m a r g i n of t ibiae p r a c t i c a l l y una rmed . T h o r a c a l and v i s ib l e abdominal s terna without outgrowths ; last (= fifth) abdominal s t e rn i t e emarginate ap ica l ly ; s t e rn i t e V I s t rong ly modi f ied , f o r m i n g 2 l o b i f o r m and 2 forceps-shaped processes.

<j>. Eyes much s m a l l e r ; i n t e r o c u l a r distance broader ; antennae shor te r ; segment X I shor t e r than 2 preceding segments combined; p rono tum s l igh t ly t ransverse ; abdominal s terna V and V I not modi f i ed .

Type species: K a i n d i l a g r i a forc ipa ta sp. n . , by present designation. R e m a r k s : T h i s genus may eas i ly be d is t inguished f r o m a l l other genera by the pecul ia r

s t ruc tu re of abdominal s te rn i te V I , the s t rongly elongate body and the papil late surface of e l y t r a .

Ka ind i l ag r i a forc ipata sp. n . (Figs 25-28)

Head, p rono tum, scu te l lum and e l y t r a brown w i t h bronzy t inge; head and p rono ta l disc somewhat d a r k e r ; v e n t r a l surface paler reddish b rown; antennae dark b rown, apex and base of segments I to X paler ; legs pale reddish b r o w n , femora somewhat darker subapical-l y ; dorsa l surface w i t h denser, v e n t r a l surface w i t h sparse r pubescence. Length: 7.5-8.5 m m .

25 26 27

Figs 25-27. 25= A b d o m i n a l apex of K a i n d i l a g r i a fo rc ipa ta gen. et sp. n . , S, i n somewhat drawn-out condi t ion. - 26-27= Aedeagus of K . forc ipata gen. et sp. n . , v e n t r a l view (26) and l a t e r a l v iew (27)

6*. Head t r ansve r se , much broader than p rono tum, densely and coarse ly punctate; with, sparse , long h a i r s ; eyes v e r y la rge , p rominen t ly convex; EI= 15: 6; antennae as long as 2/3 of body length; segment length r a t i o s : 16:7; 11: 11: 10: 11: 11.5: 11.5: 12: 11: 28; segment I longer than ha l f the distance between antennái sockets . P rono tum about of same length and w i d t h , widest at base, sides converging in s t r a igh t l ine , weakly cons t r i c ted jus t pos te r io r to the middle ; surface uneven, highest c e n t r a l l y ; punctures coarse , w e l l separated; pubescence

c o n s i s t i n g of a few s e m i e r e c t a n d m a n y l o n g e r , e r e c t h a i r s . E l y t r a v e r y n a r r o w , s u b p a r a l -l e l - s i d e d , f la t ; w i t h c o a r s e p u n c t u r e s s e p a r a t e d by d i s t a n c e s about a s b r o a d a s p u n c t u r e d i a m e t e r ; i n t e r s p a c e s l a r g e l y f lat but s u r f a c e s e t w i t h s t r a g g l i n g , f la t , s m o o t h p r o t u b e r ­a n c e s ; c l o t h e d w i t h s c a t t e r e d , v e r y f ine , m o d e r a t e l y l o n g , s e m i e r e c t p u b e s c e n c e m i x e d w i t h a n u m b e r of m u c h l o n g e r , e r e c t h a i r s . L e g s long and s l e n d e r , wi th long h a i r s ; a l l t i ­b iae n e a r l y s t r a i g h t ; m i d d l e a n d h i n d t i b i a e w i t h i n d i s t i n c t d e n t i c u l a t i o n a long i n n e r edge . A b d o m i n a l s t e r n i t e V w i d e l y n o t c h e d ; apex of m o d i f i e d s t e r n i t e V I and a e d e a g u s u s u a l l y v i s i b l e i n v e n t r a l a s p e c t ( F i g . 2 5 ) . A e d e a g u s : F i g s 2 6 - 2 7 . H a b i t u s : F i g . 28 .

F i g s 2 8 - 2 9 . 28= K a i n d i l a g r i a f o r c i p a t a gen. e t s p . n . , Ô*. - 29= C a s n o n i d e a h y s t r i x s p . n . , ^

g. N o t l a r g e r on the a v e r a g e ; e y e s m u c h s m a l l e r ; E I = 13: 11; a n t e n n á i s e g m e n t s much-s h o r t e r a n d s t o u t e r ; s e g m e n t I not l o n g e r than h a l f the d i s t a n c e between a n t e n n á i s o c k e t s ; r e l a t i v e l e n g t h s of s e g m e n t s V I I I to X I : 9 : 9 : 8 : 13; p r o n o t u m b r o a d e r ; a l l t i b i a e t o t a l l y u n ­a r m e d ; a b d o m i n a l s t e r n i t e V b r o a d l y r o u n d e d at a p e x .

H o l o t y p e , 3, l a b e l l e d a s f o l l o w s : " N E W G U I N E A : N E M t . K a i n d i 2 0 0 0 - 2 3 5 0 m . 1 0 - 1 1 . I . 1965"; " J . et M . S e d l a c e k C o l l e c t o r s B I S H O P " . I t i s d e p o s i t e d in B P B M .

P a r a t y p e s (total 9 s p e c i m e n s ) . M o r o b e P r o v i n c e : L a e , 17. V I I . 1970, H , O h l m u s (1 Ô*, A N I C ) ; M t . K a i n d i . 2350 m , 23 . I I I . 1966 , J . L . G r e s s i t t (1 ç , B P B M ) ; i d . , I I . V . 1 9 6 8 . J S

(1 o, B P B M ) ; i d . , 19.1.1970, JS (1 o, SPCB); M t . K a i n d i near Wau, 7500 f t . , 25. V I . 1974, H . F . Howden (1 S, CNCI) ; Wau, M t . K a i n d i , 24-25. V I I I . 1968 [ N G W - C . 8 . ] , J . Balogh (1 g, H N H M ) ; Wau, 1200 m , 8-14. X I I . 1976, m e r c u r y vapor l i gh t , G. F . Hevei et R. E . D ie tz (2 gg USNM). - Southern Highlands P rov ince : M u l l e r Plateau, Atea-gana-anda, V I I - V I I I . 1978, G. Smi th et a l . (1 g, A N I C ) .

Casnonidea h y s t r i x sp. n . (Figs 29, 32)

Head, p ronotum and scu te l lum y e l l o w i s h r e d , somet imes w i t h olivaceous t inge; e l y t r a reddish brown wi th expressed steely blue to olivaceous t i n t ; p ro - and mesos te rnum y e l l o w ­ish r e d , metas te rnum and abdomen redd ish b rown ; antennae and legs ye l l owi sh r e d ; las t an­tenna! segment and femora a l i t t l e da rke r . Length : 7. 8-9 m m .

S. Head not broader than p rono tum; w i t h some set igerous punctures between eyes; ve r t ex ha rd ly swol len ; EI= 12: 13; antennae r a the r s tout , segment length r a t i o s : 9 : 4 : 9 : 9 : 8 : 9 : 9 : 9 : 9 : 8 : 2 5 . P rono tum s l i gh t l y t r ansve r se , wides t i n an te r io r 1/3, base hard ly na r rower ; ' surface g labrous , copiously punctate; punctures separated by distances at least equal to puncture d i ame te r . E l y t r a w i t h a l l punc tura l rows reaching apex; s t r i a i punctures moder­ately coarse , contiguous; in t e rva l s f la t ; a l l i n t e rva l s w i th numerous (over 30) se t igerous punctures sca t tered throughout whole length; these punctures not a r ranged in r egu la r r o w s ; setae w h i t i s h , moderate ly long, somewhat b r i s t l e - l i k e ; e l y t r a l apex s i m p l y rounded. Legs wi th ha i r s shor t , suberect , f iner than e l y t r a l setae. Last abdominal s te rn i te s i m p l y rounded. Aedeagus: F i g . 32. Habi tus : F i g . 29.

g. Ve r t ex a l i t t l e m o r e swol len , long i tud ina l f u r r o w deeper; EI= 12: 13; r e l a t i ve lengths: of antennái segments V I I I to X I : 9 : 9 : 9 : 1 8 .

Holo type , S, l abe l led as fol lows : "NE W GUINEA : Haus Copper, Wau, M t . M i s s i m , 22-2 4 . I V . 1965"; " C o l l . D r . J . B A L O G H et D r . J . J . S Z E N T - I V Á N Y " . I t i s deposited in H N H M .

Paratypes (total 34 specimens) . Morobe P rov ince : label led as holotype (2 SS, 4 gg, H N H M ) ; Wau, 1100- 1800 m , 7. V I I I . 1961-30. V I I I . 1977, R. Hornabrook, R. P a r r o t t , JS, JHS, MS (10 St, 5 go, B P B M ; 1 S, C N C I ; 10, 1 g, HPCW; 2 SS, 1 g, SPCB; 2 SS, H N H M ex B P B M ) ; Wau. 8-14. X I I . 1976, m e r c u r y vapour l i g h t , G . F . Hevel , e t R . E . Die tz (2 SS, USNM); Wau, Coviak Rdg. , 763 m , 7. X I . 1963, H . C . (1 g, B P B M ) ; Wau, M t . K a i n d i , 4. V . 1971, R . E . P a r r o t t (1 S, CNCI) .

R e m a r k s : The numerous setae on a l l e l y t r a l i n t e r v a l s combined wi th the d i s t i n c t l y punc­tu red but e n t i r e l y glabrous pronotum are diagnostic for th is species.

Casnonidea punct i thorax sp. n . (Figs 30, 33)

Body usua l ly dark r edd i sh brown to black, w i t h head and pronotum and pronotum dark­e r , e l y t r a s l i gh t ly pa ler ; e l y t r a somet imes i r r e g u l a r l y mot t led wi th ochre; i n one case, p r o ­notum and basal 1/3 of e ly t r a pale ochre , res t of e l y t r a mot t led . Length: 9. 2-10. 1 m m .

S. Head not broader than pronotum; w i t h some set igerous punctures between eyes; ver tex moderate ly swol len , w i t h d i s t i nc t longi tudina l f u r r o w ; EI= 14: 12; antennae ra ther s lender , segment length r a t i o s : 10: 5: 11: 11 ; 11 : 13: 13: 14: 14: 14: 23. Pronotum s l igh t ly t rans­verse , widest in an te r io r 1/3, base near ly as wide; surface wi th few scat tered punctures (situated main ly l a t e r a l l y ) bear ing modera te ly long, w h i t i s h , semierec t setae. E l y t r a w i th a l l punc tura l rows reaching apex; s t r i a i punctures modera te ly coarse, contiguous; i n t e r v a l s less convex; a l l i n t e rva l s w i t h many (over 30) set igerous punctures sca t tered throughout whole length; these not a r ranged in two r o w s ; setae w h i t i s h , fine, moderate ly long; e l y t r a l apex s i m p l y rounded. Legs w i t h ha i rs r a t h e r shor t , nea r ly decumbent, fine. Last abdominal s te rn i te s i m p l y rounded. Aedeagus: F i g . 33. Habi tus : F i g . 30.

g. V e r t e x a l i t t l e more swol len ; EI= 15: 13; r e l a t i ve lengths of antenna! segments V I I I to X I : 14: 14: 13: 2 1 .

Holotype , t, label led as fo l lows : " N E W G U I N E A : NE K a r i m u i , 1080 m , 13. V I I . 1963"; " J . Sedlacek BISHOP". I t is deposited i n B P B M .

Paratypes (total 12 specimens) . Eas t e rn Highlands Prov ince : label led as holotype (4 SSt

2 gg, B P B M ; 1 S, H N H M ex B P B M ) ; i d . , 11-12. V I I . 1963 (1 o., B P B M ) ; i d . , 1973, R. H o r n a -

brook i l o, H P C W ) ; Okapa, I . 1968, R. Hornabrook (1 ç, HPCW). - Western Highlands : Ba iye r R i v e r , Sanctuary, 1 - 5 . I X . 1969 [ N G B - U . 2 2 . ] , J . Balogh (1 3, HNHM); i d . NGB-U. 2 i . (1 g, H N H M ) .

Remarks : The most cha rac t e r i s t i c feature of th is species is the sparsely punctate and pubescent p rono tum. Casnonides h y s t r i x also has the punctate p ronotum, but i n that spe­cies , the puntat ion is much denser, the puntures are not bear ing setae, the colour pa t te rn , the shape of body and the r e l a t i ve length of the last antennái segment of male are s t r i k i n g l y different .

F igs 3 0 - 3 1 . 30= Casnonidea punct i thorax sp. n . , 8. - 31= C. apical is sp. n . , 8

Casnonidea apica l is sp. n . (Figs 31 , 34)

Head ye l l ow , somet imes pa r t ly b lack i sh ; p ronotum ye l low; e l y t r a l a rge ly ye l low wi th apex black to a va r i ab le extent (usually f r o m l eve l of abdominal s te rn i te I V ) ; ep ip leura black­ish in pos t e r io r par t , a n t e r i o r l y to the l eve l of the middle of metas te rnum (somet imes this co loura t ion reaches e l y t r a l base); v e n t r a l surface ye l low o r meso- and metas te rnum black­i sh ; antennae black, segments I and I I y e l l o w i s h ; legs ye l low to b lackish . Length: 7. 8-8. 2 m m .

6*. Head not broader than p rono tum; w i t h two flat impress ions between eyes; ve r tex weakly swol len; EI= 12: 14; antennae r a t h e r stout, r e l a t ive length r a t ios of segments: 8:4: 9 : 9 : 9 : 1 0 : 1 0 : 1 0 : 1 0 : 9 : 2 0 . P rono tum t r ansve r se , widest in an te r io r 1/4, base much nar­r o w e r : surface impuncta te , g labrous . E l y t r a w i th a l l punc tura l rows reaching apex; s t r i a i punctures coarse , separated by distances shor te r than puncture d iamete r ; i n t e rva l s convex; pa i red in t e rva l s en t i r e l y glabrous; unpa i red in te rva l s w i t h set igerous punctures scat tered throughout whole length; i n t e r v a l I w i t h 17, I I I wi th 18, V w i t h 17, V I I w i t h 18, IX wi th 18, l a t e r a l ma rg in w i t h 12 set igerous punctures ; setae b rowni sh , modera te ly long; apex s imp ly rounded. Legs wi th ha i r s moderate ly long, suberect. Las t abdominal s t e rn i t e s i m p l y round­ed. Aedeagus: F i g . 34. Habi tus : F i g . 31 .

37

U 38 39 40

Figs 32-40. 32= Aedeagus of Casnonidea h y s t r i x sp. n . - 33= Aedeagus of C. punct i thorax sp. n . - 34= Aedeagus of C. apica l is sp. n. - 35= Aedeagus of C. flavipes sp. n . - 36 = Aedeagus of C. baloghi sp. n . - 37= Aedeagus of C. loksa i sp. n . - 38= Aedeagus of C. do-bodura sp. n . - 39= Aedeagus of C. pallens sp. n . - 40= Aedeagus of C. demet r ida sp. n.

g. Ve r t ex a l i t t l e more swol len ; EI= 12: 15; r e l a t ive lengths of antennái segments V I I I to X I : 10 :10 :10 :17 .

Holotype , Ô*, labe l led as fo l lows : " N E W GUINEA (NE) Wau, Mt . Ka ind i 24-25. V I I I . 1968"; " (No . N G W - C . 6. ) leg. D r . J . Ba logh" . I t is deposited in H N H M .

Paratypes (total 11 specimens) . Morobe Prov ince : B u l o l o - V a t u t , 700 m , 15-21 . V I . 1969. JS (1 S, B P B M ) ; Wau, 1050-1150 m , 13. X . 1961-7.1.1963, JS, JHS (4 Stf. 1 <j>, B P B M ; 1 6, 2 22> SPCB; 1 5, H N H M ex B P B M ; 1 3, H N H M ex SPCB).

R e m a r k s : Th i s species is s i m i l a r to Casnonidea v a r i a b i l i s (Macleay, 1887), but the colour pat tern of the e l y t r a c l ea r l y dis t inguishes i t .

Casnonidea f lavipes sp n . (Figs 35, 41)

Head, p ronotum, scu te l lum and v e n t r a l surface dark b lackish brown to black; e l y t r a b lackish b r o w n , r a r e l y un ico lo rous , usua l ly w i t h a va r i ab le number of i r r e g u l a r pale spots spots may p a r t l y be confluent and more numerous on humera l par t ; antennae brown wi th 2 basal segments somewhat redd i sh ; legs pale y e l l o w , often semi t rans lucen t . Length: 9. 5-11,2 m m .

S. Head not broader than pronotum; w i t h 2 shal low impress ions between eyes; EI= 15:16; antennae ra the r s lender ; segment length r a t i o s : 1 1 : 5 . 5 : 1 3 : 1 3 : 1 4 : 1 4 : 1 5 : 1 6 : 1 6 : 1 5 : 2 2 . Pronotum d i s t i nc t l y t r ansve r se , base and a n t e r i o r 1/4 of about same wid th ; surface gla­brous , impuncta te , shining, often wi th 2 ind i s t inc t l a t e r a l depress ions . E l y t r a w i t h a l l punc­t u r a l rows reaching apex; s t r i a i punctures moderate ly coarse , separated by distances at least equal to puncture d i ame te r ; in te rva l s weakly convex; pa i red in t e rva l s en t i r e ly gla­brous; unpai red in t e rva l s w i t h a few scat tered set igerous punctures; i n t e r v a l I w i th 1 sub-basal, I I I w i t h 6, V wi th 2, V I I w i t h 4, IX w i t h 5, l a t e r a l m a r g i n wi thout set igerous punc­tu res ; setae da rk , f ine, inconspicuous; e l y t r a l apex s imp ly rounded. Legs w i t h pubescence reduced; femora glabrous; t ib iae w i t h a few decumbent ha i r s along inner edge. Last ab­dominal s te rn i te s i m p l y rounded. Aedeagus: F i g . 35. Habi tus : F i g . 4 1 .

F igs 41-42. 41= Casnonidea flavipes sp. n . , Ô*. - 42= C. baloghi sp. n . , Z3

<j>. No shallow i m p r e s s i o n s between eyes but ve r tex m o r e swol len , longi tudinal fu r row more d i s t i n c t ; EI= 14: 17; r e l a t i ve lengths of antennái segments V I I I to X I : 16: 16: 15:20.

Holotype , 3, l abe l led as fo l lows : "NEW GUINEA (NE) Wau, 22. I X . -30 . I X . 1969. " ; "(No. N G W - U . 33. ) l eg . D r . J . Ba logh" . I t : i s deposited i n H N H M .

Paratypes (total 17 specimens) . Morobe P rov ince : Wau, 1200-1268 m , I . 1966-6. 1. 1971, B . B . L o w e r y , JS (1 g, A N I C ; 2 88, SPCB); Wau, B i g Wau Ck. , 1200-1500 m , I X . 1965., JS, MS (1 o_, B P B M ) ; Wau, Wau Ck. , 17. I X . 1964, JS (1 p, SPCB); 6 k m W of Wau, N a m i Creek , 1700 m , 12. V I . 1962, JS (1 8, B P B M ) ; i d . , 15. V I . 1962, JS (1 8, B P B M ) ; M t . K a i n d i , 1500-2200 m , 19. V I I I . 1965, JS, MS (1 p, B P B M ) ; M t . M i s s i m , 2000 m , JS (1 g, H N H M ex SPCB); Teptep, Huon Peninsula, 10. V I I . 1975, R . W . Hornabrook (1 o., HPCW). - Madang P r o v i n c e : F i n i s t e r r e M t s . , M o r o c. 5550 f t . , 30. X . -15 . X I . 1964, M . E . Bacchus (1 6, B M N H ) . - West­e r n Highlands Prov ince : M t . Hägen area , Ba iye r R i v e r , Sanctuary, 4 0 0 0 ' , 6 -11 . V I I . 1974, H . Howden (1 8, CNCI) . - Eastern Highlands P rov ince : Okapa, 17. X . 1964, at l i gh t , R. H o r ­nabrook (1 p, HPCW); i d . , 1. 1968, R. Hornabrook (1 8, H N H M ex H P C W ) ; i d . . 1800 m . 25. V I L , JS (1 3, SPCB); Purosa , 1700 m , 17-25. V . 1966, Gres s i t t et T a w i (1 g, B P B M ) . - Cen­t r a l P r o v i n c e : Owen Stanley Range, G o i l a l a , Borne, 1950 m , 16-30. I V . 1954, W . W . B r a n d t (1 o_, B P B M ) .

R e m a r k s : The most cha rac t e r i s t i c feature of this species is the pale yel low co lour which is i n sharp contrast to the b lack i sh res t of body. Specimens w i t h mot t led e l y t r a some­what resemble Casnonidea punct i thorax , but the p rono tum is glabrous and smooth (hairy and punctate i n punct i thorax) , only unpaired in t e rva l s have a s m a l l number of set igerous punc­tures ( a l l i n t e rva l s have a lo t of them in punct i thorax) and legs ye l low (dark brown i n punc­ t i t h o r a x ) .

Casnonidea baloghi sp. n . (Figs 36, 42)

Body dark blackish b rown . Length: 10 .5 -12 .2 m m . 3. Head not broader than p rono tum; w i t h 2 or 3 flat depressions between eyes; EI=

16: 16; antennae ra ther s lender ; segment length r a t i o s : 12: 6: 14: 12: 14: 14: 15: 15: 14: 13: 20. P r o n o t u m as long as broad o r a l i t t l e t r ansve r se ; surface impunctate , g labrous , sh in ing . E l y t r a w i t h most punctura l rows reaching apex but IV to IX d i s in tegra t ing p reap ica l ly ; most rows becoming f u r r o w - l i k e i n pos te r io r 1/3; s t r i a i punctures coarse , contiguous, i n l a t e r a l rows wide ly separated; i n t e rva l s weakly convex; pa i red in te rva l s e n t i r e l y glabrous; unpa i r ­ed i n t e r v a l s w i t h a few set igerous punctures ; i n t e r v a l I w i th 6, I I I w i t h 6, V wi th 7, V I I w i t h 4, IX., w i t h 5, l a t e r a l m a r g i n wi th 3 (preapical) set igerous punctures; setae pale, shor t ; e l y t r a l apex s i m p l y rounded. Legs w i t h few semierec t h a i r s . Las t abdominal s te rn i te s i m p l y rounded. Aedeagus: F i g . 36; Habi tus: F i g . 42.

o. Depressions between eyes v e r y s m a l l , punc tu re - l i ke ; EI= 18: 18; length ra t ios of antennái segments V I I I to X I : 16: 16: 14:22; p ronota l sides w i t h an ovoid , rugulosely punctate area.

Holo type , S, label led as fo l lows : " N E W GUINEA: NE Wau, 1150-1600 m , 9. I I . 1968"; " J . Sedlacek Co l l ec to r BISHOP". I t i s deposited in B P B M .

Paratypes (total 17 specimens) . Morobe Prov ince : labe l led as holotype (1 8, H N H M ex B P B M ) ; Wau, 1150-1550 m , 6. I I . 1963-27. DC. 1969. JS, MS, R. Hornabrook (2 3 8 , 1 p , B P B M ; 1 8 , HPCW; 4 8 8 , 2 oo., SPCB; 1 8, 1 o., H N H M ex SPCB); Wau, Haus Copper, M t . M i s s i m , 22-24 . I V . 1965, J . Balogh et J . J . Szent-Ivány (1 p, H N H M ) ; Wau, K u j e r u , 1500 m j 27. I X . 1969, A . B . M i r z a (1 p. B P B M ) ; Wau. Wau Ck. , 29. I I I . 1963, JS (1 3 , SPCB); i d . , I 1200-1500 m , 6. V . 1963, JS (1 Q_, B P B M ) .

R e m a r k s : The most c h a r a c t e r i s t i c feature of this species is the loose ly si tuated s t r i a i i punctures , ma in ly in the l a t e r a l r o w s . The rugulose p ronota l side of female is found in Cas-! nonidea loksa i as w e l l , but the combinat ion of unicolorous blackish body, the reduced e l y t r a l ves t i tu re and the fo rmat ion of the p reap ica l par t of e l y t r a l s t r iae r ead i ly d is t inguish C. ba- j l ogh i .

The species is dedicated to D r . J . B A L O G H (Zoosys temat ica l and Eco log i ca l InstituteT Eötvös Loránd U n i v e r s i t y , Budapest).

Casnonidea loksa i sp. n . (Figs 37, 43)

Head dark reddish b rown; pronotum and scu te l lum pale reddish b r o w n ; e l y t r a b lack i sh brown; v e n t r a l surface reddish brown; antennae and legs b rown, f emora l base and 2 basal segments of antennae somewhat paler . Length : 10 .2-10. 5 m m .

S. Head not broader than p rono tum; wi th 2 shallow depressions between eyes ; EI= 14: 14; ve r tex weakly swol len , longi tudinal f u r r o w faintly i m p r e s s e d ; antennae ra ther s lender ; segment length ra t ios : 1 1 : 5 : 1 3 : 1 2 : 1 3 : 1 2 : 1 4 : 1 4 : 1 5 : 1 4 : 2 1 . Pronotum a l i t t l e t rans ve r se ; widest in a n t e r i o r 1/3; surface glabrous, sh in ing , w i th v e r y ind is t inc t punctures on sides around a n t e r i o r angles. E l y t r a wi th a l l punc tu ra l rows reaching apex; rows s t rongly impres sed ; s t r i a ! punctures modera te ly coarse, contiguous; i n t e rva l s convex; pa i red inter-

Figs 43-44. 43= Casnonidea loksai sp. n . , S. - 44= C. pallens sp. n . , 5*

vais g labrous , except i n t e r v a l I I which bears 4 subbasal setae; unpai red in t e rva l s w i th sev­e r a l set igerous punctures sca t te red throughout whole length; these punctures ar ranged in 2 loose ser ies i n inner unpaired i n t e r v a l s ; i n t e r v a l I w i th 26, I I I w i t h 39, V w i t h 30, V I I wi th 20, IX wi th 25, l a t e r a l m a r g i n w i t h 32 set igerous punctures; setae pale, r a ther long; e l y t r a l apex s imply rounded. Legs w i t h moderately long , semie rec t h a i r s . Last abdominal s te rn i te s imply rounded. Aedeagus: F i g . 37. Habitus: F i g . 43.

<j>. Depress ions between eyes ve ry s m a l l ; ve r t ex s t rongly swol len , longi tud ina l fur row

deeply impressed ; EI= 15: 15; length r a t ios of antenna! segments V I I I to X I : 14: 15: 14: 19. 5; p rono tum coarse ly rugose l a t e r a l l y .

Holotype , 8, l abe l led as fo l lows : "NEW G U I N E A : NE Wau, 12- 1600 m 9 - I I 1968"; " J . Sedlacek C o l l e c t o r BISHOP". I t is deposited in B P B M .

Paratypes ( total 3 specimens): Morobe P rov ince : Wau, 8. I X . 1968 [ N G W - R . 1 9 ] , I . Loksa (1 o., H N H M ) ; i d . , 4000 f t . , 1 - 3. V I I . 1974, H . F . Howden (1 8, CNCI) . - Madang P r o ­v ince : F i n i s t e r r e M t s . , Damant i , 3500 f t . , 2 - 1 1 , X . 1964, M . E . Bacchus (1 c-, B M N H ) .

R e m a r k s : On account of the rugose pronota l sides of female, this species is somewhat re la ted to C. baloghi . The pronotum and v e n t r a l surface of C. loksai are reddish (black in baloghi) , setae of i n t e r v a l s are numerous and long (few and short in baloghi) , punc tura l rows n o r m a l (d is in tegra t ing before apex in baloghi) .

T h i s species is dedicated to D r . I . LOKSA (Zoosys temat ica l and E c o l o g i c a l Ins t i tu te , Eötvös Loránd U n i v e r s i t y , Budapest).

Casnonidea dobodura sp. n . (Figs 38, 45)

Body dark b lack ish brown; antennae, palps, t ib iae and t a r s i somewhat paler . Length: 10. 0- 10. 2 m m .

45 46 47

Figs 45-47. 45= Head of Casnonidea dobodura sp. n . , 8. - 46= E l y t r a l apex of C. caudata sp. n . , o.. - 47= A b d o m i n a l apex of C. caudata sp. n . , <j>, v e n t r o l a t e r a l v iew

8. Head (F ig . 45) not broader than p rono tum, l a r g e l y impunctate ; ve r tex moderate ly swol len , w i th long i tud ina l fu r row d i s t inc t ; EI= 18:7; antennae ra ther s lender; segment length r a t i o s : 1 0 : 5 : 1 1 : 1 1 : 1 2 : 1 3 : 1 2 : 1 2 : 1 3 : 1 2 : 2 3 . P rono tum near ly as long as broad, wides t in a n t e r i o r 1/3; base a l i t t l e n a r r o w e r ; surface g labrous , impunctate . E l y t r a w i t h a l l punc­t u r a l rows reaching apex; s t r i a i punctures r a the r coarse , contiguous; i n t e rva l s fa in t ly con­vex; pa i red in t e rva l s e n t i r e l y glabrous; unpai red in t e rva l s w i t h setigerous punctures scat­t e red throughout whole length; i n t e r v a l I w i t h 21 , I I I w i t h 19, V wi th 20, V I I w i t h 17, IX w i t h 16. l a t e r a l side w i t h 19 set igerous punctures; setae black, moderate ly long; e l y t r a l apex s i m p l y rounded. Legs w i t h ha i r s shor t , suberect , a l i t t l e f i ne r than e l y t r a l setae. Las t ab­dominal s te rn i te s i m p l y rounded. Aedeagus: F i g . 38.

g. EI= 18:8 (antenna! segments p a r t l y lost, on the s ingle female).

Holotype, Ó1, label led as fo l lows : "Dobodura , Papua, N . G . M a r - J u l y , 1944 Dar l ing ton" . I t is deposited i n M C Z C . i

Paratype. Western P rov ince : Kiunga, 28-30. V I I . 1969 [ N G K - U . 3 . ] , J . Balogh (1 o_, H N H M ) .

R e m a r k s : The ex t r eme ly n a r r o w in t e rocu l a r distance makes this species eas i ly d i s ­t inguishable . The male holotype f rom Dobodura (Nor thern Prov ince) has a l l body par ts i n ­tac t , though the e l y t r a l ve s t i t u r e has l a r g e l y been w o r n down. The female paratype lacks the left middle and hind legs as w e l l as a l l but left basal 4 antennái segments, wh i l e the e l y t r a l ve s t i t u r e is a lmost in tac t .

Casnonidea caudata sp. n . (Figs 46, 47)

Head, p rono tum, scu te l lum and v e n t r a l surface reddish ; e l y t r a black w i t h me ta l l i c g reen i sh -b lu i sh sheen; antennae and legs black, segments I and I I of antennae as w e l l as femora l base redd i sh . Length: 11.2 m m .

Ç. Head a l i t t l e n a r r o w e r than p rono tum; w i t h 2 s m a l l punctures between eyes; EI= 17: 17; ve r t ex s t rong ly swol len , w i th wel l -deve loped long i tud ina l fu r row; antennae ra the r stout; segment length r a t i o s : 1 1 : 5 : 1 1 : 1 3 : 1 3 : 1 3 : 1 4 : 1 4 : 1 5 : 1 3 : 2 2 . Pronotum weakly t r ans ­ve r se , widest at base; surface g labrous , impuncta te , sh in ing. E l y t r a wi th only punctura l rows I and X reaching apex, so e l y t r a w i t h a smooth p reap ica l area; rows v e r y weakly i m ­pressed; s t r i a i punctures minute , subcontiguous; i n t e rva l s broad and f la t ; p a i r e d in t e rva l s e n t i r e l y g labrous ; unpaired in t e rva l s w i t h a few scat tered set igerous punctures ; i n t e r v a l I w i t h 8, I I I w i th 5, V wi th 6, V I I w i t h 6, IX w i t h 6, l a t e r a l m a r g i n wi th 11 se t igerous punc­tu res ; setae black, shor t ; e l y t r a l apex bicuspidate , i . e . e l y t r a having an outer subrectan-gular and an inne r rounded angle ( F i g . 46). Legs w i t h ha i r s shor t , sparse, s e m i e r e c t . Las t abdominal s t e rn i t e w i th a process d i r ec t ed upwards (F ig . 47).

S. Unknown. Holotype, p , label led as fo l lows : " N E W GUINEA (NE) B a i y e r R i v e r Sanctuary 1-5. DC.

1969"; " (No. N G B - U . 24J leg. D r . J . Ba logh" . I t i s deposited in H N H M . R e m a r k s : Though known by the sole female specimen, Casnonidea caudata i s c l e a r l y

dis t inguished f r o m a l l other descr ibed congeners by i t s bicuspidate e l y t r a l apex and by hav­ing a process on the last abdominal s t e rn i t e . In other features i t much resembles C. s e m i - violacea P ic .

Casnonidea pallens sp. n . (Figs 39, 44)

Body pale chestnut in co lour ; e l y t r a somet imes wi th weak violaceous sheen, main ly i n basal ha l f or in humera l par t . Length: 8 .2 -10 .0 m m .

<?. Head not broader than p rono tum; wi th 2 flat punctures between eyes (sometimes confluent and f o r m i n g a t r ansve r sa l f u r r o w ) ; EI= 15: 15; antennae ra ther s lender ; segment length r a t i o s : 10: 5: 10: 10: 11 : 10: 10: 10 :9 :9 : 29. P rono tum as long as or a l i t t l e longer than broad, widest in an te r io r 1/4, base of same wid th ; surface en t i r e ly g labrous , impuncta te , shining. E l y t r a w i t h only punctura l rows I and X reaching apex, so e l y t r a w i t h a smooth preapica l area; s t r i a i punctures modera te ly coarse , contiguous; in t e rva l s f la t ; pa i red i n ­te rva l s e n t i r e l y glabrous; unpai red in t e rva l s w i t h ves t i tu re ex t remely reduced; i n t e r v a l I w i t h 1 subbasal, I I I w i th 1 subbasal and 2 p reap ica l , V wi thout , V I I wi th 2 subbasal , IX w i t h 2 p reap ica l , l a t e r a l m a r g i n without se t igerous punctures; setae dark, f ine, inconspicuous; e l y t r a l apex s i m p l y rounded; weakly caudate in do r so l a t e r a l aspect. Legs w i t h pubescence reduced; femora glabrous; t ibiae wi th f ine, decumbent ha i r s along inner edge. Las t abdom­ina l s te rn i te s i m p l y rounded. Aedeagus: F i g . 39. Habi tus : F i g . 44.

o.. V e r t e x a l i t t l e more swol len ; EI= 13: 15; length r a t ios of antennái segments V I I I to X I : 9 : 1 1 : 10: 22.

Holotype , S, l abe l led as fo l lows : " N . | Guinea: NE B u l o l o , 700 m 26. X I . 1969"; " J . et M . Sedlacek Co l l ec to r s BISHOP M U S E U M " , I t i s deposited in B P B M .

Paratypes (total 60 specimens) , Morobe P rov ince : labe l led as holotype (4 #<?, 4 PP, B P B M ; 1 Î , 1 p . H N H M ex B P B M ) ; B u l o l o , 700 m (1 S, SBCB); i d . , 6. X I . 1969, JS (3 &S, . 1 o_, B P B M ; 1 S, H N H M ex B P B M ) ; i d . , 1130 m , 17. I X . 1969, J . et M . Sedlacek (1 S, BPBM);

Pind iu , 20. I V . 1963, JS (1 3, B P B M ) ; i d . , 870-1300 m . 21-22. I V . 1963, JS (1 g, B P B M ) ; Watut R. , 900-1900 m . X . 1969, A . B . M I R Z A (1 8, B P B M ) ; Wau, 1050- 1300 m , 2. I X . 1 9 6 1 -X I I . 1968, H . Oh lmus , JS, JHS, MS (1 g, A N I C ; 9 88, 8 gg, B P B M ; 1 8, 2 gg, SPCB; 1 8, 3 gg, H N H M ex B P B M ) ; id . , 1200 m , 8-14. X I I . 1976, m e r c u r y vapor l i gh t , G. F . Heve i et i R. E . Die tz (4 Í3, USNM; 1 3, H N H M ex USNM); Wau, K i l o l o Creek . 26. V I I I . 1968 [ N G W - C j 14 . J , J . Balogh (1 g, H N H M ) ; Wau, M t . M i s s i m , 950-1300 m . 8-9 .1 .1966, J . et M . Sedla­cek (1 S, B P B M ) . - Madang P rov ince : Madang, 25. I X . 1970, R. Hornabrook (1 ô*. H P C W ) . -Eas te rn Highlands P rov ince : K a r i m u i , I I I . 1973, R. Hornabrook (1 S, HPCW); Okapa, 3500 f t . , 6 . I X . 1965, R. Hornabrook (1 8, HPCW). - Wes t e rn Highlands P rov ince : B a i y e r R i v e r . Sanctuary, 1 -5 . IX . 1969 [ N G B - U . 2 3 . ] , J . Balogh (1 g, H N H M ) . - C e n t r a l P rov ince : B i s i a -numu, E of Po r t Moresby , 500~m. 22. I X . 1955, C r o t a l a r i a f l o w e r s , J . L . Gre s s i t t (1 8, B P B M ) ; i d . , 2 3 . I X . 1955 (1 3, B P B M ) , - Western P r o v i n c e : O r i o m o Govt . Sta., 2 6 - 2 8 . X . 1960, shelf fungus, J . L . G r e s s i t t (1 g, B P B M ) . - N o r t h e r n P r o v i n c e : Dobodura, I I I - V I I , 1944. Da r l i ng ton (1 6, M C Z C ) .

R e m a r k s : Th i s species is r e a d i l y d is t inguishable f r o m the descr ibed congeners by the fo l lowing combina t ion of cha rac te r s : pale b rownish co lou r , smooth p reap ica l area of e l y t r a and the v e r y reduced e l y t r a l v e s t i t u r e .

Casnonidea demet r ida sp. n . (Figs 40, 48)

Dorsa l surface reddish b r o w n , often w i t h h u m e r a l par t of e l y t r a b lack ish ; this dark coloura t ion extending to middle of e l y t r a ; v e n t r a l surface reddish b r o w n , metas te rnum somet imes b lackish ; antennae b rowni sh , somewhat d a r k e r t oward apex; legs reddish b rown; d i s t a l half of femora and p r o x i m a l hal f of t ib iae b lack i sh (much more expressed on fore and middle legs) . Length: 7 . 5 - 9 . 0 m m .

3. Head a l i t t l e broader than pronotum; w i t h 3 s m a l l round pi ts between eyes; EI= 11 : 16; ve r t ex weakly swol l en , long i tud ina l fu r row i n d i s t i n c t ; antennae ra the r s lender; seg­ment length r a t i o s : 9: 4: 8: 9: 10: 10: 11 : 11 : 11: 11 : 17. P rono tum a l i t t l e longer than b road , widest in an t e r io r 1/4; surface impuncta te , sh in ing , l a rge ly glabrous but w i t h a few setae in an t e r io r angles. E l y t r a d i s t i n c t l y flattened; only punc tu ra l rows I and X reaching apex, so e l y t r a w i t h a smooth p reap ica l area; s t r i a i punctures modera te ly coarse , contiguous; in-j t e rva l s f la t ; pa i red in t e rva l s e n t i r e l y glabrous; unpai red in t e rva l s w i t h set igerous punctures scat tered throughout whole length; i n t e r v a l I w i t h 14, I I I w i t h 16, V w i t h 17, V I I w i t h 12, IX w i t h 13, l a t e r a l m a r g i n w i t h 19 set igerous punctures; setae black, f ine, ra ther long; e l y t r a l apex s i m p l y rounded. Legs w i t h pubescence f ine , r a t h e r long, semie rec t . Las t ab­domina l s t e rn i t e s imply rounded. Aedeagus: F i g . 40. Habi tus : F i g . 48.

g. Pi ts between eyes much s m a l l e r ; middle p i t t o t a l l y i n d i s t i n c t ; EI= 10: 14; length ra t ios of antennái segments V I I I to X I : 10: 11: 10: 15.

Holo type , 8, label led as fo l lows : Papua New Guinea, Wau, 19. I X . 1972., Malaise t r a p , leg . I - , Móczár". I t i s deposited in H N H M .

Paratypes (total 13 specimens) . Morobe P rov ince : B u l o l o R. , 1130 m , 17. DC. 1969, A . B . M i r z a (1 3. B P B M ) ; M t . K a i n d i , 1500-2200 m , 19. V I I I . 1965, J . et M . Sedlacek (1 o, B P B M ) ; Wau, 1100- 1400 m , 27. V I I I . 1961-4. I I . 1968, R. Hornabrook , JS, MS (2 33, 4 gg, B P B M ; 1 3, HPCW); Wau, M c A d a m park , 18-21 . I V . 1965, J . Balogh et J . J . Szent-Ivány ( 1 Í , H N H M ) ; Wau, Haus Copper , M t . M i s e i m , 22-24. I V . 1965, J . Balogh et J . J . Szent-Ivány (1 8, H N H M ) ; Wau, M t . M i s s i m , 1200 m,: 28. V I I I . 1968 [ N G W - C . l l ] , J . Balogh (1 g , H N H M ) ; 6 k m W of Wau, N a m i Creek , 1200 m , 15. V I . 1962, JS (1 g, B P B M ) . ,

R e m a r k s : Th i s species may be recognized by the fo l lowing combinat ion of charac te r s large but not v e r y prominent eyes, by which the head is broader than p rono tum; flat tened e l y t r a wi th smooth p reap ica l area; a number of soft, fine ha i r s i n unpai red e l y t r a l i n t e r ­va ls ; r a the r long, f rag i le legs w i t h modera te , s emie rec t pubescence; an t e r io r angles of pronotum w i t h few h a i r s .

Casnonidea setosa sp. n . (F ig . 49)

Body reddish b rown, metas te rnum and basal ha l f of e l y t r a a l i t t l e da rke r . Length: 8. 7 m m .

g. Head d i s t i nc t l y broader than pronotum; w i t h 2 confluent pits between eyes; eyes

conspicuously prominent ; EI= 13: 15; ver tex weakly swol len , long i tud ina l fu r row Ind i s t inc t ; antennae ra the r slender; segment length r a t i o s : 1 0 : 4 : 9 : 1 0 : 1 1 : 1 1 : 1 2 : 1 1 : 1 2 : 1 1 : 1 7 . P r o ­notum longer than broad, widest i n an t e r io r 1/3; surface impuncta te , sh in ing , w i t h a few setae in an t e r io r angles. E l y t r a d i s t i n c t l y f la t tened; a l l punc tura l rows reaching apex; s t r i a i punctures modera te ly coarse , not contiguous; i n t e r v a l s fa int ly convex; pa i red in t e rva l s en-t i r e l y a g l a b r o u s ; unpaired i n t e r v a l s w i th se t igerous punctures sca t te red throughout whole length; i n t e r v a l I w i th 12, I I I w i t h 2 1 , V w i t h 17, V I I wi th 18, I X w i t h 15, l a t e r a l m a r g i n w i t h 25 set igerous punctures; setae dark , f ine , v e r y long; e l y t r a l apex s imply rounded. Legs wi th conspicuously long, s e m i e r e c t to e rec t ha i r s . Last abdominal segment s i m p l y rounded. Habi tus : F i g . 49.

F ig s 48-49. 48= Casnonidea demet r ida sp. n . , <3. - 49= C. setosa sp. n . , <j>

S. Unknown. Holotype , o, label led as fo l lows : " N E W GUINEA (NE) B a i y e r R i v e r Sanctuary 1-5. D i .

1969"; " (No. NGB-U.24.) l eg . D r . J . Ba logh" . I t is deposited in H N H M . R e m a r k s : This species i s v e r y s i m i l a r to Casnonidea demet r ida but a l l the punctura l

rows of e l y t r a reach the apex (only I and X j i n demet r ida) . M o r e o v e r , the eyes are more prominent , the ha i rs on the e l y t r a and p a r t i c u l a r l y on the legs are v e r y long.

REFERENCES

B O R C H M A N N , F r . (1936): Coleoptera F a m . L a g r i i d a e . - I n : W Y T S M A N , P. (ed.): Genera Insec to rum, fasc. 204. Louis D e s m e t - V e r t e n e u i l , B r u x e l l e s , 561* pp.

M E R K L , O. (1988): O r e o g r i a gen. n . f r o m New Guina (Coleoptera, Tenebr ionidae : L a g r i i n i ) . Ac ta zool . hung. 34 (2-3) : 247-271. I

P IC, M . (1956): Nouveaux Coléoptères de d iverses f a m i l l e s . - Annls h i s t . - n a t . Mus . natn . hung. (ser. n.) 7: 71-92.

A u t h o r ' s address: D r . O. M E R K L Zoo log ica l Depar tment Hungar ian N a t u r a l H i s t o r y Museum Baross u . 13 H - 1088 Budapest HUNGARY