UNIVERSITY OF WISCONSIN
STEVENS POINT STEVENS POINT WISCONSIN
THE MUSEUM OF NATURAL
HISTORY ATLAS OF WISCONSIN BRYOPHYTES
Report No 16
Reports on the Fauna and Flora of Wisconsin
Patt 1
ATLAS OF WISCONSIN BRYOPHYTES
November 21 1979
THE MUSEUM OF NATURAL HISTORY Stevens Point Wisconsin
Reports on the Fauna and Flora of Wisconsin No 16
Part I
Climacium dendroides (Tree moss)
ATLAS OF WISCONSIN BRYOPHYTES
By Frank D Bowers
Curator of Non-Vascular Plants-Herbarium Museum of Natural History and
Department of Biology and
Sally Kauss Freckmann Research Associate
Museum of Natural History Universit y of Wisconsin
Stevens Point Wisconsin 54481 U S A
2
CONTENTS
ABSTRACT 4
ACKNOWLEDCEMENTS 4
INTRODUCTION 6
GEOGRAPHIC CLIMATIC AND BOTANICAL fEATURES OF WISCONSIN 7
SYSTEMATIC ARLNGEMENT OF WISCONSIN BRYOPHYTES 11
MAPS OF BRYOPH) iE TAXA REPORTED FOR WISCONSIN 29
HORNWORTS AND LIVERWORTS 30
MUSCI 54
FOSSIL AND SUBFOSSIL MOSSES OF WISCONSIN 119
PALEONTOLOGICAL Rb J3RENCES 119
SYNONYMS AND EXCLUDED TAXA 120
PIONEER BRYOLOGISTS OF WISCONSIN 126
LITERATURE CITED 127
3
ABSTRACf
117This atlas of Wisconsin bryophytes shows the distribution of)30 taxa (137 hepatics 389 mosses)
on state maps with a dot in each county for which each taxon has been reported in the literature A list of synonyms and excluded taxa is included along with an extensive bibliography Pertinent features of the natural history and history of bryology in the state are discussed
ACKNOWLEDGEMENTS
We want to thank Drs Fred Hermann and John Thomson of the University of Wisconsin-Madison herbarium and Botany department for locating and lending packets and for references to aid us in accumushylating a more accurate complete report The illustration of Climacium dendroides on the cover was drawn by Robert Freckmann Of course we are grateful to all the people who have collected specimens of Wisshyconsin bryophytes and preserved them in herbariums making this distributional study possible
4
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5
INTRODUCfION
This atlas of the bryophyte flora of Wisconsin is an attempt to show the distribution of all bryoshyphyte species known to occur in the state by county occurrenoes based on published reports in the literashyture It is not an annot~ted check list (only a feW problematic taxa were examined) but is a literature search intended as a foundation for a complete flora or for otherbryological research
The literature of btYo1ogy is large and spread over numerous journals reports monographs and manuals It would be almost impossible to search all sources for every reference attributing bryophyte species to Wisconsin and to particular localities in the state Wehav checked many inlportant references and paid special attention to works dealing with Wisconsiit botany and the bryologyof the upper Midwest Several bibliographies and indices were checked thoroughly including Green an~ Curtis (1955) the Bry ologist indices (Crum 1959 and Miller 1976) andt)le Wisconsin Academy of Sciences Arts and Letters indices
Only those published repots which app~red to)e substanti2ted by herbarium specimens and which gave- ~fficient location dat to P~CetheW(On in amiddot particula(county were-used in the initial compilation Species described in manuals in generalized way implying their occurrence in Wisconsin (Without menmiddot tioning Wisconsin specifically or citing a particular location in the state) were excluded eg Gnmmia raui (Wynne 1943) and Pohlia annotinll var decipiens (Crum 1976) Ecological papers which were either too general in taxonomic determinations (eg Sphagnum sp Polytrichum sp) or in location data were also excluded Some other taxa attributed to Wisconsin which have been excluded are
Cylindrothectum compr~ (Cheney 189tmiddotDane Co) identified later by Grout as Entodon seducshytrtx
Dicranell4 humilis (Holzjne~ 1925middotBuffalo Co) probablyD vllJia since Dhumilis is not found in North Arnerl~
Didymodon triarium (Foote 1 966-SW Visconsin) identified by Fred Hennann as Barbulaacuta Funoria microstoma (anonymous 1975middotDane Co) excluded because it is outside of its geographic
range and the packe~ QaMot be located Phil(motis calcarea (Cheney and Evans 1944-Dane Co) excluded in Crum Steere and Anderson
(1973) Plagiobryum demissum and P zierl (Cheney and Evans 1944-various counties) are both Arcticshy
alpine species not likely to be found in isconsin Two of Cheneys collections of Pl1giobyrum have been referred to Bryum
Pogonatum alpinum var septentrionale (Che~ey and Evans 1944-Chippewa Co) was misidentified (Ireland 1960) and is P umigerum
Rhizomnium fAPpoJachIanum and R punctalu1fl ssp chlorophyDosum (Koponen 1973middotseveral counshy ties) excluded North American bryologiflts have not recognized these taxa in America
Sphagnum tenellum (Cheney and Evans 1944-Waukesha Co)an Atlantic species not occurring in Wisconsin (Cruro 1975)
The nomenclature and systematic arrangement of hornwort 00 hepatic species in the atlas follow Stotler and Crandall-5totler (1977) and of moss species fonow Crum Steere and Anderson (1973) except in such instances as the JreptlIfockIdus fonns whiGh follow Wijk et aL (19S9middot1973) Species reported in the literature under names which are now listed as synonyms on either of these lists were converted to the accepted name A dot is placed in each county of Wisconsin from which the taxon has been reported The numbers after the name of the taxon correspond to the numbered references in the bibliography which giye ~act location data OEat least location data -s~fficient to place it in-particular counties in the state The numbers in parentheses indicate references which attribute the taxon to Wisconsin without providing exact loltation data
Fossil and subfossil reports are summarized Slfparately in our atlas only six species (Aulacomnium turgidum lJ1achythecium turgidum Ditrichum Jlelficaule Hypnum bambergti Scorpidium 6fJisities and Tortel14 1rIgi1i$) are known exclUsively from the fossil record ~
6
GEOGRAPHIC CLIMATIC AND BOTANICAL FEATURES OF WISCONSIN
Wisconsin is a north central state bounded on the east by Lake Michigan on the north by Michigan and Lake Superior on the south by Illinois and on the west by the Mississippi and St Croix rivers It is located midway between the Prime Meridian (0deg) of Longitude and 1800 Longitude and midway between the equator and the north pole thus placing it in the center of the northern half of the western hemisphere
The topography of the state is relatively flat a few hills exceed 1900 feet with Thns Hill at 1952 feet forming the highest point in Price County Lake Michigan represents the lowest point at 580 feet The northern and eastern two-thirds of the state were heavily glaciated during the last stage of the Pleistocene and now contain many of the best examples of continental glaciation topOgraphy including considerable ground moraine large terminal recessional and intetlobate moraines kames eskers drumlins and small depressions or kettle-holes Most of the nearly 9000 lakes of 20 acres or more lie within this part of the state The southwestern third of the state is the Driftless Area It includes most of the large flat generally sandy bed of Glacial Lake Wisconsin plus the highly dissected ridge and valley section with numerous cliffs and rock-outcroppings (Figure 1)
Much of the bedrock in the state lies below glacial till but exposures include igneous and metamiddot morphic rocks primarily granitic especially in the north as well as sedimentary layers primarily sandshystones and dolomitic limestones especially along Lake Michigan and in the DriftlesS Area
The climate of Wisconsin is classified as humid-temperate with cold winters The annual precipimiddot tation averages approximately 30 inches which is sufficiently even in its seasonal distribution to support forest vegetation throughout most of the state Nevertheless according to Curtis (I 959) the pre-settlement vegetation of the state (J 830-1850) included about 27 prairie and oak-savanna plus about 64 forests
The state is presently divided into seventy-two counties Marathon County is the largest (1586 sq mi) the smallest counties include Pepin (235 sq mi) Ozaukee (236 sq mi) andMilw8ukee (237 sq mi) We included Menominee County though its future status as a county is in doubt since it regained its Indian reservation status and rights in 1973 This county was established April 30 1961 from the Menomishynee Indian reservation which consisted of parts of Shawano and Oconto counties Wisconsin ranks 26th in the United States in area with 54464 square miles and 16th in population with 4651000
More ecological research is needed on the bryophyte flora of Wisconsin especially to relate it to the vegetation and to such other environmental features as soils climate and glaCial deposits Plant communishyties in Wisconsin are many and diverse these are summarized on Figure 2 from studies conducted by Curtis (1959) The state is divided into northern and southern floristic provinces by a tension zone deshyscribed by Curtis (1959) This tension zone is an eeotonal region vatying from 10middot31 mt wide where the ranges of many vascular plants overlap or terminatej it runs in an S-curve northeast from Milwaukee to Hudson in St Croix County (Figure 2) The principal plant communities of the northern province apshyproaching their southern boundaries within the zone include the northern conifermiddothardwood forests of sugar maple white pine hemlock and yellow birch and the Sphagnum bogs with tamarack and black spruce Southern communities at or near their northern limits within this zone include prairies oak and jack pine savannas and hardwood forests of maples oaks ash ehn and cottonwood
We hope to encourage research on cryptogamic ecology with our checklist We know that some species are northern while others are southern (eg cimacium) fits the tension zone concept Freckmann (I977) while Bryoxiphium is only found in the Driftless Area of the state Steere (1937) and Love and Love (1953) The tension zone theory may prove to be acceptable for bryophytes We speculate that certain bryophytes can indicate particular plant communities since many bryophytes are restricted to preshycise environmental requirements Many papers or theses have dealt with cryptogamic bark and rock ecology in Wisconsin (Culberson 1955 Foote 1962 Hale 1952 1955 and Cole 1975)
We hope that this atlas will stimulate collecting in counties poorly sampled so that our knowledge of the bryophytes can be increased It should aid anyone interested in floras for preservation environmental impact statements and distributional studies
7
A numerical summary of the bryophyte flora of the state based on our literature search is presented in Table 1 (below) This includes 517 taxa 70 families and 176 genera This numerical summary is comshypared with the published floras of Michigan mosses (Crum 1976) and Minnesota hepatics (Schuster 1953) Thus we surmise that further collecting could lead to the same approximate numbers for Wisconsin bryoshyphytes
Table 1 Comparison of Recorded Bryophytes of Wisconsin with Michigan and Minnesota and Summary of Wisconsin Taxa
Wisconsin Michigan (Crum 1976)
Minnesota (Schuster 1955)
Hepaticsamp Hornworts
Taxa Families Genera
137 28 47
174 32 52
Mosses
Taxa Families Genera
380 42
129
503 42
149
8
-
~
~ tIt
Figure 1 The Driftless Area and Tension Zone The counties are 01$0 shown on map
ca location of tension zone as deecfibed bv Curtis (1959)~ The Orittless Area
9
Agn 2 Major Plant Communitiee of Wsconsln aaed on surwvor recorda bull co 18laquo) After Curtismiddot 1959
tit beol Forest e Phillie
Northernmiddot hardwood forest reg Pine 5cMInna
copy Southernmiddot hordwood forest CD Oak Savanna
10
SYSTEMATIC ARRANGEMENT OF WISCONSIN BRYOPHYTES
ANTHOCEROTAE (Hornworts)
1 Anthocerotaceae Anthoceros 1 emnd Prosk
macounii M A Howe punctatum 1
Phaeoceros Prosk laevis (1) Prosk subsp laevis
Notothylas Sul1 orbicularis (Schwein) Sullo
HEPATICAE (liverworts)
2 PseudolepicoJeaceae Blephllrostoma (Dum emend Undb) Dum
trlchophyllum (L) Dum subsp trlchophylluni
3 Ptilidiaceae Ptilidium Nees
ci1iare (L) Hampe pulcherrimum (G Web) Hampe
4 Trichocoleaceae Trichocolea Dum nom cons
tomentella (Ehrh) Dum
5 Lepidoziaceae Kunia von Martens
setacea (Web) GroUe
Lepidozia (Dum) Dum reptans (1) Dum
Bazzania S Gray trilobata (1) S Gray var triZobata
6 Calypogejaceae 011ypogeja Raddi
flSsa (L) Raddi subsp issa integristipula Steph muelleriana (Schiffn) K Mull subsp muelleriana neesiana (Mass et Carest) K Mull sphagnicola (H Arnell et J Perss) Warnst et loeske trichomanis (1) Corda
11
7 Cephaloziaceae Cepluzlozla (Dum emend Schiffn) Dum
biscwpidata (L) Dum subsp biclupidIIta catenulllttl (Hub) Lindb coMivens (Dicks) Lindb var connivens lacinultzta Jack ampX Spruce lunulifolilz (Dum) Dum macounii (Aust) Aust pleniceps (Aust) Lindh var pleniceps
Noweilia Mitt cunifolilz (Dicks) Mitt
8 Adelanthaceae Odontoschimuz (Dwn) Dum
denudatum (Nees ampX Mart) Dum Vir denudatum mtlCount (Aust) Underw
c
9 Cephaloziellaceae CephtlJoziellll (Spruce) Schiffn nom C()II$
divarlctzt4 (Sm) sclurfn val divarlcat(l elachhta (bdc) Schiffn 1uImpetuw (Nees) Schirfn IUbellll (Nees) Warnst vat IUbeIl4rube (Nees) Warnst var biid4 (Lindb) Dquinrube (Nees) WarDSt Var sulliwmtii (Aust) K MuD
10 Geo~c~ larptuttllus Nees
scutatus (Web et Mohr) Spruce
Geocalyx Nees grGVeolens (Schrad) Nees
11 Lophocoleaceae Lophocolea (Dum) Dum
heteropllyll4 (Schrad) Own minorNees
Chiloscypbus Corda om cons pallescens (Ehrh ampX Hoffm) Dum varfragllis (Roth) K Mun polyantitos (L) Corda var poiyanthos pol)llaquo1lthos (L) Corda var rlvulllris (Schrld) Nees
12 PlagiochiJaceae Plagio(hila Dum nom cons
tlBpienioides (L) Dum var asplenioidu
13 Jungermanniaceae Btriilophoril Loeske
attenutlta (Mart) Loeske bIubtlta (Schmid ampX Scltreb) Loeske
12
Lophom (Dum) Dum alpestris (Schleich ~ Web) Evans ascendens (Warnst) Schust badenais (Gott ex Rabenh) Schiffn var bademii bierenata (Schmid ex H)ffm) Dum eollam (Nees) Dum exci8a (Dicks) Dum var excisa gllimanii (Aust) Schust varglllmanii guttulata (Lindb et H Arnell) Evans heterocolpos (Thed) M A Howe var heterocolpos ineistl (Schrad) Dum var tnelsa longidens (Lindb) Macoun subsplongidens ventricosa (Dicks) Dum var ventricosa ventricolfl (Dicks) Dum varlontifIotrl (Nees) Macoun ventricosa (Dicks) Dum var silvlcola (Bueh) Jones
Tritomarill Schiffn ex Loeske exseeta (Schrad) Loeske exsecttformi$ (Breidl) Loeske subsp exsectiormfs quinquedentata (Huds) Buch var quinquedentata scitula (rayl) Joerg
Anastrophyllum (Spruce) Schiffn hellemnum (Noes) Schust michtzuxii (Web) Buch ex Evans minutum (Schrob) Schust var minutum
Mylill S Gray nom cons anomala (Hook) S Gray
Jamesoniella (Spruce) Carring autumnalis (DC) Steph var autumnalis
Jungemumnia L emend Dum eonfertissimtt Nees crenuliformis Aust exsertijolill Steph subsp cordifolia (Dum) Vana graciUima Sm hyalina Lyell leillnthll Grolle polaris Lindb pumila With sphaerocarpa Hook
14 Scapaniaceae Diplophyllum (Dum) Dum nom cons
apiculatum (Evans) Steph var apiculatum
Scapanill (Dum) Dum nom cons apkuata Spruce carinthillca Jack ex Lindb curta (Mart) Dum var curta
13
cuspiduligera (Nees) K Mull var cuspiduiigera degenii Schlffn ex K Mull var degenii glllucocephaJa (Tayt) Aust gymnostomophila Kaal irrigwJ (Nees) Gott et al subsp irrigua lingulata Buch varlingulata mucronata Buch subsp mucronata nemorosa (l) Dum paludicola Toeske et K Mun var paludicola paludosa (K Mull) K Mull saxicola Schust scandiea (H Arnell et Buch) Macv var scandiea 8Ubalp~ (Nees) Dum umbroSll (SchIad) Dum undulata (L) Dum var undulata
15 Radulaceae Raduia Dum nom COllS
compianata (L) Dum obconica Sull
16 Porellaceae Parella L
pinTlllta L patyphylla (L) Pfeiff platyphylloidt(l (Schwein) lindb
17 Jubulaceae FruUanill Raddi
bolanderi Aust brittoniae Evans eboracemis Gott inlllta Gott oakesiana Aust plana Sull rlparia Hampe ex Lehm selwyniana Pears taTnl1iJci (L) Dum subsp asagmyana (Mont) Hatt
18 lejeuneaceae Calolejeunea (Spruce) Schlffn
biddlecomiae (Aust) Evans
Lejeunea Ub nom cons cavifolia (Ehrh) Lindb emend Buch
19 Codoniaceae Fossombronia Raddi
foveolata Lindb
20 Pa11aviciniaceae Paklvieinlil S Gray nom cons
lyeiii (Hook) Carruth
14
Moerckia Gott hibemica (Hook) Gott
21 PeWaceae Pellfa Raddi nom cons propos
endivlifolfa (Dicks) Dwn epiphylllz (L) Corda neeBiana (GottlUmpr
22 Metzgeriaceae Metzgerla Raddi
conjugata Undb
23 Blasiacae B L
pulilm L
24 Aneurareae AneumDum
pinguis (L) Dum
Riccfl1dia S Gray nom cons latifrons Undb multiftda (L) S Gray var multifida palmata (Hedw) Carruth
25 Aytoniaceae Reboulill Raddi nom conI
hemllphaerica (L) Radeli
ManniaOpiz fragrrms (Balbis) Frye el ClarK pilosa (Hornem) Frye et Clark trltmdra (Scop) GroDe
26 Conocephalaceae Conocephalum Wigg nom cons
conicum (L) Lindb
27 Marchantiaceae Preiuia Corda
quadrata (Scop) Nees
Marchantia L polymorpha L
28 Ricciaceae Riccwcarpus Corda
natans (L) Corda
Riccia 1 beyrichiana Hampe ex Lehm bifurca Horfm
15
i
CtlJPemostI Hoffnt fuitu L rostii Aust BOIOCtI1fHI Bi~ sulltvtmtii Aust
MUSCI (Mosses)
29 Sphagnaceae SpIutgnum L
capillaceum (Weiss) Schrank var Cllpil1tlceum caplllllceum (Weiss) Vir tenellum (Schimp) Andr centrille C Jens ex Amen et C Jens compactummiddot OC ex Lam et Dc ClUpidatum Ehrh ex Hoffin var aupidatum jimbrltltum Wits ex J Hookmiddot fulCUm (Scbimp) KUngrr glrgenBOhIlll Russ imbrlctltum Hornsch ex Russ muelllmitil4m BOd 1IIIIU (RUII) C Jens przIutre L papIIIoIum Lindb pulchrum (Lindb ex braithw) Wamst recumun P-Beauv var recumun recum P-Beauv var tenue Klinggr riptlrlum Ansstr ruPQWItWamst squtl1fOlUm Crome subftltens Russ et Wamst ex Wamst IUbIeCfUldum Nees ex Sturm var subsecundum subsecundum Nees ex Sturm varrutem (Nees Hormch et sUUm) Hueb teres (Sdlimp) Anptr e C Hartm
WtlI7IItorjii Russ wuljfanum Gira
30 Andreaeaceae AndNrrerI Hedw
rupeitrlr Oedw var pest
31 Fissident~e Fluid_ lledw
trdiantoid Hedw byoidei Bedw var bryolda cristaM Wils ex Mitt exiguus sun VIr exigUw onttmUi (B-Pyl) Steud jtuberi Lesq et James obtusifolhu Wits var obtwifoliul ollPlUlldQldes Heel ~litSuU lubbtulkril Bed tfDdfoliul lledw llridubu (S~ ) Wahlenb vallirlIU1u
16
32 Ditrichaceae Pleuridium Brid
subulatum (Hedw) Rabenh
Ditrichum Hampe heteromallum (Hedw) Britt linellre (Sw) lindb
~ pallidum (Hedw) Hampe pusiUum (Hedw) Hampe
Saelania Lindb gIlucescens (Hedw) Bomanss et Broth
Ceratodon Brid purpureus (He(lw) Brid var prupurew
Distichium BSG capillaceum (Hedw) BSG inclinatum (Hedw) BSG
33 Bryoxipbiaceae Bryoxlphium Mitt
norvegicum (Brid) Mitt
34 Seligeriaceae Seligeria BSG
cumpylopoda Kindb ex Macoun et Kindb pusillll (Hedw) BSG recurvata (Hedw) BSG
Bindia BSG acuta (Hedw) BSG
35 Dicranaceae BlUehia Schwegr
lexUOSll (Sw ~ Schwaegr) C MueU sullivantii Aust
Trematodon Michx ambiguus (Hedw) Hornsch
Dicranellll (C Muell) Schimp cerviculata (Hedw) Schimp val cillPicuiata
heteromalla (Hedw) Schimp ~ rufescens (With) Schimp
schreberiana (Hedw) Schhnp var schreberiana schreberitJna (Hedw) Schimp var robusta Schimp ex Braithw subulata (Hedw) Schimp varia (Hedw) Schimp
Dicranodontium BSG denudatum (Brid) Britt ex Williams
Panzeucobryum (Lindh) Loeske Iongifolium (Hedw) oelke varlongifolium
17
Tortella (Undb) Limpr fragilis (Drumm) Umpr humilis (Hedw) JeM torlUoSfl (Hedw) Umpr
Hyophila Brid involuta (Hoole) Jaeg et Sauerb
Didymodon Hedw rigidulus Hedw
BryoerythrophyUum Chen recurvirostnon (Hedw) Chen
Btubula Hedw acula (Brid) Brid var acuta convoiuta Hedw fallox Hedw unguicultltll Hedw
Acaulon C MueD muticum (Uedw) C Muell var rufeseens (Iaeg) Crum
PJl~lm Hedw CUSPidtltum Hedw
De8mlltodon Brid cernuu (Hueb) BSG oblUsifoliw (Schwaegr) Schimp
T01IUla Hedw mueronifolia Schwaegr rurall (Hedw) Gaertn Meyer et Scherb var ruralis
39 Grimmiaceae Grlmmill Hedw
alpicola Hedw var alpicola alpicola Hedw var dupretii (Ther) Crum alpicola Hedw var rlvularl8 (Brid) Wahlenb apocarpa Hedw var apocarpa apocarpa Hedw var ambipa (Sull) Jonea apocarpa Hedw var conjerttl (Funck) Spreng apocarpa Hedw var strleta (Tum) Hook et Tayl CiIIyptrrlta Hook ex Drumm donniana Sm var donn_ hemumti Crum ltlevla (Brid) Brid montana BSG oineyi SuD oVllll8 (Hedw) lindb piUfe P-Beauv pltJgiopodill Hedw pulvinata (Hedw) Sm varbullpulvinata teretinervl Limpr
18
Orothera C Muell leana (Sull) C Muell
Rfwbdoweisia crispata (With) Undb
Cynodontium BSG ex Schimp alpestre (Wahlenb) Milde strumiferum (Hedw) Undb
Dichodontium Schimp pellucidum (Hedw) Schimp
Oncophorus (Slid) Brid vuens (Hedw) Brid wahlenbergii (Brid)
Dicranum Hedw bonjeanii DeNot ex Usa drummondii C Muell lagellare Hedw varj1ogellare fulvum Hook fuscescens Turn var fuscescens montanum Hedw muehlenbeckii BSG var muehlenbeckii polysetum Sw sabuletorum Ren et Card scoparium Hedw undulatum Brid viride (Sull et Lesq ex Sull) lindb
36 Leucobryaceae Leucobryum Hampe
glaucum (Hedw) Angstr ex Fr
37 Encalyptaceae Encalypta Hedw
ciliata Hedw var ciliata procera Bruch
38 Pottiaceae Astomum Hampe
muhlenbergianum (Sw) Grout
Weissia Hedw controversa Hedw var controversa
Gymnostomum Nees Homsch et Stunn aeruginosum Sm recurvirostrum Hedw var recurvirostrum
Trichostomum Bruch spirale Prout
19
40 Ephemeraceae Ephemerum Hampe
cohaerens (Hedw) Hampe crassineium (Schwaegr) Hampe val crassinervium spinulosum Bruch et Schimp ex Schimp
41 Funariaceae AphflllolThegma SuD
serratum (1 Hook et Wils ex Drumm) Sullo
Physcomitrium (Brid) Fuernr hookeri Hampe immersum Sullo pyriforme (Hedw) Hampe
Funaria Hedw americana Undb hygrometrica Hedw vaT hygrometrica
42 Splachnaceae Splachnum Hedw
ampullaceum Hedw
43 Schistostegaceae Schistostega Mohr
pennata (Hedw) Web el Mohr
44 Bryaceae Pohlia Hedw
annolina (Hedw) Lindb var annotina camea (Tum) Lindb cruda (Hedw) Undb elongata Hedw iliformis (Dicks) Andr lescurlana (Sull) Grout nutans (Hedw) Lindb obrusifolia (Brid) L Koch proligera (Undb ex Umpr) Undb ex Amell wahlenbergii (Web et Mohr) Andr
Leptobryum (B S G) Wits pyriforme (Hedw) Wils
Bryum Hedw algovicum Sendtnex C Muell argenteum Hedw caespiticium Hedw capillare Hedw var capillare creberrlmum Tayt lonchocaulon C Muell pallens (Brid) Sw ex Roehl pseudotrlquetrum (Hedw) Gaertn Meyer et Scherb varpudotriquetrum
20
pseudotriquetrum (Hedw) GaertnMeyer et Scherb var bimum (Schreb) Ulj stenotrichum C Muell var stenotrichum tortiolium Funck ex Brid uliginosum (Brid) B S G violaceum Crundw et Nyh weigelii Spreng
Rhodobryum (Schimp) Hampe roseum (Hedw) Limpr
45 Mniaceae Mnium Hedw
ciliare (C Muell) Schimp cinclidioides Hueb cuspidarum Hedw drummondii Bruch et Schimp longirostrum Brid lycopodioides Schwaegr marginatum (With) Brid ex P Beauv mediumB~S G var medium onhorrhynchum Brid pseudopunctatum Bruch et Schimp punctalUm Hedw var punctatum punctatum Hedw var elatum Schirnp rugicum Laur spinufosum B S G stellare Hedw
46 Aulacomruaceae Au1acomnium Schwaegr
androgynum (Hedw) Schwaegr heterostichum (Hedw) B S G paJustre (Hedw) Schwaegr var paJustre
47 Meesiaceae Amblyodon B S G
deillbotus (Hedw) B S G
Meesia Hedw triquetra (Richt) Angstr uJiginosa Hedw
48 Bartramiaceae Plagiopus Brid
oederiana (Sw) Umpr
Bartramia Hedw pomiformis Hedw var pomiformis
Philonotis Brid fontana (Hedw) Brid varfontana longseta (Michx) Britt varlongseta marchica (Hedw) Brid
21
~
~
49 Timmiaceae Timmia Hedw
megapolitana Hedw var megapolitana
50 Orthotrichaceae Zygodon Hook et Tayl
viridissimus (Dicks) Brid var rupestris lindb ex C J Hartm
Amphidium Schimp lapponicum (Hedw) Schimp mougeotii (B S G) Schimp
Orthotrichum Hedw anomalum Hedw obtuaifolium Brid ohioense Sullo et Lesq ex Aust pumilumSw sordidum Sullo et Lesq ex Aust speciosum Neesax Sturm var eegam (Schwaegr ex Hook et Grev) W~rnst stellDtwn Brid strrmgulatum P Beauv
Uleta Mohr ex Web cOfIICtata (p Beallv) Hamrn crispa (Hedw) Brid
Drummondill Hook ex Drumm prorepens (Hedw) Brltt
51 Fontinalaceae Fantinalis Hedw
antipyretica Hedw var antipyrettca antipyretica Hedw forma rabusta (Card) Card dalecarlica Schimp ex B S G var dalecarlica flqccidtJ Ren et Card hypnoides C J Hartrn var hypnoides hypnoides C J Hartm var durillei (Schimp) HU$ll
missourlcaCard novae-angliae Sullo var novae-angliae splulgrUlolill (C Moen) Wijket M~S
DichelY1l1ll Myr capilloceum (With) Myr falcatum (Bedw) Myr HIllescens B S G
52 Climaciaceae aimecium Web et Mohr ex Mohr
americllnum Brid dedroides (Hedw) Web et Mohr
22
S3 Hedwigiaceae Hedwigia P Beauv
ciliata (Hedw) P Beauv
54 Cryphaeaceae Forsstroemia lindh
ohioense (Sull) lindh
55 Leucodontaceae Leucodon Schwaegr
brachypus Blid var brachypus brachypus Brid var aniJrewsitlnus Crum et Anderson julaceus (Hedw) SuU
56 Neckeraceae Neckera Hedw
pennata Hedw var pennata
1fomalia (Brid) B S G trichomanoides (Hedw) B S G
57 Theliaceae Thelia Sull
asprellll Sullo hirtella (Hedw) Sullo lescurii SuU
Myurella B SG julacea (Schwaegr) B S G
sibrica (C MueU) Reim var sibrica
58 Fabroniaceae Fabronio Raddi
ciliaris (Brid) Bnd
Anacamptodon Brid splachnoides (Groet ex Brid) Brid
Schwetschkeopsis Broth fabronia (Schwaegr) Broth
59 Leskeaceae Lindbergia Kindb
brachyptera (Mitt) Kindb
Leskea Hedw gracilescens Hedw obscura Hedw poiycarpa Hedw
Leskeella (limpr) Loeske nervosa (Brid) Loeske
23
Pterlgy1lllndrum Hedw filiforme Hedw
60 Thuidiaceae Haplohymenfum Dozy et MoIk
trlste (Ces ex De Not) Kiodb
Anomodon Hook et Tayl attenuatus (Hedw) Hueb minor (Hedw) Fuemr rostratus (Hedw) Schimp viticulosus (Hedw) Hook et Tayl
Haplocladium (C Mucn) C MueU microphyllum (Hedw) Broth virginillnum (Brld) Broth ~bull
Thuidium B S G abietinum (Hedw) B S G delicatulum (Hedw) B S G vardelicatulum delicatulum (Hedw) B S G var radicans (Kindb) Crum Steere et Anderson minutulum (Hedw) B S G pygmaeum B S G recognftum (Hedw) Lindb scitum (p Beauv) Aust
Helodium (Sull) Warnst blandowli (Web et Mohr) Warnst var blandowli blandowli (Web et Mohr) Wamst var helodioides (Ren et Card ex Roell) Crum
Steere et Anderson paludomm (Sull) AuS
61 Amblystegiaceae Cratoneuron (Sull) Spruce
commulatum (Hedw) Roth var commutfltum commutatum (Hedw) Roth var fakatum (Brid) Moenk ilicinum (Hedw) Spruce var filcinum
Campylium (Sun) Mitt chrysophyllum (Brid) 1 Lange var chrysophyllum hispidulum (Brid) Mitt var hllJidulum polygamum (B S G)C lens varpolygtlmUm stellatum (Hedw) C1ens vat stelltum
Leptodictyum (Schimp) Warnst laxirete (Card et Ther) Broth riparium (Hedw) Wamat trichopodium (Schultz) Warns l1acillans (Sull) Broth
Hygroamblystegium Loelke jlul1iatile (Hedw) oeske var luViatile
24
fluviatile (Hedw) Loeske var oWltum Grout noterophilum (Sun et Lesq ex Sull) Warnst tenox (Hedw) Jenn var tenax tenox (Hedw) Jenn var spinilolium Schimp) Jenn
Amblystegium BSG juratzkanum Schimp serpens (Hedw) BSG varlum (Hedw) Undb var varium
Platydictya Berk confervoides (Brid) Crum jungermanniodes (Brid) Crum minutissimum (Sull et Lesq ex Sull) Crum subtile (Hedw) Crum
Drepanocladus (C Muell) Roth aduncus (Hedw) Warnst var aduncus aduncus (Hedw) Warnst var capillilolius (Wamst)Riebm aduncus (Hedw) Warnst var kneiffii (BSG) Moenk aduncus (Hedw) Warnst var polycorpus (Bland ex Voit) Roth exannulatus (BSG) Warnst fluitans (Hedw) Warnst var fluitans revolvens (Sw) Wamst var revolvens sendtneri (Schimp) Warnst var sendtnerl uncinatus (Hedw) Warnst var uncinatus uncinatus ffiedw) Warnst var piwnosus (Schimp) Wamst vernicoms (Lindb ex C Hartm) Warnst
Hygrohypnum Lindb eugyrium (BSG) loeske luridum (Hedw Jenn varluridum ochraecum (Tum ex Wils) Loeske var ochraceunt
Calliergon (Sull) Kindb cordilolium Hedw Kindb glganteum (Schimp) Kindb var giganteum richardsonii (Mitt) Kindb ex Wamst stramineum (Brid) Kindb trilarium (Web et Mohr) Kindb
Calliergonella Loeske cuspidata (Hedw) Loeske
Calliergidium (Ren) Grout bakeri (Ren) Grout
Scorpidium (Schimp) Urnpr scorpiodides (Hedw) limpr
62 Brachytbeciaceae Tomenthypnum loeske
nitens (Hedw) Loeske var mtens
25
Homalotheciella (Card) Broth subcapillata (Hedw) Broth
Brachythecium BSG acuminatum (Hedw) Awt acutum (Mitt) Sullo calcareum IGndb campestTe (C MuelL) BSG cunum (Lindb) Umpr digastrum C Muell et Kindb ex Macoun et Kindb oxycladon (Brid) Jaeg et Sauerb var oxycladon plumosum (Hedw) BSG populeum (Hedw) BSG var populeum reflexum (Starke ex Web et Mohr) DsG var relexum reiexum (Starke ex Web et Mohr) DSG var pacificum Renet Card ex Roell rlvulare BSG var rivulare tabulum (Hedw) BSC var tabuum salebrosum (Web et Mohr) BSG var salebrosum velutinum (Hedw) BSG Vat velutinum
Bryhniil Kaur graminicolor (Brid) Grout var graminicolor novae-angliae (Sull et Lesq ex sun) Grout
Rhynchostegium BSG senulatum (Hedw) Ja4lg et Sauerb
Rhynchostegiella (BSG) Limpr compacta (C MueU) Loeske
EU1hynchium 8SG hians (Hedw) Sande Lac pulchellum (Hedw) Jenn var pulchelium riparloides (Hedw) Rich
Stokeriella (Kindb) Robins pmelonga (Hedw) Robins var stokesii (Tum) Crum
63 Entodontaceae Entodon C Muell
brevisetus (Hook et Wils ex Wils) Lindb cladorrhizans (Hedw) C Muell seductrlx (Hedw) C MueU var wluctrix
Pleurozium Mitt schreberl (8rid) Mitt var schreberi
64 Plagiotheciaceae Pltlglothecium BSG
CflVifolium (Brid) Iwats var CtWifolium denticulatum (Hedw) 8SG laetum BSG ltztebricola BSG
26
6S SernatophyUaceae Brotherella Loeske ex Fleisch
recumins (Michx) Fleisch tenuirostris (Bruch et Schimp ex Sull) Fleisch
Sematophyllum Mitt demissum (Wils) Mitt
66 Hypnaceae Pl(itygyrium BSG
repens (Brid) BSG
PylaisieUa Kindb intricata (Hedw) Grout polyantha (Hedw) Grout selwynii (Kindb) Crum Steere et Anderson
Homomallium (Schimp) Loeske adnatum (Hedw) Broth
Hypnum Hedw cupressiforme Hedw var cupressiforme cupressifonne Hedw varfiliforme Brid curviolium Hedw fertile Sendtn imponens Hedw lindbergii Mitt palescens (Hedw) P-Beauv pratense Koch ex Brid
Calicladium Crum haldanianum (Grev) Crum
sopterygiopsis Iwats muelleriana (Schimp) Iwats
Isopterygium Mitt e1egans (Brid) lindb pulchellum (Hedw) Jaeg et Sauerb tenerum (Sw) Mitt
Taxiphyllum Fleisch depllQtum (Bruch et Schimp ex sun) Fleisch taxirameum (MitL) Fleisch
Herzogiella Broth seligeri (Brid) Iwats striatella (Brid) Iwats ruracea (Lindh) Iwats
hilium De Not crista-castrenna (Hedw) De Not
27
67 Rhytidiaceae Rhytidium (Sull) Kindb
rugosum (Hedw) Kindb
Rhytidfodelphus (Lindb ex Iimpr) Warnst triquetrus (Hedw) Warnst var triquetrus
68 Hylocomiaceae Hylocomium BSG
pyrenaicum (Spruce) lindb splendens (Hedw) BSG varsplendens
69 Buxbaumiaceae Buxbaumia Hedw
aphyllll Hedw
Diphyscium Mohr follosum (Hedw) Mohr
70 Tetraphidaceae Tetraphi3
pellucida Hedw
71 Polytrichaceae Atrichum PmiddotBeauv
angustatum (Brid) BSG cri8pum (James) Sullo oerstedianum (C Muell) Mitt teneUum (Roehl) BSG undulatum (Hedw) P middotBeauv var unduiatum
Pogonatum P-Beauv alpinum (Hedw) Roehl var alpinum umigerum (Hedw) P-Beauv
Poiytrichum Hedw commune Hedw var commune ormosum Hedw juniperinum Hedw longisetum Brid ohioense Ren et Card piliferum Hedw var pilierum strictum Brid
28
MAPS OF BRYOPHYTE TAXA
REPORTED FOR WISCONSIN
HORNWORTS AND LIVERWORTS
29
bull bull WSHeV~N S vE II
0 ~
e~RRON ~USK
(J~ (j -
CHIPPE A DUNN
0 o0 poundAU CLAIRE
~ TREMshyPEAshyLEAU
MICE
ONEIDA
Q LINCOLN
TAYLOR
Q t-~----- MOIRA THON
CLARK
o
10
LA FAVETTE
Hornwor ts amp Liverworts
- None - 1 - 25 taxa
- 26 - 50 tRXshy
- 51 - 75 tax)
- 7c - 100 taxa
FONDOU j LAC ~
DODGE
i O JEfFERSON
WALWORTH
30
Anastrophylltlfl helleranum (Nees) Schust Anastrophyllum michauxil (We~ Buch ~
46 88 136 141 Evans 38 46 141
Anastrophyllum mlnut~ (Schreb) Schuster Aneura plnguls (L) Durn
89 21 384687 88 89 90 (138) (145)
Anthoceros macounil MA Howe ~oceros punctatus L
21 (45) (144) 34
31
Barbilophozia attenuata (Mart) Loeske Barbilophozia barbata (Schmid ~ Schre~
24 141 Loeske 28 39 46 89 90 141
Bazzania trilobata (1) S Gray
1 26 39 46 (57) (59) 79 88 89
90 119 141
BlePbarostoma trichoffiyllum (1) Dumort
1 8 26 39 46 89 90 137 141
Blasia pusilla 1
1 39 46 90
Calypogeja fissa (1) Raddi
90
32
$129geja integristipula Steph C~Ypound2geja muelleriana (Schifm) K Mull
88 89 89
CallpoundO~eja neesiana (Mass amp Carest) K Mull
39 46 89 140 (141)
~pogeja sJlapound2icola (H Arnell tl J bull
Perss) Warnst et Loeske (119) 141
CeIalozia biscuspidata (L) Dum
1 37 I 46 90 t 137 141
CalYpogej~ trichomanis (L) Corda
26 39 (119) 137
33
CeEhalozia catenulata (Hub) Lindb Cephalozia connivens (Dicks) Lindb
39 46 79 141 39 4679 88 (89) 90 119 141
Cephalozia lacinulata Jack ~ Spruce
141 Cephalozia lunulifolia (Dum~ Dum I 28 39 46 79 88 89 90 141
Cepoundhalozia macouni (Aust) Aust Cephalozia ~leniceps (Aust) Lindb 27~ 3746 (66)7987 (139)141 39 46 79 89 141
34
Ceffialoziella 9ivaricata (Sm) Schiffn Cephaloziella ~hista (Jack) Schiffn
46 90 46 (119) 141
Cephaloziella hampeana (Nees) Schiffn
46
Cephaloziella rubella (Nees) Warnst
46 39
Cephaloziella rubella var sullivantii
(Aust) K Mull 46 39
Cephaloziella rubella var bifida (Lindb) Douin
35
46
Chiloscyphus pallescens (Ehrh ~ Hof~ Dum Chiloscyphus pallescens var fragilis
39J 46 88 89J 90 (Roth) K Milll 46
Chiloscyphus E9lyanthos (L) Corda
1 24 39 46
Chiloscyphus polyanthos vax rivularis
(Schrad) Nees 39 46
poundpoundnocepha1um conicum (1) Lindb
12 28 39 46 (49) 87 88 8990 120 121 148
Cololejeunea biddlecomiae (Aust) Evans
39 46 62 87 88 89 90
36
Dipl0]hyllum apicu1atum (Evans) Steph Fossombronia foveo1ata Lindb
(35)41 46 90 140 1 t 38 (39) t 46
Frullania bolanderi Aust
13 46 62 88 90 139 140
Frullanla ari ttoniae Evans
39 46 69 (71) 87 88 89 90
Frullania eboracensis Gott
1 9 28 4 39 46 (57) (59) 69 70 (71) 87 88 89 90 92 93 97 148
Frullania inflata Gott
46 87 88 89 (139)
Frullania oakesiana Aust Frullania plana Sullo
(35) 42 46 87 (139) (140) 92
Frullania ri paria Hampe ~ Lehm
46 69 71 87 89
Frullania ~elwynianna Pears
46 89
Geocalyx SEaveolens (Schrad) Nees
394688 8990 (119) 140
Frullania tamarisci (L) Dum subsp sPagrayana
(Mont) Hatt 28 34 l~6 t (57) 88 89 90
38
Mar pan thus seutatUB ( Web amp Mohr) Spruce Jamesoniella autumnalis (DC) Steph
(35) 39 46 90 1 7 28 39 46 56 (57) 87 8889 90 (119) 137 141
Jungermannia confertissima Nees
137 141
Jungermannia crenuliform1s Aust
79 137 141
Jungermannia ~il1ima Sm
46 90 141 Jungermannia exsert1folia subsp corifo~ (Dum~
Vana (35) 46 79 141
39
Jungermannia hyalina Lyell Jungerrr6nnia 1eiantha Grolle
40 46 79 136 141 39 46 79 88 90 1)7 141
Jungermannia IXlaxis Lindb Jungermannia pumila With
46 67 79 (139) (140) 141 )9 46 90 1)6 141
Jungermannia sphaerocarpa Hook Kurzia setacea (Web) Grolle
4679136140IIH 136
Lejeunea cavifo1ia (Ehrh) Lindb Lepidozia reptans (1) Dum
1 7 27 39 46 88 89 90 137 14010 I ( 27) 46 I 69 (71) 88
lophocolea heterophylla (Schrad) Dum Lophocolea minor Nees
1 )4 39 46 (57) 87 88 89 27 39 46 88 89 90 (119) 90 (93)94 119 136
Lophozia al~stris (Schleich ~ Web) Evans Lophozia ascendens (Warnst) Schust
46 90 137 141 136 140 141
41
loIilozia badensls (Gatt ~ Rabenh)
46 141
lophozia blcrenata (Schmid sectA Hoffm)
Dum 46 141
lophozia collaris (Nees) Dum
39 46 141
lophozla excisa (Dicks) Dum
27 4679 141
lophozia gillmani (Aust) Schust lophozia gIlttulata (Undo tl H Arnell)
36 39 40 46 136 (140) 141 Evans 39 46 136 141
42
141
Lophozia heterocolEDs (Thed) MA Howe IoPhozia ipcisa (Schard) Dum
27 39 46 89 141 8 27 31 46 79 90 140 141
LoQhozia longidens Macoun Lophozia ventricosa
1 46 79 90 141
Lophozia ventricosa var silvicola
(Buch) Jones 141 LoEhozla ventricosa vax longif1ora (Nees)
Macoun 39 46 79 141
43
~~ fragrans (Balbis) Frye et Clark Mannia pi 10sa (Hornem) Frye et Clark
46 (139) (140) 9 46 79 116 139
Mannia triandra (Scop) Gro 11e
46 (140)
Marchantia pound91ymorpha L
1 13 28 3946 (49) 78 87 88 89 90 (Ill) 121 122 147
Mo~rckia hibernica (HookJ Gott
46 89
Metzgeria conjugata Lindb
44
46
Mylia anomala (Hook) S Gray Notothylas orbicularis (Schwein) Sullo
39 46 141 (35)45 46
Nowellia curvifolia (Dicks) Mitt
17 27 3846 88 89 90 141
Odontoschisma denudatuJI1 (Nees ~ Mart)
Dum 46 81 137 141
Pallavicinia lyellii (Hook~ Carruth
90
Odontoschisma macounii (Aust) Under
45 46 (66) (139) 140 141
45
=
Pellia epiphllla (L) CordaPellia endiviifolia (Dicks~ Dum
27 39 46 90 13724 39 46
Pellia neesiana Limp bull Phaeoceros
39 46 89 27 39 45 46 97
Porella pinnata L
13 27 I 46 90 Plagiochila asplenioides (L) Dum
27 I 39 1+6 88 89 I 90 137
46
~e11a platYEhylla (L) Pfeiff Porella platyphylloi~ (Schwein) Lindb
17 27 39 46 69 (71)79 87 88 89 9 34 46 (57) 59 87 88 89 92 90 92 93 77
Preissia quadrata (Seop) Nees
1 36 3946 (71) 88 89 90 121 123
137
Ptllidium ciliare (L) Hampe
1 27 39 46 132 141
Dum
(22) 28 3946 (57) 87 88 89 90 92
Ptilidium ~lcherrium (G Web) Hampe
(9) 13 394656 (57)70 87 88 89 90 93 (119) 136 141
47
Radula obconica Sullo Reboulia hemlsphaerlca (L) Raddi
39 46 88 89 (43) 46 (59) 70 90 t Cf 124 125 (139)
Riccardia ~atifrons Lindb
1 27 39 46 89
Riccardia multifi~ (L) S Gray
39 46 88 145
Riccia beyrichiana Hampe ~ Lehm
46 Riccardia palmata (Hedw) Carruth
39 46 90
48
Riccia bifurca Hoffm Riccia cavernosa Hoffm
48 13739 46 79 137
Riccia fluitans L
242731454679
Riccla frostli Aust
48 79
Riccia sullicantii Aust
79 (137) Riccia sorocarpa Bisch 46 (137)
49
Ricciocarpus natans (1) Corda Sca~ia apiculata Spruce
13 27 46 79 89 110 39 46 (139) 141
ScaJltll1ia carinthica Jack ~ Lindb
137 t 141
Scapania ~ (Mart) Durn
39467990137141
Scapania digenil Schiffn
90
ScaEsectflia cuspiduligera (Nees) K Mull
46 (139) (140) (141)
50
Scapania glaucoeerha1a (Tayl) Aust Seapania gyrrmostomophilia Kaal
39 46 (139) 141 41 44 46 64 (139) (140) 141
ScaP3poundlia irrigua Gott ~ almiddot
7 39 46 79 137 (140L 141
Seapania nemorosa (L) Dum
1 39 46 89 90 137
Scapmi~ ~~ta Smiddotlch
137 141 51
~caPnia poundludicola LDeske amp K Mull Scapania saxicola Schust
46 (66) 141 136 141
Sc~~ scandica (H Arnell ~ Buch) Macv
139 140
Scapania subalpina (Nees) Durn
39 40 46 79 137 141
Scapania undulata (L) Dum
1 42 46 79 141
Scapania umbrosa (Schrad) Dum
137 140 141
52
Trichocolea tomentella (Ehrh) Dum Triomarla exsecta (Schmid) Schiffn
27 (35) 39 46 47 88 137 141 8 28 39 46 90 140 141
Tritomaria exsectiformis (Breidl) Loe~ke Tritomaria Sluinguedentata (Huds) Buch
39 -t6 89 90 136 140 141 7 39 46 141
Trltomaria scitula (Tayl) Joerg
46 136 (139) 141
53
ATLAS OF WISCONSIN BRYOPHYTES
November 21 1979
THE MUSEUM OF NATURAL HISTORY Stevens Point Wisconsin
Reports on the Fauna and Flora of Wisconsin No 16
Part I
Climacium dendroides (Tree moss)
ATLAS OF WISCONSIN BRYOPHYTES
By Frank D Bowers
Curator of Non-Vascular Plants-Herbarium Museum of Natural History and
Department of Biology and
Sally Kauss Freckmann Research Associate
Museum of Natural History Universit y of Wisconsin
Stevens Point Wisconsin 54481 U S A
2
CONTENTS
ABSTRACT 4
ACKNOWLEDCEMENTS 4
INTRODUCTION 6
GEOGRAPHIC CLIMATIC AND BOTANICAL fEATURES OF WISCONSIN 7
SYSTEMATIC ARLNGEMENT OF WISCONSIN BRYOPHYTES 11
MAPS OF BRYOPH) iE TAXA REPORTED FOR WISCONSIN 29
HORNWORTS AND LIVERWORTS 30
MUSCI 54
FOSSIL AND SUBFOSSIL MOSSES OF WISCONSIN 119
PALEONTOLOGICAL Rb J3RENCES 119
SYNONYMS AND EXCLUDED TAXA 120
PIONEER BRYOLOGISTS OF WISCONSIN 126
LITERATURE CITED 127
3
ABSTRACf
117This atlas of Wisconsin bryophytes shows the distribution of)30 taxa (137 hepatics 389 mosses)
on state maps with a dot in each county for which each taxon has been reported in the literature A list of synonyms and excluded taxa is included along with an extensive bibliography Pertinent features of the natural history and history of bryology in the state are discussed
ACKNOWLEDGEMENTS
We want to thank Drs Fred Hermann and John Thomson of the University of Wisconsin-Madison herbarium and Botany department for locating and lending packets and for references to aid us in accumushylating a more accurate complete report The illustration of Climacium dendroides on the cover was drawn by Robert Freckmann Of course we are grateful to all the people who have collected specimens of Wisshyconsin bryophytes and preserved them in herbariums making this distributional study possible
4
ONEIDA
LINCOLN
TAYLOR
t---------- MARATHON C1l-RK
SAu~
t
~
~ C i-W=AS_-r7-1
l=itHLANO
)
lOCK
INGTb
lt
WALWORTH
5
INTRODUCfION
This atlas of the bryophyte flora of Wisconsin is an attempt to show the distribution of all bryoshyphyte species known to occur in the state by county occurrenoes based on published reports in the literashyture It is not an annot~ted check list (only a feW problematic taxa were examined) but is a literature search intended as a foundation for a complete flora or for otherbryological research
The literature of btYo1ogy is large and spread over numerous journals reports monographs and manuals It would be almost impossible to search all sources for every reference attributing bryophyte species to Wisconsin and to particular localities in the state Wehav checked many inlportant references and paid special attention to works dealing with Wisconsiit botany and the bryologyof the upper Midwest Several bibliographies and indices were checked thoroughly including Green an~ Curtis (1955) the Bry ologist indices (Crum 1959 and Miller 1976) andt)le Wisconsin Academy of Sciences Arts and Letters indices
Only those published repots which app~red to)e substanti2ted by herbarium specimens and which gave- ~fficient location dat to P~CetheW(On in amiddot particula(county were-used in the initial compilation Species described in manuals in generalized way implying their occurrence in Wisconsin (Without menmiddot tioning Wisconsin specifically or citing a particular location in the state) were excluded eg Gnmmia raui (Wynne 1943) and Pohlia annotinll var decipiens (Crum 1976) Ecological papers which were either too general in taxonomic determinations (eg Sphagnum sp Polytrichum sp) or in location data were also excluded Some other taxa attributed to Wisconsin which have been excluded are
Cylindrothectum compr~ (Cheney 189tmiddotDane Co) identified later by Grout as Entodon seducshytrtx
Dicranell4 humilis (Holzjne~ 1925middotBuffalo Co) probablyD vllJia since Dhumilis is not found in North Arnerl~
Didymodon triarium (Foote 1 966-SW Visconsin) identified by Fred Hennann as Barbulaacuta Funoria microstoma (anonymous 1975middotDane Co) excluded because it is outside of its geographic
range and the packe~ QaMot be located Phil(motis calcarea (Cheney and Evans 1944-Dane Co) excluded in Crum Steere and Anderson
(1973) Plagiobryum demissum and P zierl (Cheney and Evans 1944-various counties) are both Arcticshy
alpine species not likely to be found in isconsin Two of Cheneys collections of Pl1giobyrum have been referred to Bryum
Pogonatum alpinum var septentrionale (Che~ey and Evans 1944-Chippewa Co) was misidentified (Ireland 1960) and is P umigerum
Rhizomnium fAPpoJachIanum and R punctalu1fl ssp chlorophyDosum (Koponen 1973middotseveral counshy ties) excluded North American bryologiflts have not recognized these taxa in America
Sphagnum tenellum (Cheney and Evans 1944-Waukesha Co)an Atlantic species not occurring in Wisconsin (Cruro 1975)
The nomenclature and systematic arrangement of hornwort 00 hepatic species in the atlas follow Stotler and Crandall-5totler (1977) and of moss species fonow Crum Steere and Anderson (1973) except in such instances as the JreptlIfockIdus fonns whiGh follow Wijk et aL (19S9middot1973) Species reported in the literature under names which are now listed as synonyms on either of these lists were converted to the accepted name A dot is placed in each county of Wisconsin from which the taxon has been reported The numbers after the name of the taxon correspond to the numbered references in the bibliography which giye ~act location data OEat least location data -s~fficient to place it in-particular counties in the state The numbers in parentheses indicate references which attribute the taxon to Wisconsin without providing exact loltation data
Fossil and subfossil reports are summarized Slfparately in our atlas only six species (Aulacomnium turgidum lJ1achythecium turgidum Ditrichum Jlelficaule Hypnum bambergti Scorpidium 6fJisities and Tortel14 1rIgi1i$) are known exclUsively from the fossil record ~
6
GEOGRAPHIC CLIMATIC AND BOTANICAL FEATURES OF WISCONSIN
Wisconsin is a north central state bounded on the east by Lake Michigan on the north by Michigan and Lake Superior on the south by Illinois and on the west by the Mississippi and St Croix rivers It is located midway between the Prime Meridian (0deg) of Longitude and 1800 Longitude and midway between the equator and the north pole thus placing it in the center of the northern half of the western hemisphere
The topography of the state is relatively flat a few hills exceed 1900 feet with Thns Hill at 1952 feet forming the highest point in Price County Lake Michigan represents the lowest point at 580 feet The northern and eastern two-thirds of the state were heavily glaciated during the last stage of the Pleistocene and now contain many of the best examples of continental glaciation topOgraphy including considerable ground moraine large terminal recessional and intetlobate moraines kames eskers drumlins and small depressions or kettle-holes Most of the nearly 9000 lakes of 20 acres or more lie within this part of the state The southwestern third of the state is the Driftless Area It includes most of the large flat generally sandy bed of Glacial Lake Wisconsin plus the highly dissected ridge and valley section with numerous cliffs and rock-outcroppings (Figure 1)
Much of the bedrock in the state lies below glacial till but exposures include igneous and metamiddot morphic rocks primarily granitic especially in the north as well as sedimentary layers primarily sandshystones and dolomitic limestones especially along Lake Michigan and in the DriftlesS Area
The climate of Wisconsin is classified as humid-temperate with cold winters The annual precipimiddot tation averages approximately 30 inches which is sufficiently even in its seasonal distribution to support forest vegetation throughout most of the state Nevertheless according to Curtis (I 959) the pre-settlement vegetation of the state (J 830-1850) included about 27 prairie and oak-savanna plus about 64 forests
The state is presently divided into seventy-two counties Marathon County is the largest (1586 sq mi) the smallest counties include Pepin (235 sq mi) Ozaukee (236 sq mi) andMilw8ukee (237 sq mi) We included Menominee County though its future status as a county is in doubt since it regained its Indian reservation status and rights in 1973 This county was established April 30 1961 from the Menomishynee Indian reservation which consisted of parts of Shawano and Oconto counties Wisconsin ranks 26th in the United States in area with 54464 square miles and 16th in population with 4651000
More ecological research is needed on the bryophyte flora of Wisconsin especially to relate it to the vegetation and to such other environmental features as soils climate and glaCial deposits Plant communishyties in Wisconsin are many and diverse these are summarized on Figure 2 from studies conducted by Curtis (1959) The state is divided into northern and southern floristic provinces by a tension zone deshyscribed by Curtis (1959) This tension zone is an eeotonal region vatying from 10middot31 mt wide where the ranges of many vascular plants overlap or terminatej it runs in an S-curve northeast from Milwaukee to Hudson in St Croix County (Figure 2) The principal plant communities of the northern province apshyproaching their southern boundaries within the zone include the northern conifermiddothardwood forests of sugar maple white pine hemlock and yellow birch and the Sphagnum bogs with tamarack and black spruce Southern communities at or near their northern limits within this zone include prairies oak and jack pine savannas and hardwood forests of maples oaks ash ehn and cottonwood
We hope to encourage research on cryptogamic ecology with our checklist We know that some species are northern while others are southern (eg cimacium) fits the tension zone concept Freckmann (I977) while Bryoxiphium is only found in the Driftless Area of the state Steere (1937) and Love and Love (1953) The tension zone theory may prove to be acceptable for bryophytes We speculate that certain bryophytes can indicate particular plant communities since many bryophytes are restricted to preshycise environmental requirements Many papers or theses have dealt with cryptogamic bark and rock ecology in Wisconsin (Culberson 1955 Foote 1962 Hale 1952 1955 and Cole 1975)
We hope that this atlas will stimulate collecting in counties poorly sampled so that our knowledge of the bryophytes can be increased It should aid anyone interested in floras for preservation environmental impact statements and distributional studies
7
A numerical summary of the bryophyte flora of the state based on our literature search is presented in Table 1 (below) This includes 517 taxa 70 families and 176 genera This numerical summary is comshypared with the published floras of Michigan mosses (Crum 1976) and Minnesota hepatics (Schuster 1953) Thus we surmise that further collecting could lead to the same approximate numbers for Wisconsin bryoshyphytes
Table 1 Comparison of Recorded Bryophytes of Wisconsin with Michigan and Minnesota and Summary of Wisconsin Taxa
Wisconsin Michigan (Crum 1976)
Minnesota (Schuster 1955)
Hepaticsamp Hornworts
Taxa Families Genera
137 28 47
174 32 52
Mosses
Taxa Families Genera
380 42
129
503 42
149
8
-
~
~ tIt
Figure 1 The Driftless Area and Tension Zone The counties are 01$0 shown on map
ca location of tension zone as deecfibed bv Curtis (1959)~ The Orittless Area
9
Agn 2 Major Plant Communitiee of Wsconsln aaed on surwvor recorda bull co 18laquo) After Curtismiddot 1959
tit beol Forest e Phillie
Northernmiddot hardwood forest reg Pine 5cMInna
copy Southernmiddot hordwood forest CD Oak Savanna
10
SYSTEMATIC ARRANGEMENT OF WISCONSIN BRYOPHYTES
ANTHOCEROTAE (Hornworts)
1 Anthocerotaceae Anthoceros 1 emnd Prosk
macounii M A Howe punctatum 1
Phaeoceros Prosk laevis (1) Prosk subsp laevis
Notothylas Sul1 orbicularis (Schwein) Sullo
HEPATICAE (liverworts)
2 PseudolepicoJeaceae Blephllrostoma (Dum emend Undb) Dum
trlchophyllum (L) Dum subsp trlchophylluni
3 Ptilidiaceae Ptilidium Nees
ci1iare (L) Hampe pulcherrimum (G Web) Hampe
4 Trichocoleaceae Trichocolea Dum nom cons
tomentella (Ehrh) Dum
5 Lepidoziaceae Kunia von Martens
setacea (Web) GroUe
Lepidozia (Dum) Dum reptans (1) Dum
Bazzania S Gray trilobata (1) S Gray var triZobata
6 Calypogejaceae 011ypogeja Raddi
flSsa (L) Raddi subsp issa integristipula Steph muelleriana (Schiffn) K Mull subsp muelleriana neesiana (Mass et Carest) K Mull sphagnicola (H Arnell et J Perss) Warnst et loeske trichomanis (1) Corda
11
7 Cephaloziaceae Cepluzlozla (Dum emend Schiffn) Dum
biscwpidata (L) Dum subsp biclupidIIta catenulllttl (Hub) Lindb coMivens (Dicks) Lindb var connivens lacinultzta Jack ampX Spruce lunulifolilz (Dum) Dum macounii (Aust) Aust pleniceps (Aust) Lindh var pleniceps
Noweilia Mitt cunifolilz (Dicks) Mitt
8 Adelanthaceae Odontoschimuz (Dwn) Dum
denudatum (Nees ampX Mart) Dum Vir denudatum mtlCount (Aust) Underw
c
9 Cephaloziellaceae CephtlJoziellll (Spruce) Schiffn nom C()II$
divarlctzt4 (Sm) sclurfn val divarlcat(l elachhta (bdc) Schiffn 1uImpetuw (Nees) Schirfn IUbellll (Nees) Warnst vat IUbeIl4rube (Nees) Warnst var biid4 (Lindb) Dquinrube (Nees) WarDSt Var sulliwmtii (Aust) K MuD
10 Geo~c~ larptuttllus Nees
scutatus (Web et Mohr) Spruce
Geocalyx Nees grGVeolens (Schrad) Nees
11 Lophocoleaceae Lophocolea (Dum) Dum
heteropllyll4 (Schrad) Own minorNees
Chiloscypbus Corda om cons pallescens (Ehrh ampX Hoffm) Dum varfragllis (Roth) K Mun polyantitos (L) Corda var poiyanthos pol)llaquo1lthos (L) Corda var rlvulllris (Schrld) Nees
12 PlagiochiJaceae Plagio(hila Dum nom cons
tlBpienioides (L) Dum var asplenioidu
13 Jungermanniaceae Btriilophoril Loeske
attenutlta (Mart) Loeske bIubtlta (Schmid ampX Scltreb) Loeske
12
Lophom (Dum) Dum alpestris (Schleich ~ Web) Evans ascendens (Warnst) Schust badenais (Gott ex Rabenh) Schiffn var bademii bierenata (Schmid ex H)ffm) Dum eollam (Nees) Dum exci8a (Dicks) Dum var excisa gllimanii (Aust) Schust varglllmanii guttulata (Lindb et H Arnell) Evans heterocolpos (Thed) M A Howe var heterocolpos ineistl (Schrad) Dum var tnelsa longidens (Lindb) Macoun subsplongidens ventricosa (Dicks) Dum var ventricosa ventricolfl (Dicks) Dum varlontifIotrl (Nees) Macoun ventricosa (Dicks) Dum var silvlcola (Bueh) Jones
Tritomarill Schiffn ex Loeske exseeta (Schrad) Loeske exsecttformi$ (Breidl) Loeske subsp exsectiormfs quinquedentata (Huds) Buch var quinquedentata scitula (rayl) Joerg
Anastrophyllum (Spruce) Schiffn hellemnum (Noes) Schust michtzuxii (Web) Buch ex Evans minutum (Schrob) Schust var minutum
Mylill S Gray nom cons anomala (Hook) S Gray
Jamesoniella (Spruce) Carring autumnalis (DC) Steph var autumnalis
Jungemumnia L emend Dum eonfertissimtt Nees crenuliformis Aust exsertijolill Steph subsp cordifolia (Dum) Vana graciUima Sm hyalina Lyell leillnthll Grolle polaris Lindb pumila With sphaerocarpa Hook
14 Scapaniaceae Diplophyllum (Dum) Dum nom cons
apiculatum (Evans) Steph var apiculatum
Scapanill (Dum) Dum nom cons apkuata Spruce carinthillca Jack ex Lindb curta (Mart) Dum var curta
13
cuspiduligera (Nees) K Mull var cuspiduiigera degenii Schlffn ex K Mull var degenii glllucocephaJa (Tayt) Aust gymnostomophila Kaal irrigwJ (Nees) Gott et al subsp irrigua lingulata Buch varlingulata mucronata Buch subsp mucronata nemorosa (l) Dum paludicola Toeske et K Mun var paludicola paludosa (K Mull) K Mull saxicola Schust scandiea (H Arnell et Buch) Macv var scandiea 8Ubalp~ (Nees) Dum umbroSll (SchIad) Dum undulata (L) Dum var undulata
15 Radulaceae Raduia Dum nom COllS
compianata (L) Dum obconica Sull
16 Porellaceae Parella L
pinTlllta L patyphylla (L) Pfeiff platyphylloidt(l (Schwein) lindb
17 Jubulaceae FruUanill Raddi
bolanderi Aust brittoniae Evans eboracemis Gott inlllta Gott oakesiana Aust plana Sull rlparia Hampe ex Lehm selwyniana Pears taTnl1iJci (L) Dum subsp asagmyana (Mont) Hatt
18 lejeuneaceae Calolejeunea (Spruce) Schlffn
biddlecomiae (Aust) Evans
Lejeunea Ub nom cons cavifolia (Ehrh) Lindb emend Buch
19 Codoniaceae Fossombronia Raddi
foveolata Lindb
20 Pa11aviciniaceae Paklvieinlil S Gray nom cons
lyeiii (Hook) Carruth
14
Moerckia Gott hibemica (Hook) Gott
21 PeWaceae Pellfa Raddi nom cons propos
endivlifolfa (Dicks) Dwn epiphylllz (L) Corda neeBiana (GottlUmpr
22 Metzgeriaceae Metzgerla Raddi
conjugata Undb
23 Blasiacae B L
pulilm L
24 Aneurareae AneumDum
pinguis (L) Dum
Riccfl1dia S Gray nom cons latifrons Undb multiftda (L) S Gray var multifida palmata (Hedw) Carruth
25 Aytoniaceae Reboulill Raddi nom conI
hemllphaerica (L) Radeli
ManniaOpiz fragrrms (Balbis) Frye el ClarK pilosa (Hornem) Frye et Clark trltmdra (Scop) GroDe
26 Conocephalaceae Conocephalum Wigg nom cons
conicum (L) Lindb
27 Marchantiaceae Preiuia Corda
quadrata (Scop) Nees
Marchantia L polymorpha L
28 Ricciaceae Riccwcarpus Corda
natans (L) Corda
Riccia 1 beyrichiana Hampe ex Lehm bifurca Horfm
15
i
CtlJPemostI Hoffnt fuitu L rostii Aust BOIOCtI1fHI Bi~ sulltvtmtii Aust
MUSCI (Mosses)
29 Sphagnaceae SpIutgnum L
capillaceum (Weiss) Schrank var Cllpil1tlceum caplllllceum (Weiss) Vir tenellum (Schimp) Andr centrille C Jens ex Amen et C Jens compactummiddot OC ex Lam et Dc ClUpidatum Ehrh ex Hoffin var aupidatum jimbrltltum Wits ex J Hookmiddot fulCUm (Scbimp) KUngrr glrgenBOhIlll Russ imbrlctltum Hornsch ex Russ muelllmitil4m BOd 1IIIIU (RUII) C Jens przIutre L papIIIoIum Lindb pulchrum (Lindb ex braithw) Wamst recumun P-Beauv var recumun recum P-Beauv var tenue Klinggr riptlrlum Ansstr ruPQWItWamst squtl1fOlUm Crome subftltens Russ et Wamst ex Wamst IUbIeCfUldum Nees ex Sturm var subsecundum subsecundum Nees ex Sturm varrutem (Nees Hormch et sUUm) Hueb teres (Sdlimp) Anptr e C Hartm
WtlI7IItorjii Russ wuljfanum Gira
30 Andreaeaceae AndNrrerI Hedw
rupeitrlr Oedw var pest
31 Fissident~e Fluid_ lledw
trdiantoid Hedw byoidei Bedw var bryolda cristaM Wils ex Mitt exiguus sun VIr exigUw onttmUi (B-Pyl) Steud jtuberi Lesq et James obtusifolhu Wits var obtwifoliul ollPlUlldQldes Heel ~litSuU lubbtulkril Bed tfDdfoliul lledw llridubu (S~ ) Wahlenb vallirlIU1u
16
32 Ditrichaceae Pleuridium Brid
subulatum (Hedw) Rabenh
Ditrichum Hampe heteromallum (Hedw) Britt linellre (Sw) lindb
~ pallidum (Hedw) Hampe pusiUum (Hedw) Hampe
Saelania Lindb gIlucescens (Hedw) Bomanss et Broth
Ceratodon Brid purpureus (He(lw) Brid var prupurew
Distichium BSG capillaceum (Hedw) BSG inclinatum (Hedw) BSG
33 Bryoxipbiaceae Bryoxlphium Mitt
norvegicum (Brid) Mitt
34 Seligeriaceae Seligeria BSG
cumpylopoda Kindb ex Macoun et Kindb pusillll (Hedw) BSG recurvata (Hedw) BSG
Bindia BSG acuta (Hedw) BSG
35 Dicranaceae BlUehia Schwegr
lexUOSll (Sw ~ Schwaegr) C MueU sullivantii Aust
Trematodon Michx ambiguus (Hedw) Hornsch
Dicranellll (C Muell) Schimp cerviculata (Hedw) Schimp val cillPicuiata
heteromalla (Hedw) Schimp ~ rufescens (With) Schimp
schreberiana (Hedw) Schhnp var schreberiana schreberitJna (Hedw) Schimp var robusta Schimp ex Braithw subulata (Hedw) Schimp varia (Hedw) Schimp
Dicranodontium BSG denudatum (Brid) Britt ex Williams
Panzeucobryum (Lindh) Loeske Iongifolium (Hedw) oelke varlongifolium
17
Tortella (Undb) Limpr fragilis (Drumm) Umpr humilis (Hedw) JeM torlUoSfl (Hedw) Umpr
Hyophila Brid involuta (Hoole) Jaeg et Sauerb
Didymodon Hedw rigidulus Hedw
BryoerythrophyUum Chen recurvirostnon (Hedw) Chen
Btubula Hedw acula (Brid) Brid var acuta convoiuta Hedw fallox Hedw unguicultltll Hedw
Acaulon C MueD muticum (Uedw) C Muell var rufeseens (Iaeg) Crum
PJl~lm Hedw CUSPidtltum Hedw
De8mlltodon Brid cernuu (Hueb) BSG oblUsifoliw (Schwaegr) Schimp
T01IUla Hedw mueronifolia Schwaegr rurall (Hedw) Gaertn Meyer et Scherb var ruralis
39 Grimmiaceae Grlmmill Hedw
alpicola Hedw var alpicola alpicola Hedw var dupretii (Ther) Crum alpicola Hedw var rlvularl8 (Brid) Wahlenb apocarpa Hedw var apocarpa apocarpa Hedw var ambipa (Sull) Jonea apocarpa Hedw var conjerttl (Funck) Spreng apocarpa Hedw var strleta (Tum) Hook et Tayl CiIIyptrrlta Hook ex Drumm donniana Sm var donn_ hemumti Crum ltlevla (Brid) Brid montana BSG oineyi SuD oVllll8 (Hedw) lindb piUfe P-Beauv pltJgiopodill Hedw pulvinata (Hedw) Sm varbullpulvinata teretinervl Limpr
18
Orothera C Muell leana (Sull) C Muell
Rfwbdoweisia crispata (With) Undb
Cynodontium BSG ex Schimp alpestre (Wahlenb) Milde strumiferum (Hedw) Undb
Dichodontium Schimp pellucidum (Hedw) Schimp
Oncophorus (Slid) Brid vuens (Hedw) Brid wahlenbergii (Brid)
Dicranum Hedw bonjeanii DeNot ex Usa drummondii C Muell lagellare Hedw varj1ogellare fulvum Hook fuscescens Turn var fuscescens montanum Hedw muehlenbeckii BSG var muehlenbeckii polysetum Sw sabuletorum Ren et Card scoparium Hedw undulatum Brid viride (Sull et Lesq ex Sull) lindb
36 Leucobryaceae Leucobryum Hampe
glaucum (Hedw) Angstr ex Fr
37 Encalyptaceae Encalypta Hedw
ciliata Hedw var ciliata procera Bruch
38 Pottiaceae Astomum Hampe
muhlenbergianum (Sw) Grout
Weissia Hedw controversa Hedw var controversa
Gymnostomum Nees Homsch et Stunn aeruginosum Sm recurvirostrum Hedw var recurvirostrum
Trichostomum Bruch spirale Prout
19
40 Ephemeraceae Ephemerum Hampe
cohaerens (Hedw) Hampe crassineium (Schwaegr) Hampe val crassinervium spinulosum Bruch et Schimp ex Schimp
41 Funariaceae AphflllolThegma SuD
serratum (1 Hook et Wils ex Drumm) Sullo
Physcomitrium (Brid) Fuernr hookeri Hampe immersum Sullo pyriforme (Hedw) Hampe
Funaria Hedw americana Undb hygrometrica Hedw vaT hygrometrica
42 Splachnaceae Splachnum Hedw
ampullaceum Hedw
43 Schistostegaceae Schistostega Mohr
pennata (Hedw) Web el Mohr
44 Bryaceae Pohlia Hedw
annolina (Hedw) Lindb var annotina camea (Tum) Lindb cruda (Hedw) Undb elongata Hedw iliformis (Dicks) Andr lescurlana (Sull) Grout nutans (Hedw) Lindb obrusifolia (Brid) L Koch proligera (Undb ex Umpr) Undb ex Amell wahlenbergii (Web et Mohr) Andr
Leptobryum (B S G) Wits pyriforme (Hedw) Wils
Bryum Hedw algovicum Sendtnex C Muell argenteum Hedw caespiticium Hedw capillare Hedw var capillare creberrlmum Tayt lonchocaulon C Muell pallens (Brid) Sw ex Roehl pseudotrlquetrum (Hedw) Gaertn Meyer et Scherb varpudotriquetrum
20
pseudotriquetrum (Hedw) GaertnMeyer et Scherb var bimum (Schreb) Ulj stenotrichum C Muell var stenotrichum tortiolium Funck ex Brid uliginosum (Brid) B S G violaceum Crundw et Nyh weigelii Spreng
Rhodobryum (Schimp) Hampe roseum (Hedw) Limpr
45 Mniaceae Mnium Hedw
ciliare (C Muell) Schimp cinclidioides Hueb cuspidarum Hedw drummondii Bruch et Schimp longirostrum Brid lycopodioides Schwaegr marginatum (With) Brid ex P Beauv mediumB~S G var medium onhorrhynchum Brid pseudopunctatum Bruch et Schimp punctalUm Hedw var punctatum punctatum Hedw var elatum Schirnp rugicum Laur spinufosum B S G stellare Hedw
46 Aulacomruaceae Au1acomnium Schwaegr
androgynum (Hedw) Schwaegr heterostichum (Hedw) B S G paJustre (Hedw) Schwaegr var paJustre
47 Meesiaceae Amblyodon B S G
deillbotus (Hedw) B S G
Meesia Hedw triquetra (Richt) Angstr uJiginosa Hedw
48 Bartramiaceae Plagiopus Brid
oederiana (Sw) Umpr
Bartramia Hedw pomiformis Hedw var pomiformis
Philonotis Brid fontana (Hedw) Brid varfontana longseta (Michx) Britt varlongseta marchica (Hedw) Brid
21
~
~
49 Timmiaceae Timmia Hedw
megapolitana Hedw var megapolitana
50 Orthotrichaceae Zygodon Hook et Tayl
viridissimus (Dicks) Brid var rupestris lindb ex C J Hartm
Amphidium Schimp lapponicum (Hedw) Schimp mougeotii (B S G) Schimp
Orthotrichum Hedw anomalum Hedw obtuaifolium Brid ohioense Sullo et Lesq ex Aust pumilumSw sordidum Sullo et Lesq ex Aust speciosum Neesax Sturm var eegam (Schwaegr ex Hook et Grev) W~rnst stellDtwn Brid strrmgulatum P Beauv
Uleta Mohr ex Web cOfIICtata (p Beallv) Hamrn crispa (Hedw) Brid
Drummondill Hook ex Drumm prorepens (Hedw) Brltt
51 Fontinalaceae Fantinalis Hedw
antipyretica Hedw var antipyrettca antipyretica Hedw forma rabusta (Card) Card dalecarlica Schimp ex B S G var dalecarlica flqccidtJ Ren et Card hypnoides C J Hartrn var hypnoides hypnoides C J Hartm var durillei (Schimp) HU$ll
missourlcaCard novae-angliae Sullo var novae-angliae splulgrUlolill (C Moen) Wijket M~S
DichelY1l1ll Myr capilloceum (With) Myr falcatum (Bedw) Myr HIllescens B S G
52 Climaciaceae aimecium Web et Mohr ex Mohr
americllnum Brid dedroides (Hedw) Web et Mohr
22
S3 Hedwigiaceae Hedwigia P Beauv
ciliata (Hedw) P Beauv
54 Cryphaeaceae Forsstroemia lindh
ohioense (Sull) lindh
55 Leucodontaceae Leucodon Schwaegr
brachypus Blid var brachypus brachypus Brid var aniJrewsitlnus Crum et Anderson julaceus (Hedw) SuU
56 Neckeraceae Neckera Hedw
pennata Hedw var pennata
1fomalia (Brid) B S G trichomanoides (Hedw) B S G
57 Theliaceae Thelia Sull
asprellll Sullo hirtella (Hedw) Sullo lescurii SuU
Myurella B SG julacea (Schwaegr) B S G
sibrica (C MueU) Reim var sibrica
58 Fabroniaceae Fabronio Raddi
ciliaris (Brid) Bnd
Anacamptodon Brid splachnoides (Groet ex Brid) Brid
Schwetschkeopsis Broth fabronia (Schwaegr) Broth
59 Leskeaceae Lindbergia Kindb
brachyptera (Mitt) Kindb
Leskea Hedw gracilescens Hedw obscura Hedw poiycarpa Hedw
Leskeella (limpr) Loeske nervosa (Brid) Loeske
23
Pterlgy1lllndrum Hedw filiforme Hedw
60 Thuidiaceae Haplohymenfum Dozy et MoIk
trlste (Ces ex De Not) Kiodb
Anomodon Hook et Tayl attenuatus (Hedw) Hueb minor (Hedw) Fuemr rostratus (Hedw) Schimp viticulosus (Hedw) Hook et Tayl
Haplocladium (C Mucn) C MueU microphyllum (Hedw) Broth virginillnum (Brld) Broth ~bull
Thuidium B S G abietinum (Hedw) B S G delicatulum (Hedw) B S G vardelicatulum delicatulum (Hedw) B S G var radicans (Kindb) Crum Steere et Anderson minutulum (Hedw) B S G pygmaeum B S G recognftum (Hedw) Lindb scitum (p Beauv) Aust
Helodium (Sull) Warnst blandowli (Web et Mohr) Warnst var blandowli blandowli (Web et Mohr) Wamst var helodioides (Ren et Card ex Roell) Crum
Steere et Anderson paludomm (Sull) AuS
61 Amblystegiaceae Cratoneuron (Sull) Spruce
commulatum (Hedw) Roth var commutfltum commutatum (Hedw) Roth var fakatum (Brid) Moenk ilicinum (Hedw) Spruce var filcinum
Campylium (Sun) Mitt chrysophyllum (Brid) 1 Lange var chrysophyllum hispidulum (Brid) Mitt var hllJidulum polygamum (B S G)C lens varpolygtlmUm stellatum (Hedw) C1ens vat stelltum
Leptodictyum (Schimp) Warnst laxirete (Card et Ther) Broth riparium (Hedw) Wamat trichopodium (Schultz) Warns l1acillans (Sull) Broth
Hygroamblystegium Loelke jlul1iatile (Hedw) oeske var luViatile
24
fluviatile (Hedw) Loeske var oWltum Grout noterophilum (Sun et Lesq ex Sull) Warnst tenox (Hedw) Jenn var tenax tenox (Hedw) Jenn var spinilolium Schimp) Jenn
Amblystegium BSG juratzkanum Schimp serpens (Hedw) BSG varlum (Hedw) Undb var varium
Platydictya Berk confervoides (Brid) Crum jungermanniodes (Brid) Crum minutissimum (Sull et Lesq ex Sull) Crum subtile (Hedw) Crum
Drepanocladus (C Muell) Roth aduncus (Hedw) Warnst var aduncus aduncus (Hedw) Warnst var capillilolius (Wamst)Riebm aduncus (Hedw) Warnst var kneiffii (BSG) Moenk aduncus (Hedw) Warnst var polycorpus (Bland ex Voit) Roth exannulatus (BSG) Warnst fluitans (Hedw) Warnst var fluitans revolvens (Sw) Wamst var revolvens sendtneri (Schimp) Warnst var sendtnerl uncinatus (Hedw) Warnst var uncinatus uncinatus ffiedw) Warnst var piwnosus (Schimp) Wamst vernicoms (Lindb ex C Hartm) Warnst
Hygrohypnum Lindb eugyrium (BSG) loeske luridum (Hedw Jenn varluridum ochraecum (Tum ex Wils) Loeske var ochraceunt
Calliergon (Sull) Kindb cordilolium Hedw Kindb glganteum (Schimp) Kindb var giganteum richardsonii (Mitt) Kindb ex Wamst stramineum (Brid) Kindb trilarium (Web et Mohr) Kindb
Calliergonella Loeske cuspidata (Hedw) Loeske
Calliergidium (Ren) Grout bakeri (Ren) Grout
Scorpidium (Schimp) Urnpr scorpiodides (Hedw) limpr
62 Brachytbeciaceae Tomenthypnum loeske
nitens (Hedw) Loeske var mtens
25
Homalotheciella (Card) Broth subcapillata (Hedw) Broth
Brachythecium BSG acuminatum (Hedw) Awt acutum (Mitt) Sullo calcareum IGndb campestTe (C MuelL) BSG cunum (Lindb) Umpr digastrum C Muell et Kindb ex Macoun et Kindb oxycladon (Brid) Jaeg et Sauerb var oxycladon plumosum (Hedw) BSG populeum (Hedw) BSG var populeum reflexum (Starke ex Web et Mohr) DsG var relexum reiexum (Starke ex Web et Mohr) DSG var pacificum Renet Card ex Roell rlvulare BSG var rivulare tabulum (Hedw) BSC var tabuum salebrosum (Web et Mohr) BSG var salebrosum velutinum (Hedw) BSG Vat velutinum
Bryhniil Kaur graminicolor (Brid) Grout var graminicolor novae-angliae (Sull et Lesq ex sun) Grout
Rhynchostegium BSG senulatum (Hedw) Ja4lg et Sauerb
Rhynchostegiella (BSG) Limpr compacta (C MueU) Loeske
EU1hynchium 8SG hians (Hedw) Sande Lac pulchellum (Hedw) Jenn var pulchelium riparloides (Hedw) Rich
Stokeriella (Kindb) Robins pmelonga (Hedw) Robins var stokesii (Tum) Crum
63 Entodontaceae Entodon C Muell
brevisetus (Hook et Wils ex Wils) Lindb cladorrhizans (Hedw) C Muell seductrlx (Hedw) C MueU var wluctrix
Pleurozium Mitt schreberl (8rid) Mitt var schreberi
64 Plagiotheciaceae Pltlglothecium BSG
CflVifolium (Brid) Iwats var CtWifolium denticulatum (Hedw) 8SG laetum BSG ltztebricola BSG
26
6S SernatophyUaceae Brotherella Loeske ex Fleisch
recumins (Michx) Fleisch tenuirostris (Bruch et Schimp ex Sull) Fleisch
Sematophyllum Mitt demissum (Wils) Mitt
66 Hypnaceae Pl(itygyrium BSG
repens (Brid) BSG
PylaisieUa Kindb intricata (Hedw) Grout polyantha (Hedw) Grout selwynii (Kindb) Crum Steere et Anderson
Homomallium (Schimp) Loeske adnatum (Hedw) Broth
Hypnum Hedw cupressiforme Hedw var cupressiforme cupressifonne Hedw varfiliforme Brid curviolium Hedw fertile Sendtn imponens Hedw lindbergii Mitt palescens (Hedw) P-Beauv pratense Koch ex Brid
Calicladium Crum haldanianum (Grev) Crum
sopterygiopsis Iwats muelleriana (Schimp) Iwats
Isopterygium Mitt e1egans (Brid) lindb pulchellum (Hedw) Jaeg et Sauerb tenerum (Sw) Mitt
Taxiphyllum Fleisch depllQtum (Bruch et Schimp ex sun) Fleisch taxirameum (MitL) Fleisch
Herzogiella Broth seligeri (Brid) Iwats striatella (Brid) Iwats ruracea (Lindh) Iwats
hilium De Not crista-castrenna (Hedw) De Not
27
67 Rhytidiaceae Rhytidium (Sull) Kindb
rugosum (Hedw) Kindb
Rhytidfodelphus (Lindb ex Iimpr) Warnst triquetrus (Hedw) Warnst var triquetrus
68 Hylocomiaceae Hylocomium BSG
pyrenaicum (Spruce) lindb splendens (Hedw) BSG varsplendens
69 Buxbaumiaceae Buxbaumia Hedw
aphyllll Hedw
Diphyscium Mohr follosum (Hedw) Mohr
70 Tetraphidaceae Tetraphi3
pellucida Hedw
71 Polytrichaceae Atrichum PmiddotBeauv
angustatum (Brid) BSG cri8pum (James) Sullo oerstedianum (C Muell) Mitt teneUum (Roehl) BSG undulatum (Hedw) P middotBeauv var unduiatum
Pogonatum P-Beauv alpinum (Hedw) Roehl var alpinum umigerum (Hedw) P-Beauv
Poiytrichum Hedw commune Hedw var commune ormosum Hedw juniperinum Hedw longisetum Brid ohioense Ren et Card piliferum Hedw var pilierum strictum Brid
28
MAPS OF BRYOPHYTE TAXA
REPORTED FOR WISCONSIN
HORNWORTS AND LIVERWORTS
29
bull bull WSHeV~N S vE II
0 ~
e~RRON ~USK
(J~ (j -
CHIPPE A DUNN
0 o0 poundAU CLAIRE
~ TREMshyPEAshyLEAU
MICE
ONEIDA
Q LINCOLN
TAYLOR
Q t-~----- MOIRA THON
CLARK
o
10
LA FAVETTE
Hornwor ts amp Liverworts
- None - 1 - 25 taxa
- 26 - 50 tRXshy
- 51 - 75 tax)
- 7c - 100 taxa
FONDOU j LAC ~
DODGE
i O JEfFERSON
WALWORTH
30
Anastrophylltlfl helleranum (Nees) Schust Anastrophyllum michauxil (We~ Buch ~
46 88 136 141 Evans 38 46 141
Anastrophyllum mlnut~ (Schreb) Schuster Aneura plnguls (L) Durn
89 21 384687 88 89 90 (138) (145)
Anthoceros macounil MA Howe ~oceros punctatus L
21 (45) (144) 34
31
Barbilophozia attenuata (Mart) Loeske Barbilophozia barbata (Schmid ~ Schre~
24 141 Loeske 28 39 46 89 90 141
Bazzania trilobata (1) S Gray
1 26 39 46 (57) (59) 79 88 89
90 119 141
BlePbarostoma trichoffiyllum (1) Dumort
1 8 26 39 46 89 90 137 141
Blasia pusilla 1
1 39 46 90
Calypogeja fissa (1) Raddi
90
32
$129geja integristipula Steph C~Ypound2geja muelleriana (Schifm) K Mull
88 89 89
CallpoundO~eja neesiana (Mass amp Carest) K Mull
39 46 89 140 (141)
~pogeja sJlapound2icola (H Arnell tl J bull
Perss) Warnst et Loeske (119) 141
CeIalozia biscuspidata (L) Dum
1 37 I 46 90 t 137 141
CalYpogej~ trichomanis (L) Corda
26 39 (119) 137
33
CeEhalozia catenulata (Hub) Lindb Cephalozia connivens (Dicks) Lindb
39 46 79 141 39 4679 88 (89) 90 119 141
Cephalozia lacinulata Jack ~ Spruce
141 Cephalozia lunulifolia (Dum~ Dum I 28 39 46 79 88 89 90 141
Cepoundhalozia macouni (Aust) Aust Cephalozia ~leniceps (Aust) Lindb 27~ 3746 (66)7987 (139)141 39 46 79 89 141
34
Ceffialoziella 9ivaricata (Sm) Schiffn Cephaloziella ~hista (Jack) Schiffn
46 90 46 (119) 141
Cephaloziella hampeana (Nees) Schiffn
46
Cephaloziella rubella (Nees) Warnst
46 39
Cephaloziella rubella var sullivantii
(Aust) K Mull 46 39
Cephaloziella rubella var bifida (Lindb) Douin
35
46
Chiloscyphus pallescens (Ehrh ~ Hof~ Dum Chiloscyphus pallescens var fragilis
39J 46 88 89J 90 (Roth) K Milll 46
Chiloscyphus E9lyanthos (L) Corda
1 24 39 46
Chiloscyphus polyanthos vax rivularis
(Schrad) Nees 39 46
poundpoundnocepha1um conicum (1) Lindb
12 28 39 46 (49) 87 88 8990 120 121 148
Cololejeunea biddlecomiae (Aust) Evans
39 46 62 87 88 89 90
36
Dipl0]hyllum apicu1atum (Evans) Steph Fossombronia foveo1ata Lindb
(35)41 46 90 140 1 t 38 (39) t 46
Frullania bolanderi Aust
13 46 62 88 90 139 140
Frullanla ari ttoniae Evans
39 46 69 (71) 87 88 89 90
Frullania eboracensis Gott
1 9 28 4 39 46 (57) (59) 69 70 (71) 87 88 89 90 92 93 97 148
Frullania inflata Gott
46 87 88 89 (139)
Frullania oakesiana Aust Frullania plana Sullo
(35) 42 46 87 (139) (140) 92
Frullania ri paria Hampe ~ Lehm
46 69 71 87 89
Frullania ~elwynianna Pears
46 89
Geocalyx SEaveolens (Schrad) Nees
394688 8990 (119) 140
Frullania tamarisci (L) Dum subsp sPagrayana
(Mont) Hatt 28 34 l~6 t (57) 88 89 90
38
Mar pan thus seutatUB ( Web amp Mohr) Spruce Jamesoniella autumnalis (DC) Steph
(35) 39 46 90 1 7 28 39 46 56 (57) 87 8889 90 (119) 137 141
Jungermannia confertissima Nees
137 141
Jungermannia crenuliform1s Aust
79 137 141
Jungermannia ~il1ima Sm
46 90 141 Jungermannia exsert1folia subsp corifo~ (Dum~
Vana (35) 46 79 141
39
Jungermannia hyalina Lyell Jungerrr6nnia 1eiantha Grolle
40 46 79 136 141 39 46 79 88 90 1)7 141
Jungermannia IXlaxis Lindb Jungermannia pumila With
46 67 79 (139) (140) 141 )9 46 90 1)6 141
Jungermannia sphaerocarpa Hook Kurzia setacea (Web) Grolle
4679136140IIH 136
Lejeunea cavifo1ia (Ehrh) Lindb Lepidozia reptans (1) Dum
1 7 27 39 46 88 89 90 137 14010 I ( 27) 46 I 69 (71) 88
lophocolea heterophylla (Schrad) Dum Lophocolea minor Nees
1 )4 39 46 (57) 87 88 89 27 39 46 88 89 90 (119) 90 (93)94 119 136
Lophozia al~stris (Schleich ~ Web) Evans Lophozia ascendens (Warnst) Schust
46 90 137 141 136 140 141
41
loIilozia badensls (Gatt ~ Rabenh)
46 141
lophozia blcrenata (Schmid sectA Hoffm)
Dum 46 141
lophozia collaris (Nees) Dum
39 46 141
lophozla excisa (Dicks) Dum
27 4679 141
lophozia gillmani (Aust) Schust lophozia gIlttulata (Undo tl H Arnell)
36 39 40 46 136 (140) 141 Evans 39 46 136 141
42
141
Lophozia heterocolEDs (Thed) MA Howe IoPhozia ipcisa (Schard) Dum
27 39 46 89 141 8 27 31 46 79 90 140 141
LoQhozia longidens Macoun Lophozia ventricosa
1 46 79 90 141
Lophozia ventricosa var silvicola
(Buch) Jones 141 LoEhozla ventricosa vax longif1ora (Nees)
Macoun 39 46 79 141
43
~~ fragrans (Balbis) Frye et Clark Mannia pi 10sa (Hornem) Frye et Clark
46 (139) (140) 9 46 79 116 139
Mannia triandra (Scop) Gro 11e
46 (140)
Marchantia pound91ymorpha L
1 13 28 3946 (49) 78 87 88 89 90 (Ill) 121 122 147
Mo~rckia hibernica (HookJ Gott
46 89
Metzgeria conjugata Lindb
44
46
Mylia anomala (Hook) S Gray Notothylas orbicularis (Schwein) Sullo
39 46 141 (35)45 46
Nowellia curvifolia (Dicks) Mitt
17 27 3846 88 89 90 141
Odontoschisma denudatuJI1 (Nees ~ Mart)
Dum 46 81 137 141
Pallavicinia lyellii (Hook~ Carruth
90
Odontoschisma macounii (Aust) Under
45 46 (66) (139) 140 141
45
=
Pellia epiphllla (L) CordaPellia endiviifolia (Dicks~ Dum
27 39 46 90 13724 39 46
Pellia neesiana Limp bull Phaeoceros
39 46 89 27 39 45 46 97
Porella pinnata L
13 27 I 46 90 Plagiochila asplenioides (L) Dum
27 I 39 1+6 88 89 I 90 137
46
~e11a platYEhylla (L) Pfeiff Porella platyphylloi~ (Schwein) Lindb
17 27 39 46 69 (71)79 87 88 89 9 34 46 (57) 59 87 88 89 92 90 92 93 77
Preissia quadrata (Seop) Nees
1 36 3946 (71) 88 89 90 121 123
137
Ptllidium ciliare (L) Hampe
1 27 39 46 132 141
Dum
(22) 28 3946 (57) 87 88 89 90 92
Ptilidium ~lcherrium (G Web) Hampe
(9) 13 394656 (57)70 87 88 89 90 93 (119) 136 141
47
Radula obconica Sullo Reboulia hemlsphaerlca (L) Raddi
39 46 88 89 (43) 46 (59) 70 90 t Cf 124 125 (139)
Riccardia ~atifrons Lindb
1 27 39 46 89
Riccardia multifi~ (L) S Gray
39 46 88 145
Riccia beyrichiana Hampe ~ Lehm
46 Riccardia palmata (Hedw) Carruth
39 46 90
48
Riccia bifurca Hoffm Riccia cavernosa Hoffm
48 13739 46 79 137
Riccia fluitans L
242731454679
Riccla frostli Aust
48 79
Riccia sullicantii Aust
79 (137) Riccia sorocarpa Bisch 46 (137)
49
Ricciocarpus natans (1) Corda Sca~ia apiculata Spruce
13 27 46 79 89 110 39 46 (139) 141
ScaJltll1ia carinthica Jack ~ Lindb
137 t 141
Scapania ~ (Mart) Durn
39467990137141
Scapania digenil Schiffn
90
ScaEsectflia cuspiduligera (Nees) K Mull
46 (139) (140) (141)
50
Scapania glaucoeerha1a (Tayl) Aust Seapania gyrrmostomophilia Kaal
39 46 (139) 141 41 44 46 64 (139) (140) 141
ScaP3poundlia irrigua Gott ~ almiddot
7 39 46 79 137 (140L 141
Seapania nemorosa (L) Dum
1 39 46 89 90 137
Scapmi~ ~~ta Smiddotlch
137 141 51
~caPnia poundludicola LDeske amp K Mull Scapania saxicola Schust
46 (66) 141 136 141
Sc~~ scandica (H Arnell ~ Buch) Macv
139 140
Scapania subalpina (Nees) Durn
39 40 46 79 137 141
Scapania undulata (L) Dum
1 42 46 79 141
Scapania umbrosa (Schrad) Dum
137 140 141
52
Trichocolea tomentella (Ehrh) Dum Triomarla exsecta (Schmid) Schiffn
27 (35) 39 46 47 88 137 141 8 28 39 46 90 140 141
Tritomaria exsectiformis (Breidl) Loe~ke Tritomaria Sluinguedentata (Huds) Buch
39 -t6 89 90 136 140 141 7 39 46 141
Trltomaria scitula (Tayl) Joerg
46 136 (139) 141
53
Climacium dendroides (Tree moss)
ATLAS OF WISCONSIN BRYOPHYTES
By Frank D Bowers
Curator of Non-Vascular Plants-Herbarium Museum of Natural History and
Department of Biology and
Sally Kauss Freckmann Research Associate
Museum of Natural History Universit y of Wisconsin
Stevens Point Wisconsin 54481 U S A
2
CONTENTS
ABSTRACT 4
ACKNOWLEDCEMENTS 4
INTRODUCTION 6
GEOGRAPHIC CLIMATIC AND BOTANICAL fEATURES OF WISCONSIN 7
SYSTEMATIC ARLNGEMENT OF WISCONSIN BRYOPHYTES 11
MAPS OF BRYOPH) iE TAXA REPORTED FOR WISCONSIN 29
HORNWORTS AND LIVERWORTS 30
MUSCI 54
FOSSIL AND SUBFOSSIL MOSSES OF WISCONSIN 119
PALEONTOLOGICAL Rb J3RENCES 119
SYNONYMS AND EXCLUDED TAXA 120
PIONEER BRYOLOGISTS OF WISCONSIN 126
LITERATURE CITED 127
3
ABSTRACf
117This atlas of Wisconsin bryophytes shows the distribution of)30 taxa (137 hepatics 389 mosses)
on state maps with a dot in each county for which each taxon has been reported in the literature A list of synonyms and excluded taxa is included along with an extensive bibliography Pertinent features of the natural history and history of bryology in the state are discussed
ACKNOWLEDGEMENTS
We want to thank Drs Fred Hermann and John Thomson of the University of Wisconsin-Madison herbarium and Botany department for locating and lending packets and for references to aid us in accumushylating a more accurate complete report The illustration of Climacium dendroides on the cover was drawn by Robert Freckmann Of course we are grateful to all the people who have collected specimens of Wisshyconsin bryophytes and preserved them in herbariums making this distributional study possible
4
ONEIDA
LINCOLN
TAYLOR
t---------- MARATHON C1l-RK
SAu~
t
~
~ C i-W=AS_-r7-1
l=itHLANO
)
lOCK
INGTb
lt
WALWORTH
5
INTRODUCfION
This atlas of the bryophyte flora of Wisconsin is an attempt to show the distribution of all bryoshyphyte species known to occur in the state by county occurrenoes based on published reports in the literashyture It is not an annot~ted check list (only a feW problematic taxa were examined) but is a literature search intended as a foundation for a complete flora or for otherbryological research
The literature of btYo1ogy is large and spread over numerous journals reports monographs and manuals It would be almost impossible to search all sources for every reference attributing bryophyte species to Wisconsin and to particular localities in the state Wehav checked many inlportant references and paid special attention to works dealing with Wisconsiit botany and the bryologyof the upper Midwest Several bibliographies and indices were checked thoroughly including Green an~ Curtis (1955) the Bry ologist indices (Crum 1959 and Miller 1976) andt)le Wisconsin Academy of Sciences Arts and Letters indices
Only those published repots which app~red to)e substanti2ted by herbarium specimens and which gave- ~fficient location dat to P~CetheW(On in amiddot particula(county were-used in the initial compilation Species described in manuals in generalized way implying their occurrence in Wisconsin (Without menmiddot tioning Wisconsin specifically or citing a particular location in the state) were excluded eg Gnmmia raui (Wynne 1943) and Pohlia annotinll var decipiens (Crum 1976) Ecological papers which were either too general in taxonomic determinations (eg Sphagnum sp Polytrichum sp) or in location data were also excluded Some other taxa attributed to Wisconsin which have been excluded are
Cylindrothectum compr~ (Cheney 189tmiddotDane Co) identified later by Grout as Entodon seducshytrtx
Dicranell4 humilis (Holzjne~ 1925middotBuffalo Co) probablyD vllJia since Dhumilis is not found in North Arnerl~
Didymodon triarium (Foote 1 966-SW Visconsin) identified by Fred Hennann as Barbulaacuta Funoria microstoma (anonymous 1975middotDane Co) excluded because it is outside of its geographic
range and the packe~ QaMot be located Phil(motis calcarea (Cheney and Evans 1944-Dane Co) excluded in Crum Steere and Anderson
(1973) Plagiobryum demissum and P zierl (Cheney and Evans 1944-various counties) are both Arcticshy
alpine species not likely to be found in isconsin Two of Cheneys collections of Pl1giobyrum have been referred to Bryum
Pogonatum alpinum var septentrionale (Che~ey and Evans 1944-Chippewa Co) was misidentified (Ireland 1960) and is P umigerum
Rhizomnium fAPpoJachIanum and R punctalu1fl ssp chlorophyDosum (Koponen 1973middotseveral counshy ties) excluded North American bryologiflts have not recognized these taxa in America
Sphagnum tenellum (Cheney and Evans 1944-Waukesha Co)an Atlantic species not occurring in Wisconsin (Cruro 1975)
The nomenclature and systematic arrangement of hornwort 00 hepatic species in the atlas follow Stotler and Crandall-5totler (1977) and of moss species fonow Crum Steere and Anderson (1973) except in such instances as the JreptlIfockIdus fonns whiGh follow Wijk et aL (19S9middot1973) Species reported in the literature under names which are now listed as synonyms on either of these lists were converted to the accepted name A dot is placed in each county of Wisconsin from which the taxon has been reported The numbers after the name of the taxon correspond to the numbered references in the bibliography which giye ~act location data OEat least location data -s~fficient to place it in-particular counties in the state The numbers in parentheses indicate references which attribute the taxon to Wisconsin without providing exact loltation data
Fossil and subfossil reports are summarized Slfparately in our atlas only six species (Aulacomnium turgidum lJ1achythecium turgidum Ditrichum Jlelficaule Hypnum bambergti Scorpidium 6fJisities and Tortel14 1rIgi1i$) are known exclUsively from the fossil record ~
6
GEOGRAPHIC CLIMATIC AND BOTANICAL FEATURES OF WISCONSIN
Wisconsin is a north central state bounded on the east by Lake Michigan on the north by Michigan and Lake Superior on the south by Illinois and on the west by the Mississippi and St Croix rivers It is located midway between the Prime Meridian (0deg) of Longitude and 1800 Longitude and midway between the equator and the north pole thus placing it in the center of the northern half of the western hemisphere
The topography of the state is relatively flat a few hills exceed 1900 feet with Thns Hill at 1952 feet forming the highest point in Price County Lake Michigan represents the lowest point at 580 feet The northern and eastern two-thirds of the state were heavily glaciated during the last stage of the Pleistocene and now contain many of the best examples of continental glaciation topOgraphy including considerable ground moraine large terminal recessional and intetlobate moraines kames eskers drumlins and small depressions or kettle-holes Most of the nearly 9000 lakes of 20 acres or more lie within this part of the state The southwestern third of the state is the Driftless Area It includes most of the large flat generally sandy bed of Glacial Lake Wisconsin plus the highly dissected ridge and valley section with numerous cliffs and rock-outcroppings (Figure 1)
Much of the bedrock in the state lies below glacial till but exposures include igneous and metamiddot morphic rocks primarily granitic especially in the north as well as sedimentary layers primarily sandshystones and dolomitic limestones especially along Lake Michigan and in the DriftlesS Area
The climate of Wisconsin is classified as humid-temperate with cold winters The annual precipimiddot tation averages approximately 30 inches which is sufficiently even in its seasonal distribution to support forest vegetation throughout most of the state Nevertheless according to Curtis (I 959) the pre-settlement vegetation of the state (J 830-1850) included about 27 prairie and oak-savanna plus about 64 forests
The state is presently divided into seventy-two counties Marathon County is the largest (1586 sq mi) the smallest counties include Pepin (235 sq mi) Ozaukee (236 sq mi) andMilw8ukee (237 sq mi) We included Menominee County though its future status as a county is in doubt since it regained its Indian reservation status and rights in 1973 This county was established April 30 1961 from the Menomishynee Indian reservation which consisted of parts of Shawano and Oconto counties Wisconsin ranks 26th in the United States in area with 54464 square miles and 16th in population with 4651000
More ecological research is needed on the bryophyte flora of Wisconsin especially to relate it to the vegetation and to such other environmental features as soils climate and glaCial deposits Plant communishyties in Wisconsin are many and diverse these are summarized on Figure 2 from studies conducted by Curtis (1959) The state is divided into northern and southern floristic provinces by a tension zone deshyscribed by Curtis (1959) This tension zone is an eeotonal region vatying from 10middot31 mt wide where the ranges of many vascular plants overlap or terminatej it runs in an S-curve northeast from Milwaukee to Hudson in St Croix County (Figure 2) The principal plant communities of the northern province apshyproaching their southern boundaries within the zone include the northern conifermiddothardwood forests of sugar maple white pine hemlock and yellow birch and the Sphagnum bogs with tamarack and black spruce Southern communities at or near their northern limits within this zone include prairies oak and jack pine savannas and hardwood forests of maples oaks ash ehn and cottonwood
We hope to encourage research on cryptogamic ecology with our checklist We know that some species are northern while others are southern (eg cimacium) fits the tension zone concept Freckmann (I977) while Bryoxiphium is only found in the Driftless Area of the state Steere (1937) and Love and Love (1953) The tension zone theory may prove to be acceptable for bryophytes We speculate that certain bryophytes can indicate particular plant communities since many bryophytes are restricted to preshycise environmental requirements Many papers or theses have dealt with cryptogamic bark and rock ecology in Wisconsin (Culberson 1955 Foote 1962 Hale 1952 1955 and Cole 1975)
We hope that this atlas will stimulate collecting in counties poorly sampled so that our knowledge of the bryophytes can be increased It should aid anyone interested in floras for preservation environmental impact statements and distributional studies
7
A numerical summary of the bryophyte flora of the state based on our literature search is presented in Table 1 (below) This includes 517 taxa 70 families and 176 genera This numerical summary is comshypared with the published floras of Michigan mosses (Crum 1976) and Minnesota hepatics (Schuster 1953) Thus we surmise that further collecting could lead to the same approximate numbers for Wisconsin bryoshyphytes
Table 1 Comparison of Recorded Bryophytes of Wisconsin with Michigan and Minnesota and Summary of Wisconsin Taxa
Wisconsin Michigan (Crum 1976)
Minnesota (Schuster 1955)
Hepaticsamp Hornworts
Taxa Families Genera
137 28 47
174 32 52
Mosses
Taxa Families Genera
380 42
129
503 42
149
8
-
~
~ tIt
Figure 1 The Driftless Area and Tension Zone The counties are 01$0 shown on map
ca location of tension zone as deecfibed bv Curtis (1959)~ The Orittless Area
9
Agn 2 Major Plant Communitiee of Wsconsln aaed on surwvor recorda bull co 18laquo) After Curtismiddot 1959
tit beol Forest e Phillie
Northernmiddot hardwood forest reg Pine 5cMInna
copy Southernmiddot hordwood forest CD Oak Savanna
10
SYSTEMATIC ARRANGEMENT OF WISCONSIN BRYOPHYTES
ANTHOCEROTAE (Hornworts)
1 Anthocerotaceae Anthoceros 1 emnd Prosk
macounii M A Howe punctatum 1
Phaeoceros Prosk laevis (1) Prosk subsp laevis
Notothylas Sul1 orbicularis (Schwein) Sullo
HEPATICAE (liverworts)
2 PseudolepicoJeaceae Blephllrostoma (Dum emend Undb) Dum
trlchophyllum (L) Dum subsp trlchophylluni
3 Ptilidiaceae Ptilidium Nees
ci1iare (L) Hampe pulcherrimum (G Web) Hampe
4 Trichocoleaceae Trichocolea Dum nom cons
tomentella (Ehrh) Dum
5 Lepidoziaceae Kunia von Martens
setacea (Web) GroUe
Lepidozia (Dum) Dum reptans (1) Dum
Bazzania S Gray trilobata (1) S Gray var triZobata
6 Calypogejaceae 011ypogeja Raddi
flSsa (L) Raddi subsp issa integristipula Steph muelleriana (Schiffn) K Mull subsp muelleriana neesiana (Mass et Carest) K Mull sphagnicola (H Arnell et J Perss) Warnst et loeske trichomanis (1) Corda
11
7 Cephaloziaceae Cepluzlozla (Dum emend Schiffn) Dum
biscwpidata (L) Dum subsp biclupidIIta catenulllttl (Hub) Lindb coMivens (Dicks) Lindb var connivens lacinultzta Jack ampX Spruce lunulifolilz (Dum) Dum macounii (Aust) Aust pleniceps (Aust) Lindh var pleniceps
Noweilia Mitt cunifolilz (Dicks) Mitt
8 Adelanthaceae Odontoschimuz (Dwn) Dum
denudatum (Nees ampX Mart) Dum Vir denudatum mtlCount (Aust) Underw
c
9 Cephaloziellaceae CephtlJoziellll (Spruce) Schiffn nom C()II$
divarlctzt4 (Sm) sclurfn val divarlcat(l elachhta (bdc) Schiffn 1uImpetuw (Nees) Schirfn IUbellll (Nees) Warnst vat IUbeIl4rube (Nees) Warnst var biid4 (Lindb) Dquinrube (Nees) WarDSt Var sulliwmtii (Aust) K MuD
10 Geo~c~ larptuttllus Nees
scutatus (Web et Mohr) Spruce
Geocalyx Nees grGVeolens (Schrad) Nees
11 Lophocoleaceae Lophocolea (Dum) Dum
heteropllyll4 (Schrad) Own minorNees
Chiloscypbus Corda om cons pallescens (Ehrh ampX Hoffm) Dum varfragllis (Roth) K Mun polyantitos (L) Corda var poiyanthos pol)llaquo1lthos (L) Corda var rlvulllris (Schrld) Nees
12 PlagiochiJaceae Plagio(hila Dum nom cons
tlBpienioides (L) Dum var asplenioidu
13 Jungermanniaceae Btriilophoril Loeske
attenutlta (Mart) Loeske bIubtlta (Schmid ampX Scltreb) Loeske
12
Lophom (Dum) Dum alpestris (Schleich ~ Web) Evans ascendens (Warnst) Schust badenais (Gott ex Rabenh) Schiffn var bademii bierenata (Schmid ex H)ffm) Dum eollam (Nees) Dum exci8a (Dicks) Dum var excisa gllimanii (Aust) Schust varglllmanii guttulata (Lindb et H Arnell) Evans heterocolpos (Thed) M A Howe var heterocolpos ineistl (Schrad) Dum var tnelsa longidens (Lindb) Macoun subsplongidens ventricosa (Dicks) Dum var ventricosa ventricolfl (Dicks) Dum varlontifIotrl (Nees) Macoun ventricosa (Dicks) Dum var silvlcola (Bueh) Jones
Tritomarill Schiffn ex Loeske exseeta (Schrad) Loeske exsecttformi$ (Breidl) Loeske subsp exsectiormfs quinquedentata (Huds) Buch var quinquedentata scitula (rayl) Joerg
Anastrophyllum (Spruce) Schiffn hellemnum (Noes) Schust michtzuxii (Web) Buch ex Evans minutum (Schrob) Schust var minutum
Mylill S Gray nom cons anomala (Hook) S Gray
Jamesoniella (Spruce) Carring autumnalis (DC) Steph var autumnalis
Jungemumnia L emend Dum eonfertissimtt Nees crenuliformis Aust exsertijolill Steph subsp cordifolia (Dum) Vana graciUima Sm hyalina Lyell leillnthll Grolle polaris Lindb pumila With sphaerocarpa Hook
14 Scapaniaceae Diplophyllum (Dum) Dum nom cons
apiculatum (Evans) Steph var apiculatum
Scapanill (Dum) Dum nom cons apkuata Spruce carinthillca Jack ex Lindb curta (Mart) Dum var curta
13
cuspiduligera (Nees) K Mull var cuspiduiigera degenii Schlffn ex K Mull var degenii glllucocephaJa (Tayt) Aust gymnostomophila Kaal irrigwJ (Nees) Gott et al subsp irrigua lingulata Buch varlingulata mucronata Buch subsp mucronata nemorosa (l) Dum paludicola Toeske et K Mun var paludicola paludosa (K Mull) K Mull saxicola Schust scandiea (H Arnell et Buch) Macv var scandiea 8Ubalp~ (Nees) Dum umbroSll (SchIad) Dum undulata (L) Dum var undulata
15 Radulaceae Raduia Dum nom COllS
compianata (L) Dum obconica Sull
16 Porellaceae Parella L
pinTlllta L patyphylla (L) Pfeiff platyphylloidt(l (Schwein) lindb
17 Jubulaceae FruUanill Raddi
bolanderi Aust brittoniae Evans eboracemis Gott inlllta Gott oakesiana Aust plana Sull rlparia Hampe ex Lehm selwyniana Pears taTnl1iJci (L) Dum subsp asagmyana (Mont) Hatt
18 lejeuneaceae Calolejeunea (Spruce) Schlffn
biddlecomiae (Aust) Evans
Lejeunea Ub nom cons cavifolia (Ehrh) Lindb emend Buch
19 Codoniaceae Fossombronia Raddi
foveolata Lindb
20 Pa11aviciniaceae Paklvieinlil S Gray nom cons
lyeiii (Hook) Carruth
14
Moerckia Gott hibemica (Hook) Gott
21 PeWaceae Pellfa Raddi nom cons propos
endivlifolfa (Dicks) Dwn epiphylllz (L) Corda neeBiana (GottlUmpr
22 Metzgeriaceae Metzgerla Raddi
conjugata Undb
23 Blasiacae B L
pulilm L
24 Aneurareae AneumDum
pinguis (L) Dum
Riccfl1dia S Gray nom cons latifrons Undb multiftda (L) S Gray var multifida palmata (Hedw) Carruth
25 Aytoniaceae Reboulill Raddi nom conI
hemllphaerica (L) Radeli
ManniaOpiz fragrrms (Balbis) Frye el ClarK pilosa (Hornem) Frye et Clark trltmdra (Scop) GroDe
26 Conocephalaceae Conocephalum Wigg nom cons
conicum (L) Lindb
27 Marchantiaceae Preiuia Corda
quadrata (Scop) Nees
Marchantia L polymorpha L
28 Ricciaceae Riccwcarpus Corda
natans (L) Corda
Riccia 1 beyrichiana Hampe ex Lehm bifurca Horfm
15
i
CtlJPemostI Hoffnt fuitu L rostii Aust BOIOCtI1fHI Bi~ sulltvtmtii Aust
MUSCI (Mosses)
29 Sphagnaceae SpIutgnum L
capillaceum (Weiss) Schrank var Cllpil1tlceum caplllllceum (Weiss) Vir tenellum (Schimp) Andr centrille C Jens ex Amen et C Jens compactummiddot OC ex Lam et Dc ClUpidatum Ehrh ex Hoffin var aupidatum jimbrltltum Wits ex J Hookmiddot fulCUm (Scbimp) KUngrr glrgenBOhIlll Russ imbrlctltum Hornsch ex Russ muelllmitil4m BOd 1IIIIU (RUII) C Jens przIutre L papIIIoIum Lindb pulchrum (Lindb ex braithw) Wamst recumun P-Beauv var recumun recum P-Beauv var tenue Klinggr riptlrlum Ansstr ruPQWItWamst squtl1fOlUm Crome subftltens Russ et Wamst ex Wamst IUbIeCfUldum Nees ex Sturm var subsecundum subsecundum Nees ex Sturm varrutem (Nees Hormch et sUUm) Hueb teres (Sdlimp) Anptr e C Hartm
WtlI7IItorjii Russ wuljfanum Gira
30 Andreaeaceae AndNrrerI Hedw
rupeitrlr Oedw var pest
31 Fissident~e Fluid_ lledw
trdiantoid Hedw byoidei Bedw var bryolda cristaM Wils ex Mitt exiguus sun VIr exigUw onttmUi (B-Pyl) Steud jtuberi Lesq et James obtusifolhu Wits var obtwifoliul ollPlUlldQldes Heel ~litSuU lubbtulkril Bed tfDdfoliul lledw llridubu (S~ ) Wahlenb vallirlIU1u
16
32 Ditrichaceae Pleuridium Brid
subulatum (Hedw) Rabenh
Ditrichum Hampe heteromallum (Hedw) Britt linellre (Sw) lindb
~ pallidum (Hedw) Hampe pusiUum (Hedw) Hampe
Saelania Lindb gIlucescens (Hedw) Bomanss et Broth
Ceratodon Brid purpureus (He(lw) Brid var prupurew
Distichium BSG capillaceum (Hedw) BSG inclinatum (Hedw) BSG
33 Bryoxipbiaceae Bryoxlphium Mitt
norvegicum (Brid) Mitt
34 Seligeriaceae Seligeria BSG
cumpylopoda Kindb ex Macoun et Kindb pusillll (Hedw) BSG recurvata (Hedw) BSG
Bindia BSG acuta (Hedw) BSG
35 Dicranaceae BlUehia Schwegr
lexUOSll (Sw ~ Schwaegr) C MueU sullivantii Aust
Trematodon Michx ambiguus (Hedw) Hornsch
Dicranellll (C Muell) Schimp cerviculata (Hedw) Schimp val cillPicuiata
heteromalla (Hedw) Schimp ~ rufescens (With) Schimp
schreberiana (Hedw) Schhnp var schreberiana schreberitJna (Hedw) Schimp var robusta Schimp ex Braithw subulata (Hedw) Schimp varia (Hedw) Schimp
Dicranodontium BSG denudatum (Brid) Britt ex Williams
Panzeucobryum (Lindh) Loeske Iongifolium (Hedw) oelke varlongifolium
17
Tortella (Undb) Limpr fragilis (Drumm) Umpr humilis (Hedw) JeM torlUoSfl (Hedw) Umpr
Hyophila Brid involuta (Hoole) Jaeg et Sauerb
Didymodon Hedw rigidulus Hedw
BryoerythrophyUum Chen recurvirostnon (Hedw) Chen
Btubula Hedw acula (Brid) Brid var acuta convoiuta Hedw fallox Hedw unguicultltll Hedw
Acaulon C MueD muticum (Uedw) C Muell var rufeseens (Iaeg) Crum
PJl~lm Hedw CUSPidtltum Hedw
De8mlltodon Brid cernuu (Hueb) BSG oblUsifoliw (Schwaegr) Schimp
T01IUla Hedw mueronifolia Schwaegr rurall (Hedw) Gaertn Meyer et Scherb var ruralis
39 Grimmiaceae Grlmmill Hedw
alpicola Hedw var alpicola alpicola Hedw var dupretii (Ther) Crum alpicola Hedw var rlvularl8 (Brid) Wahlenb apocarpa Hedw var apocarpa apocarpa Hedw var ambipa (Sull) Jonea apocarpa Hedw var conjerttl (Funck) Spreng apocarpa Hedw var strleta (Tum) Hook et Tayl CiIIyptrrlta Hook ex Drumm donniana Sm var donn_ hemumti Crum ltlevla (Brid) Brid montana BSG oineyi SuD oVllll8 (Hedw) lindb piUfe P-Beauv pltJgiopodill Hedw pulvinata (Hedw) Sm varbullpulvinata teretinervl Limpr
18
Orothera C Muell leana (Sull) C Muell
Rfwbdoweisia crispata (With) Undb
Cynodontium BSG ex Schimp alpestre (Wahlenb) Milde strumiferum (Hedw) Undb
Dichodontium Schimp pellucidum (Hedw) Schimp
Oncophorus (Slid) Brid vuens (Hedw) Brid wahlenbergii (Brid)
Dicranum Hedw bonjeanii DeNot ex Usa drummondii C Muell lagellare Hedw varj1ogellare fulvum Hook fuscescens Turn var fuscescens montanum Hedw muehlenbeckii BSG var muehlenbeckii polysetum Sw sabuletorum Ren et Card scoparium Hedw undulatum Brid viride (Sull et Lesq ex Sull) lindb
36 Leucobryaceae Leucobryum Hampe
glaucum (Hedw) Angstr ex Fr
37 Encalyptaceae Encalypta Hedw
ciliata Hedw var ciliata procera Bruch
38 Pottiaceae Astomum Hampe
muhlenbergianum (Sw) Grout
Weissia Hedw controversa Hedw var controversa
Gymnostomum Nees Homsch et Stunn aeruginosum Sm recurvirostrum Hedw var recurvirostrum
Trichostomum Bruch spirale Prout
19
40 Ephemeraceae Ephemerum Hampe
cohaerens (Hedw) Hampe crassineium (Schwaegr) Hampe val crassinervium spinulosum Bruch et Schimp ex Schimp
41 Funariaceae AphflllolThegma SuD
serratum (1 Hook et Wils ex Drumm) Sullo
Physcomitrium (Brid) Fuernr hookeri Hampe immersum Sullo pyriforme (Hedw) Hampe
Funaria Hedw americana Undb hygrometrica Hedw vaT hygrometrica
42 Splachnaceae Splachnum Hedw
ampullaceum Hedw
43 Schistostegaceae Schistostega Mohr
pennata (Hedw) Web el Mohr
44 Bryaceae Pohlia Hedw
annolina (Hedw) Lindb var annotina camea (Tum) Lindb cruda (Hedw) Undb elongata Hedw iliformis (Dicks) Andr lescurlana (Sull) Grout nutans (Hedw) Lindb obrusifolia (Brid) L Koch proligera (Undb ex Umpr) Undb ex Amell wahlenbergii (Web et Mohr) Andr
Leptobryum (B S G) Wits pyriforme (Hedw) Wils
Bryum Hedw algovicum Sendtnex C Muell argenteum Hedw caespiticium Hedw capillare Hedw var capillare creberrlmum Tayt lonchocaulon C Muell pallens (Brid) Sw ex Roehl pseudotrlquetrum (Hedw) Gaertn Meyer et Scherb varpudotriquetrum
20
pseudotriquetrum (Hedw) GaertnMeyer et Scherb var bimum (Schreb) Ulj stenotrichum C Muell var stenotrichum tortiolium Funck ex Brid uliginosum (Brid) B S G violaceum Crundw et Nyh weigelii Spreng
Rhodobryum (Schimp) Hampe roseum (Hedw) Limpr
45 Mniaceae Mnium Hedw
ciliare (C Muell) Schimp cinclidioides Hueb cuspidarum Hedw drummondii Bruch et Schimp longirostrum Brid lycopodioides Schwaegr marginatum (With) Brid ex P Beauv mediumB~S G var medium onhorrhynchum Brid pseudopunctatum Bruch et Schimp punctalUm Hedw var punctatum punctatum Hedw var elatum Schirnp rugicum Laur spinufosum B S G stellare Hedw
46 Aulacomruaceae Au1acomnium Schwaegr
androgynum (Hedw) Schwaegr heterostichum (Hedw) B S G paJustre (Hedw) Schwaegr var paJustre
47 Meesiaceae Amblyodon B S G
deillbotus (Hedw) B S G
Meesia Hedw triquetra (Richt) Angstr uJiginosa Hedw
48 Bartramiaceae Plagiopus Brid
oederiana (Sw) Umpr
Bartramia Hedw pomiformis Hedw var pomiformis
Philonotis Brid fontana (Hedw) Brid varfontana longseta (Michx) Britt varlongseta marchica (Hedw) Brid
21
~
~
49 Timmiaceae Timmia Hedw
megapolitana Hedw var megapolitana
50 Orthotrichaceae Zygodon Hook et Tayl
viridissimus (Dicks) Brid var rupestris lindb ex C J Hartm
Amphidium Schimp lapponicum (Hedw) Schimp mougeotii (B S G) Schimp
Orthotrichum Hedw anomalum Hedw obtuaifolium Brid ohioense Sullo et Lesq ex Aust pumilumSw sordidum Sullo et Lesq ex Aust speciosum Neesax Sturm var eegam (Schwaegr ex Hook et Grev) W~rnst stellDtwn Brid strrmgulatum P Beauv
Uleta Mohr ex Web cOfIICtata (p Beallv) Hamrn crispa (Hedw) Brid
Drummondill Hook ex Drumm prorepens (Hedw) Brltt
51 Fontinalaceae Fantinalis Hedw
antipyretica Hedw var antipyrettca antipyretica Hedw forma rabusta (Card) Card dalecarlica Schimp ex B S G var dalecarlica flqccidtJ Ren et Card hypnoides C J Hartrn var hypnoides hypnoides C J Hartm var durillei (Schimp) HU$ll
missourlcaCard novae-angliae Sullo var novae-angliae splulgrUlolill (C Moen) Wijket M~S
DichelY1l1ll Myr capilloceum (With) Myr falcatum (Bedw) Myr HIllescens B S G
52 Climaciaceae aimecium Web et Mohr ex Mohr
americllnum Brid dedroides (Hedw) Web et Mohr
22
S3 Hedwigiaceae Hedwigia P Beauv
ciliata (Hedw) P Beauv
54 Cryphaeaceae Forsstroemia lindh
ohioense (Sull) lindh
55 Leucodontaceae Leucodon Schwaegr
brachypus Blid var brachypus brachypus Brid var aniJrewsitlnus Crum et Anderson julaceus (Hedw) SuU
56 Neckeraceae Neckera Hedw
pennata Hedw var pennata
1fomalia (Brid) B S G trichomanoides (Hedw) B S G
57 Theliaceae Thelia Sull
asprellll Sullo hirtella (Hedw) Sullo lescurii SuU
Myurella B SG julacea (Schwaegr) B S G
sibrica (C MueU) Reim var sibrica
58 Fabroniaceae Fabronio Raddi
ciliaris (Brid) Bnd
Anacamptodon Brid splachnoides (Groet ex Brid) Brid
Schwetschkeopsis Broth fabronia (Schwaegr) Broth
59 Leskeaceae Lindbergia Kindb
brachyptera (Mitt) Kindb
Leskea Hedw gracilescens Hedw obscura Hedw poiycarpa Hedw
Leskeella (limpr) Loeske nervosa (Brid) Loeske
23
Pterlgy1lllndrum Hedw filiforme Hedw
60 Thuidiaceae Haplohymenfum Dozy et MoIk
trlste (Ces ex De Not) Kiodb
Anomodon Hook et Tayl attenuatus (Hedw) Hueb minor (Hedw) Fuemr rostratus (Hedw) Schimp viticulosus (Hedw) Hook et Tayl
Haplocladium (C Mucn) C MueU microphyllum (Hedw) Broth virginillnum (Brld) Broth ~bull
Thuidium B S G abietinum (Hedw) B S G delicatulum (Hedw) B S G vardelicatulum delicatulum (Hedw) B S G var radicans (Kindb) Crum Steere et Anderson minutulum (Hedw) B S G pygmaeum B S G recognftum (Hedw) Lindb scitum (p Beauv) Aust
Helodium (Sull) Warnst blandowli (Web et Mohr) Warnst var blandowli blandowli (Web et Mohr) Wamst var helodioides (Ren et Card ex Roell) Crum
Steere et Anderson paludomm (Sull) AuS
61 Amblystegiaceae Cratoneuron (Sull) Spruce
commulatum (Hedw) Roth var commutfltum commutatum (Hedw) Roth var fakatum (Brid) Moenk ilicinum (Hedw) Spruce var filcinum
Campylium (Sun) Mitt chrysophyllum (Brid) 1 Lange var chrysophyllum hispidulum (Brid) Mitt var hllJidulum polygamum (B S G)C lens varpolygtlmUm stellatum (Hedw) C1ens vat stelltum
Leptodictyum (Schimp) Warnst laxirete (Card et Ther) Broth riparium (Hedw) Wamat trichopodium (Schultz) Warns l1acillans (Sull) Broth
Hygroamblystegium Loelke jlul1iatile (Hedw) oeske var luViatile
24
fluviatile (Hedw) Loeske var oWltum Grout noterophilum (Sun et Lesq ex Sull) Warnst tenox (Hedw) Jenn var tenax tenox (Hedw) Jenn var spinilolium Schimp) Jenn
Amblystegium BSG juratzkanum Schimp serpens (Hedw) BSG varlum (Hedw) Undb var varium
Platydictya Berk confervoides (Brid) Crum jungermanniodes (Brid) Crum minutissimum (Sull et Lesq ex Sull) Crum subtile (Hedw) Crum
Drepanocladus (C Muell) Roth aduncus (Hedw) Warnst var aduncus aduncus (Hedw) Warnst var capillilolius (Wamst)Riebm aduncus (Hedw) Warnst var kneiffii (BSG) Moenk aduncus (Hedw) Warnst var polycorpus (Bland ex Voit) Roth exannulatus (BSG) Warnst fluitans (Hedw) Warnst var fluitans revolvens (Sw) Wamst var revolvens sendtneri (Schimp) Warnst var sendtnerl uncinatus (Hedw) Warnst var uncinatus uncinatus ffiedw) Warnst var piwnosus (Schimp) Wamst vernicoms (Lindb ex C Hartm) Warnst
Hygrohypnum Lindb eugyrium (BSG) loeske luridum (Hedw Jenn varluridum ochraecum (Tum ex Wils) Loeske var ochraceunt
Calliergon (Sull) Kindb cordilolium Hedw Kindb glganteum (Schimp) Kindb var giganteum richardsonii (Mitt) Kindb ex Wamst stramineum (Brid) Kindb trilarium (Web et Mohr) Kindb
Calliergonella Loeske cuspidata (Hedw) Loeske
Calliergidium (Ren) Grout bakeri (Ren) Grout
Scorpidium (Schimp) Urnpr scorpiodides (Hedw) limpr
62 Brachytbeciaceae Tomenthypnum loeske
nitens (Hedw) Loeske var mtens
25
Homalotheciella (Card) Broth subcapillata (Hedw) Broth
Brachythecium BSG acuminatum (Hedw) Awt acutum (Mitt) Sullo calcareum IGndb campestTe (C MuelL) BSG cunum (Lindb) Umpr digastrum C Muell et Kindb ex Macoun et Kindb oxycladon (Brid) Jaeg et Sauerb var oxycladon plumosum (Hedw) BSG populeum (Hedw) BSG var populeum reflexum (Starke ex Web et Mohr) DsG var relexum reiexum (Starke ex Web et Mohr) DSG var pacificum Renet Card ex Roell rlvulare BSG var rivulare tabulum (Hedw) BSC var tabuum salebrosum (Web et Mohr) BSG var salebrosum velutinum (Hedw) BSG Vat velutinum
Bryhniil Kaur graminicolor (Brid) Grout var graminicolor novae-angliae (Sull et Lesq ex sun) Grout
Rhynchostegium BSG senulatum (Hedw) Ja4lg et Sauerb
Rhynchostegiella (BSG) Limpr compacta (C MueU) Loeske
EU1hynchium 8SG hians (Hedw) Sande Lac pulchellum (Hedw) Jenn var pulchelium riparloides (Hedw) Rich
Stokeriella (Kindb) Robins pmelonga (Hedw) Robins var stokesii (Tum) Crum
63 Entodontaceae Entodon C Muell
brevisetus (Hook et Wils ex Wils) Lindb cladorrhizans (Hedw) C Muell seductrlx (Hedw) C MueU var wluctrix
Pleurozium Mitt schreberl (8rid) Mitt var schreberi
64 Plagiotheciaceae Pltlglothecium BSG
CflVifolium (Brid) Iwats var CtWifolium denticulatum (Hedw) 8SG laetum BSG ltztebricola BSG
26
6S SernatophyUaceae Brotherella Loeske ex Fleisch
recumins (Michx) Fleisch tenuirostris (Bruch et Schimp ex Sull) Fleisch
Sematophyllum Mitt demissum (Wils) Mitt
66 Hypnaceae Pl(itygyrium BSG
repens (Brid) BSG
PylaisieUa Kindb intricata (Hedw) Grout polyantha (Hedw) Grout selwynii (Kindb) Crum Steere et Anderson
Homomallium (Schimp) Loeske adnatum (Hedw) Broth
Hypnum Hedw cupressiforme Hedw var cupressiforme cupressifonne Hedw varfiliforme Brid curviolium Hedw fertile Sendtn imponens Hedw lindbergii Mitt palescens (Hedw) P-Beauv pratense Koch ex Brid
Calicladium Crum haldanianum (Grev) Crum
sopterygiopsis Iwats muelleriana (Schimp) Iwats
Isopterygium Mitt e1egans (Brid) lindb pulchellum (Hedw) Jaeg et Sauerb tenerum (Sw) Mitt
Taxiphyllum Fleisch depllQtum (Bruch et Schimp ex sun) Fleisch taxirameum (MitL) Fleisch
Herzogiella Broth seligeri (Brid) Iwats striatella (Brid) Iwats ruracea (Lindh) Iwats
hilium De Not crista-castrenna (Hedw) De Not
27
67 Rhytidiaceae Rhytidium (Sull) Kindb
rugosum (Hedw) Kindb
Rhytidfodelphus (Lindb ex Iimpr) Warnst triquetrus (Hedw) Warnst var triquetrus
68 Hylocomiaceae Hylocomium BSG
pyrenaicum (Spruce) lindb splendens (Hedw) BSG varsplendens
69 Buxbaumiaceae Buxbaumia Hedw
aphyllll Hedw
Diphyscium Mohr follosum (Hedw) Mohr
70 Tetraphidaceae Tetraphi3
pellucida Hedw
71 Polytrichaceae Atrichum PmiddotBeauv
angustatum (Brid) BSG cri8pum (James) Sullo oerstedianum (C Muell) Mitt teneUum (Roehl) BSG undulatum (Hedw) P middotBeauv var unduiatum
Pogonatum P-Beauv alpinum (Hedw) Roehl var alpinum umigerum (Hedw) P-Beauv
Poiytrichum Hedw commune Hedw var commune ormosum Hedw juniperinum Hedw longisetum Brid ohioense Ren et Card piliferum Hedw var pilierum strictum Brid
28
MAPS OF BRYOPHYTE TAXA
REPORTED FOR WISCONSIN
HORNWORTS AND LIVERWORTS
29
bull bull WSHeV~N S vE II
0 ~
e~RRON ~USK
(J~ (j -
CHIPPE A DUNN
0 o0 poundAU CLAIRE
~ TREMshyPEAshyLEAU
MICE
ONEIDA
Q LINCOLN
TAYLOR
Q t-~----- MOIRA THON
CLARK
o
10
LA FAVETTE
Hornwor ts amp Liverworts
- None - 1 - 25 taxa
- 26 - 50 tRXshy
- 51 - 75 tax)
- 7c - 100 taxa
FONDOU j LAC ~
DODGE
i O JEfFERSON
WALWORTH
30
Anastrophylltlfl helleranum (Nees) Schust Anastrophyllum michauxil (We~ Buch ~
46 88 136 141 Evans 38 46 141
Anastrophyllum mlnut~ (Schreb) Schuster Aneura plnguls (L) Durn
89 21 384687 88 89 90 (138) (145)
Anthoceros macounil MA Howe ~oceros punctatus L
21 (45) (144) 34
31
Barbilophozia attenuata (Mart) Loeske Barbilophozia barbata (Schmid ~ Schre~
24 141 Loeske 28 39 46 89 90 141
Bazzania trilobata (1) S Gray
1 26 39 46 (57) (59) 79 88 89
90 119 141
BlePbarostoma trichoffiyllum (1) Dumort
1 8 26 39 46 89 90 137 141
Blasia pusilla 1
1 39 46 90
Calypogeja fissa (1) Raddi
90
32
$129geja integristipula Steph C~Ypound2geja muelleriana (Schifm) K Mull
88 89 89
CallpoundO~eja neesiana (Mass amp Carest) K Mull
39 46 89 140 (141)
~pogeja sJlapound2icola (H Arnell tl J bull
Perss) Warnst et Loeske (119) 141
CeIalozia biscuspidata (L) Dum
1 37 I 46 90 t 137 141
CalYpogej~ trichomanis (L) Corda
26 39 (119) 137
33
CeEhalozia catenulata (Hub) Lindb Cephalozia connivens (Dicks) Lindb
39 46 79 141 39 4679 88 (89) 90 119 141
Cephalozia lacinulata Jack ~ Spruce
141 Cephalozia lunulifolia (Dum~ Dum I 28 39 46 79 88 89 90 141
Cepoundhalozia macouni (Aust) Aust Cephalozia ~leniceps (Aust) Lindb 27~ 3746 (66)7987 (139)141 39 46 79 89 141
34
Ceffialoziella 9ivaricata (Sm) Schiffn Cephaloziella ~hista (Jack) Schiffn
46 90 46 (119) 141
Cephaloziella hampeana (Nees) Schiffn
46
Cephaloziella rubella (Nees) Warnst
46 39
Cephaloziella rubella var sullivantii
(Aust) K Mull 46 39
Cephaloziella rubella var bifida (Lindb) Douin
35
46
Chiloscyphus pallescens (Ehrh ~ Hof~ Dum Chiloscyphus pallescens var fragilis
39J 46 88 89J 90 (Roth) K Milll 46
Chiloscyphus E9lyanthos (L) Corda
1 24 39 46
Chiloscyphus polyanthos vax rivularis
(Schrad) Nees 39 46
poundpoundnocepha1um conicum (1) Lindb
12 28 39 46 (49) 87 88 8990 120 121 148
Cololejeunea biddlecomiae (Aust) Evans
39 46 62 87 88 89 90
36
Dipl0]hyllum apicu1atum (Evans) Steph Fossombronia foveo1ata Lindb
(35)41 46 90 140 1 t 38 (39) t 46
Frullania bolanderi Aust
13 46 62 88 90 139 140
Frullanla ari ttoniae Evans
39 46 69 (71) 87 88 89 90
Frullania eboracensis Gott
1 9 28 4 39 46 (57) (59) 69 70 (71) 87 88 89 90 92 93 97 148
Frullania inflata Gott
46 87 88 89 (139)
Frullania oakesiana Aust Frullania plana Sullo
(35) 42 46 87 (139) (140) 92
Frullania ri paria Hampe ~ Lehm
46 69 71 87 89
Frullania ~elwynianna Pears
46 89
Geocalyx SEaveolens (Schrad) Nees
394688 8990 (119) 140
Frullania tamarisci (L) Dum subsp sPagrayana
(Mont) Hatt 28 34 l~6 t (57) 88 89 90
38
Mar pan thus seutatUB ( Web amp Mohr) Spruce Jamesoniella autumnalis (DC) Steph
(35) 39 46 90 1 7 28 39 46 56 (57) 87 8889 90 (119) 137 141
Jungermannia confertissima Nees
137 141
Jungermannia crenuliform1s Aust
79 137 141
Jungermannia ~il1ima Sm
46 90 141 Jungermannia exsert1folia subsp corifo~ (Dum~
Vana (35) 46 79 141
39
Jungermannia hyalina Lyell Jungerrr6nnia 1eiantha Grolle
40 46 79 136 141 39 46 79 88 90 1)7 141
Jungermannia IXlaxis Lindb Jungermannia pumila With
46 67 79 (139) (140) 141 )9 46 90 1)6 141
Jungermannia sphaerocarpa Hook Kurzia setacea (Web) Grolle
4679136140IIH 136
Lejeunea cavifo1ia (Ehrh) Lindb Lepidozia reptans (1) Dum
1 7 27 39 46 88 89 90 137 14010 I ( 27) 46 I 69 (71) 88
lophocolea heterophylla (Schrad) Dum Lophocolea minor Nees
1 )4 39 46 (57) 87 88 89 27 39 46 88 89 90 (119) 90 (93)94 119 136
Lophozia al~stris (Schleich ~ Web) Evans Lophozia ascendens (Warnst) Schust
46 90 137 141 136 140 141
41
loIilozia badensls (Gatt ~ Rabenh)
46 141
lophozia blcrenata (Schmid sectA Hoffm)
Dum 46 141
lophozia collaris (Nees) Dum
39 46 141
lophozla excisa (Dicks) Dum
27 4679 141
lophozia gillmani (Aust) Schust lophozia gIlttulata (Undo tl H Arnell)
36 39 40 46 136 (140) 141 Evans 39 46 136 141
42
141
Lophozia heterocolEDs (Thed) MA Howe IoPhozia ipcisa (Schard) Dum
27 39 46 89 141 8 27 31 46 79 90 140 141
LoQhozia longidens Macoun Lophozia ventricosa
1 46 79 90 141
Lophozia ventricosa var silvicola
(Buch) Jones 141 LoEhozla ventricosa vax longif1ora (Nees)
Macoun 39 46 79 141
43
~~ fragrans (Balbis) Frye et Clark Mannia pi 10sa (Hornem) Frye et Clark
46 (139) (140) 9 46 79 116 139
Mannia triandra (Scop) Gro 11e
46 (140)
Marchantia pound91ymorpha L
1 13 28 3946 (49) 78 87 88 89 90 (Ill) 121 122 147
Mo~rckia hibernica (HookJ Gott
46 89
Metzgeria conjugata Lindb
44
46
Mylia anomala (Hook) S Gray Notothylas orbicularis (Schwein) Sullo
39 46 141 (35)45 46
Nowellia curvifolia (Dicks) Mitt
17 27 3846 88 89 90 141
Odontoschisma denudatuJI1 (Nees ~ Mart)
Dum 46 81 137 141
Pallavicinia lyellii (Hook~ Carruth
90
Odontoschisma macounii (Aust) Under
45 46 (66) (139) 140 141
45
=
Pellia epiphllla (L) CordaPellia endiviifolia (Dicks~ Dum
27 39 46 90 13724 39 46
Pellia neesiana Limp bull Phaeoceros
39 46 89 27 39 45 46 97
Porella pinnata L
13 27 I 46 90 Plagiochila asplenioides (L) Dum
27 I 39 1+6 88 89 I 90 137
46
~e11a platYEhylla (L) Pfeiff Porella platyphylloi~ (Schwein) Lindb
17 27 39 46 69 (71)79 87 88 89 9 34 46 (57) 59 87 88 89 92 90 92 93 77
Preissia quadrata (Seop) Nees
1 36 3946 (71) 88 89 90 121 123
137
Ptllidium ciliare (L) Hampe
1 27 39 46 132 141
Dum
(22) 28 3946 (57) 87 88 89 90 92
Ptilidium ~lcherrium (G Web) Hampe
(9) 13 394656 (57)70 87 88 89 90 93 (119) 136 141
47
Radula obconica Sullo Reboulia hemlsphaerlca (L) Raddi
39 46 88 89 (43) 46 (59) 70 90 t Cf 124 125 (139)
Riccardia ~atifrons Lindb
1 27 39 46 89
Riccardia multifi~ (L) S Gray
39 46 88 145
Riccia beyrichiana Hampe ~ Lehm
46 Riccardia palmata (Hedw) Carruth
39 46 90
48
Riccia bifurca Hoffm Riccia cavernosa Hoffm
48 13739 46 79 137
Riccia fluitans L
242731454679
Riccla frostli Aust
48 79
Riccia sullicantii Aust
79 (137) Riccia sorocarpa Bisch 46 (137)
49
Ricciocarpus natans (1) Corda Sca~ia apiculata Spruce
13 27 46 79 89 110 39 46 (139) 141
ScaJltll1ia carinthica Jack ~ Lindb
137 t 141
Scapania ~ (Mart) Durn
39467990137141
Scapania digenil Schiffn
90
ScaEsectflia cuspiduligera (Nees) K Mull
46 (139) (140) (141)
50
Scapania glaucoeerha1a (Tayl) Aust Seapania gyrrmostomophilia Kaal
39 46 (139) 141 41 44 46 64 (139) (140) 141
ScaP3poundlia irrigua Gott ~ almiddot
7 39 46 79 137 (140L 141
Seapania nemorosa (L) Dum
1 39 46 89 90 137
Scapmi~ ~~ta Smiddotlch
137 141 51
~caPnia poundludicola LDeske amp K Mull Scapania saxicola Schust
46 (66) 141 136 141
Sc~~ scandica (H Arnell ~ Buch) Macv
139 140
Scapania subalpina (Nees) Durn
39 40 46 79 137 141
Scapania undulata (L) Dum
1 42 46 79 141
Scapania umbrosa (Schrad) Dum
137 140 141
52
Trichocolea tomentella (Ehrh) Dum Triomarla exsecta (Schmid) Schiffn
27 (35) 39 46 47 88 137 141 8 28 39 46 90 140 141
Tritomaria exsectiformis (Breidl) Loe~ke Tritomaria Sluinguedentata (Huds) Buch
39 -t6 89 90 136 140 141 7 39 46 141
Trltomaria scitula (Tayl) Joerg
46 136 (139) 141
53
CONTENTS
ABSTRACT 4
ACKNOWLEDCEMENTS 4
INTRODUCTION 6
GEOGRAPHIC CLIMATIC AND BOTANICAL fEATURES OF WISCONSIN 7
SYSTEMATIC ARLNGEMENT OF WISCONSIN BRYOPHYTES 11
MAPS OF BRYOPH) iE TAXA REPORTED FOR WISCONSIN 29
HORNWORTS AND LIVERWORTS 30
MUSCI 54
FOSSIL AND SUBFOSSIL MOSSES OF WISCONSIN 119
PALEONTOLOGICAL Rb J3RENCES 119
SYNONYMS AND EXCLUDED TAXA 120
PIONEER BRYOLOGISTS OF WISCONSIN 126
LITERATURE CITED 127
3
ABSTRACf
117This atlas of Wisconsin bryophytes shows the distribution of)30 taxa (137 hepatics 389 mosses)
on state maps with a dot in each county for which each taxon has been reported in the literature A list of synonyms and excluded taxa is included along with an extensive bibliography Pertinent features of the natural history and history of bryology in the state are discussed
ACKNOWLEDGEMENTS
We want to thank Drs Fred Hermann and John Thomson of the University of Wisconsin-Madison herbarium and Botany department for locating and lending packets and for references to aid us in accumushylating a more accurate complete report The illustration of Climacium dendroides on the cover was drawn by Robert Freckmann Of course we are grateful to all the people who have collected specimens of Wisshyconsin bryophytes and preserved them in herbariums making this distributional study possible
4
ONEIDA
LINCOLN
TAYLOR
t---------- MARATHON C1l-RK
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t
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~ C i-W=AS_-r7-1
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WALWORTH
5
INTRODUCfION
This atlas of the bryophyte flora of Wisconsin is an attempt to show the distribution of all bryoshyphyte species known to occur in the state by county occurrenoes based on published reports in the literashyture It is not an annot~ted check list (only a feW problematic taxa were examined) but is a literature search intended as a foundation for a complete flora or for otherbryological research
The literature of btYo1ogy is large and spread over numerous journals reports monographs and manuals It would be almost impossible to search all sources for every reference attributing bryophyte species to Wisconsin and to particular localities in the state Wehav checked many inlportant references and paid special attention to works dealing with Wisconsiit botany and the bryologyof the upper Midwest Several bibliographies and indices were checked thoroughly including Green an~ Curtis (1955) the Bry ologist indices (Crum 1959 and Miller 1976) andt)le Wisconsin Academy of Sciences Arts and Letters indices
Only those published repots which app~red to)e substanti2ted by herbarium specimens and which gave- ~fficient location dat to P~CetheW(On in amiddot particula(county were-used in the initial compilation Species described in manuals in generalized way implying their occurrence in Wisconsin (Without menmiddot tioning Wisconsin specifically or citing a particular location in the state) were excluded eg Gnmmia raui (Wynne 1943) and Pohlia annotinll var decipiens (Crum 1976) Ecological papers which were either too general in taxonomic determinations (eg Sphagnum sp Polytrichum sp) or in location data were also excluded Some other taxa attributed to Wisconsin which have been excluded are
Cylindrothectum compr~ (Cheney 189tmiddotDane Co) identified later by Grout as Entodon seducshytrtx
Dicranell4 humilis (Holzjne~ 1925middotBuffalo Co) probablyD vllJia since Dhumilis is not found in North Arnerl~
Didymodon triarium (Foote 1 966-SW Visconsin) identified by Fred Hennann as Barbulaacuta Funoria microstoma (anonymous 1975middotDane Co) excluded because it is outside of its geographic
range and the packe~ QaMot be located Phil(motis calcarea (Cheney and Evans 1944-Dane Co) excluded in Crum Steere and Anderson
(1973) Plagiobryum demissum and P zierl (Cheney and Evans 1944-various counties) are both Arcticshy
alpine species not likely to be found in isconsin Two of Cheneys collections of Pl1giobyrum have been referred to Bryum
Pogonatum alpinum var septentrionale (Che~ey and Evans 1944-Chippewa Co) was misidentified (Ireland 1960) and is P umigerum
Rhizomnium fAPpoJachIanum and R punctalu1fl ssp chlorophyDosum (Koponen 1973middotseveral counshy ties) excluded North American bryologiflts have not recognized these taxa in America
Sphagnum tenellum (Cheney and Evans 1944-Waukesha Co)an Atlantic species not occurring in Wisconsin (Cruro 1975)
The nomenclature and systematic arrangement of hornwort 00 hepatic species in the atlas follow Stotler and Crandall-5totler (1977) and of moss species fonow Crum Steere and Anderson (1973) except in such instances as the JreptlIfockIdus fonns whiGh follow Wijk et aL (19S9middot1973) Species reported in the literature under names which are now listed as synonyms on either of these lists were converted to the accepted name A dot is placed in each county of Wisconsin from which the taxon has been reported The numbers after the name of the taxon correspond to the numbered references in the bibliography which giye ~act location data OEat least location data -s~fficient to place it in-particular counties in the state The numbers in parentheses indicate references which attribute the taxon to Wisconsin without providing exact loltation data
Fossil and subfossil reports are summarized Slfparately in our atlas only six species (Aulacomnium turgidum lJ1achythecium turgidum Ditrichum Jlelficaule Hypnum bambergti Scorpidium 6fJisities and Tortel14 1rIgi1i$) are known exclUsively from the fossil record ~
6
GEOGRAPHIC CLIMATIC AND BOTANICAL FEATURES OF WISCONSIN
Wisconsin is a north central state bounded on the east by Lake Michigan on the north by Michigan and Lake Superior on the south by Illinois and on the west by the Mississippi and St Croix rivers It is located midway between the Prime Meridian (0deg) of Longitude and 1800 Longitude and midway between the equator and the north pole thus placing it in the center of the northern half of the western hemisphere
The topography of the state is relatively flat a few hills exceed 1900 feet with Thns Hill at 1952 feet forming the highest point in Price County Lake Michigan represents the lowest point at 580 feet The northern and eastern two-thirds of the state were heavily glaciated during the last stage of the Pleistocene and now contain many of the best examples of continental glaciation topOgraphy including considerable ground moraine large terminal recessional and intetlobate moraines kames eskers drumlins and small depressions or kettle-holes Most of the nearly 9000 lakes of 20 acres or more lie within this part of the state The southwestern third of the state is the Driftless Area It includes most of the large flat generally sandy bed of Glacial Lake Wisconsin plus the highly dissected ridge and valley section with numerous cliffs and rock-outcroppings (Figure 1)
Much of the bedrock in the state lies below glacial till but exposures include igneous and metamiddot morphic rocks primarily granitic especially in the north as well as sedimentary layers primarily sandshystones and dolomitic limestones especially along Lake Michigan and in the DriftlesS Area
The climate of Wisconsin is classified as humid-temperate with cold winters The annual precipimiddot tation averages approximately 30 inches which is sufficiently even in its seasonal distribution to support forest vegetation throughout most of the state Nevertheless according to Curtis (I 959) the pre-settlement vegetation of the state (J 830-1850) included about 27 prairie and oak-savanna plus about 64 forests
The state is presently divided into seventy-two counties Marathon County is the largest (1586 sq mi) the smallest counties include Pepin (235 sq mi) Ozaukee (236 sq mi) andMilw8ukee (237 sq mi) We included Menominee County though its future status as a county is in doubt since it regained its Indian reservation status and rights in 1973 This county was established April 30 1961 from the Menomishynee Indian reservation which consisted of parts of Shawano and Oconto counties Wisconsin ranks 26th in the United States in area with 54464 square miles and 16th in population with 4651000
More ecological research is needed on the bryophyte flora of Wisconsin especially to relate it to the vegetation and to such other environmental features as soils climate and glaCial deposits Plant communishyties in Wisconsin are many and diverse these are summarized on Figure 2 from studies conducted by Curtis (1959) The state is divided into northern and southern floristic provinces by a tension zone deshyscribed by Curtis (1959) This tension zone is an eeotonal region vatying from 10middot31 mt wide where the ranges of many vascular plants overlap or terminatej it runs in an S-curve northeast from Milwaukee to Hudson in St Croix County (Figure 2) The principal plant communities of the northern province apshyproaching their southern boundaries within the zone include the northern conifermiddothardwood forests of sugar maple white pine hemlock and yellow birch and the Sphagnum bogs with tamarack and black spruce Southern communities at or near their northern limits within this zone include prairies oak and jack pine savannas and hardwood forests of maples oaks ash ehn and cottonwood
We hope to encourage research on cryptogamic ecology with our checklist We know that some species are northern while others are southern (eg cimacium) fits the tension zone concept Freckmann (I977) while Bryoxiphium is only found in the Driftless Area of the state Steere (1937) and Love and Love (1953) The tension zone theory may prove to be acceptable for bryophytes We speculate that certain bryophytes can indicate particular plant communities since many bryophytes are restricted to preshycise environmental requirements Many papers or theses have dealt with cryptogamic bark and rock ecology in Wisconsin (Culberson 1955 Foote 1962 Hale 1952 1955 and Cole 1975)
We hope that this atlas will stimulate collecting in counties poorly sampled so that our knowledge of the bryophytes can be increased It should aid anyone interested in floras for preservation environmental impact statements and distributional studies
7
A numerical summary of the bryophyte flora of the state based on our literature search is presented in Table 1 (below) This includes 517 taxa 70 families and 176 genera This numerical summary is comshypared with the published floras of Michigan mosses (Crum 1976) and Minnesota hepatics (Schuster 1953) Thus we surmise that further collecting could lead to the same approximate numbers for Wisconsin bryoshyphytes
Table 1 Comparison of Recorded Bryophytes of Wisconsin with Michigan and Minnesota and Summary of Wisconsin Taxa
Wisconsin Michigan (Crum 1976)
Minnesota (Schuster 1955)
Hepaticsamp Hornworts
Taxa Families Genera
137 28 47
174 32 52
Mosses
Taxa Families Genera
380 42
129
503 42
149
8
-
~
~ tIt
Figure 1 The Driftless Area and Tension Zone The counties are 01$0 shown on map
ca location of tension zone as deecfibed bv Curtis (1959)~ The Orittless Area
9
Agn 2 Major Plant Communitiee of Wsconsln aaed on surwvor recorda bull co 18laquo) After Curtismiddot 1959
tit beol Forest e Phillie
Northernmiddot hardwood forest reg Pine 5cMInna
copy Southernmiddot hordwood forest CD Oak Savanna
10
SYSTEMATIC ARRANGEMENT OF WISCONSIN BRYOPHYTES
ANTHOCEROTAE (Hornworts)
1 Anthocerotaceae Anthoceros 1 emnd Prosk
macounii M A Howe punctatum 1
Phaeoceros Prosk laevis (1) Prosk subsp laevis
Notothylas Sul1 orbicularis (Schwein) Sullo
HEPATICAE (liverworts)
2 PseudolepicoJeaceae Blephllrostoma (Dum emend Undb) Dum
trlchophyllum (L) Dum subsp trlchophylluni
3 Ptilidiaceae Ptilidium Nees
ci1iare (L) Hampe pulcherrimum (G Web) Hampe
4 Trichocoleaceae Trichocolea Dum nom cons
tomentella (Ehrh) Dum
5 Lepidoziaceae Kunia von Martens
setacea (Web) GroUe
Lepidozia (Dum) Dum reptans (1) Dum
Bazzania S Gray trilobata (1) S Gray var triZobata
6 Calypogejaceae 011ypogeja Raddi
flSsa (L) Raddi subsp issa integristipula Steph muelleriana (Schiffn) K Mull subsp muelleriana neesiana (Mass et Carest) K Mull sphagnicola (H Arnell et J Perss) Warnst et loeske trichomanis (1) Corda
11
7 Cephaloziaceae Cepluzlozla (Dum emend Schiffn) Dum
biscwpidata (L) Dum subsp biclupidIIta catenulllttl (Hub) Lindb coMivens (Dicks) Lindb var connivens lacinultzta Jack ampX Spruce lunulifolilz (Dum) Dum macounii (Aust) Aust pleniceps (Aust) Lindh var pleniceps
Noweilia Mitt cunifolilz (Dicks) Mitt
8 Adelanthaceae Odontoschimuz (Dwn) Dum
denudatum (Nees ampX Mart) Dum Vir denudatum mtlCount (Aust) Underw
c
9 Cephaloziellaceae CephtlJoziellll (Spruce) Schiffn nom C()II$
divarlctzt4 (Sm) sclurfn val divarlcat(l elachhta (bdc) Schiffn 1uImpetuw (Nees) Schirfn IUbellll (Nees) Warnst vat IUbeIl4rube (Nees) Warnst var biid4 (Lindb) Dquinrube (Nees) WarDSt Var sulliwmtii (Aust) K MuD
10 Geo~c~ larptuttllus Nees
scutatus (Web et Mohr) Spruce
Geocalyx Nees grGVeolens (Schrad) Nees
11 Lophocoleaceae Lophocolea (Dum) Dum
heteropllyll4 (Schrad) Own minorNees
Chiloscypbus Corda om cons pallescens (Ehrh ampX Hoffm) Dum varfragllis (Roth) K Mun polyantitos (L) Corda var poiyanthos pol)llaquo1lthos (L) Corda var rlvulllris (Schrld) Nees
12 PlagiochiJaceae Plagio(hila Dum nom cons
tlBpienioides (L) Dum var asplenioidu
13 Jungermanniaceae Btriilophoril Loeske
attenutlta (Mart) Loeske bIubtlta (Schmid ampX Scltreb) Loeske
12
Lophom (Dum) Dum alpestris (Schleich ~ Web) Evans ascendens (Warnst) Schust badenais (Gott ex Rabenh) Schiffn var bademii bierenata (Schmid ex H)ffm) Dum eollam (Nees) Dum exci8a (Dicks) Dum var excisa gllimanii (Aust) Schust varglllmanii guttulata (Lindb et H Arnell) Evans heterocolpos (Thed) M A Howe var heterocolpos ineistl (Schrad) Dum var tnelsa longidens (Lindb) Macoun subsplongidens ventricosa (Dicks) Dum var ventricosa ventricolfl (Dicks) Dum varlontifIotrl (Nees) Macoun ventricosa (Dicks) Dum var silvlcola (Bueh) Jones
Tritomarill Schiffn ex Loeske exseeta (Schrad) Loeske exsecttformi$ (Breidl) Loeske subsp exsectiormfs quinquedentata (Huds) Buch var quinquedentata scitula (rayl) Joerg
Anastrophyllum (Spruce) Schiffn hellemnum (Noes) Schust michtzuxii (Web) Buch ex Evans minutum (Schrob) Schust var minutum
Mylill S Gray nom cons anomala (Hook) S Gray
Jamesoniella (Spruce) Carring autumnalis (DC) Steph var autumnalis
Jungemumnia L emend Dum eonfertissimtt Nees crenuliformis Aust exsertijolill Steph subsp cordifolia (Dum) Vana graciUima Sm hyalina Lyell leillnthll Grolle polaris Lindb pumila With sphaerocarpa Hook
14 Scapaniaceae Diplophyllum (Dum) Dum nom cons
apiculatum (Evans) Steph var apiculatum
Scapanill (Dum) Dum nom cons apkuata Spruce carinthillca Jack ex Lindb curta (Mart) Dum var curta
13
cuspiduligera (Nees) K Mull var cuspiduiigera degenii Schlffn ex K Mull var degenii glllucocephaJa (Tayt) Aust gymnostomophila Kaal irrigwJ (Nees) Gott et al subsp irrigua lingulata Buch varlingulata mucronata Buch subsp mucronata nemorosa (l) Dum paludicola Toeske et K Mun var paludicola paludosa (K Mull) K Mull saxicola Schust scandiea (H Arnell et Buch) Macv var scandiea 8Ubalp~ (Nees) Dum umbroSll (SchIad) Dum undulata (L) Dum var undulata
15 Radulaceae Raduia Dum nom COllS
compianata (L) Dum obconica Sull
16 Porellaceae Parella L
pinTlllta L patyphylla (L) Pfeiff platyphylloidt(l (Schwein) lindb
17 Jubulaceae FruUanill Raddi
bolanderi Aust brittoniae Evans eboracemis Gott inlllta Gott oakesiana Aust plana Sull rlparia Hampe ex Lehm selwyniana Pears taTnl1iJci (L) Dum subsp asagmyana (Mont) Hatt
18 lejeuneaceae Calolejeunea (Spruce) Schlffn
biddlecomiae (Aust) Evans
Lejeunea Ub nom cons cavifolia (Ehrh) Lindb emend Buch
19 Codoniaceae Fossombronia Raddi
foveolata Lindb
20 Pa11aviciniaceae Paklvieinlil S Gray nom cons
lyeiii (Hook) Carruth
14
Moerckia Gott hibemica (Hook) Gott
21 PeWaceae Pellfa Raddi nom cons propos
endivlifolfa (Dicks) Dwn epiphylllz (L) Corda neeBiana (GottlUmpr
22 Metzgeriaceae Metzgerla Raddi
conjugata Undb
23 Blasiacae B L
pulilm L
24 Aneurareae AneumDum
pinguis (L) Dum
Riccfl1dia S Gray nom cons latifrons Undb multiftda (L) S Gray var multifida palmata (Hedw) Carruth
25 Aytoniaceae Reboulill Raddi nom conI
hemllphaerica (L) Radeli
ManniaOpiz fragrrms (Balbis) Frye el ClarK pilosa (Hornem) Frye et Clark trltmdra (Scop) GroDe
26 Conocephalaceae Conocephalum Wigg nom cons
conicum (L) Lindb
27 Marchantiaceae Preiuia Corda
quadrata (Scop) Nees
Marchantia L polymorpha L
28 Ricciaceae Riccwcarpus Corda
natans (L) Corda
Riccia 1 beyrichiana Hampe ex Lehm bifurca Horfm
15
i
CtlJPemostI Hoffnt fuitu L rostii Aust BOIOCtI1fHI Bi~ sulltvtmtii Aust
MUSCI (Mosses)
29 Sphagnaceae SpIutgnum L
capillaceum (Weiss) Schrank var Cllpil1tlceum caplllllceum (Weiss) Vir tenellum (Schimp) Andr centrille C Jens ex Amen et C Jens compactummiddot OC ex Lam et Dc ClUpidatum Ehrh ex Hoffin var aupidatum jimbrltltum Wits ex J Hookmiddot fulCUm (Scbimp) KUngrr glrgenBOhIlll Russ imbrlctltum Hornsch ex Russ muelllmitil4m BOd 1IIIIU (RUII) C Jens przIutre L papIIIoIum Lindb pulchrum (Lindb ex braithw) Wamst recumun P-Beauv var recumun recum P-Beauv var tenue Klinggr riptlrlum Ansstr ruPQWItWamst squtl1fOlUm Crome subftltens Russ et Wamst ex Wamst IUbIeCfUldum Nees ex Sturm var subsecundum subsecundum Nees ex Sturm varrutem (Nees Hormch et sUUm) Hueb teres (Sdlimp) Anptr e C Hartm
WtlI7IItorjii Russ wuljfanum Gira
30 Andreaeaceae AndNrrerI Hedw
rupeitrlr Oedw var pest
31 Fissident~e Fluid_ lledw
trdiantoid Hedw byoidei Bedw var bryolda cristaM Wils ex Mitt exiguus sun VIr exigUw onttmUi (B-Pyl) Steud jtuberi Lesq et James obtusifolhu Wits var obtwifoliul ollPlUlldQldes Heel ~litSuU lubbtulkril Bed tfDdfoliul lledw llridubu (S~ ) Wahlenb vallirlIU1u
16
32 Ditrichaceae Pleuridium Brid
subulatum (Hedw) Rabenh
Ditrichum Hampe heteromallum (Hedw) Britt linellre (Sw) lindb
~ pallidum (Hedw) Hampe pusiUum (Hedw) Hampe
Saelania Lindb gIlucescens (Hedw) Bomanss et Broth
Ceratodon Brid purpureus (He(lw) Brid var prupurew
Distichium BSG capillaceum (Hedw) BSG inclinatum (Hedw) BSG
33 Bryoxipbiaceae Bryoxlphium Mitt
norvegicum (Brid) Mitt
34 Seligeriaceae Seligeria BSG
cumpylopoda Kindb ex Macoun et Kindb pusillll (Hedw) BSG recurvata (Hedw) BSG
Bindia BSG acuta (Hedw) BSG
35 Dicranaceae BlUehia Schwegr
lexUOSll (Sw ~ Schwaegr) C MueU sullivantii Aust
Trematodon Michx ambiguus (Hedw) Hornsch
Dicranellll (C Muell) Schimp cerviculata (Hedw) Schimp val cillPicuiata
heteromalla (Hedw) Schimp ~ rufescens (With) Schimp
schreberiana (Hedw) Schhnp var schreberiana schreberitJna (Hedw) Schimp var robusta Schimp ex Braithw subulata (Hedw) Schimp varia (Hedw) Schimp
Dicranodontium BSG denudatum (Brid) Britt ex Williams
Panzeucobryum (Lindh) Loeske Iongifolium (Hedw) oelke varlongifolium
17
Tortella (Undb) Limpr fragilis (Drumm) Umpr humilis (Hedw) JeM torlUoSfl (Hedw) Umpr
Hyophila Brid involuta (Hoole) Jaeg et Sauerb
Didymodon Hedw rigidulus Hedw
BryoerythrophyUum Chen recurvirostnon (Hedw) Chen
Btubula Hedw acula (Brid) Brid var acuta convoiuta Hedw fallox Hedw unguicultltll Hedw
Acaulon C MueD muticum (Uedw) C Muell var rufeseens (Iaeg) Crum
PJl~lm Hedw CUSPidtltum Hedw
De8mlltodon Brid cernuu (Hueb) BSG oblUsifoliw (Schwaegr) Schimp
T01IUla Hedw mueronifolia Schwaegr rurall (Hedw) Gaertn Meyer et Scherb var ruralis
39 Grimmiaceae Grlmmill Hedw
alpicola Hedw var alpicola alpicola Hedw var dupretii (Ther) Crum alpicola Hedw var rlvularl8 (Brid) Wahlenb apocarpa Hedw var apocarpa apocarpa Hedw var ambipa (Sull) Jonea apocarpa Hedw var conjerttl (Funck) Spreng apocarpa Hedw var strleta (Tum) Hook et Tayl CiIIyptrrlta Hook ex Drumm donniana Sm var donn_ hemumti Crum ltlevla (Brid) Brid montana BSG oineyi SuD oVllll8 (Hedw) lindb piUfe P-Beauv pltJgiopodill Hedw pulvinata (Hedw) Sm varbullpulvinata teretinervl Limpr
18
Orothera C Muell leana (Sull) C Muell
Rfwbdoweisia crispata (With) Undb
Cynodontium BSG ex Schimp alpestre (Wahlenb) Milde strumiferum (Hedw) Undb
Dichodontium Schimp pellucidum (Hedw) Schimp
Oncophorus (Slid) Brid vuens (Hedw) Brid wahlenbergii (Brid)
Dicranum Hedw bonjeanii DeNot ex Usa drummondii C Muell lagellare Hedw varj1ogellare fulvum Hook fuscescens Turn var fuscescens montanum Hedw muehlenbeckii BSG var muehlenbeckii polysetum Sw sabuletorum Ren et Card scoparium Hedw undulatum Brid viride (Sull et Lesq ex Sull) lindb
36 Leucobryaceae Leucobryum Hampe
glaucum (Hedw) Angstr ex Fr
37 Encalyptaceae Encalypta Hedw
ciliata Hedw var ciliata procera Bruch
38 Pottiaceae Astomum Hampe
muhlenbergianum (Sw) Grout
Weissia Hedw controversa Hedw var controversa
Gymnostomum Nees Homsch et Stunn aeruginosum Sm recurvirostrum Hedw var recurvirostrum
Trichostomum Bruch spirale Prout
19
40 Ephemeraceae Ephemerum Hampe
cohaerens (Hedw) Hampe crassineium (Schwaegr) Hampe val crassinervium spinulosum Bruch et Schimp ex Schimp
41 Funariaceae AphflllolThegma SuD
serratum (1 Hook et Wils ex Drumm) Sullo
Physcomitrium (Brid) Fuernr hookeri Hampe immersum Sullo pyriforme (Hedw) Hampe
Funaria Hedw americana Undb hygrometrica Hedw vaT hygrometrica
42 Splachnaceae Splachnum Hedw
ampullaceum Hedw
43 Schistostegaceae Schistostega Mohr
pennata (Hedw) Web el Mohr
44 Bryaceae Pohlia Hedw
annolina (Hedw) Lindb var annotina camea (Tum) Lindb cruda (Hedw) Undb elongata Hedw iliformis (Dicks) Andr lescurlana (Sull) Grout nutans (Hedw) Lindb obrusifolia (Brid) L Koch proligera (Undb ex Umpr) Undb ex Amell wahlenbergii (Web et Mohr) Andr
Leptobryum (B S G) Wits pyriforme (Hedw) Wils
Bryum Hedw algovicum Sendtnex C Muell argenteum Hedw caespiticium Hedw capillare Hedw var capillare creberrlmum Tayt lonchocaulon C Muell pallens (Brid) Sw ex Roehl pseudotrlquetrum (Hedw) Gaertn Meyer et Scherb varpudotriquetrum
20
pseudotriquetrum (Hedw) GaertnMeyer et Scherb var bimum (Schreb) Ulj stenotrichum C Muell var stenotrichum tortiolium Funck ex Brid uliginosum (Brid) B S G violaceum Crundw et Nyh weigelii Spreng
Rhodobryum (Schimp) Hampe roseum (Hedw) Limpr
45 Mniaceae Mnium Hedw
ciliare (C Muell) Schimp cinclidioides Hueb cuspidarum Hedw drummondii Bruch et Schimp longirostrum Brid lycopodioides Schwaegr marginatum (With) Brid ex P Beauv mediumB~S G var medium onhorrhynchum Brid pseudopunctatum Bruch et Schimp punctalUm Hedw var punctatum punctatum Hedw var elatum Schirnp rugicum Laur spinufosum B S G stellare Hedw
46 Aulacomruaceae Au1acomnium Schwaegr
androgynum (Hedw) Schwaegr heterostichum (Hedw) B S G paJustre (Hedw) Schwaegr var paJustre
47 Meesiaceae Amblyodon B S G
deillbotus (Hedw) B S G
Meesia Hedw triquetra (Richt) Angstr uJiginosa Hedw
48 Bartramiaceae Plagiopus Brid
oederiana (Sw) Umpr
Bartramia Hedw pomiformis Hedw var pomiformis
Philonotis Brid fontana (Hedw) Brid varfontana longseta (Michx) Britt varlongseta marchica (Hedw) Brid
21
~
~
49 Timmiaceae Timmia Hedw
megapolitana Hedw var megapolitana
50 Orthotrichaceae Zygodon Hook et Tayl
viridissimus (Dicks) Brid var rupestris lindb ex C J Hartm
Amphidium Schimp lapponicum (Hedw) Schimp mougeotii (B S G) Schimp
Orthotrichum Hedw anomalum Hedw obtuaifolium Brid ohioense Sullo et Lesq ex Aust pumilumSw sordidum Sullo et Lesq ex Aust speciosum Neesax Sturm var eegam (Schwaegr ex Hook et Grev) W~rnst stellDtwn Brid strrmgulatum P Beauv
Uleta Mohr ex Web cOfIICtata (p Beallv) Hamrn crispa (Hedw) Brid
Drummondill Hook ex Drumm prorepens (Hedw) Brltt
51 Fontinalaceae Fantinalis Hedw
antipyretica Hedw var antipyrettca antipyretica Hedw forma rabusta (Card) Card dalecarlica Schimp ex B S G var dalecarlica flqccidtJ Ren et Card hypnoides C J Hartrn var hypnoides hypnoides C J Hartm var durillei (Schimp) HU$ll
missourlcaCard novae-angliae Sullo var novae-angliae splulgrUlolill (C Moen) Wijket M~S
DichelY1l1ll Myr capilloceum (With) Myr falcatum (Bedw) Myr HIllescens B S G
52 Climaciaceae aimecium Web et Mohr ex Mohr
americllnum Brid dedroides (Hedw) Web et Mohr
22
S3 Hedwigiaceae Hedwigia P Beauv
ciliata (Hedw) P Beauv
54 Cryphaeaceae Forsstroemia lindh
ohioense (Sull) lindh
55 Leucodontaceae Leucodon Schwaegr
brachypus Blid var brachypus brachypus Brid var aniJrewsitlnus Crum et Anderson julaceus (Hedw) SuU
56 Neckeraceae Neckera Hedw
pennata Hedw var pennata
1fomalia (Brid) B S G trichomanoides (Hedw) B S G
57 Theliaceae Thelia Sull
asprellll Sullo hirtella (Hedw) Sullo lescurii SuU
Myurella B SG julacea (Schwaegr) B S G
sibrica (C MueU) Reim var sibrica
58 Fabroniaceae Fabronio Raddi
ciliaris (Brid) Bnd
Anacamptodon Brid splachnoides (Groet ex Brid) Brid
Schwetschkeopsis Broth fabronia (Schwaegr) Broth
59 Leskeaceae Lindbergia Kindb
brachyptera (Mitt) Kindb
Leskea Hedw gracilescens Hedw obscura Hedw poiycarpa Hedw
Leskeella (limpr) Loeske nervosa (Brid) Loeske
23
Pterlgy1lllndrum Hedw filiforme Hedw
60 Thuidiaceae Haplohymenfum Dozy et MoIk
trlste (Ces ex De Not) Kiodb
Anomodon Hook et Tayl attenuatus (Hedw) Hueb minor (Hedw) Fuemr rostratus (Hedw) Schimp viticulosus (Hedw) Hook et Tayl
Haplocladium (C Mucn) C MueU microphyllum (Hedw) Broth virginillnum (Brld) Broth ~bull
Thuidium B S G abietinum (Hedw) B S G delicatulum (Hedw) B S G vardelicatulum delicatulum (Hedw) B S G var radicans (Kindb) Crum Steere et Anderson minutulum (Hedw) B S G pygmaeum B S G recognftum (Hedw) Lindb scitum (p Beauv) Aust
Helodium (Sull) Warnst blandowli (Web et Mohr) Warnst var blandowli blandowli (Web et Mohr) Wamst var helodioides (Ren et Card ex Roell) Crum
Steere et Anderson paludomm (Sull) AuS
61 Amblystegiaceae Cratoneuron (Sull) Spruce
commulatum (Hedw) Roth var commutfltum commutatum (Hedw) Roth var fakatum (Brid) Moenk ilicinum (Hedw) Spruce var filcinum
Campylium (Sun) Mitt chrysophyllum (Brid) 1 Lange var chrysophyllum hispidulum (Brid) Mitt var hllJidulum polygamum (B S G)C lens varpolygtlmUm stellatum (Hedw) C1ens vat stelltum
Leptodictyum (Schimp) Warnst laxirete (Card et Ther) Broth riparium (Hedw) Wamat trichopodium (Schultz) Warns l1acillans (Sull) Broth
Hygroamblystegium Loelke jlul1iatile (Hedw) oeske var luViatile
24
fluviatile (Hedw) Loeske var oWltum Grout noterophilum (Sun et Lesq ex Sull) Warnst tenox (Hedw) Jenn var tenax tenox (Hedw) Jenn var spinilolium Schimp) Jenn
Amblystegium BSG juratzkanum Schimp serpens (Hedw) BSG varlum (Hedw) Undb var varium
Platydictya Berk confervoides (Brid) Crum jungermanniodes (Brid) Crum minutissimum (Sull et Lesq ex Sull) Crum subtile (Hedw) Crum
Drepanocladus (C Muell) Roth aduncus (Hedw) Warnst var aduncus aduncus (Hedw) Warnst var capillilolius (Wamst)Riebm aduncus (Hedw) Warnst var kneiffii (BSG) Moenk aduncus (Hedw) Warnst var polycorpus (Bland ex Voit) Roth exannulatus (BSG) Warnst fluitans (Hedw) Warnst var fluitans revolvens (Sw) Wamst var revolvens sendtneri (Schimp) Warnst var sendtnerl uncinatus (Hedw) Warnst var uncinatus uncinatus ffiedw) Warnst var piwnosus (Schimp) Wamst vernicoms (Lindb ex C Hartm) Warnst
Hygrohypnum Lindb eugyrium (BSG) loeske luridum (Hedw Jenn varluridum ochraecum (Tum ex Wils) Loeske var ochraceunt
Calliergon (Sull) Kindb cordilolium Hedw Kindb glganteum (Schimp) Kindb var giganteum richardsonii (Mitt) Kindb ex Wamst stramineum (Brid) Kindb trilarium (Web et Mohr) Kindb
Calliergonella Loeske cuspidata (Hedw) Loeske
Calliergidium (Ren) Grout bakeri (Ren) Grout
Scorpidium (Schimp) Urnpr scorpiodides (Hedw) limpr
62 Brachytbeciaceae Tomenthypnum loeske
nitens (Hedw) Loeske var mtens
25
Homalotheciella (Card) Broth subcapillata (Hedw) Broth
Brachythecium BSG acuminatum (Hedw) Awt acutum (Mitt) Sullo calcareum IGndb campestTe (C MuelL) BSG cunum (Lindb) Umpr digastrum C Muell et Kindb ex Macoun et Kindb oxycladon (Brid) Jaeg et Sauerb var oxycladon plumosum (Hedw) BSG populeum (Hedw) BSG var populeum reflexum (Starke ex Web et Mohr) DsG var relexum reiexum (Starke ex Web et Mohr) DSG var pacificum Renet Card ex Roell rlvulare BSG var rivulare tabulum (Hedw) BSC var tabuum salebrosum (Web et Mohr) BSG var salebrosum velutinum (Hedw) BSG Vat velutinum
Bryhniil Kaur graminicolor (Brid) Grout var graminicolor novae-angliae (Sull et Lesq ex sun) Grout
Rhynchostegium BSG senulatum (Hedw) Ja4lg et Sauerb
Rhynchostegiella (BSG) Limpr compacta (C MueU) Loeske
EU1hynchium 8SG hians (Hedw) Sande Lac pulchellum (Hedw) Jenn var pulchelium riparloides (Hedw) Rich
Stokeriella (Kindb) Robins pmelonga (Hedw) Robins var stokesii (Tum) Crum
63 Entodontaceae Entodon C Muell
brevisetus (Hook et Wils ex Wils) Lindb cladorrhizans (Hedw) C Muell seductrlx (Hedw) C MueU var wluctrix
Pleurozium Mitt schreberl (8rid) Mitt var schreberi
64 Plagiotheciaceae Pltlglothecium BSG
CflVifolium (Brid) Iwats var CtWifolium denticulatum (Hedw) 8SG laetum BSG ltztebricola BSG
26
6S SernatophyUaceae Brotherella Loeske ex Fleisch
recumins (Michx) Fleisch tenuirostris (Bruch et Schimp ex Sull) Fleisch
Sematophyllum Mitt demissum (Wils) Mitt
66 Hypnaceae Pl(itygyrium BSG
repens (Brid) BSG
PylaisieUa Kindb intricata (Hedw) Grout polyantha (Hedw) Grout selwynii (Kindb) Crum Steere et Anderson
Homomallium (Schimp) Loeske adnatum (Hedw) Broth
Hypnum Hedw cupressiforme Hedw var cupressiforme cupressifonne Hedw varfiliforme Brid curviolium Hedw fertile Sendtn imponens Hedw lindbergii Mitt palescens (Hedw) P-Beauv pratense Koch ex Brid
Calicladium Crum haldanianum (Grev) Crum
sopterygiopsis Iwats muelleriana (Schimp) Iwats
Isopterygium Mitt e1egans (Brid) lindb pulchellum (Hedw) Jaeg et Sauerb tenerum (Sw) Mitt
Taxiphyllum Fleisch depllQtum (Bruch et Schimp ex sun) Fleisch taxirameum (MitL) Fleisch
Herzogiella Broth seligeri (Brid) Iwats striatella (Brid) Iwats ruracea (Lindh) Iwats
hilium De Not crista-castrenna (Hedw) De Not
27
67 Rhytidiaceae Rhytidium (Sull) Kindb
rugosum (Hedw) Kindb
Rhytidfodelphus (Lindb ex Iimpr) Warnst triquetrus (Hedw) Warnst var triquetrus
68 Hylocomiaceae Hylocomium BSG
pyrenaicum (Spruce) lindb splendens (Hedw) BSG varsplendens
69 Buxbaumiaceae Buxbaumia Hedw
aphyllll Hedw
Diphyscium Mohr follosum (Hedw) Mohr
70 Tetraphidaceae Tetraphi3
pellucida Hedw
71 Polytrichaceae Atrichum PmiddotBeauv
angustatum (Brid) BSG cri8pum (James) Sullo oerstedianum (C Muell) Mitt teneUum (Roehl) BSG undulatum (Hedw) P middotBeauv var unduiatum
Pogonatum P-Beauv alpinum (Hedw) Roehl var alpinum umigerum (Hedw) P-Beauv
Poiytrichum Hedw commune Hedw var commune ormosum Hedw juniperinum Hedw longisetum Brid ohioense Ren et Card piliferum Hedw var pilierum strictum Brid
28
MAPS OF BRYOPHYTE TAXA
REPORTED FOR WISCONSIN
HORNWORTS AND LIVERWORTS
29
bull bull WSHeV~N S vE II
0 ~
e~RRON ~USK
(J~ (j -
CHIPPE A DUNN
0 o0 poundAU CLAIRE
~ TREMshyPEAshyLEAU
MICE
ONEIDA
Q LINCOLN
TAYLOR
Q t-~----- MOIRA THON
CLARK
o
10
LA FAVETTE
Hornwor ts amp Liverworts
- None - 1 - 25 taxa
- 26 - 50 tRXshy
- 51 - 75 tax)
- 7c - 100 taxa
FONDOU j LAC ~
DODGE
i O JEfFERSON
WALWORTH
30
Anastrophylltlfl helleranum (Nees) Schust Anastrophyllum michauxil (We~ Buch ~
46 88 136 141 Evans 38 46 141
Anastrophyllum mlnut~ (Schreb) Schuster Aneura plnguls (L) Durn
89 21 384687 88 89 90 (138) (145)
Anthoceros macounil MA Howe ~oceros punctatus L
21 (45) (144) 34
31
Barbilophozia attenuata (Mart) Loeske Barbilophozia barbata (Schmid ~ Schre~
24 141 Loeske 28 39 46 89 90 141
Bazzania trilobata (1) S Gray
1 26 39 46 (57) (59) 79 88 89
90 119 141
BlePbarostoma trichoffiyllum (1) Dumort
1 8 26 39 46 89 90 137 141
Blasia pusilla 1
1 39 46 90
Calypogeja fissa (1) Raddi
90
32
$129geja integristipula Steph C~Ypound2geja muelleriana (Schifm) K Mull
88 89 89
CallpoundO~eja neesiana (Mass amp Carest) K Mull
39 46 89 140 (141)
~pogeja sJlapound2icola (H Arnell tl J bull
Perss) Warnst et Loeske (119) 141
CeIalozia biscuspidata (L) Dum
1 37 I 46 90 t 137 141
CalYpogej~ trichomanis (L) Corda
26 39 (119) 137
33
CeEhalozia catenulata (Hub) Lindb Cephalozia connivens (Dicks) Lindb
39 46 79 141 39 4679 88 (89) 90 119 141
Cephalozia lacinulata Jack ~ Spruce
141 Cephalozia lunulifolia (Dum~ Dum I 28 39 46 79 88 89 90 141
Cepoundhalozia macouni (Aust) Aust Cephalozia ~leniceps (Aust) Lindb 27~ 3746 (66)7987 (139)141 39 46 79 89 141
34
Ceffialoziella 9ivaricata (Sm) Schiffn Cephaloziella ~hista (Jack) Schiffn
46 90 46 (119) 141
Cephaloziella hampeana (Nees) Schiffn
46
Cephaloziella rubella (Nees) Warnst
46 39
Cephaloziella rubella var sullivantii
(Aust) K Mull 46 39
Cephaloziella rubella var bifida (Lindb) Douin
35
46
Chiloscyphus pallescens (Ehrh ~ Hof~ Dum Chiloscyphus pallescens var fragilis
39J 46 88 89J 90 (Roth) K Milll 46
Chiloscyphus E9lyanthos (L) Corda
1 24 39 46
Chiloscyphus polyanthos vax rivularis
(Schrad) Nees 39 46
poundpoundnocepha1um conicum (1) Lindb
12 28 39 46 (49) 87 88 8990 120 121 148
Cololejeunea biddlecomiae (Aust) Evans
39 46 62 87 88 89 90
36
Dipl0]hyllum apicu1atum (Evans) Steph Fossombronia foveo1ata Lindb
(35)41 46 90 140 1 t 38 (39) t 46
Frullania bolanderi Aust
13 46 62 88 90 139 140
Frullanla ari ttoniae Evans
39 46 69 (71) 87 88 89 90
Frullania eboracensis Gott
1 9 28 4 39 46 (57) (59) 69 70 (71) 87 88 89 90 92 93 97 148
Frullania inflata Gott
46 87 88 89 (139)
Frullania oakesiana Aust Frullania plana Sullo
(35) 42 46 87 (139) (140) 92
Frullania ri paria Hampe ~ Lehm
46 69 71 87 89
Frullania ~elwynianna Pears
46 89
Geocalyx SEaveolens (Schrad) Nees
394688 8990 (119) 140
Frullania tamarisci (L) Dum subsp sPagrayana
(Mont) Hatt 28 34 l~6 t (57) 88 89 90
38
Mar pan thus seutatUB ( Web amp Mohr) Spruce Jamesoniella autumnalis (DC) Steph
(35) 39 46 90 1 7 28 39 46 56 (57) 87 8889 90 (119) 137 141
Jungermannia confertissima Nees
137 141
Jungermannia crenuliform1s Aust
79 137 141
Jungermannia ~il1ima Sm
46 90 141 Jungermannia exsert1folia subsp corifo~ (Dum~
Vana (35) 46 79 141
39
Jungermannia hyalina Lyell Jungerrr6nnia 1eiantha Grolle
40 46 79 136 141 39 46 79 88 90 1)7 141
Jungermannia IXlaxis Lindb Jungermannia pumila With
46 67 79 (139) (140) 141 )9 46 90 1)6 141
Jungermannia sphaerocarpa Hook Kurzia setacea (Web) Grolle
4679136140IIH 136
Lejeunea cavifo1ia (Ehrh) Lindb Lepidozia reptans (1) Dum
1 7 27 39 46 88 89 90 137 14010 I ( 27) 46 I 69 (71) 88
lophocolea heterophylla (Schrad) Dum Lophocolea minor Nees
1 )4 39 46 (57) 87 88 89 27 39 46 88 89 90 (119) 90 (93)94 119 136
Lophozia al~stris (Schleich ~ Web) Evans Lophozia ascendens (Warnst) Schust
46 90 137 141 136 140 141
41
loIilozia badensls (Gatt ~ Rabenh)
46 141
lophozia blcrenata (Schmid sectA Hoffm)
Dum 46 141
lophozia collaris (Nees) Dum
39 46 141
lophozla excisa (Dicks) Dum
27 4679 141
lophozia gillmani (Aust) Schust lophozia gIlttulata (Undo tl H Arnell)
36 39 40 46 136 (140) 141 Evans 39 46 136 141
42
141
Lophozia heterocolEDs (Thed) MA Howe IoPhozia ipcisa (Schard) Dum
27 39 46 89 141 8 27 31 46 79 90 140 141
LoQhozia longidens Macoun Lophozia ventricosa
1 46 79 90 141
Lophozia ventricosa var silvicola
(Buch) Jones 141 LoEhozla ventricosa vax longif1ora (Nees)
Macoun 39 46 79 141
43
~~ fragrans (Balbis) Frye et Clark Mannia pi 10sa (Hornem) Frye et Clark
46 (139) (140) 9 46 79 116 139
Mannia triandra (Scop) Gro 11e
46 (140)
Marchantia pound91ymorpha L
1 13 28 3946 (49) 78 87 88 89 90 (Ill) 121 122 147
Mo~rckia hibernica (HookJ Gott
46 89
Metzgeria conjugata Lindb
44
46
Mylia anomala (Hook) S Gray Notothylas orbicularis (Schwein) Sullo
39 46 141 (35)45 46
Nowellia curvifolia (Dicks) Mitt
17 27 3846 88 89 90 141
Odontoschisma denudatuJI1 (Nees ~ Mart)
Dum 46 81 137 141
Pallavicinia lyellii (Hook~ Carruth
90
Odontoschisma macounii (Aust) Under
45 46 (66) (139) 140 141
45
=
Pellia epiphllla (L) CordaPellia endiviifolia (Dicks~ Dum
27 39 46 90 13724 39 46
Pellia neesiana Limp bull Phaeoceros
39 46 89 27 39 45 46 97
Porella pinnata L
13 27 I 46 90 Plagiochila asplenioides (L) Dum
27 I 39 1+6 88 89 I 90 137
46
~e11a platYEhylla (L) Pfeiff Porella platyphylloi~ (Schwein) Lindb
17 27 39 46 69 (71)79 87 88 89 9 34 46 (57) 59 87 88 89 92 90 92 93 77
Preissia quadrata (Seop) Nees
1 36 3946 (71) 88 89 90 121 123
137
Ptllidium ciliare (L) Hampe
1 27 39 46 132 141
Dum
(22) 28 3946 (57) 87 88 89 90 92
Ptilidium ~lcherrium (G Web) Hampe
(9) 13 394656 (57)70 87 88 89 90 93 (119) 136 141
47
Radula obconica Sullo Reboulia hemlsphaerlca (L) Raddi
39 46 88 89 (43) 46 (59) 70 90 t Cf 124 125 (139)
Riccardia ~atifrons Lindb
1 27 39 46 89
Riccardia multifi~ (L) S Gray
39 46 88 145
Riccia beyrichiana Hampe ~ Lehm
46 Riccardia palmata (Hedw) Carruth
39 46 90
48
Riccia bifurca Hoffm Riccia cavernosa Hoffm
48 13739 46 79 137
Riccia fluitans L
242731454679
Riccla frostli Aust
48 79
Riccia sullicantii Aust
79 (137) Riccia sorocarpa Bisch 46 (137)
49
Ricciocarpus natans (1) Corda Sca~ia apiculata Spruce
13 27 46 79 89 110 39 46 (139) 141
ScaJltll1ia carinthica Jack ~ Lindb
137 t 141
Scapania ~ (Mart) Durn
39467990137141
Scapania digenil Schiffn
90
ScaEsectflia cuspiduligera (Nees) K Mull
46 (139) (140) (141)
50
Scapania glaucoeerha1a (Tayl) Aust Seapania gyrrmostomophilia Kaal
39 46 (139) 141 41 44 46 64 (139) (140) 141
ScaP3poundlia irrigua Gott ~ almiddot
7 39 46 79 137 (140L 141
Seapania nemorosa (L) Dum
1 39 46 89 90 137
Scapmi~ ~~ta Smiddotlch
137 141 51
~caPnia poundludicola LDeske amp K Mull Scapania saxicola Schust
46 (66) 141 136 141
Sc~~ scandica (H Arnell ~ Buch) Macv
139 140
Scapania subalpina (Nees) Durn
39 40 46 79 137 141
Scapania undulata (L) Dum
1 42 46 79 141
Scapania umbrosa (Schrad) Dum
137 140 141
52
Trichocolea tomentella (Ehrh) Dum Triomarla exsecta (Schmid) Schiffn
27 (35) 39 46 47 88 137 141 8 28 39 46 90 140 141
Tritomaria exsectiformis (Breidl) Loe~ke Tritomaria Sluinguedentata (Huds) Buch
39 -t6 89 90 136 140 141 7 39 46 141
Trltomaria scitula (Tayl) Joerg
46 136 (139) 141
53
ABSTRACf
117This atlas of Wisconsin bryophytes shows the distribution of)30 taxa (137 hepatics 389 mosses)
on state maps with a dot in each county for which each taxon has been reported in the literature A list of synonyms and excluded taxa is included along with an extensive bibliography Pertinent features of the natural history and history of bryology in the state are discussed
ACKNOWLEDGEMENTS
We want to thank Drs Fred Hermann and John Thomson of the University of Wisconsin-Madison herbarium and Botany department for locating and lending packets and for references to aid us in accumushylating a more accurate complete report The illustration of Climacium dendroides on the cover was drawn by Robert Freckmann Of course we are grateful to all the people who have collected specimens of Wisshyconsin bryophytes and preserved them in herbariums making this distributional study possible
4
ONEIDA
LINCOLN
TAYLOR
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t
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INGTb
lt
WALWORTH
5
INTRODUCfION
This atlas of the bryophyte flora of Wisconsin is an attempt to show the distribution of all bryoshyphyte species known to occur in the state by county occurrenoes based on published reports in the literashyture It is not an annot~ted check list (only a feW problematic taxa were examined) but is a literature search intended as a foundation for a complete flora or for otherbryological research
The literature of btYo1ogy is large and spread over numerous journals reports monographs and manuals It would be almost impossible to search all sources for every reference attributing bryophyte species to Wisconsin and to particular localities in the state Wehav checked many inlportant references and paid special attention to works dealing with Wisconsiit botany and the bryologyof the upper Midwest Several bibliographies and indices were checked thoroughly including Green an~ Curtis (1955) the Bry ologist indices (Crum 1959 and Miller 1976) andt)le Wisconsin Academy of Sciences Arts and Letters indices
Only those published repots which app~red to)e substanti2ted by herbarium specimens and which gave- ~fficient location dat to P~CetheW(On in amiddot particula(county were-used in the initial compilation Species described in manuals in generalized way implying their occurrence in Wisconsin (Without menmiddot tioning Wisconsin specifically or citing a particular location in the state) were excluded eg Gnmmia raui (Wynne 1943) and Pohlia annotinll var decipiens (Crum 1976) Ecological papers which were either too general in taxonomic determinations (eg Sphagnum sp Polytrichum sp) or in location data were also excluded Some other taxa attributed to Wisconsin which have been excluded are
Cylindrothectum compr~ (Cheney 189tmiddotDane Co) identified later by Grout as Entodon seducshytrtx
Dicranell4 humilis (Holzjne~ 1925middotBuffalo Co) probablyD vllJia since Dhumilis is not found in North Arnerl~
Didymodon triarium (Foote 1 966-SW Visconsin) identified by Fred Hennann as Barbulaacuta Funoria microstoma (anonymous 1975middotDane Co) excluded because it is outside of its geographic
range and the packe~ QaMot be located Phil(motis calcarea (Cheney and Evans 1944-Dane Co) excluded in Crum Steere and Anderson
(1973) Plagiobryum demissum and P zierl (Cheney and Evans 1944-various counties) are both Arcticshy
alpine species not likely to be found in isconsin Two of Cheneys collections of Pl1giobyrum have been referred to Bryum
Pogonatum alpinum var septentrionale (Che~ey and Evans 1944-Chippewa Co) was misidentified (Ireland 1960) and is P umigerum
Rhizomnium fAPpoJachIanum and R punctalu1fl ssp chlorophyDosum (Koponen 1973middotseveral counshy ties) excluded North American bryologiflts have not recognized these taxa in America
Sphagnum tenellum (Cheney and Evans 1944-Waukesha Co)an Atlantic species not occurring in Wisconsin (Cruro 1975)
The nomenclature and systematic arrangement of hornwort 00 hepatic species in the atlas follow Stotler and Crandall-5totler (1977) and of moss species fonow Crum Steere and Anderson (1973) except in such instances as the JreptlIfockIdus fonns whiGh follow Wijk et aL (19S9middot1973) Species reported in the literature under names which are now listed as synonyms on either of these lists were converted to the accepted name A dot is placed in each county of Wisconsin from which the taxon has been reported The numbers after the name of the taxon correspond to the numbered references in the bibliography which giye ~act location data OEat least location data -s~fficient to place it in-particular counties in the state The numbers in parentheses indicate references which attribute the taxon to Wisconsin without providing exact loltation data
Fossil and subfossil reports are summarized Slfparately in our atlas only six species (Aulacomnium turgidum lJ1achythecium turgidum Ditrichum Jlelficaule Hypnum bambergti Scorpidium 6fJisities and Tortel14 1rIgi1i$) are known exclUsively from the fossil record ~
6
GEOGRAPHIC CLIMATIC AND BOTANICAL FEATURES OF WISCONSIN
Wisconsin is a north central state bounded on the east by Lake Michigan on the north by Michigan and Lake Superior on the south by Illinois and on the west by the Mississippi and St Croix rivers It is located midway between the Prime Meridian (0deg) of Longitude and 1800 Longitude and midway between the equator and the north pole thus placing it in the center of the northern half of the western hemisphere
The topography of the state is relatively flat a few hills exceed 1900 feet with Thns Hill at 1952 feet forming the highest point in Price County Lake Michigan represents the lowest point at 580 feet The northern and eastern two-thirds of the state were heavily glaciated during the last stage of the Pleistocene and now contain many of the best examples of continental glaciation topOgraphy including considerable ground moraine large terminal recessional and intetlobate moraines kames eskers drumlins and small depressions or kettle-holes Most of the nearly 9000 lakes of 20 acres or more lie within this part of the state The southwestern third of the state is the Driftless Area It includes most of the large flat generally sandy bed of Glacial Lake Wisconsin plus the highly dissected ridge and valley section with numerous cliffs and rock-outcroppings (Figure 1)
Much of the bedrock in the state lies below glacial till but exposures include igneous and metamiddot morphic rocks primarily granitic especially in the north as well as sedimentary layers primarily sandshystones and dolomitic limestones especially along Lake Michigan and in the DriftlesS Area
The climate of Wisconsin is classified as humid-temperate with cold winters The annual precipimiddot tation averages approximately 30 inches which is sufficiently even in its seasonal distribution to support forest vegetation throughout most of the state Nevertheless according to Curtis (I 959) the pre-settlement vegetation of the state (J 830-1850) included about 27 prairie and oak-savanna plus about 64 forests
The state is presently divided into seventy-two counties Marathon County is the largest (1586 sq mi) the smallest counties include Pepin (235 sq mi) Ozaukee (236 sq mi) andMilw8ukee (237 sq mi) We included Menominee County though its future status as a county is in doubt since it regained its Indian reservation status and rights in 1973 This county was established April 30 1961 from the Menomishynee Indian reservation which consisted of parts of Shawano and Oconto counties Wisconsin ranks 26th in the United States in area with 54464 square miles and 16th in population with 4651000
More ecological research is needed on the bryophyte flora of Wisconsin especially to relate it to the vegetation and to such other environmental features as soils climate and glaCial deposits Plant communishyties in Wisconsin are many and diverse these are summarized on Figure 2 from studies conducted by Curtis (1959) The state is divided into northern and southern floristic provinces by a tension zone deshyscribed by Curtis (1959) This tension zone is an eeotonal region vatying from 10middot31 mt wide where the ranges of many vascular plants overlap or terminatej it runs in an S-curve northeast from Milwaukee to Hudson in St Croix County (Figure 2) The principal plant communities of the northern province apshyproaching their southern boundaries within the zone include the northern conifermiddothardwood forests of sugar maple white pine hemlock and yellow birch and the Sphagnum bogs with tamarack and black spruce Southern communities at or near their northern limits within this zone include prairies oak and jack pine savannas and hardwood forests of maples oaks ash ehn and cottonwood
We hope to encourage research on cryptogamic ecology with our checklist We know that some species are northern while others are southern (eg cimacium) fits the tension zone concept Freckmann (I977) while Bryoxiphium is only found in the Driftless Area of the state Steere (1937) and Love and Love (1953) The tension zone theory may prove to be acceptable for bryophytes We speculate that certain bryophytes can indicate particular plant communities since many bryophytes are restricted to preshycise environmental requirements Many papers or theses have dealt with cryptogamic bark and rock ecology in Wisconsin (Culberson 1955 Foote 1962 Hale 1952 1955 and Cole 1975)
We hope that this atlas will stimulate collecting in counties poorly sampled so that our knowledge of the bryophytes can be increased It should aid anyone interested in floras for preservation environmental impact statements and distributional studies
7
A numerical summary of the bryophyte flora of the state based on our literature search is presented in Table 1 (below) This includes 517 taxa 70 families and 176 genera This numerical summary is comshypared with the published floras of Michigan mosses (Crum 1976) and Minnesota hepatics (Schuster 1953) Thus we surmise that further collecting could lead to the same approximate numbers for Wisconsin bryoshyphytes
Table 1 Comparison of Recorded Bryophytes of Wisconsin with Michigan and Minnesota and Summary of Wisconsin Taxa
Wisconsin Michigan (Crum 1976)
Minnesota (Schuster 1955)
Hepaticsamp Hornworts
Taxa Families Genera
137 28 47
174 32 52
Mosses
Taxa Families Genera
380 42
129
503 42
149
8
-
~
~ tIt
Figure 1 The Driftless Area and Tension Zone The counties are 01$0 shown on map
ca location of tension zone as deecfibed bv Curtis (1959)~ The Orittless Area
9
Agn 2 Major Plant Communitiee of Wsconsln aaed on surwvor recorda bull co 18laquo) After Curtismiddot 1959
tit beol Forest e Phillie
Northernmiddot hardwood forest reg Pine 5cMInna
copy Southernmiddot hordwood forest CD Oak Savanna
10
SYSTEMATIC ARRANGEMENT OF WISCONSIN BRYOPHYTES
ANTHOCEROTAE (Hornworts)
1 Anthocerotaceae Anthoceros 1 emnd Prosk
macounii M A Howe punctatum 1
Phaeoceros Prosk laevis (1) Prosk subsp laevis
Notothylas Sul1 orbicularis (Schwein) Sullo
HEPATICAE (liverworts)
2 PseudolepicoJeaceae Blephllrostoma (Dum emend Undb) Dum
trlchophyllum (L) Dum subsp trlchophylluni
3 Ptilidiaceae Ptilidium Nees
ci1iare (L) Hampe pulcherrimum (G Web) Hampe
4 Trichocoleaceae Trichocolea Dum nom cons
tomentella (Ehrh) Dum
5 Lepidoziaceae Kunia von Martens
setacea (Web) GroUe
Lepidozia (Dum) Dum reptans (1) Dum
Bazzania S Gray trilobata (1) S Gray var triZobata
6 Calypogejaceae 011ypogeja Raddi
flSsa (L) Raddi subsp issa integristipula Steph muelleriana (Schiffn) K Mull subsp muelleriana neesiana (Mass et Carest) K Mull sphagnicola (H Arnell et J Perss) Warnst et loeske trichomanis (1) Corda
11
7 Cephaloziaceae Cepluzlozla (Dum emend Schiffn) Dum
biscwpidata (L) Dum subsp biclupidIIta catenulllttl (Hub) Lindb coMivens (Dicks) Lindb var connivens lacinultzta Jack ampX Spruce lunulifolilz (Dum) Dum macounii (Aust) Aust pleniceps (Aust) Lindh var pleniceps
Noweilia Mitt cunifolilz (Dicks) Mitt
8 Adelanthaceae Odontoschimuz (Dwn) Dum
denudatum (Nees ampX Mart) Dum Vir denudatum mtlCount (Aust) Underw
c
9 Cephaloziellaceae CephtlJoziellll (Spruce) Schiffn nom C()II$
divarlctzt4 (Sm) sclurfn val divarlcat(l elachhta (bdc) Schiffn 1uImpetuw (Nees) Schirfn IUbellll (Nees) Warnst vat IUbeIl4rube (Nees) Warnst var biid4 (Lindb) Dquinrube (Nees) WarDSt Var sulliwmtii (Aust) K MuD
10 Geo~c~ larptuttllus Nees
scutatus (Web et Mohr) Spruce
Geocalyx Nees grGVeolens (Schrad) Nees
11 Lophocoleaceae Lophocolea (Dum) Dum
heteropllyll4 (Schrad) Own minorNees
Chiloscypbus Corda om cons pallescens (Ehrh ampX Hoffm) Dum varfragllis (Roth) K Mun polyantitos (L) Corda var poiyanthos pol)llaquo1lthos (L) Corda var rlvulllris (Schrld) Nees
12 PlagiochiJaceae Plagio(hila Dum nom cons
tlBpienioides (L) Dum var asplenioidu
13 Jungermanniaceae Btriilophoril Loeske
attenutlta (Mart) Loeske bIubtlta (Schmid ampX Scltreb) Loeske
12
Lophom (Dum) Dum alpestris (Schleich ~ Web) Evans ascendens (Warnst) Schust badenais (Gott ex Rabenh) Schiffn var bademii bierenata (Schmid ex H)ffm) Dum eollam (Nees) Dum exci8a (Dicks) Dum var excisa gllimanii (Aust) Schust varglllmanii guttulata (Lindb et H Arnell) Evans heterocolpos (Thed) M A Howe var heterocolpos ineistl (Schrad) Dum var tnelsa longidens (Lindb) Macoun subsplongidens ventricosa (Dicks) Dum var ventricosa ventricolfl (Dicks) Dum varlontifIotrl (Nees) Macoun ventricosa (Dicks) Dum var silvlcola (Bueh) Jones
Tritomarill Schiffn ex Loeske exseeta (Schrad) Loeske exsecttformi$ (Breidl) Loeske subsp exsectiormfs quinquedentata (Huds) Buch var quinquedentata scitula (rayl) Joerg
Anastrophyllum (Spruce) Schiffn hellemnum (Noes) Schust michtzuxii (Web) Buch ex Evans minutum (Schrob) Schust var minutum
Mylill S Gray nom cons anomala (Hook) S Gray
Jamesoniella (Spruce) Carring autumnalis (DC) Steph var autumnalis
Jungemumnia L emend Dum eonfertissimtt Nees crenuliformis Aust exsertijolill Steph subsp cordifolia (Dum) Vana graciUima Sm hyalina Lyell leillnthll Grolle polaris Lindb pumila With sphaerocarpa Hook
14 Scapaniaceae Diplophyllum (Dum) Dum nom cons
apiculatum (Evans) Steph var apiculatum
Scapanill (Dum) Dum nom cons apkuata Spruce carinthillca Jack ex Lindb curta (Mart) Dum var curta
13
cuspiduligera (Nees) K Mull var cuspiduiigera degenii Schlffn ex K Mull var degenii glllucocephaJa (Tayt) Aust gymnostomophila Kaal irrigwJ (Nees) Gott et al subsp irrigua lingulata Buch varlingulata mucronata Buch subsp mucronata nemorosa (l) Dum paludicola Toeske et K Mun var paludicola paludosa (K Mull) K Mull saxicola Schust scandiea (H Arnell et Buch) Macv var scandiea 8Ubalp~ (Nees) Dum umbroSll (SchIad) Dum undulata (L) Dum var undulata
15 Radulaceae Raduia Dum nom COllS
compianata (L) Dum obconica Sull
16 Porellaceae Parella L
pinTlllta L patyphylla (L) Pfeiff platyphylloidt(l (Schwein) lindb
17 Jubulaceae FruUanill Raddi
bolanderi Aust brittoniae Evans eboracemis Gott inlllta Gott oakesiana Aust plana Sull rlparia Hampe ex Lehm selwyniana Pears taTnl1iJci (L) Dum subsp asagmyana (Mont) Hatt
18 lejeuneaceae Calolejeunea (Spruce) Schlffn
biddlecomiae (Aust) Evans
Lejeunea Ub nom cons cavifolia (Ehrh) Lindb emend Buch
19 Codoniaceae Fossombronia Raddi
foveolata Lindb
20 Pa11aviciniaceae Paklvieinlil S Gray nom cons
lyeiii (Hook) Carruth
14
Moerckia Gott hibemica (Hook) Gott
21 PeWaceae Pellfa Raddi nom cons propos
endivlifolfa (Dicks) Dwn epiphylllz (L) Corda neeBiana (GottlUmpr
22 Metzgeriaceae Metzgerla Raddi
conjugata Undb
23 Blasiacae B L
pulilm L
24 Aneurareae AneumDum
pinguis (L) Dum
Riccfl1dia S Gray nom cons latifrons Undb multiftda (L) S Gray var multifida palmata (Hedw) Carruth
25 Aytoniaceae Reboulill Raddi nom conI
hemllphaerica (L) Radeli
ManniaOpiz fragrrms (Balbis) Frye el ClarK pilosa (Hornem) Frye et Clark trltmdra (Scop) GroDe
26 Conocephalaceae Conocephalum Wigg nom cons
conicum (L) Lindb
27 Marchantiaceae Preiuia Corda
quadrata (Scop) Nees
Marchantia L polymorpha L
28 Ricciaceae Riccwcarpus Corda
natans (L) Corda
Riccia 1 beyrichiana Hampe ex Lehm bifurca Horfm
15
i
CtlJPemostI Hoffnt fuitu L rostii Aust BOIOCtI1fHI Bi~ sulltvtmtii Aust
MUSCI (Mosses)
29 Sphagnaceae SpIutgnum L
capillaceum (Weiss) Schrank var Cllpil1tlceum caplllllceum (Weiss) Vir tenellum (Schimp) Andr centrille C Jens ex Amen et C Jens compactummiddot OC ex Lam et Dc ClUpidatum Ehrh ex Hoffin var aupidatum jimbrltltum Wits ex J Hookmiddot fulCUm (Scbimp) KUngrr glrgenBOhIlll Russ imbrlctltum Hornsch ex Russ muelllmitil4m BOd 1IIIIU (RUII) C Jens przIutre L papIIIoIum Lindb pulchrum (Lindb ex braithw) Wamst recumun P-Beauv var recumun recum P-Beauv var tenue Klinggr riptlrlum Ansstr ruPQWItWamst squtl1fOlUm Crome subftltens Russ et Wamst ex Wamst IUbIeCfUldum Nees ex Sturm var subsecundum subsecundum Nees ex Sturm varrutem (Nees Hormch et sUUm) Hueb teres (Sdlimp) Anptr e C Hartm
WtlI7IItorjii Russ wuljfanum Gira
30 Andreaeaceae AndNrrerI Hedw
rupeitrlr Oedw var pest
31 Fissident~e Fluid_ lledw
trdiantoid Hedw byoidei Bedw var bryolda cristaM Wils ex Mitt exiguus sun VIr exigUw onttmUi (B-Pyl) Steud jtuberi Lesq et James obtusifolhu Wits var obtwifoliul ollPlUlldQldes Heel ~litSuU lubbtulkril Bed tfDdfoliul lledw llridubu (S~ ) Wahlenb vallirlIU1u
16
32 Ditrichaceae Pleuridium Brid
subulatum (Hedw) Rabenh
Ditrichum Hampe heteromallum (Hedw) Britt linellre (Sw) lindb
~ pallidum (Hedw) Hampe pusiUum (Hedw) Hampe
Saelania Lindb gIlucescens (Hedw) Bomanss et Broth
Ceratodon Brid purpureus (He(lw) Brid var prupurew
Distichium BSG capillaceum (Hedw) BSG inclinatum (Hedw) BSG
33 Bryoxipbiaceae Bryoxlphium Mitt
norvegicum (Brid) Mitt
34 Seligeriaceae Seligeria BSG
cumpylopoda Kindb ex Macoun et Kindb pusillll (Hedw) BSG recurvata (Hedw) BSG
Bindia BSG acuta (Hedw) BSG
35 Dicranaceae BlUehia Schwegr
lexUOSll (Sw ~ Schwaegr) C MueU sullivantii Aust
Trematodon Michx ambiguus (Hedw) Hornsch
Dicranellll (C Muell) Schimp cerviculata (Hedw) Schimp val cillPicuiata
heteromalla (Hedw) Schimp ~ rufescens (With) Schimp
schreberiana (Hedw) Schhnp var schreberiana schreberitJna (Hedw) Schimp var robusta Schimp ex Braithw subulata (Hedw) Schimp varia (Hedw) Schimp
Dicranodontium BSG denudatum (Brid) Britt ex Williams
Panzeucobryum (Lindh) Loeske Iongifolium (Hedw) oelke varlongifolium
17
Tortella (Undb) Limpr fragilis (Drumm) Umpr humilis (Hedw) JeM torlUoSfl (Hedw) Umpr
Hyophila Brid involuta (Hoole) Jaeg et Sauerb
Didymodon Hedw rigidulus Hedw
BryoerythrophyUum Chen recurvirostnon (Hedw) Chen
Btubula Hedw acula (Brid) Brid var acuta convoiuta Hedw fallox Hedw unguicultltll Hedw
Acaulon C MueD muticum (Uedw) C Muell var rufeseens (Iaeg) Crum
PJl~lm Hedw CUSPidtltum Hedw
De8mlltodon Brid cernuu (Hueb) BSG oblUsifoliw (Schwaegr) Schimp
T01IUla Hedw mueronifolia Schwaegr rurall (Hedw) Gaertn Meyer et Scherb var ruralis
39 Grimmiaceae Grlmmill Hedw
alpicola Hedw var alpicola alpicola Hedw var dupretii (Ther) Crum alpicola Hedw var rlvularl8 (Brid) Wahlenb apocarpa Hedw var apocarpa apocarpa Hedw var ambipa (Sull) Jonea apocarpa Hedw var conjerttl (Funck) Spreng apocarpa Hedw var strleta (Tum) Hook et Tayl CiIIyptrrlta Hook ex Drumm donniana Sm var donn_ hemumti Crum ltlevla (Brid) Brid montana BSG oineyi SuD oVllll8 (Hedw) lindb piUfe P-Beauv pltJgiopodill Hedw pulvinata (Hedw) Sm varbullpulvinata teretinervl Limpr
18
Orothera C Muell leana (Sull) C Muell
Rfwbdoweisia crispata (With) Undb
Cynodontium BSG ex Schimp alpestre (Wahlenb) Milde strumiferum (Hedw) Undb
Dichodontium Schimp pellucidum (Hedw) Schimp
Oncophorus (Slid) Brid vuens (Hedw) Brid wahlenbergii (Brid)
Dicranum Hedw bonjeanii DeNot ex Usa drummondii C Muell lagellare Hedw varj1ogellare fulvum Hook fuscescens Turn var fuscescens montanum Hedw muehlenbeckii BSG var muehlenbeckii polysetum Sw sabuletorum Ren et Card scoparium Hedw undulatum Brid viride (Sull et Lesq ex Sull) lindb
36 Leucobryaceae Leucobryum Hampe
glaucum (Hedw) Angstr ex Fr
37 Encalyptaceae Encalypta Hedw
ciliata Hedw var ciliata procera Bruch
38 Pottiaceae Astomum Hampe
muhlenbergianum (Sw) Grout
Weissia Hedw controversa Hedw var controversa
Gymnostomum Nees Homsch et Stunn aeruginosum Sm recurvirostrum Hedw var recurvirostrum
Trichostomum Bruch spirale Prout
19
40 Ephemeraceae Ephemerum Hampe
cohaerens (Hedw) Hampe crassineium (Schwaegr) Hampe val crassinervium spinulosum Bruch et Schimp ex Schimp
41 Funariaceae AphflllolThegma SuD
serratum (1 Hook et Wils ex Drumm) Sullo
Physcomitrium (Brid) Fuernr hookeri Hampe immersum Sullo pyriforme (Hedw) Hampe
Funaria Hedw americana Undb hygrometrica Hedw vaT hygrometrica
42 Splachnaceae Splachnum Hedw
ampullaceum Hedw
43 Schistostegaceae Schistostega Mohr
pennata (Hedw) Web el Mohr
44 Bryaceae Pohlia Hedw
annolina (Hedw) Lindb var annotina camea (Tum) Lindb cruda (Hedw) Undb elongata Hedw iliformis (Dicks) Andr lescurlana (Sull) Grout nutans (Hedw) Lindb obrusifolia (Brid) L Koch proligera (Undb ex Umpr) Undb ex Amell wahlenbergii (Web et Mohr) Andr
Leptobryum (B S G) Wits pyriforme (Hedw) Wils
Bryum Hedw algovicum Sendtnex C Muell argenteum Hedw caespiticium Hedw capillare Hedw var capillare creberrlmum Tayt lonchocaulon C Muell pallens (Brid) Sw ex Roehl pseudotrlquetrum (Hedw) Gaertn Meyer et Scherb varpudotriquetrum
20
pseudotriquetrum (Hedw) GaertnMeyer et Scherb var bimum (Schreb) Ulj stenotrichum C Muell var stenotrichum tortiolium Funck ex Brid uliginosum (Brid) B S G violaceum Crundw et Nyh weigelii Spreng
Rhodobryum (Schimp) Hampe roseum (Hedw) Limpr
45 Mniaceae Mnium Hedw
ciliare (C Muell) Schimp cinclidioides Hueb cuspidarum Hedw drummondii Bruch et Schimp longirostrum Brid lycopodioides Schwaegr marginatum (With) Brid ex P Beauv mediumB~S G var medium onhorrhynchum Brid pseudopunctatum Bruch et Schimp punctalUm Hedw var punctatum punctatum Hedw var elatum Schirnp rugicum Laur spinufosum B S G stellare Hedw
46 Aulacomruaceae Au1acomnium Schwaegr
androgynum (Hedw) Schwaegr heterostichum (Hedw) B S G paJustre (Hedw) Schwaegr var paJustre
47 Meesiaceae Amblyodon B S G
deillbotus (Hedw) B S G
Meesia Hedw triquetra (Richt) Angstr uJiginosa Hedw
48 Bartramiaceae Plagiopus Brid
oederiana (Sw) Umpr
Bartramia Hedw pomiformis Hedw var pomiformis
Philonotis Brid fontana (Hedw) Brid varfontana longseta (Michx) Britt varlongseta marchica (Hedw) Brid
21
~
~
49 Timmiaceae Timmia Hedw
megapolitana Hedw var megapolitana
50 Orthotrichaceae Zygodon Hook et Tayl
viridissimus (Dicks) Brid var rupestris lindb ex C J Hartm
Amphidium Schimp lapponicum (Hedw) Schimp mougeotii (B S G) Schimp
Orthotrichum Hedw anomalum Hedw obtuaifolium Brid ohioense Sullo et Lesq ex Aust pumilumSw sordidum Sullo et Lesq ex Aust speciosum Neesax Sturm var eegam (Schwaegr ex Hook et Grev) W~rnst stellDtwn Brid strrmgulatum P Beauv
Uleta Mohr ex Web cOfIICtata (p Beallv) Hamrn crispa (Hedw) Brid
Drummondill Hook ex Drumm prorepens (Hedw) Brltt
51 Fontinalaceae Fantinalis Hedw
antipyretica Hedw var antipyrettca antipyretica Hedw forma rabusta (Card) Card dalecarlica Schimp ex B S G var dalecarlica flqccidtJ Ren et Card hypnoides C J Hartrn var hypnoides hypnoides C J Hartm var durillei (Schimp) HU$ll
missourlcaCard novae-angliae Sullo var novae-angliae splulgrUlolill (C Moen) Wijket M~S
DichelY1l1ll Myr capilloceum (With) Myr falcatum (Bedw) Myr HIllescens B S G
52 Climaciaceae aimecium Web et Mohr ex Mohr
americllnum Brid dedroides (Hedw) Web et Mohr
22
S3 Hedwigiaceae Hedwigia P Beauv
ciliata (Hedw) P Beauv
54 Cryphaeaceae Forsstroemia lindh
ohioense (Sull) lindh
55 Leucodontaceae Leucodon Schwaegr
brachypus Blid var brachypus brachypus Brid var aniJrewsitlnus Crum et Anderson julaceus (Hedw) SuU
56 Neckeraceae Neckera Hedw
pennata Hedw var pennata
1fomalia (Brid) B S G trichomanoides (Hedw) B S G
57 Theliaceae Thelia Sull
asprellll Sullo hirtella (Hedw) Sullo lescurii SuU
Myurella B SG julacea (Schwaegr) B S G
sibrica (C MueU) Reim var sibrica
58 Fabroniaceae Fabronio Raddi
ciliaris (Brid) Bnd
Anacamptodon Brid splachnoides (Groet ex Brid) Brid
Schwetschkeopsis Broth fabronia (Schwaegr) Broth
59 Leskeaceae Lindbergia Kindb
brachyptera (Mitt) Kindb
Leskea Hedw gracilescens Hedw obscura Hedw poiycarpa Hedw
Leskeella (limpr) Loeske nervosa (Brid) Loeske
23
Pterlgy1lllndrum Hedw filiforme Hedw
60 Thuidiaceae Haplohymenfum Dozy et MoIk
trlste (Ces ex De Not) Kiodb
Anomodon Hook et Tayl attenuatus (Hedw) Hueb minor (Hedw) Fuemr rostratus (Hedw) Schimp viticulosus (Hedw) Hook et Tayl
Haplocladium (C Mucn) C MueU microphyllum (Hedw) Broth virginillnum (Brld) Broth ~bull
Thuidium B S G abietinum (Hedw) B S G delicatulum (Hedw) B S G vardelicatulum delicatulum (Hedw) B S G var radicans (Kindb) Crum Steere et Anderson minutulum (Hedw) B S G pygmaeum B S G recognftum (Hedw) Lindb scitum (p Beauv) Aust
Helodium (Sull) Warnst blandowli (Web et Mohr) Warnst var blandowli blandowli (Web et Mohr) Wamst var helodioides (Ren et Card ex Roell) Crum
Steere et Anderson paludomm (Sull) AuS
61 Amblystegiaceae Cratoneuron (Sull) Spruce
commulatum (Hedw) Roth var commutfltum commutatum (Hedw) Roth var fakatum (Brid) Moenk ilicinum (Hedw) Spruce var filcinum
Campylium (Sun) Mitt chrysophyllum (Brid) 1 Lange var chrysophyllum hispidulum (Brid) Mitt var hllJidulum polygamum (B S G)C lens varpolygtlmUm stellatum (Hedw) C1ens vat stelltum
Leptodictyum (Schimp) Warnst laxirete (Card et Ther) Broth riparium (Hedw) Wamat trichopodium (Schultz) Warns l1acillans (Sull) Broth
Hygroamblystegium Loelke jlul1iatile (Hedw) oeske var luViatile
24
fluviatile (Hedw) Loeske var oWltum Grout noterophilum (Sun et Lesq ex Sull) Warnst tenox (Hedw) Jenn var tenax tenox (Hedw) Jenn var spinilolium Schimp) Jenn
Amblystegium BSG juratzkanum Schimp serpens (Hedw) BSG varlum (Hedw) Undb var varium
Platydictya Berk confervoides (Brid) Crum jungermanniodes (Brid) Crum minutissimum (Sull et Lesq ex Sull) Crum subtile (Hedw) Crum
Drepanocladus (C Muell) Roth aduncus (Hedw) Warnst var aduncus aduncus (Hedw) Warnst var capillilolius (Wamst)Riebm aduncus (Hedw) Warnst var kneiffii (BSG) Moenk aduncus (Hedw) Warnst var polycorpus (Bland ex Voit) Roth exannulatus (BSG) Warnst fluitans (Hedw) Warnst var fluitans revolvens (Sw) Wamst var revolvens sendtneri (Schimp) Warnst var sendtnerl uncinatus (Hedw) Warnst var uncinatus uncinatus ffiedw) Warnst var piwnosus (Schimp) Wamst vernicoms (Lindb ex C Hartm) Warnst
Hygrohypnum Lindb eugyrium (BSG) loeske luridum (Hedw Jenn varluridum ochraecum (Tum ex Wils) Loeske var ochraceunt
Calliergon (Sull) Kindb cordilolium Hedw Kindb glganteum (Schimp) Kindb var giganteum richardsonii (Mitt) Kindb ex Wamst stramineum (Brid) Kindb trilarium (Web et Mohr) Kindb
Calliergonella Loeske cuspidata (Hedw) Loeske
Calliergidium (Ren) Grout bakeri (Ren) Grout
Scorpidium (Schimp) Urnpr scorpiodides (Hedw) limpr
62 Brachytbeciaceae Tomenthypnum loeske
nitens (Hedw) Loeske var mtens
25
Homalotheciella (Card) Broth subcapillata (Hedw) Broth
Brachythecium BSG acuminatum (Hedw) Awt acutum (Mitt) Sullo calcareum IGndb campestTe (C MuelL) BSG cunum (Lindb) Umpr digastrum C Muell et Kindb ex Macoun et Kindb oxycladon (Brid) Jaeg et Sauerb var oxycladon plumosum (Hedw) BSG populeum (Hedw) BSG var populeum reflexum (Starke ex Web et Mohr) DsG var relexum reiexum (Starke ex Web et Mohr) DSG var pacificum Renet Card ex Roell rlvulare BSG var rivulare tabulum (Hedw) BSC var tabuum salebrosum (Web et Mohr) BSG var salebrosum velutinum (Hedw) BSG Vat velutinum
Bryhniil Kaur graminicolor (Brid) Grout var graminicolor novae-angliae (Sull et Lesq ex sun) Grout
Rhynchostegium BSG senulatum (Hedw) Ja4lg et Sauerb
Rhynchostegiella (BSG) Limpr compacta (C MueU) Loeske
EU1hynchium 8SG hians (Hedw) Sande Lac pulchellum (Hedw) Jenn var pulchelium riparloides (Hedw) Rich
Stokeriella (Kindb) Robins pmelonga (Hedw) Robins var stokesii (Tum) Crum
63 Entodontaceae Entodon C Muell
brevisetus (Hook et Wils ex Wils) Lindb cladorrhizans (Hedw) C Muell seductrlx (Hedw) C MueU var wluctrix
Pleurozium Mitt schreberl (8rid) Mitt var schreberi
64 Plagiotheciaceae Pltlglothecium BSG
CflVifolium (Brid) Iwats var CtWifolium denticulatum (Hedw) 8SG laetum BSG ltztebricola BSG
26
6S SernatophyUaceae Brotherella Loeske ex Fleisch
recumins (Michx) Fleisch tenuirostris (Bruch et Schimp ex Sull) Fleisch
Sematophyllum Mitt demissum (Wils) Mitt
66 Hypnaceae Pl(itygyrium BSG
repens (Brid) BSG
PylaisieUa Kindb intricata (Hedw) Grout polyantha (Hedw) Grout selwynii (Kindb) Crum Steere et Anderson
Homomallium (Schimp) Loeske adnatum (Hedw) Broth
Hypnum Hedw cupressiforme Hedw var cupressiforme cupressifonne Hedw varfiliforme Brid curviolium Hedw fertile Sendtn imponens Hedw lindbergii Mitt palescens (Hedw) P-Beauv pratense Koch ex Brid
Calicladium Crum haldanianum (Grev) Crum
sopterygiopsis Iwats muelleriana (Schimp) Iwats
Isopterygium Mitt e1egans (Brid) lindb pulchellum (Hedw) Jaeg et Sauerb tenerum (Sw) Mitt
Taxiphyllum Fleisch depllQtum (Bruch et Schimp ex sun) Fleisch taxirameum (MitL) Fleisch
Herzogiella Broth seligeri (Brid) Iwats striatella (Brid) Iwats ruracea (Lindh) Iwats
hilium De Not crista-castrenna (Hedw) De Not
27
67 Rhytidiaceae Rhytidium (Sull) Kindb
rugosum (Hedw) Kindb
Rhytidfodelphus (Lindb ex Iimpr) Warnst triquetrus (Hedw) Warnst var triquetrus
68 Hylocomiaceae Hylocomium BSG
pyrenaicum (Spruce) lindb splendens (Hedw) BSG varsplendens
69 Buxbaumiaceae Buxbaumia Hedw
aphyllll Hedw
Diphyscium Mohr follosum (Hedw) Mohr
70 Tetraphidaceae Tetraphi3
pellucida Hedw
71 Polytrichaceae Atrichum PmiddotBeauv
angustatum (Brid) BSG cri8pum (James) Sullo oerstedianum (C Muell) Mitt teneUum (Roehl) BSG undulatum (Hedw) P middotBeauv var unduiatum
Pogonatum P-Beauv alpinum (Hedw) Roehl var alpinum umigerum (Hedw) P-Beauv
Poiytrichum Hedw commune Hedw var commune ormosum Hedw juniperinum Hedw longisetum Brid ohioense Ren et Card piliferum Hedw var pilierum strictum Brid
28
MAPS OF BRYOPHYTE TAXA
REPORTED FOR WISCONSIN
HORNWORTS AND LIVERWORTS
29
bull bull WSHeV~N S vE II
0 ~
e~RRON ~USK
(J~ (j -
CHIPPE A DUNN
0 o0 poundAU CLAIRE
~ TREMshyPEAshyLEAU
MICE
ONEIDA
Q LINCOLN
TAYLOR
Q t-~----- MOIRA THON
CLARK
o
10
LA FAVETTE
Hornwor ts amp Liverworts
- None - 1 - 25 taxa
- 26 - 50 tRXshy
- 51 - 75 tax)
- 7c - 100 taxa
FONDOU j LAC ~
DODGE
i O JEfFERSON
WALWORTH
30
Anastrophylltlfl helleranum (Nees) Schust Anastrophyllum michauxil (We~ Buch ~
46 88 136 141 Evans 38 46 141
Anastrophyllum mlnut~ (Schreb) Schuster Aneura plnguls (L) Durn
89 21 384687 88 89 90 (138) (145)
Anthoceros macounil MA Howe ~oceros punctatus L
21 (45) (144) 34
31
Barbilophozia attenuata (Mart) Loeske Barbilophozia barbata (Schmid ~ Schre~
24 141 Loeske 28 39 46 89 90 141
Bazzania trilobata (1) S Gray
1 26 39 46 (57) (59) 79 88 89
90 119 141
BlePbarostoma trichoffiyllum (1) Dumort
1 8 26 39 46 89 90 137 141
Blasia pusilla 1
1 39 46 90
Calypogeja fissa (1) Raddi
90
32
$129geja integristipula Steph C~Ypound2geja muelleriana (Schifm) K Mull
88 89 89
CallpoundO~eja neesiana (Mass amp Carest) K Mull
39 46 89 140 (141)
~pogeja sJlapound2icola (H Arnell tl J bull
Perss) Warnst et Loeske (119) 141
CeIalozia biscuspidata (L) Dum
1 37 I 46 90 t 137 141
CalYpogej~ trichomanis (L) Corda
26 39 (119) 137
33
CeEhalozia catenulata (Hub) Lindb Cephalozia connivens (Dicks) Lindb
39 46 79 141 39 4679 88 (89) 90 119 141
Cephalozia lacinulata Jack ~ Spruce
141 Cephalozia lunulifolia (Dum~ Dum I 28 39 46 79 88 89 90 141
Cepoundhalozia macouni (Aust) Aust Cephalozia ~leniceps (Aust) Lindb 27~ 3746 (66)7987 (139)141 39 46 79 89 141
34
Ceffialoziella 9ivaricata (Sm) Schiffn Cephaloziella ~hista (Jack) Schiffn
46 90 46 (119) 141
Cephaloziella hampeana (Nees) Schiffn
46
Cephaloziella rubella (Nees) Warnst
46 39
Cephaloziella rubella var sullivantii
(Aust) K Mull 46 39
Cephaloziella rubella var bifida (Lindb) Douin
35
46
Chiloscyphus pallescens (Ehrh ~ Hof~ Dum Chiloscyphus pallescens var fragilis
39J 46 88 89J 90 (Roth) K Milll 46
Chiloscyphus E9lyanthos (L) Corda
1 24 39 46
Chiloscyphus polyanthos vax rivularis
(Schrad) Nees 39 46
poundpoundnocepha1um conicum (1) Lindb
12 28 39 46 (49) 87 88 8990 120 121 148
Cololejeunea biddlecomiae (Aust) Evans
39 46 62 87 88 89 90
36
Dipl0]hyllum apicu1atum (Evans) Steph Fossombronia foveo1ata Lindb
(35)41 46 90 140 1 t 38 (39) t 46
Frullania bolanderi Aust
13 46 62 88 90 139 140
Frullanla ari ttoniae Evans
39 46 69 (71) 87 88 89 90
Frullania eboracensis Gott
1 9 28 4 39 46 (57) (59) 69 70 (71) 87 88 89 90 92 93 97 148
Frullania inflata Gott
46 87 88 89 (139)
Frullania oakesiana Aust Frullania plana Sullo
(35) 42 46 87 (139) (140) 92
Frullania ri paria Hampe ~ Lehm
46 69 71 87 89
Frullania ~elwynianna Pears
46 89
Geocalyx SEaveolens (Schrad) Nees
394688 8990 (119) 140
Frullania tamarisci (L) Dum subsp sPagrayana
(Mont) Hatt 28 34 l~6 t (57) 88 89 90
38
Mar pan thus seutatUB ( Web amp Mohr) Spruce Jamesoniella autumnalis (DC) Steph
(35) 39 46 90 1 7 28 39 46 56 (57) 87 8889 90 (119) 137 141
Jungermannia confertissima Nees
137 141
Jungermannia crenuliform1s Aust
79 137 141
Jungermannia ~il1ima Sm
46 90 141 Jungermannia exsert1folia subsp corifo~ (Dum~
Vana (35) 46 79 141
39
Jungermannia hyalina Lyell Jungerrr6nnia 1eiantha Grolle
40 46 79 136 141 39 46 79 88 90 1)7 141
Jungermannia IXlaxis Lindb Jungermannia pumila With
46 67 79 (139) (140) 141 )9 46 90 1)6 141
Jungermannia sphaerocarpa Hook Kurzia setacea (Web) Grolle
4679136140IIH 136
Lejeunea cavifo1ia (Ehrh) Lindb Lepidozia reptans (1) Dum
1 7 27 39 46 88 89 90 137 14010 I ( 27) 46 I 69 (71) 88
lophocolea heterophylla (Schrad) Dum Lophocolea minor Nees
1 )4 39 46 (57) 87 88 89 27 39 46 88 89 90 (119) 90 (93)94 119 136
Lophozia al~stris (Schleich ~ Web) Evans Lophozia ascendens (Warnst) Schust
46 90 137 141 136 140 141
41
loIilozia badensls (Gatt ~ Rabenh)
46 141
lophozia blcrenata (Schmid sectA Hoffm)
Dum 46 141
lophozia collaris (Nees) Dum
39 46 141
lophozla excisa (Dicks) Dum
27 4679 141
lophozia gillmani (Aust) Schust lophozia gIlttulata (Undo tl H Arnell)
36 39 40 46 136 (140) 141 Evans 39 46 136 141
42
141
Lophozia heterocolEDs (Thed) MA Howe IoPhozia ipcisa (Schard) Dum
27 39 46 89 141 8 27 31 46 79 90 140 141
LoQhozia longidens Macoun Lophozia ventricosa
1 46 79 90 141
Lophozia ventricosa var silvicola
(Buch) Jones 141 LoEhozla ventricosa vax longif1ora (Nees)
Macoun 39 46 79 141
43
~~ fragrans (Balbis) Frye et Clark Mannia pi 10sa (Hornem) Frye et Clark
46 (139) (140) 9 46 79 116 139
Mannia triandra (Scop) Gro 11e
46 (140)
Marchantia pound91ymorpha L
1 13 28 3946 (49) 78 87 88 89 90 (Ill) 121 122 147
Mo~rckia hibernica (HookJ Gott
46 89
Metzgeria conjugata Lindb
44
46
Mylia anomala (Hook) S Gray Notothylas orbicularis (Schwein) Sullo
39 46 141 (35)45 46
Nowellia curvifolia (Dicks) Mitt
17 27 3846 88 89 90 141
Odontoschisma denudatuJI1 (Nees ~ Mart)
Dum 46 81 137 141
Pallavicinia lyellii (Hook~ Carruth
90
Odontoschisma macounii (Aust) Under
45 46 (66) (139) 140 141
45
=
Pellia epiphllla (L) CordaPellia endiviifolia (Dicks~ Dum
27 39 46 90 13724 39 46
Pellia neesiana Limp bull Phaeoceros
39 46 89 27 39 45 46 97
Porella pinnata L
13 27 I 46 90 Plagiochila asplenioides (L) Dum
27 I 39 1+6 88 89 I 90 137
46
~e11a platYEhylla (L) Pfeiff Porella platyphylloi~ (Schwein) Lindb
17 27 39 46 69 (71)79 87 88 89 9 34 46 (57) 59 87 88 89 92 90 92 93 77
Preissia quadrata (Seop) Nees
1 36 3946 (71) 88 89 90 121 123
137
Ptllidium ciliare (L) Hampe
1 27 39 46 132 141
Dum
(22) 28 3946 (57) 87 88 89 90 92
Ptilidium ~lcherrium (G Web) Hampe
(9) 13 394656 (57)70 87 88 89 90 93 (119) 136 141
47
Radula obconica Sullo Reboulia hemlsphaerlca (L) Raddi
39 46 88 89 (43) 46 (59) 70 90 t Cf 124 125 (139)
Riccardia ~atifrons Lindb
1 27 39 46 89
Riccardia multifi~ (L) S Gray
39 46 88 145
Riccia beyrichiana Hampe ~ Lehm
46 Riccardia palmata (Hedw) Carruth
39 46 90
48
Riccia bifurca Hoffm Riccia cavernosa Hoffm
48 13739 46 79 137
Riccia fluitans L
242731454679
Riccla frostli Aust
48 79
Riccia sullicantii Aust
79 (137) Riccia sorocarpa Bisch 46 (137)
49
Ricciocarpus natans (1) Corda Sca~ia apiculata Spruce
13 27 46 79 89 110 39 46 (139) 141
ScaJltll1ia carinthica Jack ~ Lindb
137 t 141
Scapania ~ (Mart) Durn
39467990137141
Scapania digenil Schiffn
90
ScaEsectflia cuspiduligera (Nees) K Mull
46 (139) (140) (141)
50
Scapania glaucoeerha1a (Tayl) Aust Seapania gyrrmostomophilia Kaal
39 46 (139) 141 41 44 46 64 (139) (140) 141
ScaP3poundlia irrigua Gott ~ almiddot
7 39 46 79 137 (140L 141
Seapania nemorosa (L) Dum
1 39 46 89 90 137
Scapmi~ ~~ta Smiddotlch
137 141 51
~caPnia poundludicola LDeske amp K Mull Scapania saxicola Schust
46 (66) 141 136 141
Sc~~ scandica (H Arnell ~ Buch) Macv
139 140
Scapania subalpina (Nees) Durn
39 40 46 79 137 141
Scapania undulata (L) Dum
1 42 46 79 141
Scapania umbrosa (Schrad) Dum
137 140 141
52
Trichocolea tomentella (Ehrh) Dum Triomarla exsecta (Schmid) Schiffn
27 (35) 39 46 47 88 137 141 8 28 39 46 90 140 141
Tritomaria exsectiformis (Breidl) Loe~ke Tritomaria Sluinguedentata (Huds) Buch
39 -t6 89 90 136 140 141 7 39 46 141
Trltomaria scitula (Tayl) Joerg
46 136 (139) 141
53
ONEIDA
LINCOLN
TAYLOR
t---------- MARATHON C1l-RK
SAu~
t
~
~ C i-W=AS_-r7-1
l=itHLANO
)
lOCK
INGTb
lt
WALWORTH
5
INTRODUCfION
This atlas of the bryophyte flora of Wisconsin is an attempt to show the distribution of all bryoshyphyte species known to occur in the state by county occurrenoes based on published reports in the literashyture It is not an annot~ted check list (only a feW problematic taxa were examined) but is a literature search intended as a foundation for a complete flora or for otherbryological research
The literature of btYo1ogy is large and spread over numerous journals reports monographs and manuals It would be almost impossible to search all sources for every reference attributing bryophyte species to Wisconsin and to particular localities in the state Wehav checked many inlportant references and paid special attention to works dealing with Wisconsiit botany and the bryologyof the upper Midwest Several bibliographies and indices were checked thoroughly including Green an~ Curtis (1955) the Bry ologist indices (Crum 1959 and Miller 1976) andt)le Wisconsin Academy of Sciences Arts and Letters indices
Only those published repots which app~red to)e substanti2ted by herbarium specimens and which gave- ~fficient location dat to P~CetheW(On in amiddot particula(county were-used in the initial compilation Species described in manuals in generalized way implying their occurrence in Wisconsin (Without menmiddot tioning Wisconsin specifically or citing a particular location in the state) were excluded eg Gnmmia raui (Wynne 1943) and Pohlia annotinll var decipiens (Crum 1976) Ecological papers which were either too general in taxonomic determinations (eg Sphagnum sp Polytrichum sp) or in location data were also excluded Some other taxa attributed to Wisconsin which have been excluded are
Cylindrothectum compr~ (Cheney 189tmiddotDane Co) identified later by Grout as Entodon seducshytrtx
Dicranell4 humilis (Holzjne~ 1925middotBuffalo Co) probablyD vllJia since Dhumilis is not found in North Arnerl~
Didymodon triarium (Foote 1 966-SW Visconsin) identified by Fred Hennann as Barbulaacuta Funoria microstoma (anonymous 1975middotDane Co) excluded because it is outside of its geographic
range and the packe~ QaMot be located Phil(motis calcarea (Cheney and Evans 1944-Dane Co) excluded in Crum Steere and Anderson
(1973) Plagiobryum demissum and P zierl (Cheney and Evans 1944-various counties) are both Arcticshy
alpine species not likely to be found in isconsin Two of Cheneys collections of Pl1giobyrum have been referred to Bryum
Pogonatum alpinum var septentrionale (Che~ey and Evans 1944-Chippewa Co) was misidentified (Ireland 1960) and is P umigerum
Rhizomnium fAPpoJachIanum and R punctalu1fl ssp chlorophyDosum (Koponen 1973middotseveral counshy ties) excluded North American bryologiflts have not recognized these taxa in America
Sphagnum tenellum (Cheney and Evans 1944-Waukesha Co)an Atlantic species not occurring in Wisconsin (Cruro 1975)
The nomenclature and systematic arrangement of hornwort 00 hepatic species in the atlas follow Stotler and Crandall-5totler (1977) and of moss species fonow Crum Steere and Anderson (1973) except in such instances as the JreptlIfockIdus fonns whiGh follow Wijk et aL (19S9middot1973) Species reported in the literature under names which are now listed as synonyms on either of these lists were converted to the accepted name A dot is placed in each county of Wisconsin from which the taxon has been reported The numbers after the name of the taxon correspond to the numbered references in the bibliography which giye ~act location data OEat least location data -s~fficient to place it in-particular counties in the state The numbers in parentheses indicate references which attribute the taxon to Wisconsin without providing exact loltation data
Fossil and subfossil reports are summarized Slfparately in our atlas only six species (Aulacomnium turgidum lJ1achythecium turgidum Ditrichum Jlelficaule Hypnum bambergti Scorpidium 6fJisities and Tortel14 1rIgi1i$) are known exclUsively from the fossil record ~
6
GEOGRAPHIC CLIMATIC AND BOTANICAL FEATURES OF WISCONSIN
Wisconsin is a north central state bounded on the east by Lake Michigan on the north by Michigan and Lake Superior on the south by Illinois and on the west by the Mississippi and St Croix rivers It is located midway between the Prime Meridian (0deg) of Longitude and 1800 Longitude and midway between the equator and the north pole thus placing it in the center of the northern half of the western hemisphere
The topography of the state is relatively flat a few hills exceed 1900 feet with Thns Hill at 1952 feet forming the highest point in Price County Lake Michigan represents the lowest point at 580 feet The northern and eastern two-thirds of the state were heavily glaciated during the last stage of the Pleistocene and now contain many of the best examples of continental glaciation topOgraphy including considerable ground moraine large terminal recessional and intetlobate moraines kames eskers drumlins and small depressions or kettle-holes Most of the nearly 9000 lakes of 20 acres or more lie within this part of the state The southwestern third of the state is the Driftless Area It includes most of the large flat generally sandy bed of Glacial Lake Wisconsin plus the highly dissected ridge and valley section with numerous cliffs and rock-outcroppings (Figure 1)
Much of the bedrock in the state lies below glacial till but exposures include igneous and metamiddot morphic rocks primarily granitic especially in the north as well as sedimentary layers primarily sandshystones and dolomitic limestones especially along Lake Michigan and in the DriftlesS Area
The climate of Wisconsin is classified as humid-temperate with cold winters The annual precipimiddot tation averages approximately 30 inches which is sufficiently even in its seasonal distribution to support forest vegetation throughout most of the state Nevertheless according to Curtis (I 959) the pre-settlement vegetation of the state (J 830-1850) included about 27 prairie and oak-savanna plus about 64 forests
The state is presently divided into seventy-two counties Marathon County is the largest (1586 sq mi) the smallest counties include Pepin (235 sq mi) Ozaukee (236 sq mi) andMilw8ukee (237 sq mi) We included Menominee County though its future status as a county is in doubt since it regained its Indian reservation status and rights in 1973 This county was established April 30 1961 from the Menomishynee Indian reservation which consisted of parts of Shawano and Oconto counties Wisconsin ranks 26th in the United States in area with 54464 square miles and 16th in population with 4651000
More ecological research is needed on the bryophyte flora of Wisconsin especially to relate it to the vegetation and to such other environmental features as soils climate and glaCial deposits Plant communishyties in Wisconsin are many and diverse these are summarized on Figure 2 from studies conducted by Curtis (1959) The state is divided into northern and southern floristic provinces by a tension zone deshyscribed by Curtis (1959) This tension zone is an eeotonal region vatying from 10middot31 mt wide where the ranges of many vascular plants overlap or terminatej it runs in an S-curve northeast from Milwaukee to Hudson in St Croix County (Figure 2) The principal plant communities of the northern province apshyproaching their southern boundaries within the zone include the northern conifermiddothardwood forests of sugar maple white pine hemlock and yellow birch and the Sphagnum bogs with tamarack and black spruce Southern communities at or near their northern limits within this zone include prairies oak and jack pine savannas and hardwood forests of maples oaks ash ehn and cottonwood
We hope to encourage research on cryptogamic ecology with our checklist We know that some species are northern while others are southern (eg cimacium) fits the tension zone concept Freckmann (I977) while Bryoxiphium is only found in the Driftless Area of the state Steere (1937) and Love and Love (1953) The tension zone theory may prove to be acceptable for bryophytes We speculate that certain bryophytes can indicate particular plant communities since many bryophytes are restricted to preshycise environmental requirements Many papers or theses have dealt with cryptogamic bark and rock ecology in Wisconsin (Culberson 1955 Foote 1962 Hale 1952 1955 and Cole 1975)
We hope that this atlas will stimulate collecting in counties poorly sampled so that our knowledge of the bryophytes can be increased It should aid anyone interested in floras for preservation environmental impact statements and distributional studies
7
A numerical summary of the bryophyte flora of the state based on our literature search is presented in Table 1 (below) This includes 517 taxa 70 families and 176 genera This numerical summary is comshypared with the published floras of Michigan mosses (Crum 1976) and Minnesota hepatics (Schuster 1953) Thus we surmise that further collecting could lead to the same approximate numbers for Wisconsin bryoshyphytes
Table 1 Comparison of Recorded Bryophytes of Wisconsin with Michigan and Minnesota and Summary of Wisconsin Taxa
Wisconsin Michigan (Crum 1976)
Minnesota (Schuster 1955)
Hepaticsamp Hornworts
Taxa Families Genera
137 28 47
174 32 52
Mosses
Taxa Families Genera
380 42
129
503 42
149
8
-
~
~ tIt
Figure 1 The Driftless Area and Tension Zone The counties are 01$0 shown on map
ca location of tension zone as deecfibed bv Curtis (1959)~ The Orittless Area
9
Agn 2 Major Plant Communitiee of Wsconsln aaed on surwvor recorda bull co 18laquo) After Curtismiddot 1959
tit beol Forest e Phillie
Northernmiddot hardwood forest reg Pine 5cMInna
copy Southernmiddot hordwood forest CD Oak Savanna
10
SYSTEMATIC ARRANGEMENT OF WISCONSIN BRYOPHYTES
ANTHOCEROTAE (Hornworts)
1 Anthocerotaceae Anthoceros 1 emnd Prosk
macounii M A Howe punctatum 1
Phaeoceros Prosk laevis (1) Prosk subsp laevis
Notothylas Sul1 orbicularis (Schwein) Sullo
HEPATICAE (liverworts)
2 PseudolepicoJeaceae Blephllrostoma (Dum emend Undb) Dum
trlchophyllum (L) Dum subsp trlchophylluni
3 Ptilidiaceae Ptilidium Nees
ci1iare (L) Hampe pulcherrimum (G Web) Hampe
4 Trichocoleaceae Trichocolea Dum nom cons
tomentella (Ehrh) Dum
5 Lepidoziaceae Kunia von Martens
setacea (Web) GroUe
Lepidozia (Dum) Dum reptans (1) Dum
Bazzania S Gray trilobata (1) S Gray var triZobata
6 Calypogejaceae 011ypogeja Raddi
flSsa (L) Raddi subsp issa integristipula Steph muelleriana (Schiffn) K Mull subsp muelleriana neesiana (Mass et Carest) K Mull sphagnicola (H Arnell et J Perss) Warnst et loeske trichomanis (1) Corda
11
7 Cephaloziaceae Cepluzlozla (Dum emend Schiffn) Dum
biscwpidata (L) Dum subsp biclupidIIta catenulllttl (Hub) Lindb coMivens (Dicks) Lindb var connivens lacinultzta Jack ampX Spruce lunulifolilz (Dum) Dum macounii (Aust) Aust pleniceps (Aust) Lindh var pleniceps
Noweilia Mitt cunifolilz (Dicks) Mitt
8 Adelanthaceae Odontoschimuz (Dwn) Dum
denudatum (Nees ampX Mart) Dum Vir denudatum mtlCount (Aust) Underw
c
9 Cephaloziellaceae CephtlJoziellll (Spruce) Schiffn nom C()II$
divarlctzt4 (Sm) sclurfn val divarlcat(l elachhta (bdc) Schiffn 1uImpetuw (Nees) Schirfn IUbellll (Nees) Warnst vat IUbeIl4rube (Nees) Warnst var biid4 (Lindb) Dquinrube (Nees) WarDSt Var sulliwmtii (Aust) K MuD
10 Geo~c~ larptuttllus Nees
scutatus (Web et Mohr) Spruce
Geocalyx Nees grGVeolens (Schrad) Nees
11 Lophocoleaceae Lophocolea (Dum) Dum
heteropllyll4 (Schrad) Own minorNees
Chiloscypbus Corda om cons pallescens (Ehrh ampX Hoffm) Dum varfragllis (Roth) K Mun polyantitos (L) Corda var poiyanthos pol)llaquo1lthos (L) Corda var rlvulllris (Schrld) Nees
12 PlagiochiJaceae Plagio(hila Dum nom cons
tlBpienioides (L) Dum var asplenioidu
13 Jungermanniaceae Btriilophoril Loeske
attenutlta (Mart) Loeske bIubtlta (Schmid ampX Scltreb) Loeske
12
Lophom (Dum) Dum alpestris (Schleich ~ Web) Evans ascendens (Warnst) Schust badenais (Gott ex Rabenh) Schiffn var bademii bierenata (Schmid ex H)ffm) Dum eollam (Nees) Dum exci8a (Dicks) Dum var excisa gllimanii (Aust) Schust varglllmanii guttulata (Lindb et H Arnell) Evans heterocolpos (Thed) M A Howe var heterocolpos ineistl (Schrad) Dum var tnelsa longidens (Lindb) Macoun subsplongidens ventricosa (Dicks) Dum var ventricosa ventricolfl (Dicks) Dum varlontifIotrl (Nees) Macoun ventricosa (Dicks) Dum var silvlcola (Bueh) Jones
Tritomarill Schiffn ex Loeske exseeta (Schrad) Loeske exsecttformi$ (Breidl) Loeske subsp exsectiormfs quinquedentata (Huds) Buch var quinquedentata scitula (rayl) Joerg
Anastrophyllum (Spruce) Schiffn hellemnum (Noes) Schust michtzuxii (Web) Buch ex Evans minutum (Schrob) Schust var minutum
Mylill S Gray nom cons anomala (Hook) S Gray
Jamesoniella (Spruce) Carring autumnalis (DC) Steph var autumnalis
Jungemumnia L emend Dum eonfertissimtt Nees crenuliformis Aust exsertijolill Steph subsp cordifolia (Dum) Vana graciUima Sm hyalina Lyell leillnthll Grolle polaris Lindb pumila With sphaerocarpa Hook
14 Scapaniaceae Diplophyllum (Dum) Dum nom cons
apiculatum (Evans) Steph var apiculatum
Scapanill (Dum) Dum nom cons apkuata Spruce carinthillca Jack ex Lindb curta (Mart) Dum var curta
13
cuspiduligera (Nees) K Mull var cuspiduiigera degenii Schlffn ex K Mull var degenii glllucocephaJa (Tayt) Aust gymnostomophila Kaal irrigwJ (Nees) Gott et al subsp irrigua lingulata Buch varlingulata mucronata Buch subsp mucronata nemorosa (l) Dum paludicola Toeske et K Mun var paludicola paludosa (K Mull) K Mull saxicola Schust scandiea (H Arnell et Buch) Macv var scandiea 8Ubalp~ (Nees) Dum umbroSll (SchIad) Dum undulata (L) Dum var undulata
15 Radulaceae Raduia Dum nom COllS
compianata (L) Dum obconica Sull
16 Porellaceae Parella L
pinTlllta L patyphylla (L) Pfeiff platyphylloidt(l (Schwein) lindb
17 Jubulaceae FruUanill Raddi
bolanderi Aust brittoniae Evans eboracemis Gott inlllta Gott oakesiana Aust plana Sull rlparia Hampe ex Lehm selwyniana Pears taTnl1iJci (L) Dum subsp asagmyana (Mont) Hatt
18 lejeuneaceae Calolejeunea (Spruce) Schlffn
biddlecomiae (Aust) Evans
Lejeunea Ub nom cons cavifolia (Ehrh) Lindb emend Buch
19 Codoniaceae Fossombronia Raddi
foveolata Lindb
20 Pa11aviciniaceae Paklvieinlil S Gray nom cons
lyeiii (Hook) Carruth
14
Moerckia Gott hibemica (Hook) Gott
21 PeWaceae Pellfa Raddi nom cons propos
endivlifolfa (Dicks) Dwn epiphylllz (L) Corda neeBiana (GottlUmpr
22 Metzgeriaceae Metzgerla Raddi
conjugata Undb
23 Blasiacae B L
pulilm L
24 Aneurareae AneumDum
pinguis (L) Dum
Riccfl1dia S Gray nom cons latifrons Undb multiftda (L) S Gray var multifida palmata (Hedw) Carruth
25 Aytoniaceae Reboulill Raddi nom conI
hemllphaerica (L) Radeli
ManniaOpiz fragrrms (Balbis) Frye el ClarK pilosa (Hornem) Frye et Clark trltmdra (Scop) GroDe
26 Conocephalaceae Conocephalum Wigg nom cons
conicum (L) Lindb
27 Marchantiaceae Preiuia Corda
quadrata (Scop) Nees
Marchantia L polymorpha L
28 Ricciaceae Riccwcarpus Corda
natans (L) Corda
Riccia 1 beyrichiana Hampe ex Lehm bifurca Horfm
15
i
CtlJPemostI Hoffnt fuitu L rostii Aust BOIOCtI1fHI Bi~ sulltvtmtii Aust
MUSCI (Mosses)
29 Sphagnaceae SpIutgnum L
capillaceum (Weiss) Schrank var Cllpil1tlceum caplllllceum (Weiss) Vir tenellum (Schimp) Andr centrille C Jens ex Amen et C Jens compactummiddot OC ex Lam et Dc ClUpidatum Ehrh ex Hoffin var aupidatum jimbrltltum Wits ex J Hookmiddot fulCUm (Scbimp) KUngrr glrgenBOhIlll Russ imbrlctltum Hornsch ex Russ muelllmitil4m BOd 1IIIIU (RUII) C Jens przIutre L papIIIoIum Lindb pulchrum (Lindb ex braithw) Wamst recumun P-Beauv var recumun recum P-Beauv var tenue Klinggr riptlrlum Ansstr ruPQWItWamst squtl1fOlUm Crome subftltens Russ et Wamst ex Wamst IUbIeCfUldum Nees ex Sturm var subsecundum subsecundum Nees ex Sturm varrutem (Nees Hormch et sUUm) Hueb teres (Sdlimp) Anptr e C Hartm
WtlI7IItorjii Russ wuljfanum Gira
30 Andreaeaceae AndNrrerI Hedw
rupeitrlr Oedw var pest
31 Fissident~e Fluid_ lledw
trdiantoid Hedw byoidei Bedw var bryolda cristaM Wils ex Mitt exiguus sun VIr exigUw onttmUi (B-Pyl) Steud jtuberi Lesq et James obtusifolhu Wits var obtwifoliul ollPlUlldQldes Heel ~litSuU lubbtulkril Bed tfDdfoliul lledw llridubu (S~ ) Wahlenb vallirlIU1u
16
32 Ditrichaceae Pleuridium Brid
subulatum (Hedw) Rabenh
Ditrichum Hampe heteromallum (Hedw) Britt linellre (Sw) lindb
~ pallidum (Hedw) Hampe pusiUum (Hedw) Hampe
Saelania Lindb gIlucescens (Hedw) Bomanss et Broth
Ceratodon Brid purpureus (He(lw) Brid var prupurew
Distichium BSG capillaceum (Hedw) BSG inclinatum (Hedw) BSG
33 Bryoxipbiaceae Bryoxlphium Mitt
norvegicum (Brid) Mitt
34 Seligeriaceae Seligeria BSG
cumpylopoda Kindb ex Macoun et Kindb pusillll (Hedw) BSG recurvata (Hedw) BSG
Bindia BSG acuta (Hedw) BSG
35 Dicranaceae BlUehia Schwegr
lexUOSll (Sw ~ Schwaegr) C MueU sullivantii Aust
Trematodon Michx ambiguus (Hedw) Hornsch
Dicranellll (C Muell) Schimp cerviculata (Hedw) Schimp val cillPicuiata
heteromalla (Hedw) Schimp ~ rufescens (With) Schimp
schreberiana (Hedw) Schhnp var schreberiana schreberitJna (Hedw) Schimp var robusta Schimp ex Braithw subulata (Hedw) Schimp varia (Hedw) Schimp
Dicranodontium BSG denudatum (Brid) Britt ex Williams
Panzeucobryum (Lindh) Loeske Iongifolium (Hedw) oelke varlongifolium
17
Tortella (Undb) Limpr fragilis (Drumm) Umpr humilis (Hedw) JeM torlUoSfl (Hedw) Umpr
Hyophila Brid involuta (Hoole) Jaeg et Sauerb
Didymodon Hedw rigidulus Hedw
BryoerythrophyUum Chen recurvirostnon (Hedw) Chen
Btubula Hedw acula (Brid) Brid var acuta convoiuta Hedw fallox Hedw unguicultltll Hedw
Acaulon C MueD muticum (Uedw) C Muell var rufeseens (Iaeg) Crum
PJl~lm Hedw CUSPidtltum Hedw
De8mlltodon Brid cernuu (Hueb) BSG oblUsifoliw (Schwaegr) Schimp
T01IUla Hedw mueronifolia Schwaegr rurall (Hedw) Gaertn Meyer et Scherb var ruralis
39 Grimmiaceae Grlmmill Hedw
alpicola Hedw var alpicola alpicola Hedw var dupretii (Ther) Crum alpicola Hedw var rlvularl8 (Brid) Wahlenb apocarpa Hedw var apocarpa apocarpa Hedw var ambipa (Sull) Jonea apocarpa Hedw var conjerttl (Funck) Spreng apocarpa Hedw var strleta (Tum) Hook et Tayl CiIIyptrrlta Hook ex Drumm donniana Sm var donn_ hemumti Crum ltlevla (Brid) Brid montana BSG oineyi SuD oVllll8 (Hedw) lindb piUfe P-Beauv pltJgiopodill Hedw pulvinata (Hedw) Sm varbullpulvinata teretinervl Limpr
18
Orothera C Muell leana (Sull) C Muell
Rfwbdoweisia crispata (With) Undb
Cynodontium BSG ex Schimp alpestre (Wahlenb) Milde strumiferum (Hedw) Undb
Dichodontium Schimp pellucidum (Hedw) Schimp
Oncophorus (Slid) Brid vuens (Hedw) Brid wahlenbergii (Brid)
Dicranum Hedw bonjeanii DeNot ex Usa drummondii C Muell lagellare Hedw varj1ogellare fulvum Hook fuscescens Turn var fuscescens montanum Hedw muehlenbeckii BSG var muehlenbeckii polysetum Sw sabuletorum Ren et Card scoparium Hedw undulatum Brid viride (Sull et Lesq ex Sull) lindb
36 Leucobryaceae Leucobryum Hampe
glaucum (Hedw) Angstr ex Fr
37 Encalyptaceae Encalypta Hedw
ciliata Hedw var ciliata procera Bruch
38 Pottiaceae Astomum Hampe
muhlenbergianum (Sw) Grout
Weissia Hedw controversa Hedw var controversa
Gymnostomum Nees Homsch et Stunn aeruginosum Sm recurvirostrum Hedw var recurvirostrum
Trichostomum Bruch spirale Prout
19
40 Ephemeraceae Ephemerum Hampe
cohaerens (Hedw) Hampe crassineium (Schwaegr) Hampe val crassinervium spinulosum Bruch et Schimp ex Schimp
41 Funariaceae AphflllolThegma SuD
serratum (1 Hook et Wils ex Drumm) Sullo
Physcomitrium (Brid) Fuernr hookeri Hampe immersum Sullo pyriforme (Hedw) Hampe
Funaria Hedw americana Undb hygrometrica Hedw vaT hygrometrica
42 Splachnaceae Splachnum Hedw
ampullaceum Hedw
43 Schistostegaceae Schistostega Mohr
pennata (Hedw) Web el Mohr
44 Bryaceae Pohlia Hedw
annolina (Hedw) Lindb var annotina camea (Tum) Lindb cruda (Hedw) Undb elongata Hedw iliformis (Dicks) Andr lescurlana (Sull) Grout nutans (Hedw) Lindb obrusifolia (Brid) L Koch proligera (Undb ex Umpr) Undb ex Amell wahlenbergii (Web et Mohr) Andr
Leptobryum (B S G) Wits pyriforme (Hedw) Wils
Bryum Hedw algovicum Sendtnex C Muell argenteum Hedw caespiticium Hedw capillare Hedw var capillare creberrlmum Tayt lonchocaulon C Muell pallens (Brid) Sw ex Roehl pseudotrlquetrum (Hedw) Gaertn Meyer et Scherb varpudotriquetrum
20
pseudotriquetrum (Hedw) GaertnMeyer et Scherb var bimum (Schreb) Ulj stenotrichum C Muell var stenotrichum tortiolium Funck ex Brid uliginosum (Brid) B S G violaceum Crundw et Nyh weigelii Spreng
Rhodobryum (Schimp) Hampe roseum (Hedw) Limpr
45 Mniaceae Mnium Hedw
ciliare (C Muell) Schimp cinclidioides Hueb cuspidarum Hedw drummondii Bruch et Schimp longirostrum Brid lycopodioides Schwaegr marginatum (With) Brid ex P Beauv mediumB~S G var medium onhorrhynchum Brid pseudopunctatum Bruch et Schimp punctalUm Hedw var punctatum punctatum Hedw var elatum Schirnp rugicum Laur spinufosum B S G stellare Hedw
46 Aulacomruaceae Au1acomnium Schwaegr
androgynum (Hedw) Schwaegr heterostichum (Hedw) B S G paJustre (Hedw) Schwaegr var paJustre
47 Meesiaceae Amblyodon B S G
deillbotus (Hedw) B S G
Meesia Hedw triquetra (Richt) Angstr uJiginosa Hedw
48 Bartramiaceae Plagiopus Brid
oederiana (Sw) Umpr
Bartramia Hedw pomiformis Hedw var pomiformis
Philonotis Brid fontana (Hedw) Brid varfontana longseta (Michx) Britt varlongseta marchica (Hedw) Brid
21
~
~
49 Timmiaceae Timmia Hedw
megapolitana Hedw var megapolitana
50 Orthotrichaceae Zygodon Hook et Tayl
viridissimus (Dicks) Brid var rupestris lindb ex C J Hartm
Amphidium Schimp lapponicum (Hedw) Schimp mougeotii (B S G) Schimp
Orthotrichum Hedw anomalum Hedw obtuaifolium Brid ohioense Sullo et Lesq ex Aust pumilumSw sordidum Sullo et Lesq ex Aust speciosum Neesax Sturm var eegam (Schwaegr ex Hook et Grev) W~rnst stellDtwn Brid strrmgulatum P Beauv
Uleta Mohr ex Web cOfIICtata (p Beallv) Hamrn crispa (Hedw) Brid
Drummondill Hook ex Drumm prorepens (Hedw) Brltt
51 Fontinalaceae Fantinalis Hedw
antipyretica Hedw var antipyrettca antipyretica Hedw forma rabusta (Card) Card dalecarlica Schimp ex B S G var dalecarlica flqccidtJ Ren et Card hypnoides C J Hartrn var hypnoides hypnoides C J Hartm var durillei (Schimp) HU$ll
missourlcaCard novae-angliae Sullo var novae-angliae splulgrUlolill (C Moen) Wijket M~S
DichelY1l1ll Myr capilloceum (With) Myr falcatum (Bedw) Myr HIllescens B S G
52 Climaciaceae aimecium Web et Mohr ex Mohr
americllnum Brid dedroides (Hedw) Web et Mohr
22
S3 Hedwigiaceae Hedwigia P Beauv
ciliata (Hedw) P Beauv
54 Cryphaeaceae Forsstroemia lindh
ohioense (Sull) lindh
55 Leucodontaceae Leucodon Schwaegr
brachypus Blid var brachypus brachypus Brid var aniJrewsitlnus Crum et Anderson julaceus (Hedw) SuU
56 Neckeraceae Neckera Hedw
pennata Hedw var pennata
1fomalia (Brid) B S G trichomanoides (Hedw) B S G
57 Theliaceae Thelia Sull
asprellll Sullo hirtella (Hedw) Sullo lescurii SuU
Myurella B SG julacea (Schwaegr) B S G
sibrica (C MueU) Reim var sibrica
58 Fabroniaceae Fabronio Raddi
ciliaris (Brid) Bnd
Anacamptodon Brid splachnoides (Groet ex Brid) Brid
Schwetschkeopsis Broth fabronia (Schwaegr) Broth
59 Leskeaceae Lindbergia Kindb
brachyptera (Mitt) Kindb
Leskea Hedw gracilescens Hedw obscura Hedw poiycarpa Hedw
Leskeella (limpr) Loeske nervosa (Brid) Loeske
23
Pterlgy1lllndrum Hedw filiforme Hedw
60 Thuidiaceae Haplohymenfum Dozy et MoIk
trlste (Ces ex De Not) Kiodb
Anomodon Hook et Tayl attenuatus (Hedw) Hueb minor (Hedw) Fuemr rostratus (Hedw) Schimp viticulosus (Hedw) Hook et Tayl
Haplocladium (C Mucn) C MueU microphyllum (Hedw) Broth virginillnum (Brld) Broth ~bull
Thuidium B S G abietinum (Hedw) B S G delicatulum (Hedw) B S G vardelicatulum delicatulum (Hedw) B S G var radicans (Kindb) Crum Steere et Anderson minutulum (Hedw) B S G pygmaeum B S G recognftum (Hedw) Lindb scitum (p Beauv) Aust
Helodium (Sull) Warnst blandowli (Web et Mohr) Warnst var blandowli blandowli (Web et Mohr) Wamst var helodioides (Ren et Card ex Roell) Crum
Steere et Anderson paludomm (Sull) AuS
61 Amblystegiaceae Cratoneuron (Sull) Spruce
commulatum (Hedw) Roth var commutfltum commutatum (Hedw) Roth var fakatum (Brid) Moenk ilicinum (Hedw) Spruce var filcinum
Campylium (Sun) Mitt chrysophyllum (Brid) 1 Lange var chrysophyllum hispidulum (Brid) Mitt var hllJidulum polygamum (B S G)C lens varpolygtlmUm stellatum (Hedw) C1ens vat stelltum
Leptodictyum (Schimp) Warnst laxirete (Card et Ther) Broth riparium (Hedw) Wamat trichopodium (Schultz) Warns l1acillans (Sull) Broth
Hygroamblystegium Loelke jlul1iatile (Hedw) oeske var luViatile
24
fluviatile (Hedw) Loeske var oWltum Grout noterophilum (Sun et Lesq ex Sull) Warnst tenox (Hedw) Jenn var tenax tenox (Hedw) Jenn var spinilolium Schimp) Jenn
Amblystegium BSG juratzkanum Schimp serpens (Hedw) BSG varlum (Hedw) Undb var varium
Platydictya Berk confervoides (Brid) Crum jungermanniodes (Brid) Crum minutissimum (Sull et Lesq ex Sull) Crum subtile (Hedw) Crum
Drepanocladus (C Muell) Roth aduncus (Hedw) Warnst var aduncus aduncus (Hedw) Warnst var capillilolius (Wamst)Riebm aduncus (Hedw) Warnst var kneiffii (BSG) Moenk aduncus (Hedw) Warnst var polycorpus (Bland ex Voit) Roth exannulatus (BSG) Warnst fluitans (Hedw) Warnst var fluitans revolvens (Sw) Wamst var revolvens sendtneri (Schimp) Warnst var sendtnerl uncinatus (Hedw) Warnst var uncinatus uncinatus ffiedw) Warnst var piwnosus (Schimp) Wamst vernicoms (Lindb ex C Hartm) Warnst
Hygrohypnum Lindb eugyrium (BSG) loeske luridum (Hedw Jenn varluridum ochraecum (Tum ex Wils) Loeske var ochraceunt
Calliergon (Sull) Kindb cordilolium Hedw Kindb glganteum (Schimp) Kindb var giganteum richardsonii (Mitt) Kindb ex Wamst stramineum (Brid) Kindb trilarium (Web et Mohr) Kindb
Calliergonella Loeske cuspidata (Hedw) Loeske
Calliergidium (Ren) Grout bakeri (Ren) Grout
Scorpidium (Schimp) Urnpr scorpiodides (Hedw) limpr
62 Brachytbeciaceae Tomenthypnum loeske
nitens (Hedw) Loeske var mtens
25
Homalotheciella (Card) Broth subcapillata (Hedw) Broth
Brachythecium BSG acuminatum (Hedw) Awt acutum (Mitt) Sullo calcareum IGndb campestTe (C MuelL) BSG cunum (Lindb) Umpr digastrum C Muell et Kindb ex Macoun et Kindb oxycladon (Brid) Jaeg et Sauerb var oxycladon plumosum (Hedw) BSG populeum (Hedw) BSG var populeum reflexum (Starke ex Web et Mohr) DsG var relexum reiexum (Starke ex Web et Mohr) DSG var pacificum Renet Card ex Roell rlvulare BSG var rivulare tabulum (Hedw) BSC var tabuum salebrosum (Web et Mohr) BSG var salebrosum velutinum (Hedw) BSG Vat velutinum
Bryhniil Kaur graminicolor (Brid) Grout var graminicolor novae-angliae (Sull et Lesq ex sun) Grout
Rhynchostegium BSG senulatum (Hedw) Ja4lg et Sauerb
Rhynchostegiella (BSG) Limpr compacta (C MueU) Loeske
EU1hynchium 8SG hians (Hedw) Sande Lac pulchellum (Hedw) Jenn var pulchelium riparloides (Hedw) Rich
Stokeriella (Kindb) Robins pmelonga (Hedw) Robins var stokesii (Tum) Crum
63 Entodontaceae Entodon C Muell
brevisetus (Hook et Wils ex Wils) Lindb cladorrhizans (Hedw) C Muell seductrlx (Hedw) C MueU var wluctrix
Pleurozium Mitt schreberl (8rid) Mitt var schreberi
64 Plagiotheciaceae Pltlglothecium BSG
CflVifolium (Brid) Iwats var CtWifolium denticulatum (Hedw) 8SG laetum BSG ltztebricola BSG
26
6S SernatophyUaceae Brotherella Loeske ex Fleisch
recumins (Michx) Fleisch tenuirostris (Bruch et Schimp ex Sull) Fleisch
Sematophyllum Mitt demissum (Wils) Mitt
66 Hypnaceae Pl(itygyrium BSG
repens (Brid) BSG
PylaisieUa Kindb intricata (Hedw) Grout polyantha (Hedw) Grout selwynii (Kindb) Crum Steere et Anderson
Homomallium (Schimp) Loeske adnatum (Hedw) Broth
Hypnum Hedw cupressiforme Hedw var cupressiforme cupressifonne Hedw varfiliforme Brid curviolium Hedw fertile Sendtn imponens Hedw lindbergii Mitt palescens (Hedw) P-Beauv pratense Koch ex Brid
Calicladium Crum haldanianum (Grev) Crum
sopterygiopsis Iwats muelleriana (Schimp) Iwats
Isopterygium Mitt e1egans (Brid) lindb pulchellum (Hedw) Jaeg et Sauerb tenerum (Sw) Mitt
Taxiphyllum Fleisch depllQtum (Bruch et Schimp ex sun) Fleisch taxirameum (MitL) Fleisch
Herzogiella Broth seligeri (Brid) Iwats striatella (Brid) Iwats ruracea (Lindh) Iwats
hilium De Not crista-castrenna (Hedw) De Not
27
67 Rhytidiaceae Rhytidium (Sull) Kindb
rugosum (Hedw) Kindb
Rhytidfodelphus (Lindb ex Iimpr) Warnst triquetrus (Hedw) Warnst var triquetrus
68 Hylocomiaceae Hylocomium BSG
pyrenaicum (Spruce) lindb splendens (Hedw) BSG varsplendens
69 Buxbaumiaceae Buxbaumia Hedw
aphyllll Hedw
Diphyscium Mohr follosum (Hedw) Mohr
70 Tetraphidaceae Tetraphi3
pellucida Hedw
71 Polytrichaceae Atrichum PmiddotBeauv
angustatum (Brid) BSG cri8pum (James) Sullo oerstedianum (C Muell) Mitt teneUum (Roehl) BSG undulatum (Hedw) P middotBeauv var unduiatum
Pogonatum P-Beauv alpinum (Hedw) Roehl var alpinum umigerum (Hedw) P-Beauv
Poiytrichum Hedw commune Hedw var commune ormosum Hedw juniperinum Hedw longisetum Brid ohioense Ren et Card piliferum Hedw var pilierum strictum Brid
28
MAPS OF BRYOPHYTE TAXA
REPORTED FOR WISCONSIN
HORNWORTS AND LIVERWORTS
29
bull bull WSHeV~N S vE II
0 ~
e~RRON ~USK
(J~ (j -
CHIPPE A DUNN
0 o0 poundAU CLAIRE
~ TREMshyPEAshyLEAU
MICE
ONEIDA
Q LINCOLN
TAYLOR
Q t-~----- MOIRA THON
CLARK
o
10
LA FAVETTE
Hornwor ts amp Liverworts
- None - 1 - 25 taxa
- 26 - 50 tRXshy
- 51 - 75 tax)
- 7c - 100 taxa
FONDOU j LAC ~
DODGE
i O JEfFERSON
WALWORTH
30
Anastrophylltlfl helleranum (Nees) Schust Anastrophyllum michauxil (We~ Buch ~
46 88 136 141 Evans 38 46 141
Anastrophyllum mlnut~ (Schreb) Schuster Aneura plnguls (L) Durn
89 21 384687 88 89 90 (138) (145)
Anthoceros macounil MA Howe ~oceros punctatus L
21 (45) (144) 34
31
Barbilophozia attenuata (Mart) Loeske Barbilophozia barbata (Schmid ~ Schre~
24 141 Loeske 28 39 46 89 90 141
Bazzania trilobata (1) S Gray
1 26 39 46 (57) (59) 79 88 89
90 119 141
BlePbarostoma trichoffiyllum (1) Dumort
1 8 26 39 46 89 90 137 141
Blasia pusilla 1
1 39 46 90
Calypogeja fissa (1) Raddi
90
32
$129geja integristipula Steph C~Ypound2geja muelleriana (Schifm) K Mull
88 89 89
CallpoundO~eja neesiana (Mass amp Carest) K Mull
39 46 89 140 (141)
~pogeja sJlapound2icola (H Arnell tl J bull
Perss) Warnst et Loeske (119) 141
CeIalozia biscuspidata (L) Dum
1 37 I 46 90 t 137 141
CalYpogej~ trichomanis (L) Corda
26 39 (119) 137
33
CeEhalozia catenulata (Hub) Lindb Cephalozia connivens (Dicks) Lindb
39 46 79 141 39 4679 88 (89) 90 119 141
Cephalozia lacinulata Jack ~ Spruce
141 Cephalozia lunulifolia (Dum~ Dum I 28 39 46 79 88 89 90 141
Cepoundhalozia macouni (Aust) Aust Cephalozia ~leniceps (Aust) Lindb 27~ 3746 (66)7987 (139)141 39 46 79 89 141
34
Ceffialoziella 9ivaricata (Sm) Schiffn Cephaloziella ~hista (Jack) Schiffn
46 90 46 (119) 141
Cephaloziella hampeana (Nees) Schiffn
46
Cephaloziella rubella (Nees) Warnst
46 39
Cephaloziella rubella var sullivantii
(Aust) K Mull 46 39
Cephaloziella rubella var bifida (Lindb) Douin
35
46
Chiloscyphus pallescens (Ehrh ~ Hof~ Dum Chiloscyphus pallescens var fragilis
39J 46 88 89J 90 (Roth) K Milll 46
Chiloscyphus E9lyanthos (L) Corda
1 24 39 46
Chiloscyphus polyanthos vax rivularis
(Schrad) Nees 39 46
poundpoundnocepha1um conicum (1) Lindb
12 28 39 46 (49) 87 88 8990 120 121 148
Cololejeunea biddlecomiae (Aust) Evans
39 46 62 87 88 89 90
36
Dipl0]hyllum apicu1atum (Evans) Steph Fossombronia foveo1ata Lindb
(35)41 46 90 140 1 t 38 (39) t 46
Frullania bolanderi Aust
13 46 62 88 90 139 140
Frullanla ari ttoniae Evans
39 46 69 (71) 87 88 89 90
Frullania eboracensis Gott
1 9 28 4 39 46 (57) (59) 69 70 (71) 87 88 89 90 92 93 97 148
Frullania inflata Gott
46 87 88 89 (139)
Frullania oakesiana Aust Frullania plana Sullo
(35) 42 46 87 (139) (140) 92
Frullania ri paria Hampe ~ Lehm
46 69 71 87 89
Frullania ~elwynianna Pears
46 89
Geocalyx SEaveolens (Schrad) Nees
394688 8990 (119) 140
Frullania tamarisci (L) Dum subsp sPagrayana
(Mont) Hatt 28 34 l~6 t (57) 88 89 90
38
Mar pan thus seutatUB ( Web amp Mohr) Spruce Jamesoniella autumnalis (DC) Steph
(35) 39 46 90 1 7 28 39 46 56 (57) 87 8889 90 (119) 137 141
Jungermannia confertissima Nees
137 141
Jungermannia crenuliform1s Aust
79 137 141
Jungermannia ~il1ima Sm
46 90 141 Jungermannia exsert1folia subsp corifo~ (Dum~
Vana (35) 46 79 141
39
Jungermannia hyalina Lyell Jungerrr6nnia 1eiantha Grolle
40 46 79 136 141 39 46 79 88 90 1)7 141
Jungermannia IXlaxis Lindb Jungermannia pumila With
46 67 79 (139) (140) 141 )9 46 90 1)6 141
Jungermannia sphaerocarpa Hook Kurzia setacea (Web) Grolle
4679136140IIH 136
Lejeunea cavifo1ia (Ehrh) Lindb Lepidozia reptans (1) Dum
1 7 27 39 46 88 89 90 137 14010 I ( 27) 46 I 69 (71) 88
lophocolea heterophylla (Schrad) Dum Lophocolea minor Nees
1 )4 39 46 (57) 87 88 89 27 39 46 88 89 90 (119) 90 (93)94 119 136
Lophozia al~stris (Schleich ~ Web) Evans Lophozia ascendens (Warnst) Schust
46 90 137 141 136 140 141
41
loIilozia badensls (Gatt ~ Rabenh)
46 141
lophozia blcrenata (Schmid sectA Hoffm)
Dum 46 141
lophozia collaris (Nees) Dum
39 46 141
lophozla excisa (Dicks) Dum
27 4679 141
lophozia gillmani (Aust) Schust lophozia gIlttulata (Undo tl H Arnell)
36 39 40 46 136 (140) 141 Evans 39 46 136 141
42
141
Lophozia heterocolEDs (Thed) MA Howe IoPhozia ipcisa (Schard) Dum
27 39 46 89 141 8 27 31 46 79 90 140 141
LoQhozia longidens Macoun Lophozia ventricosa
1 46 79 90 141
Lophozia ventricosa var silvicola
(Buch) Jones 141 LoEhozla ventricosa vax longif1ora (Nees)
Macoun 39 46 79 141
43
~~ fragrans (Balbis) Frye et Clark Mannia pi 10sa (Hornem) Frye et Clark
46 (139) (140) 9 46 79 116 139
Mannia triandra (Scop) Gro 11e
46 (140)
Marchantia pound91ymorpha L
1 13 28 3946 (49) 78 87 88 89 90 (Ill) 121 122 147
Mo~rckia hibernica (HookJ Gott
46 89
Metzgeria conjugata Lindb
44
46
Mylia anomala (Hook) S Gray Notothylas orbicularis (Schwein) Sullo
39 46 141 (35)45 46
Nowellia curvifolia (Dicks) Mitt
17 27 3846 88 89 90 141
Odontoschisma denudatuJI1 (Nees ~ Mart)
Dum 46 81 137 141
Pallavicinia lyellii (Hook~ Carruth
90
Odontoschisma macounii (Aust) Under
45 46 (66) (139) 140 141
45
=
Pellia epiphllla (L) CordaPellia endiviifolia (Dicks~ Dum
27 39 46 90 13724 39 46
Pellia neesiana Limp bull Phaeoceros
39 46 89 27 39 45 46 97
Porella pinnata L
13 27 I 46 90 Plagiochila asplenioides (L) Dum
27 I 39 1+6 88 89 I 90 137
46
~e11a platYEhylla (L) Pfeiff Porella platyphylloi~ (Schwein) Lindb
17 27 39 46 69 (71)79 87 88 89 9 34 46 (57) 59 87 88 89 92 90 92 93 77
Preissia quadrata (Seop) Nees
1 36 3946 (71) 88 89 90 121 123
137
Ptllidium ciliare (L) Hampe
1 27 39 46 132 141
Dum
(22) 28 3946 (57) 87 88 89 90 92
Ptilidium ~lcherrium (G Web) Hampe
(9) 13 394656 (57)70 87 88 89 90 93 (119) 136 141
47
Radula obconica Sullo Reboulia hemlsphaerlca (L) Raddi
39 46 88 89 (43) 46 (59) 70 90 t Cf 124 125 (139)
Riccardia ~atifrons Lindb
1 27 39 46 89
Riccardia multifi~ (L) S Gray
39 46 88 145
Riccia beyrichiana Hampe ~ Lehm
46 Riccardia palmata (Hedw) Carruth
39 46 90
48
Riccia bifurca Hoffm Riccia cavernosa Hoffm
48 13739 46 79 137
Riccia fluitans L
242731454679
Riccla frostli Aust
48 79
Riccia sullicantii Aust
79 (137) Riccia sorocarpa Bisch 46 (137)
49
Ricciocarpus natans (1) Corda Sca~ia apiculata Spruce
13 27 46 79 89 110 39 46 (139) 141
ScaJltll1ia carinthica Jack ~ Lindb
137 t 141
Scapania ~ (Mart) Durn
39467990137141
Scapania digenil Schiffn
90
ScaEsectflia cuspiduligera (Nees) K Mull
46 (139) (140) (141)
50
Scapania glaucoeerha1a (Tayl) Aust Seapania gyrrmostomophilia Kaal
39 46 (139) 141 41 44 46 64 (139) (140) 141
ScaP3poundlia irrigua Gott ~ almiddot
7 39 46 79 137 (140L 141
Seapania nemorosa (L) Dum
1 39 46 89 90 137
Scapmi~ ~~ta Smiddotlch
137 141 51
~caPnia poundludicola LDeske amp K Mull Scapania saxicola Schust
46 (66) 141 136 141
Sc~~ scandica (H Arnell ~ Buch) Macv
139 140
Scapania subalpina (Nees) Durn
39 40 46 79 137 141
Scapania undulata (L) Dum
1 42 46 79 141
Scapania umbrosa (Schrad) Dum
137 140 141
52
Trichocolea tomentella (Ehrh) Dum Triomarla exsecta (Schmid) Schiffn
27 (35) 39 46 47 88 137 141 8 28 39 46 90 140 141
Tritomaria exsectiformis (Breidl) Loe~ke Tritomaria Sluinguedentata (Huds) Buch
39 -t6 89 90 136 140 141 7 39 46 141
Trltomaria scitula (Tayl) Joerg
46 136 (139) 141
53
INTRODUCfION
This atlas of the bryophyte flora of Wisconsin is an attempt to show the distribution of all bryoshyphyte species known to occur in the state by county occurrenoes based on published reports in the literashyture It is not an annot~ted check list (only a feW problematic taxa were examined) but is a literature search intended as a foundation for a complete flora or for otherbryological research
The literature of btYo1ogy is large and spread over numerous journals reports monographs and manuals It would be almost impossible to search all sources for every reference attributing bryophyte species to Wisconsin and to particular localities in the state Wehav checked many inlportant references and paid special attention to works dealing with Wisconsiit botany and the bryologyof the upper Midwest Several bibliographies and indices were checked thoroughly including Green an~ Curtis (1955) the Bry ologist indices (Crum 1959 and Miller 1976) andt)le Wisconsin Academy of Sciences Arts and Letters indices
Only those published repots which app~red to)e substanti2ted by herbarium specimens and which gave- ~fficient location dat to P~CetheW(On in amiddot particula(county were-used in the initial compilation Species described in manuals in generalized way implying their occurrence in Wisconsin (Without menmiddot tioning Wisconsin specifically or citing a particular location in the state) were excluded eg Gnmmia raui (Wynne 1943) and Pohlia annotinll var decipiens (Crum 1976) Ecological papers which were either too general in taxonomic determinations (eg Sphagnum sp Polytrichum sp) or in location data were also excluded Some other taxa attributed to Wisconsin which have been excluded are
Cylindrothectum compr~ (Cheney 189tmiddotDane Co) identified later by Grout as Entodon seducshytrtx
Dicranell4 humilis (Holzjne~ 1925middotBuffalo Co) probablyD vllJia since Dhumilis is not found in North Arnerl~
Didymodon triarium (Foote 1 966-SW Visconsin) identified by Fred Hennann as Barbulaacuta Funoria microstoma (anonymous 1975middotDane Co) excluded because it is outside of its geographic
range and the packe~ QaMot be located Phil(motis calcarea (Cheney and Evans 1944-Dane Co) excluded in Crum Steere and Anderson
(1973) Plagiobryum demissum and P zierl (Cheney and Evans 1944-various counties) are both Arcticshy
alpine species not likely to be found in isconsin Two of Cheneys collections of Pl1giobyrum have been referred to Bryum
Pogonatum alpinum var septentrionale (Che~ey and Evans 1944-Chippewa Co) was misidentified (Ireland 1960) and is P umigerum
Rhizomnium fAPpoJachIanum and R punctalu1fl ssp chlorophyDosum (Koponen 1973middotseveral counshy ties) excluded North American bryologiflts have not recognized these taxa in America
Sphagnum tenellum (Cheney and Evans 1944-Waukesha Co)an Atlantic species not occurring in Wisconsin (Cruro 1975)
The nomenclature and systematic arrangement of hornwort 00 hepatic species in the atlas follow Stotler and Crandall-5totler (1977) and of moss species fonow Crum Steere and Anderson (1973) except in such instances as the JreptlIfockIdus fonns whiGh follow Wijk et aL (19S9middot1973) Species reported in the literature under names which are now listed as synonyms on either of these lists were converted to the accepted name A dot is placed in each county of Wisconsin from which the taxon has been reported The numbers after the name of the taxon correspond to the numbered references in the bibliography which giye ~act location data OEat least location data -s~fficient to place it in-particular counties in the state The numbers in parentheses indicate references which attribute the taxon to Wisconsin without providing exact loltation data
Fossil and subfossil reports are summarized Slfparately in our atlas only six species (Aulacomnium turgidum lJ1achythecium turgidum Ditrichum Jlelficaule Hypnum bambergti Scorpidium 6fJisities and Tortel14 1rIgi1i$) are known exclUsively from the fossil record ~
6
GEOGRAPHIC CLIMATIC AND BOTANICAL FEATURES OF WISCONSIN
Wisconsin is a north central state bounded on the east by Lake Michigan on the north by Michigan and Lake Superior on the south by Illinois and on the west by the Mississippi and St Croix rivers It is located midway between the Prime Meridian (0deg) of Longitude and 1800 Longitude and midway between the equator and the north pole thus placing it in the center of the northern half of the western hemisphere
The topography of the state is relatively flat a few hills exceed 1900 feet with Thns Hill at 1952 feet forming the highest point in Price County Lake Michigan represents the lowest point at 580 feet The northern and eastern two-thirds of the state were heavily glaciated during the last stage of the Pleistocene and now contain many of the best examples of continental glaciation topOgraphy including considerable ground moraine large terminal recessional and intetlobate moraines kames eskers drumlins and small depressions or kettle-holes Most of the nearly 9000 lakes of 20 acres or more lie within this part of the state The southwestern third of the state is the Driftless Area It includes most of the large flat generally sandy bed of Glacial Lake Wisconsin plus the highly dissected ridge and valley section with numerous cliffs and rock-outcroppings (Figure 1)
Much of the bedrock in the state lies below glacial till but exposures include igneous and metamiddot morphic rocks primarily granitic especially in the north as well as sedimentary layers primarily sandshystones and dolomitic limestones especially along Lake Michigan and in the DriftlesS Area
The climate of Wisconsin is classified as humid-temperate with cold winters The annual precipimiddot tation averages approximately 30 inches which is sufficiently even in its seasonal distribution to support forest vegetation throughout most of the state Nevertheless according to Curtis (I 959) the pre-settlement vegetation of the state (J 830-1850) included about 27 prairie and oak-savanna plus about 64 forests
The state is presently divided into seventy-two counties Marathon County is the largest (1586 sq mi) the smallest counties include Pepin (235 sq mi) Ozaukee (236 sq mi) andMilw8ukee (237 sq mi) We included Menominee County though its future status as a county is in doubt since it regained its Indian reservation status and rights in 1973 This county was established April 30 1961 from the Menomishynee Indian reservation which consisted of parts of Shawano and Oconto counties Wisconsin ranks 26th in the United States in area with 54464 square miles and 16th in population with 4651000
More ecological research is needed on the bryophyte flora of Wisconsin especially to relate it to the vegetation and to such other environmental features as soils climate and glaCial deposits Plant communishyties in Wisconsin are many and diverse these are summarized on Figure 2 from studies conducted by Curtis (1959) The state is divided into northern and southern floristic provinces by a tension zone deshyscribed by Curtis (1959) This tension zone is an eeotonal region vatying from 10middot31 mt wide where the ranges of many vascular plants overlap or terminatej it runs in an S-curve northeast from Milwaukee to Hudson in St Croix County (Figure 2) The principal plant communities of the northern province apshyproaching their southern boundaries within the zone include the northern conifermiddothardwood forests of sugar maple white pine hemlock and yellow birch and the Sphagnum bogs with tamarack and black spruce Southern communities at or near their northern limits within this zone include prairies oak and jack pine savannas and hardwood forests of maples oaks ash ehn and cottonwood
We hope to encourage research on cryptogamic ecology with our checklist We know that some species are northern while others are southern (eg cimacium) fits the tension zone concept Freckmann (I977) while Bryoxiphium is only found in the Driftless Area of the state Steere (1937) and Love and Love (1953) The tension zone theory may prove to be acceptable for bryophytes We speculate that certain bryophytes can indicate particular plant communities since many bryophytes are restricted to preshycise environmental requirements Many papers or theses have dealt with cryptogamic bark and rock ecology in Wisconsin (Culberson 1955 Foote 1962 Hale 1952 1955 and Cole 1975)
We hope that this atlas will stimulate collecting in counties poorly sampled so that our knowledge of the bryophytes can be increased It should aid anyone interested in floras for preservation environmental impact statements and distributional studies
7
A numerical summary of the bryophyte flora of the state based on our literature search is presented in Table 1 (below) This includes 517 taxa 70 families and 176 genera This numerical summary is comshypared with the published floras of Michigan mosses (Crum 1976) and Minnesota hepatics (Schuster 1953) Thus we surmise that further collecting could lead to the same approximate numbers for Wisconsin bryoshyphytes
Table 1 Comparison of Recorded Bryophytes of Wisconsin with Michigan and Minnesota and Summary of Wisconsin Taxa
Wisconsin Michigan (Crum 1976)
Minnesota (Schuster 1955)
Hepaticsamp Hornworts
Taxa Families Genera
137 28 47
174 32 52
Mosses
Taxa Families Genera
380 42
129
503 42
149
8
-
~
~ tIt
Figure 1 The Driftless Area and Tension Zone The counties are 01$0 shown on map
ca location of tension zone as deecfibed bv Curtis (1959)~ The Orittless Area
9
Agn 2 Major Plant Communitiee of Wsconsln aaed on surwvor recorda bull co 18laquo) After Curtismiddot 1959
tit beol Forest e Phillie
Northernmiddot hardwood forest reg Pine 5cMInna
copy Southernmiddot hordwood forest CD Oak Savanna
10
SYSTEMATIC ARRANGEMENT OF WISCONSIN BRYOPHYTES
ANTHOCEROTAE (Hornworts)
1 Anthocerotaceae Anthoceros 1 emnd Prosk
macounii M A Howe punctatum 1
Phaeoceros Prosk laevis (1) Prosk subsp laevis
Notothylas Sul1 orbicularis (Schwein) Sullo
HEPATICAE (liverworts)
2 PseudolepicoJeaceae Blephllrostoma (Dum emend Undb) Dum
trlchophyllum (L) Dum subsp trlchophylluni
3 Ptilidiaceae Ptilidium Nees
ci1iare (L) Hampe pulcherrimum (G Web) Hampe
4 Trichocoleaceae Trichocolea Dum nom cons
tomentella (Ehrh) Dum
5 Lepidoziaceae Kunia von Martens
setacea (Web) GroUe
Lepidozia (Dum) Dum reptans (1) Dum
Bazzania S Gray trilobata (1) S Gray var triZobata
6 Calypogejaceae 011ypogeja Raddi
flSsa (L) Raddi subsp issa integristipula Steph muelleriana (Schiffn) K Mull subsp muelleriana neesiana (Mass et Carest) K Mull sphagnicola (H Arnell et J Perss) Warnst et loeske trichomanis (1) Corda
11
7 Cephaloziaceae Cepluzlozla (Dum emend Schiffn) Dum
biscwpidata (L) Dum subsp biclupidIIta catenulllttl (Hub) Lindb coMivens (Dicks) Lindb var connivens lacinultzta Jack ampX Spruce lunulifolilz (Dum) Dum macounii (Aust) Aust pleniceps (Aust) Lindh var pleniceps
Noweilia Mitt cunifolilz (Dicks) Mitt
8 Adelanthaceae Odontoschimuz (Dwn) Dum
denudatum (Nees ampX Mart) Dum Vir denudatum mtlCount (Aust) Underw
c
9 Cephaloziellaceae CephtlJoziellll (Spruce) Schiffn nom C()II$
divarlctzt4 (Sm) sclurfn val divarlcat(l elachhta (bdc) Schiffn 1uImpetuw (Nees) Schirfn IUbellll (Nees) Warnst vat IUbeIl4rube (Nees) Warnst var biid4 (Lindb) Dquinrube (Nees) WarDSt Var sulliwmtii (Aust) K MuD
10 Geo~c~ larptuttllus Nees
scutatus (Web et Mohr) Spruce
Geocalyx Nees grGVeolens (Schrad) Nees
11 Lophocoleaceae Lophocolea (Dum) Dum
heteropllyll4 (Schrad) Own minorNees
Chiloscypbus Corda om cons pallescens (Ehrh ampX Hoffm) Dum varfragllis (Roth) K Mun polyantitos (L) Corda var poiyanthos pol)llaquo1lthos (L) Corda var rlvulllris (Schrld) Nees
12 PlagiochiJaceae Plagio(hila Dum nom cons
tlBpienioides (L) Dum var asplenioidu
13 Jungermanniaceae Btriilophoril Loeske
attenutlta (Mart) Loeske bIubtlta (Schmid ampX Scltreb) Loeske
12
Lophom (Dum) Dum alpestris (Schleich ~ Web) Evans ascendens (Warnst) Schust badenais (Gott ex Rabenh) Schiffn var bademii bierenata (Schmid ex H)ffm) Dum eollam (Nees) Dum exci8a (Dicks) Dum var excisa gllimanii (Aust) Schust varglllmanii guttulata (Lindb et H Arnell) Evans heterocolpos (Thed) M A Howe var heterocolpos ineistl (Schrad) Dum var tnelsa longidens (Lindb) Macoun subsplongidens ventricosa (Dicks) Dum var ventricosa ventricolfl (Dicks) Dum varlontifIotrl (Nees) Macoun ventricosa (Dicks) Dum var silvlcola (Bueh) Jones
Tritomarill Schiffn ex Loeske exseeta (Schrad) Loeske exsecttformi$ (Breidl) Loeske subsp exsectiormfs quinquedentata (Huds) Buch var quinquedentata scitula (rayl) Joerg
Anastrophyllum (Spruce) Schiffn hellemnum (Noes) Schust michtzuxii (Web) Buch ex Evans minutum (Schrob) Schust var minutum
Mylill S Gray nom cons anomala (Hook) S Gray
Jamesoniella (Spruce) Carring autumnalis (DC) Steph var autumnalis
Jungemumnia L emend Dum eonfertissimtt Nees crenuliformis Aust exsertijolill Steph subsp cordifolia (Dum) Vana graciUima Sm hyalina Lyell leillnthll Grolle polaris Lindb pumila With sphaerocarpa Hook
14 Scapaniaceae Diplophyllum (Dum) Dum nom cons
apiculatum (Evans) Steph var apiculatum
Scapanill (Dum) Dum nom cons apkuata Spruce carinthillca Jack ex Lindb curta (Mart) Dum var curta
13
cuspiduligera (Nees) K Mull var cuspiduiigera degenii Schlffn ex K Mull var degenii glllucocephaJa (Tayt) Aust gymnostomophila Kaal irrigwJ (Nees) Gott et al subsp irrigua lingulata Buch varlingulata mucronata Buch subsp mucronata nemorosa (l) Dum paludicola Toeske et K Mun var paludicola paludosa (K Mull) K Mull saxicola Schust scandiea (H Arnell et Buch) Macv var scandiea 8Ubalp~ (Nees) Dum umbroSll (SchIad) Dum undulata (L) Dum var undulata
15 Radulaceae Raduia Dum nom COllS
compianata (L) Dum obconica Sull
16 Porellaceae Parella L
pinTlllta L patyphylla (L) Pfeiff platyphylloidt(l (Schwein) lindb
17 Jubulaceae FruUanill Raddi
bolanderi Aust brittoniae Evans eboracemis Gott inlllta Gott oakesiana Aust plana Sull rlparia Hampe ex Lehm selwyniana Pears taTnl1iJci (L) Dum subsp asagmyana (Mont) Hatt
18 lejeuneaceae Calolejeunea (Spruce) Schlffn
biddlecomiae (Aust) Evans
Lejeunea Ub nom cons cavifolia (Ehrh) Lindb emend Buch
19 Codoniaceae Fossombronia Raddi
foveolata Lindb
20 Pa11aviciniaceae Paklvieinlil S Gray nom cons
lyeiii (Hook) Carruth
14
Moerckia Gott hibemica (Hook) Gott
21 PeWaceae Pellfa Raddi nom cons propos
endivlifolfa (Dicks) Dwn epiphylllz (L) Corda neeBiana (GottlUmpr
22 Metzgeriaceae Metzgerla Raddi
conjugata Undb
23 Blasiacae B L
pulilm L
24 Aneurareae AneumDum
pinguis (L) Dum
Riccfl1dia S Gray nom cons latifrons Undb multiftda (L) S Gray var multifida palmata (Hedw) Carruth
25 Aytoniaceae Reboulill Raddi nom conI
hemllphaerica (L) Radeli
ManniaOpiz fragrrms (Balbis) Frye el ClarK pilosa (Hornem) Frye et Clark trltmdra (Scop) GroDe
26 Conocephalaceae Conocephalum Wigg nom cons
conicum (L) Lindb
27 Marchantiaceae Preiuia Corda
quadrata (Scop) Nees
Marchantia L polymorpha L
28 Ricciaceae Riccwcarpus Corda
natans (L) Corda
Riccia 1 beyrichiana Hampe ex Lehm bifurca Horfm
15
i
CtlJPemostI Hoffnt fuitu L rostii Aust BOIOCtI1fHI Bi~ sulltvtmtii Aust
MUSCI (Mosses)
29 Sphagnaceae SpIutgnum L
capillaceum (Weiss) Schrank var Cllpil1tlceum caplllllceum (Weiss) Vir tenellum (Schimp) Andr centrille C Jens ex Amen et C Jens compactummiddot OC ex Lam et Dc ClUpidatum Ehrh ex Hoffin var aupidatum jimbrltltum Wits ex J Hookmiddot fulCUm (Scbimp) KUngrr glrgenBOhIlll Russ imbrlctltum Hornsch ex Russ muelllmitil4m BOd 1IIIIU (RUII) C Jens przIutre L papIIIoIum Lindb pulchrum (Lindb ex braithw) Wamst recumun P-Beauv var recumun recum P-Beauv var tenue Klinggr riptlrlum Ansstr ruPQWItWamst squtl1fOlUm Crome subftltens Russ et Wamst ex Wamst IUbIeCfUldum Nees ex Sturm var subsecundum subsecundum Nees ex Sturm varrutem (Nees Hormch et sUUm) Hueb teres (Sdlimp) Anptr e C Hartm
WtlI7IItorjii Russ wuljfanum Gira
30 Andreaeaceae AndNrrerI Hedw
rupeitrlr Oedw var pest
31 Fissident~e Fluid_ lledw
trdiantoid Hedw byoidei Bedw var bryolda cristaM Wils ex Mitt exiguus sun VIr exigUw onttmUi (B-Pyl) Steud jtuberi Lesq et James obtusifolhu Wits var obtwifoliul ollPlUlldQldes Heel ~litSuU lubbtulkril Bed tfDdfoliul lledw llridubu (S~ ) Wahlenb vallirlIU1u
16
32 Ditrichaceae Pleuridium Brid
subulatum (Hedw) Rabenh
Ditrichum Hampe heteromallum (Hedw) Britt linellre (Sw) lindb
~ pallidum (Hedw) Hampe pusiUum (Hedw) Hampe
Saelania Lindb gIlucescens (Hedw) Bomanss et Broth
Ceratodon Brid purpureus (He(lw) Brid var prupurew
Distichium BSG capillaceum (Hedw) BSG inclinatum (Hedw) BSG
33 Bryoxipbiaceae Bryoxlphium Mitt
norvegicum (Brid) Mitt
34 Seligeriaceae Seligeria BSG
cumpylopoda Kindb ex Macoun et Kindb pusillll (Hedw) BSG recurvata (Hedw) BSG
Bindia BSG acuta (Hedw) BSG
35 Dicranaceae BlUehia Schwegr
lexUOSll (Sw ~ Schwaegr) C MueU sullivantii Aust
Trematodon Michx ambiguus (Hedw) Hornsch
Dicranellll (C Muell) Schimp cerviculata (Hedw) Schimp val cillPicuiata
heteromalla (Hedw) Schimp ~ rufescens (With) Schimp
schreberiana (Hedw) Schhnp var schreberiana schreberitJna (Hedw) Schimp var robusta Schimp ex Braithw subulata (Hedw) Schimp varia (Hedw) Schimp
Dicranodontium BSG denudatum (Brid) Britt ex Williams
Panzeucobryum (Lindh) Loeske Iongifolium (Hedw) oelke varlongifolium
17
Tortella (Undb) Limpr fragilis (Drumm) Umpr humilis (Hedw) JeM torlUoSfl (Hedw) Umpr
Hyophila Brid involuta (Hoole) Jaeg et Sauerb
Didymodon Hedw rigidulus Hedw
BryoerythrophyUum Chen recurvirostnon (Hedw) Chen
Btubula Hedw acula (Brid) Brid var acuta convoiuta Hedw fallox Hedw unguicultltll Hedw
Acaulon C MueD muticum (Uedw) C Muell var rufeseens (Iaeg) Crum
PJl~lm Hedw CUSPidtltum Hedw
De8mlltodon Brid cernuu (Hueb) BSG oblUsifoliw (Schwaegr) Schimp
T01IUla Hedw mueronifolia Schwaegr rurall (Hedw) Gaertn Meyer et Scherb var ruralis
39 Grimmiaceae Grlmmill Hedw
alpicola Hedw var alpicola alpicola Hedw var dupretii (Ther) Crum alpicola Hedw var rlvularl8 (Brid) Wahlenb apocarpa Hedw var apocarpa apocarpa Hedw var ambipa (Sull) Jonea apocarpa Hedw var conjerttl (Funck) Spreng apocarpa Hedw var strleta (Tum) Hook et Tayl CiIIyptrrlta Hook ex Drumm donniana Sm var donn_ hemumti Crum ltlevla (Brid) Brid montana BSG oineyi SuD oVllll8 (Hedw) lindb piUfe P-Beauv pltJgiopodill Hedw pulvinata (Hedw) Sm varbullpulvinata teretinervl Limpr
18
Orothera C Muell leana (Sull) C Muell
Rfwbdoweisia crispata (With) Undb
Cynodontium BSG ex Schimp alpestre (Wahlenb) Milde strumiferum (Hedw) Undb
Dichodontium Schimp pellucidum (Hedw) Schimp
Oncophorus (Slid) Brid vuens (Hedw) Brid wahlenbergii (Brid)
Dicranum Hedw bonjeanii DeNot ex Usa drummondii C Muell lagellare Hedw varj1ogellare fulvum Hook fuscescens Turn var fuscescens montanum Hedw muehlenbeckii BSG var muehlenbeckii polysetum Sw sabuletorum Ren et Card scoparium Hedw undulatum Brid viride (Sull et Lesq ex Sull) lindb
36 Leucobryaceae Leucobryum Hampe
glaucum (Hedw) Angstr ex Fr
37 Encalyptaceae Encalypta Hedw
ciliata Hedw var ciliata procera Bruch
38 Pottiaceae Astomum Hampe
muhlenbergianum (Sw) Grout
Weissia Hedw controversa Hedw var controversa
Gymnostomum Nees Homsch et Stunn aeruginosum Sm recurvirostrum Hedw var recurvirostrum
Trichostomum Bruch spirale Prout
19
40 Ephemeraceae Ephemerum Hampe
cohaerens (Hedw) Hampe crassineium (Schwaegr) Hampe val crassinervium spinulosum Bruch et Schimp ex Schimp
41 Funariaceae AphflllolThegma SuD
serratum (1 Hook et Wils ex Drumm) Sullo
Physcomitrium (Brid) Fuernr hookeri Hampe immersum Sullo pyriforme (Hedw) Hampe
Funaria Hedw americana Undb hygrometrica Hedw vaT hygrometrica
42 Splachnaceae Splachnum Hedw
ampullaceum Hedw
43 Schistostegaceae Schistostega Mohr
pennata (Hedw) Web el Mohr
44 Bryaceae Pohlia Hedw
annolina (Hedw) Lindb var annotina camea (Tum) Lindb cruda (Hedw) Undb elongata Hedw iliformis (Dicks) Andr lescurlana (Sull) Grout nutans (Hedw) Lindb obrusifolia (Brid) L Koch proligera (Undb ex Umpr) Undb ex Amell wahlenbergii (Web et Mohr) Andr
Leptobryum (B S G) Wits pyriforme (Hedw) Wils
Bryum Hedw algovicum Sendtnex C Muell argenteum Hedw caespiticium Hedw capillare Hedw var capillare creberrlmum Tayt lonchocaulon C Muell pallens (Brid) Sw ex Roehl pseudotrlquetrum (Hedw) Gaertn Meyer et Scherb varpudotriquetrum
20
pseudotriquetrum (Hedw) GaertnMeyer et Scherb var bimum (Schreb) Ulj stenotrichum C Muell var stenotrichum tortiolium Funck ex Brid uliginosum (Brid) B S G violaceum Crundw et Nyh weigelii Spreng
Rhodobryum (Schimp) Hampe roseum (Hedw) Limpr
45 Mniaceae Mnium Hedw
ciliare (C Muell) Schimp cinclidioides Hueb cuspidarum Hedw drummondii Bruch et Schimp longirostrum Brid lycopodioides Schwaegr marginatum (With) Brid ex P Beauv mediumB~S G var medium onhorrhynchum Brid pseudopunctatum Bruch et Schimp punctalUm Hedw var punctatum punctatum Hedw var elatum Schirnp rugicum Laur spinufosum B S G stellare Hedw
46 Aulacomruaceae Au1acomnium Schwaegr
androgynum (Hedw) Schwaegr heterostichum (Hedw) B S G paJustre (Hedw) Schwaegr var paJustre
47 Meesiaceae Amblyodon B S G
deillbotus (Hedw) B S G
Meesia Hedw triquetra (Richt) Angstr uJiginosa Hedw
48 Bartramiaceae Plagiopus Brid
oederiana (Sw) Umpr
Bartramia Hedw pomiformis Hedw var pomiformis
Philonotis Brid fontana (Hedw) Brid varfontana longseta (Michx) Britt varlongseta marchica (Hedw) Brid
21
~
~
49 Timmiaceae Timmia Hedw
megapolitana Hedw var megapolitana
50 Orthotrichaceae Zygodon Hook et Tayl
viridissimus (Dicks) Brid var rupestris lindb ex C J Hartm
Amphidium Schimp lapponicum (Hedw) Schimp mougeotii (B S G) Schimp
Orthotrichum Hedw anomalum Hedw obtuaifolium Brid ohioense Sullo et Lesq ex Aust pumilumSw sordidum Sullo et Lesq ex Aust speciosum Neesax Sturm var eegam (Schwaegr ex Hook et Grev) W~rnst stellDtwn Brid strrmgulatum P Beauv
Uleta Mohr ex Web cOfIICtata (p Beallv) Hamrn crispa (Hedw) Brid
Drummondill Hook ex Drumm prorepens (Hedw) Brltt
51 Fontinalaceae Fantinalis Hedw
antipyretica Hedw var antipyrettca antipyretica Hedw forma rabusta (Card) Card dalecarlica Schimp ex B S G var dalecarlica flqccidtJ Ren et Card hypnoides C J Hartrn var hypnoides hypnoides C J Hartm var durillei (Schimp) HU$ll
missourlcaCard novae-angliae Sullo var novae-angliae splulgrUlolill (C Moen) Wijket M~S
DichelY1l1ll Myr capilloceum (With) Myr falcatum (Bedw) Myr HIllescens B S G
52 Climaciaceae aimecium Web et Mohr ex Mohr
americllnum Brid dedroides (Hedw) Web et Mohr
22
S3 Hedwigiaceae Hedwigia P Beauv
ciliata (Hedw) P Beauv
54 Cryphaeaceae Forsstroemia lindh
ohioense (Sull) lindh
55 Leucodontaceae Leucodon Schwaegr
brachypus Blid var brachypus brachypus Brid var aniJrewsitlnus Crum et Anderson julaceus (Hedw) SuU
56 Neckeraceae Neckera Hedw
pennata Hedw var pennata
1fomalia (Brid) B S G trichomanoides (Hedw) B S G
57 Theliaceae Thelia Sull
asprellll Sullo hirtella (Hedw) Sullo lescurii SuU
Myurella B SG julacea (Schwaegr) B S G
sibrica (C MueU) Reim var sibrica
58 Fabroniaceae Fabronio Raddi
ciliaris (Brid) Bnd
Anacamptodon Brid splachnoides (Groet ex Brid) Brid
Schwetschkeopsis Broth fabronia (Schwaegr) Broth
59 Leskeaceae Lindbergia Kindb
brachyptera (Mitt) Kindb
Leskea Hedw gracilescens Hedw obscura Hedw poiycarpa Hedw
Leskeella (limpr) Loeske nervosa (Brid) Loeske
23
Pterlgy1lllndrum Hedw filiforme Hedw
60 Thuidiaceae Haplohymenfum Dozy et MoIk
trlste (Ces ex De Not) Kiodb
Anomodon Hook et Tayl attenuatus (Hedw) Hueb minor (Hedw) Fuemr rostratus (Hedw) Schimp viticulosus (Hedw) Hook et Tayl
Haplocladium (C Mucn) C MueU microphyllum (Hedw) Broth virginillnum (Brld) Broth ~bull
Thuidium B S G abietinum (Hedw) B S G delicatulum (Hedw) B S G vardelicatulum delicatulum (Hedw) B S G var radicans (Kindb) Crum Steere et Anderson minutulum (Hedw) B S G pygmaeum B S G recognftum (Hedw) Lindb scitum (p Beauv) Aust
Helodium (Sull) Warnst blandowli (Web et Mohr) Warnst var blandowli blandowli (Web et Mohr) Wamst var helodioides (Ren et Card ex Roell) Crum
Steere et Anderson paludomm (Sull) AuS
61 Amblystegiaceae Cratoneuron (Sull) Spruce
commulatum (Hedw) Roth var commutfltum commutatum (Hedw) Roth var fakatum (Brid) Moenk ilicinum (Hedw) Spruce var filcinum
Campylium (Sun) Mitt chrysophyllum (Brid) 1 Lange var chrysophyllum hispidulum (Brid) Mitt var hllJidulum polygamum (B S G)C lens varpolygtlmUm stellatum (Hedw) C1ens vat stelltum
Leptodictyum (Schimp) Warnst laxirete (Card et Ther) Broth riparium (Hedw) Wamat trichopodium (Schultz) Warns l1acillans (Sull) Broth
Hygroamblystegium Loelke jlul1iatile (Hedw) oeske var luViatile
24
fluviatile (Hedw) Loeske var oWltum Grout noterophilum (Sun et Lesq ex Sull) Warnst tenox (Hedw) Jenn var tenax tenox (Hedw) Jenn var spinilolium Schimp) Jenn
Amblystegium BSG juratzkanum Schimp serpens (Hedw) BSG varlum (Hedw) Undb var varium
Platydictya Berk confervoides (Brid) Crum jungermanniodes (Brid) Crum minutissimum (Sull et Lesq ex Sull) Crum subtile (Hedw) Crum
Drepanocladus (C Muell) Roth aduncus (Hedw) Warnst var aduncus aduncus (Hedw) Warnst var capillilolius (Wamst)Riebm aduncus (Hedw) Warnst var kneiffii (BSG) Moenk aduncus (Hedw) Warnst var polycorpus (Bland ex Voit) Roth exannulatus (BSG) Warnst fluitans (Hedw) Warnst var fluitans revolvens (Sw) Wamst var revolvens sendtneri (Schimp) Warnst var sendtnerl uncinatus (Hedw) Warnst var uncinatus uncinatus ffiedw) Warnst var piwnosus (Schimp) Wamst vernicoms (Lindb ex C Hartm) Warnst
Hygrohypnum Lindb eugyrium (BSG) loeske luridum (Hedw Jenn varluridum ochraecum (Tum ex Wils) Loeske var ochraceunt
Calliergon (Sull) Kindb cordilolium Hedw Kindb glganteum (Schimp) Kindb var giganteum richardsonii (Mitt) Kindb ex Wamst stramineum (Brid) Kindb trilarium (Web et Mohr) Kindb
Calliergonella Loeske cuspidata (Hedw) Loeske
Calliergidium (Ren) Grout bakeri (Ren) Grout
Scorpidium (Schimp) Urnpr scorpiodides (Hedw) limpr
62 Brachytbeciaceae Tomenthypnum loeske
nitens (Hedw) Loeske var mtens
25
Homalotheciella (Card) Broth subcapillata (Hedw) Broth
Brachythecium BSG acuminatum (Hedw) Awt acutum (Mitt) Sullo calcareum IGndb campestTe (C MuelL) BSG cunum (Lindb) Umpr digastrum C Muell et Kindb ex Macoun et Kindb oxycladon (Brid) Jaeg et Sauerb var oxycladon plumosum (Hedw) BSG populeum (Hedw) BSG var populeum reflexum (Starke ex Web et Mohr) DsG var relexum reiexum (Starke ex Web et Mohr) DSG var pacificum Renet Card ex Roell rlvulare BSG var rivulare tabulum (Hedw) BSC var tabuum salebrosum (Web et Mohr) BSG var salebrosum velutinum (Hedw) BSG Vat velutinum
Bryhniil Kaur graminicolor (Brid) Grout var graminicolor novae-angliae (Sull et Lesq ex sun) Grout
Rhynchostegium BSG senulatum (Hedw) Ja4lg et Sauerb
Rhynchostegiella (BSG) Limpr compacta (C MueU) Loeske
EU1hynchium 8SG hians (Hedw) Sande Lac pulchellum (Hedw) Jenn var pulchelium riparloides (Hedw) Rich
Stokeriella (Kindb) Robins pmelonga (Hedw) Robins var stokesii (Tum) Crum
63 Entodontaceae Entodon C Muell
brevisetus (Hook et Wils ex Wils) Lindb cladorrhizans (Hedw) C Muell seductrlx (Hedw) C MueU var wluctrix
Pleurozium Mitt schreberl (8rid) Mitt var schreberi
64 Plagiotheciaceae Pltlglothecium BSG
CflVifolium (Brid) Iwats var CtWifolium denticulatum (Hedw) 8SG laetum BSG ltztebricola BSG
26
6S SernatophyUaceae Brotherella Loeske ex Fleisch
recumins (Michx) Fleisch tenuirostris (Bruch et Schimp ex Sull) Fleisch
Sematophyllum Mitt demissum (Wils) Mitt
66 Hypnaceae Pl(itygyrium BSG
repens (Brid) BSG
PylaisieUa Kindb intricata (Hedw) Grout polyantha (Hedw) Grout selwynii (Kindb) Crum Steere et Anderson
Homomallium (Schimp) Loeske adnatum (Hedw) Broth
Hypnum Hedw cupressiforme Hedw var cupressiforme cupressifonne Hedw varfiliforme Brid curviolium Hedw fertile Sendtn imponens Hedw lindbergii Mitt palescens (Hedw) P-Beauv pratense Koch ex Brid
Calicladium Crum haldanianum (Grev) Crum
sopterygiopsis Iwats muelleriana (Schimp) Iwats
Isopterygium Mitt e1egans (Brid) lindb pulchellum (Hedw) Jaeg et Sauerb tenerum (Sw) Mitt
Taxiphyllum Fleisch depllQtum (Bruch et Schimp ex sun) Fleisch taxirameum (MitL) Fleisch
Herzogiella Broth seligeri (Brid) Iwats striatella (Brid) Iwats ruracea (Lindh) Iwats
hilium De Not crista-castrenna (Hedw) De Not
27
67 Rhytidiaceae Rhytidium (Sull) Kindb
rugosum (Hedw) Kindb
Rhytidfodelphus (Lindb ex Iimpr) Warnst triquetrus (Hedw) Warnst var triquetrus
68 Hylocomiaceae Hylocomium BSG
pyrenaicum (Spruce) lindb splendens (Hedw) BSG varsplendens
69 Buxbaumiaceae Buxbaumia Hedw
aphyllll Hedw
Diphyscium Mohr follosum (Hedw) Mohr
70 Tetraphidaceae Tetraphi3
pellucida Hedw
71 Polytrichaceae Atrichum PmiddotBeauv
angustatum (Brid) BSG cri8pum (James) Sullo oerstedianum (C Muell) Mitt teneUum (Roehl) BSG undulatum (Hedw) P middotBeauv var unduiatum
Pogonatum P-Beauv alpinum (Hedw) Roehl var alpinum umigerum (Hedw) P-Beauv
Poiytrichum Hedw commune Hedw var commune ormosum Hedw juniperinum Hedw longisetum Brid ohioense Ren et Card piliferum Hedw var pilierum strictum Brid
28
MAPS OF BRYOPHYTE TAXA
REPORTED FOR WISCONSIN
HORNWORTS AND LIVERWORTS
29
bull bull WSHeV~N S vE II
0 ~
e~RRON ~USK
(J~ (j -
CHIPPE A DUNN
0 o0 poundAU CLAIRE
~ TREMshyPEAshyLEAU
MICE
ONEIDA
Q LINCOLN
TAYLOR
Q t-~----- MOIRA THON
CLARK
o
10
LA FAVETTE
Hornwor ts amp Liverworts
- None - 1 - 25 taxa
- 26 - 50 tRXshy
- 51 - 75 tax)
- 7c - 100 taxa
FONDOU j LAC ~
DODGE
i O JEfFERSON
WALWORTH
30
Anastrophylltlfl helleranum (Nees) Schust Anastrophyllum michauxil (We~ Buch ~
46 88 136 141 Evans 38 46 141
Anastrophyllum mlnut~ (Schreb) Schuster Aneura plnguls (L) Durn
89 21 384687 88 89 90 (138) (145)
Anthoceros macounil MA Howe ~oceros punctatus L
21 (45) (144) 34
31
Barbilophozia attenuata (Mart) Loeske Barbilophozia barbata (Schmid ~ Schre~
24 141 Loeske 28 39 46 89 90 141
Bazzania trilobata (1) S Gray
1 26 39 46 (57) (59) 79 88 89
90 119 141
BlePbarostoma trichoffiyllum (1) Dumort
1 8 26 39 46 89 90 137 141
Blasia pusilla 1
1 39 46 90
Calypogeja fissa (1) Raddi
90
32
$129geja integristipula Steph C~Ypound2geja muelleriana (Schifm) K Mull
88 89 89
CallpoundO~eja neesiana (Mass amp Carest) K Mull
39 46 89 140 (141)
~pogeja sJlapound2icola (H Arnell tl J bull
Perss) Warnst et Loeske (119) 141
CeIalozia biscuspidata (L) Dum
1 37 I 46 90 t 137 141
CalYpogej~ trichomanis (L) Corda
26 39 (119) 137
33
CeEhalozia catenulata (Hub) Lindb Cephalozia connivens (Dicks) Lindb
39 46 79 141 39 4679 88 (89) 90 119 141
Cephalozia lacinulata Jack ~ Spruce
141 Cephalozia lunulifolia (Dum~ Dum I 28 39 46 79 88 89 90 141
Cepoundhalozia macouni (Aust) Aust Cephalozia ~leniceps (Aust) Lindb 27~ 3746 (66)7987 (139)141 39 46 79 89 141
34
Ceffialoziella 9ivaricata (Sm) Schiffn Cephaloziella ~hista (Jack) Schiffn
46 90 46 (119) 141
Cephaloziella hampeana (Nees) Schiffn
46
Cephaloziella rubella (Nees) Warnst
46 39
Cephaloziella rubella var sullivantii
(Aust) K Mull 46 39
Cephaloziella rubella var bifida (Lindb) Douin
35
46
Chiloscyphus pallescens (Ehrh ~ Hof~ Dum Chiloscyphus pallescens var fragilis
39J 46 88 89J 90 (Roth) K Milll 46
Chiloscyphus E9lyanthos (L) Corda
1 24 39 46
Chiloscyphus polyanthos vax rivularis
(Schrad) Nees 39 46
poundpoundnocepha1um conicum (1) Lindb
12 28 39 46 (49) 87 88 8990 120 121 148
Cololejeunea biddlecomiae (Aust) Evans
39 46 62 87 88 89 90
36
Dipl0]hyllum apicu1atum (Evans) Steph Fossombronia foveo1ata Lindb
(35)41 46 90 140 1 t 38 (39) t 46
Frullania bolanderi Aust
13 46 62 88 90 139 140
Frullanla ari ttoniae Evans
39 46 69 (71) 87 88 89 90
Frullania eboracensis Gott
1 9 28 4 39 46 (57) (59) 69 70 (71) 87 88 89 90 92 93 97 148
Frullania inflata Gott
46 87 88 89 (139)
Frullania oakesiana Aust Frullania plana Sullo
(35) 42 46 87 (139) (140) 92
Frullania ri paria Hampe ~ Lehm
46 69 71 87 89
Frullania ~elwynianna Pears
46 89
Geocalyx SEaveolens (Schrad) Nees
394688 8990 (119) 140
Frullania tamarisci (L) Dum subsp sPagrayana
(Mont) Hatt 28 34 l~6 t (57) 88 89 90
38
Mar pan thus seutatUB ( Web amp Mohr) Spruce Jamesoniella autumnalis (DC) Steph
(35) 39 46 90 1 7 28 39 46 56 (57) 87 8889 90 (119) 137 141
Jungermannia confertissima Nees
137 141
Jungermannia crenuliform1s Aust
79 137 141
Jungermannia ~il1ima Sm
46 90 141 Jungermannia exsert1folia subsp corifo~ (Dum~
Vana (35) 46 79 141
39
Jungermannia hyalina Lyell Jungerrr6nnia 1eiantha Grolle
40 46 79 136 141 39 46 79 88 90 1)7 141
Jungermannia IXlaxis Lindb Jungermannia pumila With
46 67 79 (139) (140) 141 )9 46 90 1)6 141
Jungermannia sphaerocarpa Hook Kurzia setacea (Web) Grolle
4679136140IIH 136
Lejeunea cavifo1ia (Ehrh) Lindb Lepidozia reptans (1) Dum
1 7 27 39 46 88 89 90 137 14010 I ( 27) 46 I 69 (71) 88
lophocolea heterophylla (Schrad) Dum Lophocolea minor Nees
1 )4 39 46 (57) 87 88 89 27 39 46 88 89 90 (119) 90 (93)94 119 136
Lophozia al~stris (Schleich ~ Web) Evans Lophozia ascendens (Warnst) Schust
46 90 137 141 136 140 141
41
loIilozia badensls (Gatt ~ Rabenh)
46 141
lophozia blcrenata (Schmid sectA Hoffm)
Dum 46 141
lophozia collaris (Nees) Dum
39 46 141
lophozla excisa (Dicks) Dum
27 4679 141
lophozia gillmani (Aust) Schust lophozia gIlttulata (Undo tl H Arnell)
36 39 40 46 136 (140) 141 Evans 39 46 136 141
42
141
Lophozia heterocolEDs (Thed) MA Howe IoPhozia ipcisa (Schard) Dum
27 39 46 89 141 8 27 31 46 79 90 140 141
LoQhozia longidens Macoun Lophozia ventricosa
1 46 79 90 141
Lophozia ventricosa var silvicola
(Buch) Jones 141 LoEhozla ventricosa vax longif1ora (Nees)
Macoun 39 46 79 141
43
~~ fragrans (Balbis) Frye et Clark Mannia pi 10sa (Hornem) Frye et Clark
46 (139) (140) 9 46 79 116 139
Mannia triandra (Scop) Gro 11e
46 (140)
Marchantia pound91ymorpha L
1 13 28 3946 (49) 78 87 88 89 90 (Ill) 121 122 147
Mo~rckia hibernica (HookJ Gott
46 89
Metzgeria conjugata Lindb
44
46
Mylia anomala (Hook) S Gray Notothylas orbicularis (Schwein) Sullo
39 46 141 (35)45 46
Nowellia curvifolia (Dicks) Mitt
17 27 3846 88 89 90 141
Odontoschisma denudatuJI1 (Nees ~ Mart)
Dum 46 81 137 141
Pallavicinia lyellii (Hook~ Carruth
90
Odontoschisma macounii (Aust) Under
45 46 (66) (139) 140 141
45
=
Pellia epiphllla (L) CordaPellia endiviifolia (Dicks~ Dum
27 39 46 90 13724 39 46
Pellia neesiana Limp bull Phaeoceros
39 46 89 27 39 45 46 97
Porella pinnata L
13 27 I 46 90 Plagiochila asplenioides (L) Dum
27 I 39 1+6 88 89 I 90 137
46
~e11a platYEhylla (L) Pfeiff Porella platyphylloi~ (Schwein) Lindb
17 27 39 46 69 (71)79 87 88 89 9 34 46 (57) 59 87 88 89 92 90 92 93 77
Preissia quadrata (Seop) Nees
1 36 3946 (71) 88 89 90 121 123
137
Ptllidium ciliare (L) Hampe
1 27 39 46 132 141
Dum
(22) 28 3946 (57) 87 88 89 90 92
Ptilidium ~lcherrium (G Web) Hampe
(9) 13 394656 (57)70 87 88 89 90 93 (119) 136 141
47
Radula obconica Sullo Reboulia hemlsphaerlca (L) Raddi
39 46 88 89 (43) 46 (59) 70 90 t Cf 124 125 (139)
Riccardia ~atifrons Lindb
1 27 39 46 89
Riccardia multifi~ (L) S Gray
39 46 88 145
Riccia beyrichiana Hampe ~ Lehm
46 Riccardia palmata (Hedw) Carruth
39 46 90
48
Riccia bifurca Hoffm Riccia cavernosa Hoffm
48 13739 46 79 137
Riccia fluitans L
242731454679
Riccla frostli Aust
48 79
Riccia sullicantii Aust
79 (137) Riccia sorocarpa Bisch 46 (137)
49
Ricciocarpus natans (1) Corda Sca~ia apiculata Spruce
13 27 46 79 89 110 39 46 (139) 141
ScaJltll1ia carinthica Jack ~ Lindb
137 t 141
Scapania ~ (Mart) Durn
39467990137141
Scapania digenil Schiffn
90
ScaEsectflia cuspiduligera (Nees) K Mull
46 (139) (140) (141)
50
Scapania glaucoeerha1a (Tayl) Aust Seapania gyrrmostomophilia Kaal
39 46 (139) 141 41 44 46 64 (139) (140) 141
ScaP3poundlia irrigua Gott ~ almiddot
7 39 46 79 137 (140L 141
Seapania nemorosa (L) Dum
1 39 46 89 90 137
Scapmi~ ~~ta Smiddotlch
137 141 51
~caPnia poundludicola LDeske amp K Mull Scapania saxicola Schust
46 (66) 141 136 141
Sc~~ scandica (H Arnell ~ Buch) Macv
139 140
Scapania subalpina (Nees) Durn
39 40 46 79 137 141
Scapania undulata (L) Dum
1 42 46 79 141
Scapania umbrosa (Schrad) Dum
137 140 141
52
Trichocolea tomentella (Ehrh) Dum Triomarla exsecta (Schmid) Schiffn
27 (35) 39 46 47 88 137 141 8 28 39 46 90 140 141
Tritomaria exsectiformis (Breidl) Loe~ke Tritomaria Sluinguedentata (Huds) Buch
39 -t6 89 90 136 140 141 7 39 46 141
Trltomaria scitula (Tayl) Joerg
46 136 (139) 141
53
GEOGRAPHIC CLIMATIC AND BOTANICAL FEATURES OF WISCONSIN
Wisconsin is a north central state bounded on the east by Lake Michigan on the north by Michigan and Lake Superior on the south by Illinois and on the west by the Mississippi and St Croix rivers It is located midway between the Prime Meridian (0deg) of Longitude and 1800 Longitude and midway between the equator and the north pole thus placing it in the center of the northern half of the western hemisphere
The topography of the state is relatively flat a few hills exceed 1900 feet with Thns Hill at 1952 feet forming the highest point in Price County Lake Michigan represents the lowest point at 580 feet The northern and eastern two-thirds of the state were heavily glaciated during the last stage of the Pleistocene and now contain many of the best examples of continental glaciation topOgraphy including considerable ground moraine large terminal recessional and intetlobate moraines kames eskers drumlins and small depressions or kettle-holes Most of the nearly 9000 lakes of 20 acres or more lie within this part of the state The southwestern third of the state is the Driftless Area It includes most of the large flat generally sandy bed of Glacial Lake Wisconsin plus the highly dissected ridge and valley section with numerous cliffs and rock-outcroppings (Figure 1)
Much of the bedrock in the state lies below glacial till but exposures include igneous and metamiddot morphic rocks primarily granitic especially in the north as well as sedimentary layers primarily sandshystones and dolomitic limestones especially along Lake Michigan and in the DriftlesS Area
The climate of Wisconsin is classified as humid-temperate with cold winters The annual precipimiddot tation averages approximately 30 inches which is sufficiently even in its seasonal distribution to support forest vegetation throughout most of the state Nevertheless according to Curtis (I 959) the pre-settlement vegetation of the state (J 830-1850) included about 27 prairie and oak-savanna plus about 64 forests
The state is presently divided into seventy-two counties Marathon County is the largest (1586 sq mi) the smallest counties include Pepin (235 sq mi) Ozaukee (236 sq mi) andMilw8ukee (237 sq mi) We included Menominee County though its future status as a county is in doubt since it regained its Indian reservation status and rights in 1973 This county was established April 30 1961 from the Menomishynee Indian reservation which consisted of parts of Shawano and Oconto counties Wisconsin ranks 26th in the United States in area with 54464 square miles and 16th in population with 4651000
More ecological research is needed on the bryophyte flora of Wisconsin especially to relate it to the vegetation and to such other environmental features as soils climate and glaCial deposits Plant communishyties in Wisconsin are many and diverse these are summarized on Figure 2 from studies conducted by Curtis (1959) The state is divided into northern and southern floristic provinces by a tension zone deshyscribed by Curtis (1959) This tension zone is an eeotonal region vatying from 10middot31 mt wide where the ranges of many vascular plants overlap or terminatej it runs in an S-curve northeast from Milwaukee to Hudson in St Croix County (Figure 2) The principal plant communities of the northern province apshyproaching their southern boundaries within the zone include the northern conifermiddothardwood forests of sugar maple white pine hemlock and yellow birch and the Sphagnum bogs with tamarack and black spruce Southern communities at or near their northern limits within this zone include prairies oak and jack pine savannas and hardwood forests of maples oaks ash ehn and cottonwood
We hope to encourage research on cryptogamic ecology with our checklist We know that some species are northern while others are southern (eg cimacium) fits the tension zone concept Freckmann (I977) while Bryoxiphium is only found in the Driftless Area of the state Steere (1937) and Love and Love (1953) The tension zone theory may prove to be acceptable for bryophytes We speculate that certain bryophytes can indicate particular plant communities since many bryophytes are restricted to preshycise environmental requirements Many papers or theses have dealt with cryptogamic bark and rock ecology in Wisconsin (Culberson 1955 Foote 1962 Hale 1952 1955 and Cole 1975)
We hope that this atlas will stimulate collecting in counties poorly sampled so that our knowledge of the bryophytes can be increased It should aid anyone interested in floras for preservation environmental impact statements and distributional studies
7
A numerical summary of the bryophyte flora of the state based on our literature search is presented in Table 1 (below) This includes 517 taxa 70 families and 176 genera This numerical summary is comshypared with the published floras of Michigan mosses (Crum 1976) and Minnesota hepatics (Schuster 1953) Thus we surmise that further collecting could lead to the same approximate numbers for Wisconsin bryoshyphytes
Table 1 Comparison of Recorded Bryophytes of Wisconsin with Michigan and Minnesota and Summary of Wisconsin Taxa
Wisconsin Michigan (Crum 1976)
Minnesota (Schuster 1955)
Hepaticsamp Hornworts
Taxa Families Genera
137 28 47
174 32 52
Mosses
Taxa Families Genera
380 42
129
503 42
149
8
-
~
~ tIt
Figure 1 The Driftless Area and Tension Zone The counties are 01$0 shown on map
ca location of tension zone as deecfibed bv Curtis (1959)~ The Orittless Area
9
Agn 2 Major Plant Communitiee of Wsconsln aaed on surwvor recorda bull co 18laquo) After Curtismiddot 1959
tit beol Forest e Phillie
Northernmiddot hardwood forest reg Pine 5cMInna
copy Southernmiddot hordwood forest CD Oak Savanna
10
SYSTEMATIC ARRANGEMENT OF WISCONSIN BRYOPHYTES
ANTHOCEROTAE (Hornworts)
1 Anthocerotaceae Anthoceros 1 emnd Prosk
macounii M A Howe punctatum 1
Phaeoceros Prosk laevis (1) Prosk subsp laevis
Notothylas Sul1 orbicularis (Schwein) Sullo
HEPATICAE (liverworts)
2 PseudolepicoJeaceae Blephllrostoma (Dum emend Undb) Dum
trlchophyllum (L) Dum subsp trlchophylluni
3 Ptilidiaceae Ptilidium Nees
ci1iare (L) Hampe pulcherrimum (G Web) Hampe
4 Trichocoleaceae Trichocolea Dum nom cons
tomentella (Ehrh) Dum
5 Lepidoziaceae Kunia von Martens
setacea (Web) GroUe
Lepidozia (Dum) Dum reptans (1) Dum
Bazzania S Gray trilobata (1) S Gray var triZobata
6 Calypogejaceae 011ypogeja Raddi
flSsa (L) Raddi subsp issa integristipula Steph muelleriana (Schiffn) K Mull subsp muelleriana neesiana (Mass et Carest) K Mull sphagnicola (H Arnell et J Perss) Warnst et loeske trichomanis (1) Corda
11
7 Cephaloziaceae Cepluzlozla (Dum emend Schiffn) Dum
biscwpidata (L) Dum subsp biclupidIIta catenulllttl (Hub) Lindb coMivens (Dicks) Lindb var connivens lacinultzta Jack ampX Spruce lunulifolilz (Dum) Dum macounii (Aust) Aust pleniceps (Aust) Lindh var pleniceps
Noweilia Mitt cunifolilz (Dicks) Mitt
8 Adelanthaceae Odontoschimuz (Dwn) Dum
denudatum (Nees ampX Mart) Dum Vir denudatum mtlCount (Aust) Underw
c
9 Cephaloziellaceae CephtlJoziellll (Spruce) Schiffn nom C()II$
divarlctzt4 (Sm) sclurfn val divarlcat(l elachhta (bdc) Schiffn 1uImpetuw (Nees) Schirfn IUbellll (Nees) Warnst vat IUbeIl4rube (Nees) Warnst var biid4 (Lindb) Dquinrube (Nees) WarDSt Var sulliwmtii (Aust) K MuD
10 Geo~c~ larptuttllus Nees
scutatus (Web et Mohr) Spruce
Geocalyx Nees grGVeolens (Schrad) Nees
11 Lophocoleaceae Lophocolea (Dum) Dum
heteropllyll4 (Schrad) Own minorNees
Chiloscypbus Corda om cons pallescens (Ehrh ampX Hoffm) Dum varfragllis (Roth) K Mun polyantitos (L) Corda var poiyanthos pol)llaquo1lthos (L) Corda var rlvulllris (Schrld) Nees
12 PlagiochiJaceae Plagio(hila Dum nom cons
tlBpienioides (L) Dum var asplenioidu
13 Jungermanniaceae Btriilophoril Loeske
attenutlta (Mart) Loeske bIubtlta (Schmid ampX Scltreb) Loeske
12
Lophom (Dum) Dum alpestris (Schleich ~ Web) Evans ascendens (Warnst) Schust badenais (Gott ex Rabenh) Schiffn var bademii bierenata (Schmid ex H)ffm) Dum eollam (Nees) Dum exci8a (Dicks) Dum var excisa gllimanii (Aust) Schust varglllmanii guttulata (Lindb et H Arnell) Evans heterocolpos (Thed) M A Howe var heterocolpos ineistl (Schrad) Dum var tnelsa longidens (Lindb) Macoun subsplongidens ventricosa (Dicks) Dum var ventricosa ventricolfl (Dicks) Dum varlontifIotrl (Nees) Macoun ventricosa (Dicks) Dum var silvlcola (Bueh) Jones
Tritomarill Schiffn ex Loeske exseeta (Schrad) Loeske exsecttformi$ (Breidl) Loeske subsp exsectiormfs quinquedentata (Huds) Buch var quinquedentata scitula (rayl) Joerg
Anastrophyllum (Spruce) Schiffn hellemnum (Noes) Schust michtzuxii (Web) Buch ex Evans minutum (Schrob) Schust var minutum
Mylill S Gray nom cons anomala (Hook) S Gray
Jamesoniella (Spruce) Carring autumnalis (DC) Steph var autumnalis
Jungemumnia L emend Dum eonfertissimtt Nees crenuliformis Aust exsertijolill Steph subsp cordifolia (Dum) Vana graciUima Sm hyalina Lyell leillnthll Grolle polaris Lindb pumila With sphaerocarpa Hook
14 Scapaniaceae Diplophyllum (Dum) Dum nom cons
apiculatum (Evans) Steph var apiculatum
Scapanill (Dum) Dum nom cons apkuata Spruce carinthillca Jack ex Lindb curta (Mart) Dum var curta
13
cuspiduligera (Nees) K Mull var cuspiduiigera degenii Schlffn ex K Mull var degenii glllucocephaJa (Tayt) Aust gymnostomophila Kaal irrigwJ (Nees) Gott et al subsp irrigua lingulata Buch varlingulata mucronata Buch subsp mucronata nemorosa (l) Dum paludicola Toeske et K Mun var paludicola paludosa (K Mull) K Mull saxicola Schust scandiea (H Arnell et Buch) Macv var scandiea 8Ubalp~ (Nees) Dum umbroSll (SchIad) Dum undulata (L) Dum var undulata
15 Radulaceae Raduia Dum nom COllS
compianata (L) Dum obconica Sull
16 Porellaceae Parella L
pinTlllta L patyphylla (L) Pfeiff platyphylloidt(l (Schwein) lindb
17 Jubulaceae FruUanill Raddi
bolanderi Aust brittoniae Evans eboracemis Gott inlllta Gott oakesiana Aust plana Sull rlparia Hampe ex Lehm selwyniana Pears taTnl1iJci (L) Dum subsp asagmyana (Mont) Hatt
18 lejeuneaceae Calolejeunea (Spruce) Schlffn
biddlecomiae (Aust) Evans
Lejeunea Ub nom cons cavifolia (Ehrh) Lindb emend Buch
19 Codoniaceae Fossombronia Raddi
foveolata Lindb
20 Pa11aviciniaceae Paklvieinlil S Gray nom cons
lyeiii (Hook) Carruth
14
Moerckia Gott hibemica (Hook) Gott
21 PeWaceae Pellfa Raddi nom cons propos
endivlifolfa (Dicks) Dwn epiphylllz (L) Corda neeBiana (GottlUmpr
22 Metzgeriaceae Metzgerla Raddi
conjugata Undb
23 Blasiacae B L
pulilm L
24 Aneurareae AneumDum
pinguis (L) Dum
Riccfl1dia S Gray nom cons latifrons Undb multiftda (L) S Gray var multifida palmata (Hedw) Carruth
25 Aytoniaceae Reboulill Raddi nom conI
hemllphaerica (L) Radeli
ManniaOpiz fragrrms (Balbis) Frye el ClarK pilosa (Hornem) Frye et Clark trltmdra (Scop) GroDe
26 Conocephalaceae Conocephalum Wigg nom cons
conicum (L) Lindb
27 Marchantiaceae Preiuia Corda
quadrata (Scop) Nees
Marchantia L polymorpha L
28 Ricciaceae Riccwcarpus Corda
natans (L) Corda
Riccia 1 beyrichiana Hampe ex Lehm bifurca Horfm
15
i
CtlJPemostI Hoffnt fuitu L rostii Aust BOIOCtI1fHI Bi~ sulltvtmtii Aust
MUSCI (Mosses)
29 Sphagnaceae SpIutgnum L
capillaceum (Weiss) Schrank var Cllpil1tlceum caplllllceum (Weiss) Vir tenellum (Schimp) Andr centrille C Jens ex Amen et C Jens compactummiddot OC ex Lam et Dc ClUpidatum Ehrh ex Hoffin var aupidatum jimbrltltum Wits ex J Hookmiddot fulCUm (Scbimp) KUngrr glrgenBOhIlll Russ imbrlctltum Hornsch ex Russ muelllmitil4m BOd 1IIIIU (RUII) C Jens przIutre L papIIIoIum Lindb pulchrum (Lindb ex braithw) Wamst recumun P-Beauv var recumun recum P-Beauv var tenue Klinggr riptlrlum Ansstr ruPQWItWamst squtl1fOlUm Crome subftltens Russ et Wamst ex Wamst IUbIeCfUldum Nees ex Sturm var subsecundum subsecundum Nees ex Sturm varrutem (Nees Hormch et sUUm) Hueb teres (Sdlimp) Anptr e C Hartm
WtlI7IItorjii Russ wuljfanum Gira
30 Andreaeaceae AndNrrerI Hedw
rupeitrlr Oedw var pest
31 Fissident~e Fluid_ lledw
trdiantoid Hedw byoidei Bedw var bryolda cristaM Wils ex Mitt exiguus sun VIr exigUw onttmUi (B-Pyl) Steud jtuberi Lesq et James obtusifolhu Wits var obtwifoliul ollPlUlldQldes Heel ~litSuU lubbtulkril Bed tfDdfoliul lledw llridubu (S~ ) Wahlenb vallirlIU1u
16
32 Ditrichaceae Pleuridium Brid
subulatum (Hedw) Rabenh
Ditrichum Hampe heteromallum (Hedw) Britt linellre (Sw) lindb
~ pallidum (Hedw) Hampe pusiUum (Hedw) Hampe
Saelania Lindb gIlucescens (Hedw) Bomanss et Broth
Ceratodon Brid purpureus (He(lw) Brid var prupurew
Distichium BSG capillaceum (Hedw) BSG inclinatum (Hedw) BSG
33 Bryoxipbiaceae Bryoxlphium Mitt
norvegicum (Brid) Mitt
34 Seligeriaceae Seligeria BSG
cumpylopoda Kindb ex Macoun et Kindb pusillll (Hedw) BSG recurvata (Hedw) BSG
Bindia BSG acuta (Hedw) BSG
35 Dicranaceae BlUehia Schwegr
lexUOSll (Sw ~ Schwaegr) C MueU sullivantii Aust
Trematodon Michx ambiguus (Hedw) Hornsch
Dicranellll (C Muell) Schimp cerviculata (Hedw) Schimp val cillPicuiata
heteromalla (Hedw) Schimp ~ rufescens (With) Schimp
schreberiana (Hedw) Schhnp var schreberiana schreberitJna (Hedw) Schimp var robusta Schimp ex Braithw subulata (Hedw) Schimp varia (Hedw) Schimp
Dicranodontium BSG denudatum (Brid) Britt ex Williams
Panzeucobryum (Lindh) Loeske Iongifolium (Hedw) oelke varlongifolium
17
Tortella (Undb) Limpr fragilis (Drumm) Umpr humilis (Hedw) JeM torlUoSfl (Hedw) Umpr
Hyophila Brid involuta (Hoole) Jaeg et Sauerb
Didymodon Hedw rigidulus Hedw
BryoerythrophyUum Chen recurvirostnon (Hedw) Chen
Btubula Hedw acula (Brid) Brid var acuta convoiuta Hedw fallox Hedw unguicultltll Hedw
Acaulon C MueD muticum (Uedw) C Muell var rufeseens (Iaeg) Crum
PJl~lm Hedw CUSPidtltum Hedw
De8mlltodon Brid cernuu (Hueb) BSG oblUsifoliw (Schwaegr) Schimp
T01IUla Hedw mueronifolia Schwaegr rurall (Hedw) Gaertn Meyer et Scherb var ruralis
39 Grimmiaceae Grlmmill Hedw
alpicola Hedw var alpicola alpicola Hedw var dupretii (Ther) Crum alpicola Hedw var rlvularl8 (Brid) Wahlenb apocarpa Hedw var apocarpa apocarpa Hedw var ambipa (Sull) Jonea apocarpa Hedw var conjerttl (Funck) Spreng apocarpa Hedw var strleta (Tum) Hook et Tayl CiIIyptrrlta Hook ex Drumm donniana Sm var donn_ hemumti Crum ltlevla (Brid) Brid montana BSG oineyi SuD oVllll8 (Hedw) lindb piUfe P-Beauv pltJgiopodill Hedw pulvinata (Hedw) Sm varbullpulvinata teretinervl Limpr
18
Orothera C Muell leana (Sull) C Muell
Rfwbdoweisia crispata (With) Undb
Cynodontium BSG ex Schimp alpestre (Wahlenb) Milde strumiferum (Hedw) Undb
Dichodontium Schimp pellucidum (Hedw) Schimp
Oncophorus (Slid) Brid vuens (Hedw) Brid wahlenbergii (Brid)
Dicranum Hedw bonjeanii DeNot ex Usa drummondii C Muell lagellare Hedw varj1ogellare fulvum Hook fuscescens Turn var fuscescens montanum Hedw muehlenbeckii BSG var muehlenbeckii polysetum Sw sabuletorum Ren et Card scoparium Hedw undulatum Brid viride (Sull et Lesq ex Sull) lindb
36 Leucobryaceae Leucobryum Hampe
glaucum (Hedw) Angstr ex Fr
37 Encalyptaceae Encalypta Hedw
ciliata Hedw var ciliata procera Bruch
38 Pottiaceae Astomum Hampe
muhlenbergianum (Sw) Grout
Weissia Hedw controversa Hedw var controversa
Gymnostomum Nees Homsch et Stunn aeruginosum Sm recurvirostrum Hedw var recurvirostrum
Trichostomum Bruch spirale Prout
19
40 Ephemeraceae Ephemerum Hampe
cohaerens (Hedw) Hampe crassineium (Schwaegr) Hampe val crassinervium spinulosum Bruch et Schimp ex Schimp
41 Funariaceae AphflllolThegma SuD
serratum (1 Hook et Wils ex Drumm) Sullo
Physcomitrium (Brid) Fuernr hookeri Hampe immersum Sullo pyriforme (Hedw) Hampe
Funaria Hedw americana Undb hygrometrica Hedw vaT hygrometrica
42 Splachnaceae Splachnum Hedw
ampullaceum Hedw
43 Schistostegaceae Schistostega Mohr
pennata (Hedw) Web el Mohr
44 Bryaceae Pohlia Hedw
annolina (Hedw) Lindb var annotina camea (Tum) Lindb cruda (Hedw) Undb elongata Hedw iliformis (Dicks) Andr lescurlana (Sull) Grout nutans (Hedw) Lindb obrusifolia (Brid) L Koch proligera (Undb ex Umpr) Undb ex Amell wahlenbergii (Web et Mohr) Andr
Leptobryum (B S G) Wits pyriforme (Hedw) Wils
Bryum Hedw algovicum Sendtnex C Muell argenteum Hedw caespiticium Hedw capillare Hedw var capillare creberrlmum Tayt lonchocaulon C Muell pallens (Brid) Sw ex Roehl pseudotrlquetrum (Hedw) Gaertn Meyer et Scherb varpudotriquetrum
20
pseudotriquetrum (Hedw) GaertnMeyer et Scherb var bimum (Schreb) Ulj stenotrichum C Muell var stenotrichum tortiolium Funck ex Brid uliginosum (Brid) B S G violaceum Crundw et Nyh weigelii Spreng
Rhodobryum (Schimp) Hampe roseum (Hedw) Limpr
45 Mniaceae Mnium Hedw
ciliare (C Muell) Schimp cinclidioides Hueb cuspidarum Hedw drummondii Bruch et Schimp longirostrum Brid lycopodioides Schwaegr marginatum (With) Brid ex P Beauv mediumB~S G var medium onhorrhynchum Brid pseudopunctatum Bruch et Schimp punctalUm Hedw var punctatum punctatum Hedw var elatum Schirnp rugicum Laur spinufosum B S G stellare Hedw
46 Aulacomruaceae Au1acomnium Schwaegr
androgynum (Hedw) Schwaegr heterostichum (Hedw) B S G paJustre (Hedw) Schwaegr var paJustre
47 Meesiaceae Amblyodon B S G
deillbotus (Hedw) B S G
Meesia Hedw triquetra (Richt) Angstr uJiginosa Hedw
48 Bartramiaceae Plagiopus Brid
oederiana (Sw) Umpr
Bartramia Hedw pomiformis Hedw var pomiformis
Philonotis Brid fontana (Hedw) Brid varfontana longseta (Michx) Britt varlongseta marchica (Hedw) Brid
21
~
~
49 Timmiaceae Timmia Hedw
megapolitana Hedw var megapolitana
50 Orthotrichaceae Zygodon Hook et Tayl
viridissimus (Dicks) Brid var rupestris lindb ex C J Hartm
Amphidium Schimp lapponicum (Hedw) Schimp mougeotii (B S G) Schimp
Orthotrichum Hedw anomalum Hedw obtuaifolium Brid ohioense Sullo et Lesq ex Aust pumilumSw sordidum Sullo et Lesq ex Aust speciosum Neesax Sturm var eegam (Schwaegr ex Hook et Grev) W~rnst stellDtwn Brid strrmgulatum P Beauv
Uleta Mohr ex Web cOfIICtata (p Beallv) Hamrn crispa (Hedw) Brid
Drummondill Hook ex Drumm prorepens (Hedw) Brltt
51 Fontinalaceae Fantinalis Hedw
antipyretica Hedw var antipyrettca antipyretica Hedw forma rabusta (Card) Card dalecarlica Schimp ex B S G var dalecarlica flqccidtJ Ren et Card hypnoides C J Hartrn var hypnoides hypnoides C J Hartm var durillei (Schimp) HU$ll
missourlcaCard novae-angliae Sullo var novae-angliae splulgrUlolill (C Moen) Wijket M~S
DichelY1l1ll Myr capilloceum (With) Myr falcatum (Bedw) Myr HIllescens B S G
52 Climaciaceae aimecium Web et Mohr ex Mohr
americllnum Brid dedroides (Hedw) Web et Mohr
22
S3 Hedwigiaceae Hedwigia P Beauv
ciliata (Hedw) P Beauv
54 Cryphaeaceae Forsstroemia lindh
ohioense (Sull) lindh
55 Leucodontaceae Leucodon Schwaegr
brachypus Blid var brachypus brachypus Brid var aniJrewsitlnus Crum et Anderson julaceus (Hedw) SuU
56 Neckeraceae Neckera Hedw
pennata Hedw var pennata
1fomalia (Brid) B S G trichomanoides (Hedw) B S G
57 Theliaceae Thelia Sull
asprellll Sullo hirtella (Hedw) Sullo lescurii SuU
Myurella B SG julacea (Schwaegr) B S G
sibrica (C MueU) Reim var sibrica
58 Fabroniaceae Fabronio Raddi
ciliaris (Brid) Bnd
Anacamptodon Brid splachnoides (Groet ex Brid) Brid
Schwetschkeopsis Broth fabronia (Schwaegr) Broth
59 Leskeaceae Lindbergia Kindb
brachyptera (Mitt) Kindb
Leskea Hedw gracilescens Hedw obscura Hedw poiycarpa Hedw
Leskeella (limpr) Loeske nervosa (Brid) Loeske
23
Pterlgy1lllndrum Hedw filiforme Hedw
60 Thuidiaceae Haplohymenfum Dozy et MoIk
trlste (Ces ex De Not) Kiodb
Anomodon Hook et Tayl attenuatus (Hedw) Hueb minor (Hedw) Fuemr rostratus (Hedw) Schimp viticulosus (Hedw) Hook et Tayl
Haplocladium (C Mucn) C MueU microphyllum (Hedw) Broth virginillnum (Brld) Broth ~bull
Thuidium B S G abietinum (Hedw) B S G delicatulum (Hedw) B S G vardelicatulum delicatulum (Hedw) B S G var radicans (Kindb) Crum Steere et Anderson minutulum (Hedw) B S G pygmaeum B S G recognftum (Hedw) Lindb scitum (p Beauv) Aust
Helodium (Sull) Warnst blandowli (Web et Mohr) Warnst var blandowli blandowli (Web et Mohr) Wamst var helodioides (Ren et Card ex Roell) Crum
Steere et Anderson paludomm (Sull) AuS
61 Amblystegiaceae Cratoneuron (Sull) Spruce
commulatum (Hedw) Roth var commutfltum commutatum (Hedw) Roth var fakatum (Brid) Moenk ilicinum (Hedw) Spruce var filcinum
Campylium (Sun) Mitt chrysophyllum (Brid) 1 Lange var chrysophyllum hispidulum (Brid) Mitt var hllJidulum polygamum (B S G)C lens varpolygtlmUm stellatum (Hedw) C1ens vat stelltum
Leptodictyum (Schimp) Warnst laxirete (Card et Ther) Broth riparium (Hedw) Wamat trichopodium (Schultz) Warns l1acillans (Sull) Broth
Hygroamblystegium Loelke jlul1iatile (Hedw) oeske var luViatile
24
fluviatile (Hedw) Loeske var oWltum Grout noterophilum (Sun et Lesq ex Sull) Warnst tenox (Hedw) Jenn var tenax tenox (Hedw) Jenn var spinilolium Schimp) Jenn
Amblystegium BSG juratzkanum Schimp serpens (Hedw) BSG varlum (Hedw) Undb var varium
Platydictya Berk confervoides (Brid) Crum jungermanniodes (Brid) Crum minutissimum (Sull et Lesq ex Sull) Crum subtile (Hedw) Crum
Drepanocladus (C Muell) Roth aduncus (Hedw) Warnst var aduncus aduncus (Hedw) Warnst var capillilolius (Wamst)Riebm aduncus (Hedw) Warnst var kneiffii (BSG) Moenk aduncus (Hedw) Warnst var polycorpus (Bland ex Voit) Roth exannulatus (BSG) Warnst fluitans (Hedw) Warnst var fluitans revolvens (Sw) Wamst var revolvens sendtneri (Schimp) Warnst var sendtnerl uncinatus (Hedw) Warnst var uncinatus uncinatus ffiedw) Warnst var piwnosus (Schimp) Wamst vernicoms (Lindb ex C Hartm) Warnst
Hygrohypnum Lindb eugyrium (BSG) loeske luridum (Hedw Jenn varluridum ochraecum (Tum ex Wils) Loeske var ochraceunt
Calliergon (Sull) Kindb cordilolium Hedw Kindb glganteum (Schimp) Kindb var giganteum richardsonii (Mitt) Kindb ex Wamst stramineum (Brid) Kindb trilarium (Web et Mohr) Kindb
Calliergonella Loeske cuspidata (Hedw) Loeske
Calliergidium (Ren) Grout bakeri (Ren) Grout
Scorpidium (Schimp) Urnpr scorpiodides (Hedw) limpr
62 Brachytbeciaceae Tomenthypnum loeske
nitens (Hedw) Loeske var mtens
25
Homalotheciella (Card) Broth subcapillata (Hedw) Broth
Brachythecium BSG acuminatum (Hedw) Awt acutum (Mitt) Sullo calcareum IGndb campestTe (C MuelL) BSG cunum (Lindb) Umpr digastrum C Muell et Kindb ex Macoun et Kindb oxycladon (Brid) Jaeg et Sauerb var oxycladon plumosum (Hedw) BSG populeum (Hedw) BSG var populeum reflexum (Starke ex Web et Mohr) DsG var relexum reiexum (Starke ex Web et Mohr) DSG var pacificum Renet Card ex Roell rlvulare BSG var rivulare tabulum (Hedw) BSC var tabuum salebrosum (Web et Mohr) BSG var salebrosum velutinum (Hedw) BSG Vat velutinum
Bryhniil Kaur graminicolor (Brid) Grout var graminicolor novae-angliae (Sull et Lesq ex sun) Grout
Rhynchostegium BSG senulatum (Hedw) Ja4lg et Sauerb
Rhynchostegiella (BSG) Limpr compacta (C MueU) Loeske
EU1hynchium 8SG hians (Hedw) Sande Lac pulchellum (Hedw) Jenn var pulchelium riparloides (Hedw) Rich
Stokeriella (Kindb) Robins pmelonga (Hedw) Robins var stokesii (Tum) Crum
63 Entodontaceae Entodon C Muell
brevisetus (Hook et Wils ex Wils) Lindb cladorrhizans (Hedw) C Muell seductrlx (Hedw) C MueU var wluctrix
Pleurozium Mitt schreberl (8rid) Mitt var schreberi
64 Plagiotheciaceae Pltlglothecium BSG
CflVifolium (Brid) Iwats var CtWifolium denticulatum (Hedw) 8SG laetum BSG ltztebricola BSG
26
6S SernatophyUaceae Brotherella Loeske ex Fleisch
recumins (Michx) Fleisch tenuirostris (Bruch et Schimp ex Sull) Fleisch
Sematophyllum Mitt demissum (Wils) Mitt
66 Hypnaceae Pl(itygyrium BSG
repens (Brid) BSG
PylaisieUa Kindb intricata (Hedw) Grout polyantha (Hedw) Grout selwynii (Kindb) Crum Steere et Anderson
Homomallium (Schimp) Loeske adnatum (Hedw) Broth
Hypnum Hedw cupressiforme Hedw var cupressiforme cupressifonne Hedw varfiliforme Brid curviolium Hedw fertile Sendtn imponens Hedw lindbergii Mitt palescens (Hedw) P-Beauv pratense Koch ex Brid
Calicladium Crum haldanianum (Grev) Crum
sopterygiopsis Iwats muelleriana (Schimp) Iwats
Isopterygium Mitt e1egans (Brid) lindb pulchellum (Hedw) Jaeg et Sauerb tenerum (Sw) Mitt
Taxiphyllum Fleisch depllQtum (Bruch et Schimp ex sun) Fleisch taxirameum (MitL) Fleisch
Herzogiella Broth seligeri (Brid) Iwats striatella (Brid) Iwats ruracea (Lindh) Iwats
hilium De Not crista-castrenna (Hedw) De Not
27
67 Rhytidiaceae Rhytidium (Sull) Kindb
rugosum (Hedw) Kindb
Rhytidfodelphus (Lindb ex Iimpr) Warnst triquetrus (Hedw) Warnst var triquetrus
68 Hylocomiaceae Hylocomium BSG
pyrenaicum (Spruce) lindb splendens (Hedw) BSG varsplendens
69 Buxbaumiaceae Buxbaumia Hedw
aphyllll Hedw
Diphyscium Mohr follosum (Hedw) Mohr
70 Tetraphidaceae Tetraphi3
pellucida Hedw
71 Polytrichaceae Atrichum PmiddotBeauv
angustatum (Brid) BSG cri8pum (James) Sullo oerstedianum (C Muell) Mitt teneUum (Roehl) BSG undulatum (Hedw) P middotBeauv var unduiatum
Pogonatum P-Beauv alpinum (Hedw) Roehl var alpinum umigerum (Hedw) P-Beauv
Poiytrichum Hedw commune Hedw var commune ormosum Hedw juniperinum Hedw longisetum Brid ohioense Ren et Card piliferum Hedw var pilierum strictum Brid
28
MAPS OF BRYOPHYTE TAXA
REPORTED FOR WISCONSIN
HORNWORTS AND LIVERWORTS
29
bull bull WSHeV~N S vE II
0 ~
e~RRON ~USK
(J~ (j -
CHIPPE A DUNN
0 o0 poundAU CLAIRE
~ TREMshyPEAshyLEAU
MICE
ONEIDA
Q LINCOLN
TAYLOR
Q t-~----- MOIRA THON
CLARK
o
10
LA FAVETTE
Hornwor ts amp Liverworts
- None - 1 - 25 taxa
- 26 - 50 tRXshy
- 51 - 75 tax)
- 7c - 100 taxa
FONDOU j LAC ~
DODGE
i O JEfFERSON
WALWORTH
30
Anastrophylltlfl helleranum (Nees) Schust Anastrophyllum michauxil (We~ Buch ~
46 88 136 141 Evans 38 46 141
Anastrophyllum mlnut~ (Schreb) Schuster Aneura plnguls (L) Durn
89 21 384687 88 89 90 (138) (145)
Anthoceros macounil MA Howe ~oceros punctatus L
21 (45) (144) 34
31
Barbilophozia attenuata (Mart) Loeske Barbilophozia barbata (Schmid ~ Schre~
24 141 Loeske 28 39 46 89 90 141
Bazzania trilobata (1) S Gray
1 26 39 46 (57) (59) 79 88 89
90 119 141
BlePbarostoma trichoffiyllum (1) Dumort
1 8 26 39 46 89 90 137 141
Blasia pusilla 1
1 39 46 90
Calypogeja fissa (1) Raddi
90
32
$129geja integristipula Steph C~Ypound2geja muelleriana (Schifm) K Mull
88 89 89
CallpoundO~eja neesiana (Mass amp Carest) K Mull
39 46 89 140 (141)
~pogeja sJlapound2icola (H Arnell tl J bull
Perss) Warnst et Loeske (119) 141
CeIalozia biscuspidata (L) Dum
1 37 I 46 90 t 137 141
CalYpogej~ trichomanis (L) Corda
26 39 (119) 137
33
CeEhalozia catenulata (Hub) Lindb Cephalozia connivens (Dicks) Lindb
39 46 79 141 39 4679 88 (89) 90 119 141
Cephalozia lacinulata Jack ~ Spruce
141 Cephalozia lunulifolia (Dum~ Dum I 28 39 46 79 88 89 90 141
Cepoundhalozia macouni (Aust) Aust Cephalozia ~leniceps (Aust) Lindb 27~ 3746 (66)7987 (139)141 39 46 79 89 141
34
Ceffialoziella 9ivaricata (Sm) Schiffn Cephaloziella ~hista (Jack) Schiffn
46 90 46 (119) 141
Cephaloziella hampeana (Nees) Schiffn
46
Cephaloziella rubella (Nees) Warnst
46 39
Cephaloziella rubella var sullivantii
(Aust) K Mull 46 39
Cephaloziella rubella var bifida (Lindb) Douin
35
46
Chiloscyphus pallescens (Ehrh ~ Hof~ Dum Chiloscyphus pallescens var fragilis
39J 46 88 89J 90 (Roth) K Milll 46
Chiloscyphus E9lyanthos (L) Corda
1 24 39 46
Chiloscyphus polyanthos vax rivularis
(Schrad) Nees 39 46
poundpoundnocepha1um conicum (1) Lindb
12 28 39 46 (49) 87 88 8990 120 121 148
Cololejeunea biddlecomiae (Aust) Evans
39 46 62 87 88 89 90
36
Dipl0]hyllum apicu1atum (Evans) Steph Fossombronia foveo1ata Lindb
(35)41 46 90 140 1 t 38 (39) t 46
Frullania bolanderi Aust
13 46 62 88 90 139 140
Frullanla ari ttoniae Evans
39 46 69 (71) 87 88 89 90
Frullania eboracensis Gott
1 9 28 4 39 46 (57) (59) 69 70 (71) 87 88 89 90 92 93 97 148
Frullania inflata Gott
46 87 88 89 (139)
Frullania oakesiana Aust Frullania plana Sullo
(35) 42 46 87 (139) (140) 92
Frullania ri paria Hampe ~ Lehm
46 69 71 87 89
Frullania ~elwynianna Pears
46 89
Geocalyx SEaveolens (Schrad) Nees
394688 8990 (119) 140
Frullania tamarisci (L) Dum subsp sPagrayana
(Mont) Hatt 28 34 l~6 t (57) 88 89 90
38
Mar pan thus seutatUB ( Web amp Mohr) Spruce Jamesoniella autumnalis (DC) Steph
(35) 39 46 90 1 7 28 39 46 56 (57) 87 8889 90 (119) 137 141
Jungermannia confertissima Nees
137 141
Jungermannia crenuliform1s Aust
79 137 141
Jungermannia ~il1ima Sm
46 90 141 Jungermannia exsert1folia subsp corifo~ (Dum~
Vana (35) 46 79 141
39
Jungermannia hyalina Lyell Jungerrr6nnia 1eiantha Grolle
40 46 79 136 141 39 46 79 88 90 1)7 141
Jungermannia IXlaxis Lindb Jungermannia pumila With
46 67 79 (139) (140) 141 )9 46 90 1)6 141
Jungermannia sphaerocarpa Hook Kurzia setacea (Web) Grolle
4679136140IIH 136
Lejeunea cavifo1ia (Ehrh) Lindb Lepidozia reptans (1) Dum
1 7 27 39 46 88 89 90 137 14010 I ( 27) 46 I 69 (71) 88
lophocolea heterophylla (Schrad) Dum Lophocolea minor Nees
1 )4 39 46 (57) 87 88 89 27 39 46 88 89 90 (119) 90 (93)94 119 136
Lophozia al~stris (Schleich ~ Web) Evans Lophozia ascendens (Warnst) Schust
46 90 137 141 136 140 141
41
loIilozia badensls (Gatt ~ Rabenh)
46 141
lophozia blcrenata (Schmid sectA Hoffm)
Dum 46 141
lophozia collaris (Nees) Dum
39 46 141
lophozla excisa (Dicks) Dum
27 4679 141
lophozia gillmani (Aust) Schust lophozia gIlttulata (Undo tl H Arnell)
36 39 40 46 136 (140) 141 Evans 39 46 136 141
42
141
Lophozia heterocolEDs (Thed) MA Howe IoPhozia ipcisa (Schard) Dum
27 39 46 89 141 8 27 31 46 79 90 140 141
LoQhozia longidens Macoun Lophozia ventricosa
1 46 79 90 141
Lophozia ventricosa var silvicola
(Buch) Jones 141 LoEhozla ventricosa vax longif1ora (Nees)
Macoun 39 46 79 141
43
~~ fragrans (Balbis) Frye et Clark Mannia pi 10sa (Hornem) Frye et Clark
46 (139) (140) 9 46 79 116 139
Mannia triandra (Scop) Gro 11e
46 (140)
Marchantia pound91ymorpha L
1 13 28 3946 (49) 78 87 88 89 90 (Ill) 121 122 147
Mo~rckia hibernica (HookJ Gott
46 89
Metzgeria conjugata Lindb
44
46
Mylia anomala (Hook) S Gray Notothylas orbicularis (Schwein) Sullo
39 46 141 (35)45 46
Nowellia curvifolia (Dicks) Mitt
17 27 3846 88 89 90 141
Odontoschisma denudatuJI1 (Nees ~ Mart)
Dum 46 81 137 141
Pallavicinia lyellii (Hook~ Carruth
90
Odontoschisma macounii (Aust) Under
45 46 (66) (139) 140 141
45
=
Pellia epiphllla (L) CordaPellia endiviifolia (Dicks~ Dum
27 39 46 90 13724 39 46
Pellia neesiana Limp bull Phaeoceros
39 46 89 27 39 45 46 97
Porella pinnata L
13 27 I 46 90 Plagiochila asplenioides (L) Dum
27 I 39 1+6 88 89 I 90 137
46
~e11a platYEhylla (L) Pfeiff Porella platyphylloi~ (Schwein) Lindb
17 27 39 46 69 (71)79 87 88 89 9 34 46 (57) 59 87 88 89 92 90 92 93 77
Preissia quadrata (Seop) Nees
1 36 3946 (71) 88 89 90 121 123
137
Ptllidium ciliare (L) Hampe
1 27 39 46 132 141
Dum
(22) 28 3946 (57) 87 88 89 90 92
Ptilidium ~lcherrium (G Web) Hampe
(9) 13 394656 (57)70 87 88 89 90 93 (119) 136 141
47
Radula obconica Sullo Reboulia hemlsphaerlca (L) Raddi
39 46 88 89 (43) 46 (59) 70 90 t Cf 124 125 (139)
Riccardia ~atifrons Lindb
1 27 39 46 89
Riccardia multifi~ (L) S Gray
39 46 88 145
Riccia beyrichiana Hampe ~ Lehm
46 Riccardia palmata (Hedw) Carruth
39 46 90
48
Riccia bifurca Hoffm Riccia cavernosa Hoffm
48 13739 46 79 137
Riccia fluitans L
242731454679
Riccla frostli Aust
48 79
Riccia sullicantii Aust
79 (137) Riccia sorocarpa Bisch 46 (137)
49
Ricciocarpus natans (1) Corda Sca~ia apiculata Spruce
13 27 46 79 89 110 39 46 (139) 141
ScaJltll1ia carinthica Jack ~ Lindb
137 t 141
Scapania ~ (Mart) Durn
39467990137141
Scapania digenil Schiffn
90
ScaEsectflia cuspiduligera (Nees) K Mull
46 (139) (140) (141)
50
Scapania glaucoeerha1a (Tayl) Aust Seapania gyrrmostomophilia Kaal
39 46 (139) 141 41 44 46 64 (139) (140) 141
ScaP3poundlia irrigua Gott ~ almiddot
7 39 46 79 137 (140L 141
Seapania nemorosa (L) Dum
1 39 46 89 90 137
Scapmi~ ~~ta Smiddotlch
137 141 51
~caPnia poundludicola LDeske amp K Mull Scapania saxicola Schust
46 (66) 141 136 141
Sc~~ scandica (H Arnell ~ Buch) Macv
139 140
Scapania subalpina (Nees) Durn
39 40 46 79 137 141
Scapania undulata (L) Dum
1 42 46 79 141
Scapania umbrosa (Schrad) Dum
137 140 141
52
Trichocolea tomentella (Ehrh) Dum Triomarla exsecta (Schmid) Schiffn
27 (35) 39 46 47 88 137 141 8 28 39 46 90 140 141
Tritomaria exsectiformis (Breidl) Loe~ke Tritomaria Sluinguedentata (Huds) Buch
39 -t6 89 90 136 140 141 7 39 46 141
Trltomaria scitula (Tayl) Joerg
46 136 (139) 141
53
A numerical summary of the bryophyte flora of the state based on our literature search is presented in Table 1 (below) This includes 517 taxa 70 families and 176 genera This numerical summary is comshypared with the published floras of Michigan mosses (Crum 1976) and Minnesota hepatics (Schuster 1953) Thus we surmise that further collecting could lead to the same approximate numbers for Wisconsin bryoshyphytes
Table 1 Comparison of Recorded Bryophytes of Wisconsin with Michigan and Minnesota and Summary of Wisconsin Taxa
Wisconsin Michigan (Crum 1976)
Minnesota (Schuster 1955)
Hepaticsamp Hornworts
Taxa Families Genera
137 28 47
174 32 52
Mosses
Taxa Families Genera
380 42
129
503 42
149
8
-
~
~ tIt
Figure 1 The Driftless Area and Tension Zone The counties are 01$0 shown on map
ca location of tension zone as deecfibed bv Curtis (1959)~ The Orittless Area
9
Agn 2 Major Plant Communitiee of Wsconsln aaed on surwvor recorda bull co 18laquo) After Curtismiddot 1959
tit beol Forest e Phillie
Northernmiddot hardwood forest reg Pine 5cMInna
copy Southernmiddot hordwood forest CD Oak Savanna
10
SYSTEMATIC ARRANGEMENT OF WISCONSIN BRYOPHYTES
ANTHOCEROTAE (Hornworts)
1 Anthocerotaceae Anthoceros 1 emnd Prosk
macounii M A Howe punctatum 1
Phaeoceros Prosk laevis (1) Prosk subsp laevis
Notothylas Sul1 orbicularis (Schwein) Sullo
HEPATICAE (liverworts)
2 PseudolepicoJeaceae Blephllrostoma (Dum emend Undb) Dum
trlchophyllum (L) Dum subsp trlchophylluni
3 Ptilidiaceae Ptilidium Nees
ci1iare (L) Hampe pulcherrimum (G Web) Hampe
4 Trichocoleaceae Trichocolea Dum nom cons
tomentella (Ehrh) Dum
5 Lepidoziaceae Kunia von Martens
setacea (Web) GroUe
Lepidozia (Dum) Dum reptans (1) Dum
Bazzania S Gray trilobata (1) S Gray var triZobata
6 Calypogejaceae 011ypogeja Raddi
flSsa (L) Raddi subsp issa integristipula Steph muelleriana (Schiffn) K Mull subsp muelleriana neesiana (Mass et Carest) K Mull sphagnicola (H Arnell et J Perss) Warnst et loeske trichomanis (1) Corda
11
7 Cephaloziaceae Cepluzlozla (Dum emend Schiffn) Dum
biscwpidata (L) Dum subsp biclupidIIta catenulllttl (Hub) Lindb coMivens (Dicks) Lindb var connivens lacinultzta Jack ampX Spruce lunulifolilz (Dum) Dum macounii (Aust) Aust pleniceps (Aust) Lindh var pleniceps
Noweilia Mitt cunifolilz (Dicks) Mitt
8 Adelanthaceae Odontoschimuz (Dwn) Dum
denudatum (Nees ampX Mart) Dum Vir denudatum mtlCount (Aust) Underw
c
9 Cephaloziellaceae CephtlJoziellll (Spruce) Schiffn nom C()II$
divarlctzt4 (Sm) sclurfn val divarlcat(l elachhta (bdc) Schiffn 1uImpetuw (Nees) Schirfn IUbellll (Nees) Warnst vat IUbeIl4rube (Nees) Warnst var biid4 (Lindb) Dquinrube (Nees) WarDSt Var sulliwmtii (Aust) K MuD
10 Geo~c~ larptuttllus Nees
scutatus (Web et Mohr) Spruce
Geocalyx Nees grGVeolens (Schrad) Nees
11 Lophocoleaceae Lophocolea (Dum) Dum
heteropllyll4 (Schrad) Own minorNees
Chiloscypbus Corda om cons pallescens (Ehrh ampX Hoffm) Dum varfragllis (Roth) K Mun polyantitos (L) Corda var poiyanthos pol)llaquo1lthos (L) Corda var rlvulllris (Schrld) Nees
12 PlagiochiJaceae Plagio(hila Dum nom cons
tlBpienioides (L) Dum var asplenioidu
13 Jungermanniaceae Btriilophoril Loeske
attenutlta (Mart) Loeske bIubtlta (Schmid ampX Scltreb) Loeske
12
Lophom (Dum) Dum alpestris (Schleich ~ Web) Evans ascendens (Warnst) Schust badenais (Gott ex Rabenh) Schiffn var bademii bierenata (Schmid ex H)ffm) Dum eollam (Nees) Dum exci8a (Dicks) Dum var excisa gllimanii (Aust) Schust varglllmanii guttulata (Lindb et H Arnell) Evans heterocolpos (Thed) M A Howe var heterocolpos ineistl (Schrad) Dum var tnelsa longidens (Lindb) Macoun subsplongidens ventricosa (Dicks) Dum var ventricosa ventricolfl (Dicks) Dum varlontifIotrl (Nees) Macoun ventricosa (Dicks) Dum var silvlcola (Bueh) Jones
Tritomarill Schiffn ex Loeske exseeta (Schrad) Loeske exsecttformi$ (Breidl) Loeske subsp exsectiormfs quinquedentata (Huds) Buch var quinquedentata scitula (rayl) Joerg
Anastrophyllum (Spruce) Schiffn hellemnum (Noes) Schust michtzuxii (Web) Buch ex Evans minutum (Schrob) Schust var minutum
Mylill S Gray nom cons anomala (Hook) S Gray
Jamesoniella (Spruce) Carring autumnalis (DC) Steph var autumnalis
Jungemumnia L emend Dum eonfertissimtt Nees crenuliformis Aust exsertijolill Steph subsp cordifolia (Dum) Vana graciUima Sm hyalina Lyell leillnthll Grolle polaris Lindb pumila With sphaerocarpa Hook
14 Scapaniaceae Diplophyllum (Dum) Dum nom cons
apiculatum (Evans) Steph var apiculatum
Scapanill (Dum) Dum nom cons apkuata Spruce carinthillca Jack ex Lindb curta (Mart) Dum var curta
13
cuspiduligera (Nees) K Mull var cuspiduiigera degenii Schlffn ex K Mull var degenii glllucocephaJa (Tayt) Aust gymnostomophila Kaal irrigwJ (Nees) Gott et al subsp irrigua lingulata Buch varlingulata mucronata Buch subsp mucronata nemorosa (l) Dum paludicola Toeske et K Mun var paludicola paludosa (K Mull) K Mull saxicola Schust scandiea (H Arnell et Buch) Macv var scandiea 8Ubalp~ (Nees) Dum umbroSll (SchIad) Dum undulata (L) Dum var undulata
15 Radulaceae Raduia Dum nom COllS
compianata (L) Dum obconica Sull
16 Porellaceae Parella L
pinTlllta L patyphylla (L) Pfeiff platyphylloidt(l (Schwein) lindb
17 Jubulaceae FruUanill Raddi
bolanderi Aust brittoniae Evans eboracemis Gott inlllta Gott oakesiana Aust plana Sull rlparia Hampe ex Lehm selwyniana Pears taTnl1iJci (L) Dum subsp asagmyana (Mont) Hatt
18 lejeuneaceae Calolejeunea (Spruce) Schlffn
biddlecomiae (Aust) Evans
Lejeunea Ub nom cons cavifolia (Ehrh) Lindb emend Buch
19 Codoniaceae Fossombronia Raddi
foveolata Lindb
20 Pa11aviciniaceae Paklvieinlil S Gray nom cons
lyeiii (Hook) Carruth
14
Moerckia Gott hibemica (Hook) Gott
21 PeWaceae Pellfa Raddi nom cons propos
endivlifolfa (Dicks) Dwn epiphylllz (L) Corda neeBiana (GottlUmpr
22 Metzgeriaceae Metzgerla Raddi
conjugata Undb
23 Blasiacae B L
pulilm L
24 Aneurareae AneumDum
pinguis (L) Dum
Riccfl1dia S Gray nom cons latifrons Undb multiftda (L) S Gray var multifida palmata (Hedw) Carruth
25 Aytoniaceae Reboulill Raddi nom conI
hemllphaerica (L) Radeli
ManniaOpiz fragrrms (Balbis) Frye el ClarK pilosa (Hornem) Frye et Clark trltmdra (Scop) GroDe
26 Conocephalaceae Conocephalum Wigg nom cons
conicum (L) Lindb
27 Marchantiaceae Preiuia Corda
quadrata (Scop) Nees
Marchantia L polymorpha L
28 Ricciaceae Riccwcarpus Corda
natans (L) Corda
Riccia 1 beyrichiana Hampe ex Lehm bifurca Horfm
15
i
CtlJPemostI Hoffnt fuitu L rostii Aust BOIOCtI1fHI Bi~ sulltvtmtii Aust
MUSCI (Mosses)
29 Sphagnaceae SpIutgnum L
capillaceum (Weiss) Schrank var Cllpil1tlceum caplllllceum (Weiss) Vir tenellum (Schimp) Andr centrille C Jens ex Amen et C Jens compactummiddot OC ex Lam et Dc ClUpidatum Ehrh ex Hoffin var aupidatum jimbrltltum Wits ex J Hookmiddot fulCUm (Scbimp) KUngrr glrgenBOhIlll Russ imbrlctltum Hornsch ex Russ muelllmitil4m BOd 1IIIIU (RUII) C Jens przIutre L papIIIoIum Lindb pulchrum (Lindb ex braithw) Wamst recumun P-Beauv var recumun recum P-Beauv var tenue Klinggr riptlrlum Ansstr ruPQWItWamst squtl1fOlUm Crome subftltens Russ et Wamst ex Wamst IUbIeCfUldum Nees ex Sturm var subsecundum subsecundum Nees ex Sturm varrutem (Nees Hormch et sUUm) Hueb teres (Sdlimp) Anptr e C Hartm
WtlI7IItorjii Russ wuljfanum Gira
30 Andreaeaceae AndNrrerI Hedw
rupeitrlr Oedw var pest
31 Fissident~e Fluid_ lledw
trdiantoid Hedw byoidei Bedw var bryolda cristaM Wils ex Mitt exiguus sun VIr exigUw onttmUi (B-Pyl) Steud jtuberi Lesq et James obtusifolhu Wits var obtwifoliul ollPlUlldQldes Heel ~litSuU lubbtulkril Bed tfDdfoliul lledw llridubu (S~ ) Wahlenb vallirlIU1u
16
32 Ditrichaceae Pleuridium Brid
subulatum (Hedw) Rabenh
Ditrichum Hampe heteromallum (Hedw) Britt linellre (Sw) lindb
~ pallidum (Hedw) Hampe pusiUum (Hedw) Hampe
Saelania Lindb gIlucescens (Hedw) Bomanss et Broth
Ceratodon Brid purpureus (He(lw) Brid var prupurew
Distichium BSG capillaceum (Hedw) BSG inclinatum (Hedw) BSG
33 Bryoxipbiaceae Bryoxlphium Mitt
norvegicum (Brid) Mitt
34 Seligeriaceae Seligeria BSG
cumpylopoda Kindb ex Macoun et Kindb pusillll (Hedw) BSG recurvata (Hedw) BSG
Bindia BSG acuta (Hedw) BSG
35 Dicranaceae BlUehia Schwegr
lexUOSll (Sw ~ Schwaegr) C MueU sullivantii Aust
Trematodon Michx ambiguus (Hedw) Hornsch
Dicranellll (C Muell) Schimp cerviculata (Hedw) Schimp val cillPicuiata
heteromalla (Hedw) Schimp ~ rufescens (With) Schimp
schreberiana (Hedw) Schhnp var schreberiana schreberitJna (Hedw) Schimp var robusta Schimp ex Braithw subulata (Hedw) Schimp varia (Hedw) Schimp
Dicranodontium BSG denudatum (Brid) Britt ex Williams
Panzeucobryum (Lindh) Loeske Iongifolium (Hedw) oelke varlongifolium
17
Tortella (Undb) Limpr fragilis (Drumm) Umpr humilis (Hedw) JeM torlUoSfl (Hedw) Umpr
Hyophila Brid involuta (Hoole) Jaeg et Sauerb
Didymodon Hedw rigidulus Hedw
BryoerythrophyUum Chen recurvirostnon (Hedw) Chen
Btubula Hedw acula (Brid) Brid var acuta convoiuta Hedw fallox Hedw unguicultltll Hedw
Acaulon C MueD muticum (Uedw) C Muell var rufeseens (Iaeg) Crum
PJl~lm Hedw CUSPidtltum Hedw
De8mlltodon Brid cernuu (Hueb) BSG oblUsifoliw (Schwaegr) Schimp
T01IUla Hedw mueronifolia Schwaegr rurall (Hedw) Gaertn Meyer et Scherb var ruralis
39 Grimmiaceae Grlmmill Hedw
alpicola Hedw var alpicola alpicola Hedw var dupretii (Ther) Crum alpicola Hedw var rlvularl8 (Brid) Wahlenb apocarpa Hedw var apocarpa apocarpa Hedw var ambipa (Sull) Jonea apocarpa Hedw var conjerttl (Funck) Spreng apocarpa Hedw var strleta (Tum) Hook et Tayl CiIIyptrrlta Hook ex Drumm donniana Sm var donn_ hemumti Crum ltlevla (Brid) Brid montana BSG oineyi SuD oVllll8 (Hedw) lindb piUfe P-Beauv pltJgiopodill Hedw pulvinata (Hedw) Sm varbullpulvinata teretinervl Limpr
18
Orothera C Muell leana (Sull) C Muell
Rfwbdoweisia crispata (With) Undb
Cynodontium BSG ex Schimp alpestre (Wahlenb) Milde strumiferum (Hedw) Undb
Dichodontium Schimp pellucidum (Hedw) Schimp
Oncophorus (Slid) Brid vuens (Hedw) Brid wahlenbergii (Brid)
Dicranum Hedw bonjeanii DeNot ex Usa drummondii C Muell lagellare Hedw varj1ogellare fulvum Hook fuscescens Turn var fuscescens montanum Hedw muehlenbeckii BSG var muehlenbeckii polysetum Sw sabuletorum Ren et Card scoparium Hedw undulatum Brid viride (Sull et Lesq ex Sull) lindb
36 Leucobryaceae Leucobryum Hampe
glaucum (Hedw) Angstr ex Fr
37 Encalyptaceae Encalypta Hedw
ciliata Hedw var ciliata procera Bruch
38 Pottiaceae Astomum Hampe
muhlenbergianum (Sw) Grout
Weissia Hedw controversa Hedw var controversa
Gymnostomum Nees Homsch et Stunn aeruginosum Sm recurvirostrum Hedw var recurvirostrum
Trichostomum Bruch spirale Prout
19
40 Ephemeraceae Ephemerum Hampe
cohaerens (Hedw) Hampe crassineium (Schwaegr) Hampe val crassinervium spinulosum Bruch et Schimp ex Schimp
41 Funariaceae AphflllolThegma SuD
serratum (1 Hook et Wils ex Drumm) Sullo
Physcomitrium (Brid) Fuernr hookeri Hampe immersum Sullo pyriforme (Hedw) Hampe
Funaria Hedw americana Undb hygrometrica Hedw vaT hygrometrica
42 Splachnaceae Splachnum Hedw
ampullaceum Hedw
43 Schistostegaceae Schistostega Mohr
pennata (Hedw) Web el Mohr
44 Bryaceae Pohlia Hedw
annolina (Hedw) Lindb var annotina camea (Tum) Lindb cruda (Hedw) Undb elongata Hedw iliformis (Dicks) Andr lescurlana (Sull) Grout nutans (Hedw) Lindb obrusifolia (Brid) L Koch proligera (Undb ex Umpr) Undb ex Amell wahlenbergii (Web et Mohr) Andr
Leptobryum (B S G) Wits pyriforme (Hedw) Wils
Bryum Hedw algovicum Sendtnex C Muell argenteum Hedw caespiticium Hedw capillare Hedw var capillare creberrlmum Tayt lonchocaulon C Muell pallens (Brid) Sw ex Roehl pseudotrlquetrum (Hedw) Gaertn Meyer et Scherb varpudotriquetrum
20
pseudotriquetrum (Hedw) GaertnMeyer et Scherb var bimum (Schreb) Ulj stenotrichum C Muell var stenotrichum tortiolium Funck ex Brid uliginosum (Brid) B S G violaceum Crundw et Nyh weigelii Spreng
Rhodobryum (Schimp) Hampe roseum (Hedw) Limpr
45 Mniaceae Mnium Hedw
ciliare (C Muell) Schimp cinclidioides Hueb cuspidarum Hedw drummondii Bruch et Schimp longirostrum Brid lycopodioides Schwaegr marginatum (With) Brid ex P Beauv mediumB~S G var medium onhorrhynchum Brid pseudopunctatum Bruch et Schimp punctalUm Hedw var punctatum punctatum Hedw var elatum Schirnp rugicum Laur spinufosum B S G stellare Hedw
46 Aulacomruaceae Au1acomnium Schwaegr
androgynum (Hedw) Schwaegr heterostichum (Hedw) B S G paJustre (Hedw) Schwaegr var paJustre
47 Meesiaceae Amblyodon B S G
deillbotus (Hedw) B S G
Meesia Hedw triquetra (Richt) Angstr uJiginosa Hedw
48 Bartramiaceae Plagiopus Brid
oederiana (Sw) Umpr
Bartramia Hedw pomiformis Hedw var pomiformis
Philonotis Brid fontana (Hedw) Brid varfontana longseta (Michx) Britt varlongseta marchica (Hedw) Brid
21
~
~
49 Timmiaceae Timmia Hedw
megapolitana Hedw var megapolitana
50 Orthotrichaceae Zygodon Hook et Tayl
viridissimus (Dicks) Brid var rupestris lindb ex C J Hartm
Amphidium Schimp lapponicum (Hedw) Schimp mougeotii (B S G) Schimp
Orthotrichum Hedw anomalum Hedw obtuaifolium Brid ohioense Sullo et Lesq ex Aust pumilumSw sordidum Sullo et Lesq ex Aust speciosum Neesax Sturm var eegam (Schwaegr ex Hook et Grev) W~rnst stellDtwn Brid strrmgulatum P Beauv
Uleta Mohr ex Web cOfIICtata (p Beallv) Hamrn crispa (Hedw) Brid
Drummondill Hook ex Drumm prorepens (Hedw) Brltt
51 Fontinalaceae Fantinalis Hedw
antipyretica Hedw var antipyrettca antipyretica Hedw forma rabusta (Card) Card dalecarlica Schimp ex B S G var dalecarlica flqccidtJ Ren et Card hypnoides C J Hartrn var hypnoides hypnoides C J Hartm var durillei (Schimp) HU$ll
missourlcaCard novae-angliae Sullo var novae-angliae splulgrUlolill (C Moen) Wijket M~S
DichelY1l1ll Myr capilloceum (With) Myr falcatum (Bedw) Myr HIllescens B S G
52 Climaciaceae aimecium Web et Mohr ex Mohr
americllnum Brid dedroides (Hedw) Web et Mohr
22
S3 Hedwigiaceae Hedwigia P Beauv
ciliata (Hedw) P Beauv
54 Cryphaeaceae Forsstroemia lindh
ohioense (Sull) lindh
55 Leucodontaceae Leucodon Schwaegr
brachypus Blid var brachypus brachypus Brid var aniJrewsitlnus Crum et Anderson julaceus (Hedw) SuU
56 Neckeraceae Neckera Hedw
pennata Hedw var pennata
1fomalia (Brid) B S G trichomanoides (Hedw) B S G
57 Theliaceae Thelia Sull
asprellll Sullo hirtella (Hedw) Sullo lescurii SuU
Myurella B SG julacea (Schwaegr) B S G
sibrica (C MueU) Reim var sibrica
58 Fabroniaceae Fabronio Raddi
ciliaris (Brid) Bnd
Anacamptodon Brid splachnoides (Groet ex Brid) Brid
Schwetschkeopsis Broth fabronia (Schwaegr) Broth
59 Leskeaceae Lindbergia Kindb
brachyptera (Mitt) Kindb
Leskea Hedw gracilescens Hedw obscura Hedw poiycarpa Hedw
Leskeella (limpr) Loeske nervosa (Brid) Loeske
23
Pterlgy1lllndrum Hedw filiforme Hedw
60 Thuidiaceae Haplohymenfum Dozy et MoIk
trlste (Ces ex De Not) Kiodb
Anomodon Hook et Tayl attenuatus (Hedw) Hueb minor (Hedw) Fuemr rostratus (Hedw) Schimp viticulosus (Hedw) Hook et Tayl
Haplocladium (C Mucn) C MueU microphyllum (Hedw) Broth virginillnum (Brld) Broth ~bull
Thuidium B S G abietinum (Hedw) B S G delicatulum (Hedw) B S G vardelicatulum delicatulum (Hedw) B S G var radicans (Kindb) Crum Steere et Anderson minutulum (Hedw) B S G pygmaeum B S G recognftum (Hedw) Lindb scitum (p Beauv) Aust
Helodium (Sull) Warnst blandowli (Web et Mohr) Warnst var blandowli blandowli (Web et Mohr) Wamst var helodioides (Ren et Card ex Roell) Crum
Steere et Anderson paludomm (Sull) AuS
61 Amblystegiaceae Cratoneuron (Sull) Spruce
commulatum (Hedw) Roth var commutfltum commutatum (Hedw) Roth var fakatum (Brid) Moenk ilicinum (Hedw) Spruce var filcinum
Campylium (Sun) Mitt chrysophyllum (Brid) 1 Lange var chrysophyllum hispidulum (Brid) Mitt var hllJidulum polygamum (B S G)C lens varpolygtlmUm stellatum (Hedw) C1ens vat stelltum
Leptodictyum (Schimp) Warnst laxirete (Card et Ther) Broth riparium (Hedw) Wamat trichopodium (Schultz) Warns l1acillans (Sull) Broth
Hygroamblystegium Loelke jlul1iatile (Hedw) oeske var luViatile
24
fluviatile (Hedw) Loeske var oWltum Grout noterophilum (Sun et Lesq ex Sull) Warnst tenox (Hedw) Jenn var tenax tenox (Hedw) Jenn var spinilolium Schimp) Jenn
Amblystegium BSG juratzkanum Schimp serpens (Hedw) BSG varlum (Hedw) Undb var varium
Platydictya Berk confervoides (Brid) Crum jungermanniodes (Brid) Crum minutissimum (Sull et Lesq ex Sull) Crum subtile (Hedw) Crum
Drepanocladus (C Muell) Roth aduncus (Hedw) Warnst var aduncus aduncus (Hedw) Warnst var capillilolius (Wamst)Riebm aduncus (Hedw) Warnst var kneiffii (BSG) Moenk aduncus (Hedw) Warnst var polycorpus (Bland ex Voit) Roth exannulatus (BSG) Warnst fluitans (Hedw) Warnst var fluitans revolvens (Sw) Wamst var revolvens sendtneri (Schimp) Warnst var sendtnerl uncinatus (Hedw) Warnst var uncinatus uncinatus ffiedw) Warnst var piwnosus (Schimp) Wamst vernicoms (Lindb ex C Hartm) Warnst
Hygrohypnum Lindb eugyrium (BSG) loeske luridum (Hedw Jenn varluridum ochraecum (Tum ex Wils) Loeske var ochraceunt
Calliergon (Sull) Kindb cordilolium Hedw Kindb glganteum (Schimp) Kindb var giganteum richardsonii (Mitt) Kindb ex Wamst stramineum (Brid) Kindb trilarium (Web et Mohr) Kindb
Calliergonella Loeske cuspidata (Hedw) Loeske
Calliergidium (Ren) Grout bakeri (Ren) Grout
Scorpidium (Schimp) Urnpr scorpiodides (Hedw) limpr
62 Brachytbeciaceae Tomenthypnum loeske
nitens (Hedw) Loeske var mtens
25
Homalotheciella (Card) Broth subcapillata (Hedw) Broth
Brachythecium BSG acuminatum (Hedw) Awt acutum (Mitt) Sullo calcareum IGndb campestTe (C MuelL) BSG cunum (Lindb) Umpr digastrum C Muell et Kindb ex Macoun et Kindb oxycladon (Brid) Jaeg et Sauerb var oxycladon plumosum (Hedw) BSG populeum (Hedw) BSG var populeum reflexum (Starke ex Web et Mohr) DsG var relexum reiexum (Starke ex Web et Mohr) DSG var pacificum Renet Card ex Roell rlvulare BSG var rivulare tabulum (Hedw) BSC var tabuum salebrosum (Web et Mohr) BSG var salebrosum velutinum (Hedw) BSG Vat velutinum
Bryhniil Kaur graminicolor (Brid) Grout var graminicolor novae-angliae (Sull et Lesq ex sun) Grout
Rhynchostegium BSG senulatum (Hedw) Ja4lg et Sauerb
Rhynchostegiella (BSG) Limpr compacta (C MueU) Loeske
EU1hynchium 8SG hians (Hedw) Sande Lac pulchellum (Hedw) Jenn var pulchelium riparloides (Hedw) Rich
Stokeriella (Kindb) Robins pmelonga (Hedw) Robins var stokesii (Tum) Crum
63 Entodontaceae Entodon C Muell
brevisetus (Hook et Wils ex Wils) Lindb cladorrhizans (Hedw) C Muell seductrlx (Hedw) C MueU var wluctrix
Pleurozium Mitt schreberl (8rid) Mitt var schreberi
64 Plagiotheciaceae Pltlglothecium BSG
CflVifolium (Brid) Iwats var CtWifolium denticulatum (Hedw) 8SG laetum BSG ltztebricola BSG
26
6S SernatophyUaceae Brotherella Loeske ex Fleisch
recumins (Michx) Fleisch tenuirostris (Bruch et Schimp ex Sull) Fleisch
Sematophyllum Mitt demissum (Wils) Mitt
66 Hypnaceae Pl(itygyrium BSG
repens (Brid) BSG
PylaisieUa Kindb intricata (Hedw) Grout polyantha (Hedw) Grout selwynii (Kindb) Crum Steere et Anderson
Homomallium (Schimp) Loeske adnatum (Hedw) Broth
Hypnum Hedw cupressiforme Hedw var cupressiforme cupressifonne Hedw varfiliforme Brid curviolium Hedw fertile Sendtn imponens Hedw lindbergii Mitt palescens (Hedw) P-Beauv pratense Koch ex Brid
Calicladium Crum haldanianum (Grev) Crum
sopterygiopsis Iwats muelleriana (Schimp) Iwats
Isopterygium Mitt e1egans (Brid) lindb pulchellum (Hedw) Jaeg et Sauerb tenerum (Sw) Mitt
Taxiphyllum Fleisch depllQtum (Bruch et Schimp ex sun) Fleisch taxirameum (MitL) Fleisch
Herzogiella Broth seligeri (Brid) Iwats striatella (Brid) Iwats ruracea (Lindh) Iwats
hilium De Not crista-castrenna (Hedw) De Not
27
67 Rhytidiaceae Rhytidium (Sull) Kindb
rugosum (Hedw) Kindb
Rhytidfodelphus (Lindb ex Iimpr) Warnst triquetrus (Hedw) Warnst var triquetrus
68 Hylocomiaceae Hylocomium BSG
pyrenaicum (Spruce) lindb splendens (Hedw) BSG varsplendens
69 Buxbaumiaceae Buxbaumia Hedw
aphyllll Hedw
Diphyscium Mohr follosum (Hedw) Mohr
70 Tetraphidaceae Tetraphi3
pellucida Hedw
71 Polytrichaceae Atrichum PmiddotBeauv
angustatum (Brid) BSG cri8pum (James) Sullo oerstedianum (C Muell) Mitt teneUum (Roehl) BSG undulatum (Hedw) P middotBeauv var unduiatum
Pogonatum P-Beauv alpinum (Hedw) Roehl var alpinum umigerum (Hedw) P-Beauv
Poiytrichum Hedw commune Hedw var commune ormosum Hedw juniperinum Hedw longisetum Brid ohioense Ren et Card piliferum Hedw var pilierum strictum Brid
28
MAPS OF BRYOPHYTE TAXA
REPORTED FOR WISCONSIN
HORNWORTS AND LIVERWORTS
29
bull bull WSHeV~N S vE II
0 ~
e~RRON ~USK
(J~ (j -
CHIPPE A DUNN
0 o0 poundAU CLAIRE
~ TREMshyPEAshyLEAU
MICE
ONEIDA
Q LINCOLN
TAYLOR
Q t-~----- MOIRA THON
CLARK
o
10
LA FAVETTE
Hornwor ts amp Liverworts
- None - 1 - 25 taxa
- 26 - 50 tRXshy
- 51 - 75 tax)
- 7c - 100 taxa
FONDOU j LAC ~
DODGE
i O JEfFERSON
WALWORTH
30
Anastrophylltlfl helleranum (Nees) Schust Anastrophyllum michauxil (We~ Buch ~
46 88 136 141 Evans 38 46 141
Anastrophyllum mlnut~ (Schreb) Schuster Aneura plnguls (L) Durn
89 21 384687 88 89 90 (138) (145)
Anthoceros macounil MA Howe ~oceros punctatus L
21 (45) (144) 34
31
Barbilophozia attenuata (Mart) Loeske Barbilophozia barbata (Schmid ~ Schre~
24 141 Loeske 28 39 46 89 90 141
Bazzania trilobata (1) S Gray
1 26 39 46 (57) (59) 79 88 89
90 119 141
BlePbarostoma trichoffiyllum (1) Dumort
1 8 26 39 46 89 90 137 141
Blasia pusilla 1
1 39 46 90
Calypogeja fissa (1) Raddi
90
32
$129geja integristipula Steph C~Ypound2geja muelleriana (Schifm) K Mull
88 89 89
CallpoundO~eja neesiana (Mass amp Carest) K Mull
39 46 89 140 (141)
~pogeja sJlapound2icola (H Arnell tl J bull
Perss) Warnst et Loeske (119) 141
CeIalozia biscuspidata (L) Dum
1 37 I 46 90 t 137 141
CalYpogej~ trichomanis (L) Corda
26 39 (119) 137
33
CeEhalozia catenulata (Hub) Lindb Cephalozia connivens (Dicks) Lindb
39 46 79 141 39 4679 88 (89) 90 119 141
Cephalozia lacinulata Jack ~ Spruce
141 Cephalozia lunulifolia (Dum~ Dum I 28 39 46 79 88 89 90 141
Cepoundhalozia macouni (Aust) Aust Cephalozia ~leniceps (Aust) Lindb 27~ 3746 (66)7987 (139)141 39 46 79 89 141
34
Ceffialoziella 9ivaricata (Sm) Schiffn Cephaloziella ~hista (Jack) Schiffn
46 90 46 (119) 141
Cephaloziella hampeana (Nees) Schiffn
46
Cephaloziella rubella (Nees) Warnst
46 39
Cephaloziella rubella var sullivantii
(Aust) K Mull 46 39
Cephaloziella rubella var bifida (Lindb) Douin
35
46
Chiloscyphus pallescens (Ehrh ~ Hof~ Dum Chiloscyphus pallescens var fragilis
39J 46 88 89J 90 (Roth) K Milll 46
Chiloscyphus E9lyanthos (L) Corda
1 24 39 46
Chiloscyphus polyanthos vax rivularis
(Schrad) Nees 39 46
poundpoundnocepha1um conicum (1) Lindb
12 28 39 46 (49) 87 88 8990 120 121 148
Cololejeunea biddlecomiae (Aust) Evans
39 46 62 87 88 89 90
36
Dipl0]hyllum apicu1atum (Evans) Steph Fossombronia foveo1ata Lindb
(35)41 46 90 140 1 t 38 (39) t 46
Frullania bolanderi Aust
13 46 62 88 90 139 140
Frullanla ari ttoniae Evans
39 46 69 (71) 87 88 89 90
Frullania eboracensis Gott
1 9 28 4 39 46 (57) (59) 69 70 (71) 87 88 89 90 92 93 97 148
Frullania inflata Gott
46 87 88 89 (139)
Frullania oakesiana Aust Frullania plana Sullo
(35) 42 46 87 (139) (140) 92
Frullania ri paria Hampe ~ Lehm
46 69 71 87 89
Frullania ~elwynianna Pears
46 89
Geocalyx SEaveolens (Schrad) Nees
394688 8990 (119) 140
Frullania tamarisci (L) Dum subsp sPagrayana
(Mont) Hatt 28 34 l~6 t (57) 88 89 90
38
Mar pan thus seutatUB ( Web amp Mohr) Spruce Jamesoniella autumnalis (DC) Steph
(35) 39 46 90 1 7 28 39 46 56 (57) 87 8889 90 (119) 137 141
Jungermannia confertissima Nees
137 141
Jungermannia crenuliform1s Aust
79 137 141
Jungermannia ~il1ima Sm
46 90 141 Jungermannia exsert1folia subsp corifo~ (Dum~
Vana (35) 46 79 141
39
Jungermannia hyalina Lyell Jungerrr6nnia 1eiantha Grolle
40 46 79 136 141 39 46 79 88 90 1)7 141
Jungermannia IXlaxis Lindb Jungermannia pumila With
46 67 79 (139) (140) 141 )9 46 90 1)6 141
Jungermannia sphaerocarpa Hook Kurzia setacea (Web) Grolle
4679136140IIH 136
Lejeunea cavifo1ia (Ehrh) Lindb Lepidozia reptans (1) Dum
1 7 27 39 46 88 89 90 137 14010 I ( 27) 46 I 69 (71) 88
lophocolea heterophylla (Schrad) Dum Lophocolea minor Nees
1 )4 39 46 (57) 87 88 89 27 39 46 88 89 90 (119) 90 (93)94 119 136
Lophozia al~stris (Schleich ~ Web) Evans Lophozia ascendens (Warnst) Schust
46 90 137 141 136 140 141
41
loIilozia badensls (Gatt ~ Rabenh)
46 141
lophozia blcrenata (Schmid sectA Hoffm)
Dum 46 141
lophozia collaris (Nees) Dum
39 46 141
lophozla excisa (Dicks) Dum
27 4679 141
lophozia gillmani (Aust) Schust lophozia gIlttulata (Undo tl H Arnell)
36 39 40 46 136 (140) 141 Evans 39 46 136 141
42
141
Lophozia heterocolEDs (Thed) MA Howe IoPhozia ipcisa (Schard) Dum
27 39 46 89 141 8 27 31 46 79 90 140 141
LoQhozia longidens Macoun Lophozia ventricosa
1 46 79 90 141
Lophozia ventricosa var silvicola
(Buch) Jones 141 LoEhozla ventricosa vax longif1ora (Nees)
Macoun 39 46 79 141
43
~~ fragrans (Balbis) Frye et Clark Mannia pi 10sa (Hornem) Frye et Clark
46 (139) (140) 9 46 79 116 139
Mannia triandra (Scop) Gro 11e
46 (140)
Marchantia pound91ymorpha L
1 13 28 3946 (49) 78 87 88 89 90 (Ill) 121 122 147
Mo~rckia hibernica (HookJ Gott
46 89
Metzgeria conjugata Lindb
44
46
Mylia anomala (Hook) S Gray Notothylas orbicularis (Schwein) Sullo
39 46 141 (35)45 46
Nowellia curvifolia (Dicks) Mitt
17 27 3846 88 89 90 141
Odontoschisma denudatuJI1 (Nees ~ Mart)
Dum 46 81 137 141
Pallavicinia lyellii (Hook~ Carruth
90
Odontoschisma macounii (Aust) Under
45 46 (66) (139) 140 141
45
=
Pellia epiphllla (L) CordaPellia endiviifolia (Dicks~ Dum
27 39 46 90 13724 39 46
Pellia neesiana Limp bull Phaeoceros
39 46 89 27 39 45 46 97
Porella pinnata L
13 27 I 46 90 Plagiochila asplenioides (L) Dum
27 I 39 1+6 88 89 I 90 137
46
~e11a platYEhylla (L) Pfeiff Porella platyphylloi~ (Schwein) Lindb
17 27 39 46 69 (71)79 87 88 89 9 34 46 (57) 59 87 88 89 92 90 92 93 77
Preissia quadrata (Seop) Nees
1 36 3946 (71) 88 89 90 121 123
137
Ptllidium ciliare (L) Hampe
1 27 39 46 132 141
Dum
(22) 28 3946 (57) 87 88 89 90 92
Ptilidium ~lcherrium (G Web) Hampe
(9) 13 394656 (57)70 87 88 89 90 93 (119) 136 141
47
Radula obconica Sullo Reboulia hemlsphaerlca (L) Raddi
39 46 88 89 (43) 46 (59) 70 90 t Cf 124 125 (139)
Riccardia ~atifrons Lindb
1 27 39 46 89
Riccardia multifi~ (L) S Gray
39 46 88 145
Riccia beyrichiana Hampe ~ Lehm
46 Riccardia palmata (Hedw) Carruth
39 46 90
48
Riccia bifurca Hoffm Riccia cavernosa Hoffm
48 13739 46 79 137
Riccia fluitans L
242731454679
Riccla frostli Aust
48 79
Riccia sullicantii Aust
79 (137) Riccia sorocarpa Bisch 46 (137)
49
Ricciocarpus natans (1) Corda Sca~ia apiculata Spruce
13 27 46 79 89 110 39 46 (139) 141
ScaJltll1ia carinthica Jack ~ Lindb
137 t 141
Scapania ~ (Mart) Durn
39467990137141
Scapania digenil Schiffn
90
ScaEsectflia cuspiduligera (Nees) K Mull
46 (139) (140) (141)
50
Scapania glaucoeerha1a (Tayl) Aust Seapania gyrrmostomophilia Kaal
39 46 (139) 141 41 44 46 64 (139) (140) 141
ScaP3poundlia irrigua Gott ~ almiddot
7 39 46 79 137 (140L 141
Seapania nemorosa (L) Dum
1 39 46 89 90 137
Scapmi~ ~~ta Smiddotlch
137 141 51
~caPnia poundludicola LDeske amp K Mull Scapania saxicola Schust
46 (66) 141 136 141
Sc~~ scandica (H Arnell ~ Buch) Macv
139 140
Scapania subalpina (Nees) Durn
39 40 46 79 137 141
Scapania undulata (L) Dum
1 42 46 79 141
Scapania umbrosa (Schrad) Dum
137 140 141
52
Trichocolea tomentella (Ehrh) Dum Triomarla exsecta (Schmid) Schiffn
27 (35) 39 46 47 88 137 141 8 28 39 46 90 140 141
Tritomaria exsectiformis (Breidl) Loe~ke Tritomaria Sluinguedentata (Huds) Buch
39 -t6 89 90 136 140 141 7 39 46 141
Trltomaria scitula (Tayl) Joerg
46 136 (139) 141
53
-
~
~ tIt
Figure 1 The Driftless Area and Tension Zone The counties are 01$0 shown on map
ca location of tension zone as deecfibed bv Curtis (1959)~ The Orittless Area
9
Agn 2 Major Plant Communitiee of Wsconsln aaed on surwvor recorda bull co 18laquo) After Curtismiddot 1959
tit beol Forest e Phillie
Northernmiddot hardwood forest reg Pine 5cMInna
copy Southernmiddot hordwood forest CD Oak Savanna
10
SYSTEMATIC ARRANGEMENT OF WISCONSIN BRYOPHYTES
ANTHOCEROTAE (Hornworts)
1 Anthocerotaceae Anthoceros 1 emnd Prosk
macounii M A Howe punctatum 1
Phaeoceros Prosk laevis (1) Prosk subsp laevis
Notothylas Sul1 orbicularis (Schwein) Sullo
HEPATICAE (liverworts)
2 PseudolepicoJeaceae Blephllrostoma (Dum emend Undb) Dum
trlchophyllum (L) Dum subsp trlchophylluni
3 Ptilidiaceae Ptilidium Nees
ci1iare (L) Hampe pulcherrimum (G Web) Hampe
4 Trichocoleaceae Trichocolea Dum nom cons
tomentella (Ehrh) Dum
5 Lepidoziaceae Kunia von Martens
setacea (Web) GroUe
Lepidozia (Dum) Dum reptans (1) Dum
Bazzania S Gray trilobata (1) S Gray var triZobata
6 Calypogejaceae 011ypogeja Raddi
flSsa (L) Raddi subsp issa integristipula Steph muelleriana (Schiffn) K Mull subsp muelleriana neesiana (Mass et Carest) K Mull sphagnicola (H Arnell et J Perss) Warnst et loeske trichomanis (1) Corda
11
7 Cephaloziaceae Cepluzlozla (Dum emend Schiffn) Dum
biscwpidata (L) Dum subsp biclupidIIta catenulllttl (Hub) Lindb coMivens (Dicks) Lindb var connivens lacinultzta Jack ampX Spruce lunulifolilz (Dum) Dum macounii (Aust) Aust pleniceps (Aust) Lindh var pleniceps
Noweilia Mitt cunifolilz (Dicks) Mitt
8 Adelanthaceae Odontoschimuz (Dwn) Dum
denudatum (Nees ampX Mart) Dum Vir denudatum mtlCount (Aust) Underw
c
9 Cephaloziellaceae CephtlJoziellll (Spruce) Schiffn nom C()II$
divarlctzt4 (Sm) sclurfn val divarlcat(l elachhta (bdc) Schiffn 1uImpetuw (Nees) Schirfn IUbellll (Nees) Warnst vat IUbeIl4rube (Nees) Warnst var biid4 (Lindb) Dquinrube (Nees) WarDSt Var sulliwmtii (Aust) K MuD
10 Geo~c~ larptuttllus Nees
scutatus (Web et Mohr) Spruce
Geocalyx Nees grGVeolens (Schrad) Nees
11 Lophocoleaceae Lophocolea (Dum) Dum
heteropllyll4 (Schrad) Own minorNees
Chiloscypbus Corda om cons pallescens (Ehrh ampX Hoffm) Dum varfragllis (Roth) K Mun polyantitos (L) Corda var poiyanthos pol)llaquo1lthos (L) Corda var rlvulllris (Schrld) Nees
12 PlagiochiJaceae Plagio(hila Dum nom cons
tlBpienioides (L) Dum var asplenioidu
13 Jungermanniaceae Btriilophoril Loeske
attenutlta (Mart) Loeske bIubtlta (Schmid ampX Scltreb) Loeske
12
Lophom (Dum) Dum alpestris (Schleich ~ Web) Evans ascendens (Warnst) Schust badenais (Gott ex Rabenh) Schiffn var bademii bierenata (Schmid ex H)ffm) Dum eollam (Nees) Dum exci8a (Dicks) Dum var excisa gllimanii (Aust) Schust varglllmanii guttulata (Lindb et H Arnell) Evans heterocolpos (Thed) M A Howe var heterocolpos ineistl (Schrad) Dum var tnelsa longidens (Lindb) Macoun subsplongidens ventricosa (Dicks) Dum var ventricosa ventricolfl (Dicks) Dum varlontifIotrl (Nees) Macoun ventricosa (Dicks) Dum var silvlcola (Bueh) Jones
Tritomarill Schiffn ex Loeske exseeta (Schrad) Loeske exsecttformi$ (Breidl) Loeske subsp exsectiormfs quinquedentata (Huds) Buch var quinquedentata scitula (rayl) Joerg
Anastrophyllum (Spruce) Schiffn hellemnum (Noes) Schust michtzuxii (Web) Buch ex Evans minutum (Schrob) Schust var minutum
Mylill S Gray nom cons anomala (Hook) S Gray
Jamesoniella (Spruce) Carring autumnalis (DC) Steph var autumnalis
Jungemumnia L emend Dum eonfertissimtt Nees crenuliformis Aust exsertijolill Steph subsp cordifolia (Dum) Vana graciUima Sm hyalina Lyell leillnthll Grolle polaris Lindb pumila With sphaerocarpa Hook
14 Scapaniaceae Diplophyllum (Dum) Dum nom cons
apiculatum (Evans) Steph var apiculatum
Scapanill (Dum) Dum nom cons apkuata Spruce carinthillca Jack ex Lindb curta (Mart) Dum var curta
13
cuspiduligera (Nees) K Mull var cuspiduiigera degenii Schlffn ex K Mull var degenii glllucocephaJa (Tayt) Aust gymnostomophila Kaal irrigwJ (Nees) Gott et al subsp irrigua lingulata Buch varlingulata mucronata Buch subsp mucronata nemorosa (l) Dum paludicola Toeske et K Mun var paludicola paludosa (K Mull) K Mull saxicola Schust scandiea (H Arnell et Buch) Macv var scandiea 8Ubalp~ (Nees) Dum umbroSll (SchIad) Dum undulata (L) Dum var undulata
15 Radulaceae Raduia Dum nom COllS
compianata (L) Dum obconica Sull
16 Porellaceae Parella L
pinTlllta L patyphylla (L) Pfeiff platyphylloidt(l (Schwein) lindb
17 Jubulaceae FruUanill Raddi
bolanderi Aust brittoniae Evans eboracemis Gott inlllta Gott oakesiana Aust plana Sull rlparia Hampe ex Lehm selwyniana Pears taTnl1iJci (L) Dum subsp asagmyana (Mont) Hatt
18 lejeuneaceae Calolejeunea (Spruce) Schlffn
biddlecomiae (Aust) Evans
Lejeunea Ub nom cons cavifolia (Ehrh) Lindb emend Buch
19 Codoniaceae Fossombronia Raddi
foveolata Lindb
20 Pa11aviciniaceae Paklvieinlil S Gray nom cons
lyeiii (Hook) Carruth
14
Moerckia Gott hibemica (Hook) Gott
21 PeWaceae Pellfa Raddi nom cons propos
endivlifolfa (Dicks) Dwn epiphylllz (L) Corda neeBiana (GottlUmpr
22 Metzgeriaceae Metzgerla Raddi
conjugata Undb
23 Blasiacae B L
pulilm L
24 Aneurareae AneumDum
pinguis (L) Dum
Riccfl1dia S Gray nom cons latifrons Undb multiftda (L) S Gray var multifida palmata (Hedw) Carruth
25 Aytoniaceae Reboulill Raddi nom conI
hemllphaerica (L) Radeli
ManniaOpiz fragrrms (Balbis) Frye el ClarK pilosa (Hornem) Frye et Clark trltmdra (Scop) GroDe
26 Conocephalaceae Conocephalum Wigg nom cons
conicum (L) Lindb
27 Marchantiaceae Preiuia Corda
quadrata (Scop) Nees
Marchantia L polymorpha L
28 Ricciaceae Riccwcarpus Corda
natans (L) Corda
Riccia 1 beyrichiana Hampe ex Lehm bifurca Horfm
15
i
CtlJPemostI Hoffnt fuitu L rostii Aust BOIOCtI1fHI Bi~ sulltvtmtii Aust
MUSCI (Mosses)
29 Sphagnaceae SpIutgnum L
capillaceum (Weiss) Schrank var Cllpil1tlceum caplllllceum (Weiss) Vir tenellum (Schimp) Andr centrille C Jens ex Amen et C Jens compactummiddot OC ex Lam et Dc ClUpidatum Ehrh ex Hoffin var aupidatum jimbrltltum Wits ex J Hookmiddot fulCUm (Scbimp) KUngrr glrgenBOhIlll Russ imbrlctltum Hornsch ex Russ muelllmitil4m BOd 1IIIIU (RUII) C Jens przIutre L papIIIoIum Lindb pulchrum (Lindb ex braithw) Wamst recumun P-Beauv var recumun recum P-Beauv var tenue Klinggr riptlrlum Ansstr ruPQWItWamst squtl1fOlUm Crome subftltens Russ et Wamst ex Wamst IUbIeCfUldum Nees ex Sturm var subsecundum subsecundum Nees ex Sturm varrutem (Nees Hormch et sUUm) Hueb teres (Sdlimp) Anptr e C Hartm
WtlI7IItorjii Russ wuljfanum Gira
30 Andreaeaceae AndNrrerI Hedw
rupeitrlr Oedw var pest
31 Fissident~e Fluid_ lledw
trdiantoid Hedw byoidei Bedw var bryolda cristaM Wils ex Mitt exiguus sun VIr exigUw onttmUi (B-Pyl) Steud jtuberi Lesq et James obtusifolhu Wits var obtwifoliul ollPlUlldQldes Heel ~litSuU lubbtulkril Bed tfDdfoliul lledw llridubu (S~ ) Wahlenb vallirlIU1u
16
32 Ditrichaceae Pleuridium Brid
subulatum (Hedw) Rabenh
Ditrichum Hampe heteromallum (Hedw) Britt linellre (Sw) lindb
~ pallidum (Hedw) Hampe pusiUum (Hedw) Hampe
Saelania Lindb gIlucescens (Hedw) Bomanss et Broth
Ceratodon Brid purpureus (He(lw) Brid var prupurew
Distichium BSG capillaceum (Hedw) BSG inclinatum (Hedw) BSG
33 Bryoxipbiaceae Bryoxlphium Mitt
norvegicum (Brid) Mitt
34 Seligeriaceae Seligeria BSG
cumpylopoda Kindb ex Macoun et Kindb pusillll (Hedw) BSG recurvata (Hedw) BSG
Bindia BSG acuta (Hedw) BSG
35 Dicranaceae BlUehia Schwegr
lexUOSll (Sw ~ Schwaegr) C MueU sullivantii Aust
Trematodon Michx ambiguus (Hedw) Hornsch
Dicranellll (C Muell) Schimp cerviculata (Hedw) Schimp val cillPicuiata
heteromalla (Hedw) Schimp ~ rufescens (With) Schimp
schreberiana (Hedw) Schhnp var schreberiana schreberitJna (Hedw) Schimp var robusta Schimp ex Braithw subulata (Hedw) Schimp varia (Hedw) Schimp
Dicranodontium BSG denudatum (Brid) Britt ex Williams
Panzeucobryum (Lindh) Loeske Iongifolium (Hedw) oelke varlongifolium
17
Tortella (Undb) Limpr fragilis (Drumm) Umpr humilis (Hedw) JeM torlUoSfl (Hedw) Umpr
Hyophila Brid involuta (Hoole) Jaeg et Sauerb
Didymodon Hedw rigidulus Hedw
BryoerythrophyUum Chen recurvirostnon (Hedw) Chen
Btubula Hedw acula (Brid) Brid var acuta convoiuta Hedw fallox Hedw unguicultltll Hedw
Acaulon C MueD muticum (Uedw) C Muell var rufeseens (Iaeg) Crum
PJl~lm Hedw CUSPidtltum Hedw
De8mlltodon Brid cernuu (Hueb) BSG oblUsifoliw (Schwaegr) Schimp
T01IUla Hedw mueronifolia Schwaegr rurall (Hedw) Gaertn Meyer et Scherb var ruralis
39 Grimmiaceae Grlmmill Hedw
alpicola Hedw var alpicola alpicola Hedw var dupretii (Ther) Crum alpicola Hedw var rlvularl8 (Brid) Wahlenb apocarpa Hedw var apocarpa apocarpa Hedw var ambipa (Sull) Jonea apocarpa Hedw var conjerttl (Funck) Spreng apocarpa Hedw var strleta (Tum) Hook et Tayl CiIIyptrrlta Hook ex Drumm donniana Sm var donn_ hemumti Crum ltlevla (Brid) Brid montana BSG oineyi SuD oVllll8 (Hedw) lindb piUfe P-Beauv pltJgiopodill Hedw pulvinata (Hedw) Sm varbullpulvinata teretinervl Limpr
18
Orothera C Muell leana (Sull) C Muell
Rfwbdoweisia crispata (With) Undb
Cynodontium BSG ex Schimp alpestre (Wahlenb) Milde strumiferum (Hedw) Undb
Dichodontium Schimp pellucidum (Hedw) Schimp
Oncophorus (Slid) Brid vuens (Hedw) Brid wahlenbergii (Brid)
Dicranum Hedw bonjeanii DeNot ex Usa drummondii C Muell lagellare Hedw varj1ogellare fulvum Hook fuscescens Turn var fuscescens montanum Hedw muehlenbeckii BSG var muehlenbeckii polysetum Sw sabuletorum Ren et Card scoparium Hedw undulatum Brid viride (Sull et Lesq ex Sull) lindb
36 Leucobryaceae Leucobryum Hampe
glaucum (Hedw) Angstr ex Fr
37 Encalyptaceae Encalypta Hedw
ciliata Hedw var ciliata procera Bruch
38 Pottiaceae Astomum Hampe
muhlenbergianum (Sw) Grout
Weissia Hedw controversa Hedw var controversa
Gymnostomum Nees Homsch et Stunn aeruginosum Sm recurvirostrum Hedw var recurvirostrum
Trichostomum Bruch spirale Prout
19
40 Ephemeraceae Ephemerum Hampe
cohaerens (Hedw) Hampe crassineium (Schwaegr) Hampe val crassinervium spinulosum Bruch et Schimp ex Schimp
41 Funariaceae AphflllolThegma SuD
serratum (1 Hook et Wils ex Drumm) Sullo
Physcomitrium (Brid) Fuernr hookeri Hampe immersum Sullo pyriforme (Hedw) Hampe
Funaria Hedw americana Undb hygrometrica Hedw vaT hygrometrica
42 Splachnaceae Splachnum Hedw
ampullaceum Hedw
43 Schistostegaceae Schistostega Mohr
pennata (Hedw) Web el Mohr
44 Bryaceae Pohlia Hedw
annolina (Hedw) Lindb var annotina camea (Tum) Lindb cruda (Hedw) Undb elongata Hedw iliformis (Dicks) Andr lescurlana (Sull) Grout nutans (Hedw) Lindb obrusifolia (Brid) L Koch proligera (Undb ex Umpr) Undb ex Amell wahlenbergii (Web et Mohr) Andr
Leptobryum (B S G) Wits pyriforme (Hedw) Wils
Bryum Hedw algovicum Sendtnex C Muell argenteum Hedw caespiticium Hedw capillare Hedw var capillare creberrlmum Tayt lonchocaulon C Muell pallens (Brid) Sw ex Roehl pseudotrlquetrum (Hedw) Gaertn Meyer et Scherb varpudotriquetrum
20
pseudotriquetrum (Hedw) GaertnMeyer et Scherb var bimum (Schreb) Ulj stenotrichum C Muell var stenotrichum tortiolium Funck ex Brid uliginosum (Brid) B S G violaceum Crundw et Nyh weigelii Spreng
Rhodobryum (Schimp) Hampe roseum (Hedw) Limpr
45 Mniaceae Mnium Hedw
ciliare (C Muell) Schimp cinclidioides Hueb cuspidarum Hedw drummondii Bruch et Schimp longirostrum Brid lycopodioides Schwaegr marginatum (With) Brid ex P Beauv mediumB~S G var medium onhorrhynchum Brid pseudopunctatum Bruch et Schimp punctalUm Hedw var punctatum punctatum Hedw var elatum Schirnp rugicum Laur spinufosum B S G stellare Hedw
46 Aulacomruaceae Au1acomnium Schwaegr
androgynum (Hedw) Schwaegr heterostichum (Hedw) B S G paJustre (Hedw) Schwaegr var paJustre
47 Meesiaceae Amblyodon B S G
deillbotus (Hedw) B S G
Meesia Hedw triquetra (Richt) Angstr uJiginosa Hedw
48 Bartramiaceae Plagiopus Brid
oederiana (Sw) Umpr
Bartramia Hedw pomiformis Hedw var pomiformis
Philonotis Brid fontana (Hedw) Brid varfontana longseta (Michx) Britt varlongseta marchica (Hedw) Brid
21
~
~
49 Timmiaceae Timmia Hedw
megapolitana Hedw var megapolitana
50 Orthotrichaceae Zygodon Hook et Tayl
viridissimus (Dicks) Brid var rupestris lindb ex C J Hartm
Amphidium Schimp lapponicum (Hedw) Schimp mougeotii (B S G) Schimp
Orthotrichum Hedw anomalum Hedw obtuaifolium Brid ohioense Sullo et Lesq ex Aust pumilumSw sordidum Sullo et Lesq ex Aust speciosum Neesax Sturm var eegam (Schwaegr ex Hook et Grev) W~rnst stellDtwn Brid strrmgulatum P Beauv
Uleta Mohr ex Web cOfIICtata (p Beallv) Hamrn crispa (Hedw) Brid
Drummondill Hook ex Drumm prorepens (Hedw) Brltt
51 Fontinalaceae Fantinalis Hedw
antipyretica Hedw var antipyrettca antipyretica Hedw forma rabusta (Card) Card dalecarlica Schimp ex B S G var dalecarlica flqccidtJ Ren et Card hypnoides C J Hartrn var hypnoides hypnoides C J Hartm var durillei (Schimp) HU$ll
missourlcaCard novae-angliae Sullo var novae-angliae splulgrUlolill (C Moen) Wijket M~S
DichelY1l1ll Myr capilloceum (With) Myr falcatum (Bedw) Myr HIllescens B S G
52 Climaciaceae aimecium Web et Mohr ex Mohr
americllnum Brid dedroides (Hedw) Web et Mohr
22
S3 Hedwigiaceae Hedwigia P Beauv
ciliata (Hedw) P Beauv
54 Cryphaeaceae Forsstroemia lindh
ohioense (Sull) lindh
55 Leucodontaceae Leucodon Schwaegr
brachypus Blid var brachypus brachypus Brid var aniJrewsitlnus Crum et Anderson julaceus (Hedw) SuU
56 Neckeraceae Neckera Hedw
pennata Hedw var pennata
1fomalia (Brid) B S G trichomanoides (Hedw) B S G
57 Theliaceae Thelia Sull
asprellll Sullo hirtella (Hedw) Sullo lescurii SuU
Myurella B SG julacea (Schwaegr) B S G
sibrica (C MueU) Reim var sibrica
58 Fabroniaceae Fabronio Raddi
ciliaris (Brid) Bnd
Anacamptodon Brid splachnoides (Groet ex Brid) Brid
Schwetschkeopsis Broth fabronia (Schwaegr) Broth
59 Leskeaceae Lindbergia Kindb
brachyptera (Mitt) Kindb
Leskea Hedw gracilescens Hedw obscura Hedw poiycarpa Hedw
Leskeella (limpr) Loeske nervosa (Brid) Loeske
23
Pterlgy1lllndrum Hedw filiforme Hedw
60 Thuidiaceae Haplohymenfum Dozy et MoIk
trlste (Ces ex De Not) Kiodb
Anomodon Hook et Tayl attenuatus (Hedw) Hueb minor (Hedw) Fuemr rostratus (Hedw) Schimp viticulosus (Hedw) Hook et Tayl
Haplocladium (C Mucn) C MueU microphyllum (Hedw) Broth virginillnum (Brld) Broth ~bull
Thuidium B S G abietinum (Hedw) B S G delicatulum (Hedw) B S G vardelicatulum delicatulum (Hedw) B S G var radicans (Kindb) Crum Steere et Anderson minutulum (Hedw) B S G pygmaeum B S G recognftum (Hedw) Lindb scitum (p Beauv) Aust
Helodium (Sull) Warnst blandowli (Web et Mohr) Warnst var blandowli blandowli (Web et Mohr) Wamst var helodioides (Ren et Card ex Roell) Crum
Steere et Anderson paludomm (Sull) AuS
61 Amblystegiaceae Cratoneuron (Sull) Spruce
commulatum (Hedw) Roth var commutfltum commutatum (Hedw) Roth var fakatum (Brid) Moenk ilicinum (Hedw) Spruce var filcinum
Campylium (Sun) Mitt chrysophyllum (Brid) 1 Lange var chrysophyllum hispidulum (Brid) Mitt var hllJidulum polygamum (B S G)C lens varpolygtlmUm stellatum (Hedw) C1ens vat stelltum
Leptodictyum (Schimp) Warnst laxirete (Card et Ther) Broth riparium (Hedw) Wamat trichopodium (Schultz) Warns l1acillans (Sull) Broth
Hygroamblystegium Loelke jlul1iatile (Hedw) oeske var luViatile
24
fluviatile (Hedw) Loeske var oWltum Grout noterophilum (Sun et Lesq ex Sull) Warnst tenox (Hedw) Jenn var tenax tenox (Hedw) Jenn var spinilolium Schimp) Jenn
Amblystegium BSG juratzkanum Schimp serpens (Hedw) BSG varlum (Hedw) Undb var varium
Platydictya Berk confervoides (Brid) Crum jungermanniodes (Brid) Crum minutissimum (Sull et Lesq ex Sull) Crum subtile (Hedw) Crum
Drepanocladus (C Muell) Roth aduncus (Hedw) Warnst var aduncus aduncus (Hedw) Warnst var capillilolius (Wamst)Riebm aduncus (Hedw) Warnst var kneiffii (BSG) Moenk aduncus (Hedw) Warnst var polycorpus (Bland ex Voit) Roth exannulatus (BSG) Warnst fluitans (Hedw) Warnst var fluitans revolvens (Sw) Wamst var revolvens sendtneri (Schimp) Warnst var sendtnerl uncinatus (Hedw) Warnst var uncinatus uncinatus ffiedw) Warnst var piwnosus (Schimp) Wamst vernicoms (Lindb ex C Hartm) Warnst
Hygrohypnum Lindb eugyrium (BSG) loeske luridum (Hedw Jenn varluridum ochraecum (Tum ex Wils) Loeske var ochraceunt
Calliergon (Sull) Kindb cordilolium Hedw Kindb glganteum (Schimp) Kindb var giganteum richardsonii (Mitt) Kindb ex Wamst stramineum (Brid) Kindb trilarium (Web et Mohr) Kindb
Calliergonella Loeske cuspidata (Hedw) Loeske
Calliergidium (Ren) Grout bakeri (Ren) Grout
Scorpidium (Schimp) Urnpr scorpiodides (Hedw) limpr
62 Brachytbeciaceae Tomenthypnum loeske
nitens (Hedw) Loeske var mtens
25
Homalotheciella (Card) Broth subcapillata (Hedw) Broth
Brachythecium BSG acuminatum (Hedw) Awt acutum (Mitt) Sullo calcareum IGndb campestTe (C MuelL) BSG cunum (Lindb) Umpr digastrum C Muell et Kindb ex Macoun et Kindb oxycladon (Brid) Jaeg et Sauerb var oxycladon plumosum (Hedw) BSG populeum (Hedw) BSG var populeum reflexum (Starke ex Web et Mohr) DsG var relexum reiexum (Starke ex Web et Mohr) DSG var pacificum Renet Card ex Roell rlvulare BSG var rivulare tabulum (Hedw) BSC var tabuum salebrosum (Web et Mohr) BSG var salebrosum velutinum (Hedw) BSG Vat velutinum
Bryhniil Kaur graminicolor (Brid) Grout var graminicolor novae-angliae (Sull et Lesq ex sun) Grout
Rhynchostegium BSG senulatum (Hedw) Ja4lg et Sauerb
Rhynchostegiella (BSG) Limpr compacta (C MueU) Loeske
EU1hynchium 8SG hians (Hedw) Sande Lac pulchellum (Hedw) Jenn var pulchelium riparloides (Hedw) Rich
Stokeriella (Kindb) Robins pmelonga (Hedw) Robins var stokesii (Tum) Crum
63 Entodontaceae Entodon C Muell
brevisetus (Hook et Wils ex Wils) Lindb cladorrhizans (Hedw) C Muell seductrlx (Hedw) C MueU var wluctrix
Pleurozium Mitt schreberl (8rid) Mitt var schreberi
64 Plagiotheciaceae Pltlglothecium BSG
CflVifolium (Brid) Iwats var CtWifolium denticulatum (Hedw) 8SG laetum BSG ltztebricola BSG
26
6S SernatophyUaceae Brotherella Loeske ex Fleisch
recumins (Michx) Fleisch tenuirostris (Bruch et Schimp ex Sull) Fleisch
Sematophyllum Mitt demissum (Wils) Mitt
66 Hypnaceae Pl(itygyrium BSG
repens (Brid) BSG
PylaisieUa Kindb intricata (Hedw) Grout polyantha (Hedw) Grout selwynii (Kindb) Crum Steere et Anderson
Homomallium (Schimp) Loeske adnatum (Hedw) Broth
Hypnum Hedw cupressiforme Hedw var cupressiforme cupressifonne Hedw varfiliforme Brid curviolium Hedw fertile Sendtn imponens Hedw lindbergii Mitt palescens (Hedw) P-Beauv pratense Koch ex Brid
Calicladium Crum haldanianum (Grev) Crum
sopterygiopsis Iwats muelleriana (Schimp) Iwats
Isopterygium Mitt e1egans (Brid) lindb pulchellum (Hedw) Jaeg et Sauerb tenerum (Sw) Mitt
Taxiphyllum Fleisch depllQtum (Bruch et Schimp ex sun) Fleisch taxirameum (MitL) Fleisch
Herzogiella Broth seligeri (Brid) Iwats striatella (Brid) Iwats ruracea (Lindh) Iwats
hilium De Not crista-castrenna (Hedw) De Not
27
67 Rhytidiaceae Rhytidium (Sull) Kindb
rugosum (Hedw) Kindb
Rhytidfodelphus (Lindb ex Iimpr) Warnst triquetrus (Hedw) Warnst var triquetrus
68 Hylocomiaceae Hylocomium BSG
pyrenaicum (Spruce) lindb splendens (Hedw) BSG varsplendens
69 Buxbaumiaceae Buxbaumia Hedw
aphyllll Hedw
Diphyscium Mohr follosum (Hedw) Mohr
70 Tetraphidaceae Tetraphi3
pellucida Hedw
71 Polytrichaceae Atrichum PmiddotBeauv
angustatum (Brid) BSG cri8pum (James) Sullo oerstedianum (C Muell) Mitt teneUum (Roehl) BSG undulatum (Hedw) P middotBeauv var unduiatum
Pogonatum P-Beauv alpinum (Hedw) Roehl var alpinum umigerum (Hedw) P-Beauv
Poiytrichum Hedw commune Hedw var commune ormosum Hedw juniperinum Hedw longisetum Brid ohioense Ren et Card piliferum Hedw var pilierum strictum Brid
28
MAPS OF BRYOPHYTE TAXA
REPORTED FOR WISCONSIN
HORNWORTS AND LIVERWORTS
29
bull bull WSHeV~N S vE II
0 ~
e~RRON ~USK
(J~ (j -
CHIPPE A DUNN
0 o0 poundAU CLAIRE
~ TREMshyPEAshyLEAU
MICE
ONEIDA
Q LINCOLN
TAYLOR
Q t-~----- MOIRA THON
CLARK
o
10
LA FAVETTE
Hornwor ts amp Liverworts
- None - 1 - 25 taxa
- 26 - 50 tRXshy
- 51 - 75 tax)
- 7c - 100 taxa
FONDOU j LAC ~
DODGE
i O JEfFERSON
WALWORTH
30
Anastrophylltlfl helleranum (Nees) Schust Anastrophyllum michauxil (We~ Buch ~
46 88 136 141 Evans 38 46 141
Anastrophyllum mlnut~ (Schreb) Schuster Aneura plnguls (L) Durn
89 21 384687 88 89 90 (138) (145)
Anthoceros macounil MA Howe ~oceros punctatus L
21 (45) (144) 34
31
Barbilophozia attenuata (Mart) Loeske Barbilophozia barbata (Schmid ~ Schre~
24 141 Loeske 28 39 46 89 90 141
Bazzania trilobata (1) S Gray
1 26 39 46 (57) (59) 79 88 89
90 119 141
BlePbarostoma trichoffiyllum (1) Dumort
1 8 26 39 46 89 90 137 141
Blasia pusilla 1
1 39 46 90
Calypogeja fissa (1) Raddi
90
32
$129geja integristipula Steph C~Ypound2geja muelleriana (Schifm) K Mull
88 89 89
CallpoundO~eja neesiana (Mass amp Carest) K Mull
39 46 89 140 (141)
~pogeja sJlapound2icola (H Arnell tl J bull
Perss) Warnst et Loeske (119) 141
CeIalozia biscuspidata (L) Dum
1 37 I 46 90 t 137 141
CalYpogej~ trichomanis (L) Corda
26 39 (119) 137
33
CeEhalozia catenulata (Hub) Lindb Cephalozia connivens (Dicks) Lindb
39 46 79 141 39 4679 88 (89) 90 119 141
Cephalozia lacinulata Jack ~ Spruce
141 Cephalozia lunulifolia (Dum~ Dum I 28 39 46 79 88 89 90 141
Cepoundhalozia macouni (Aust) Aust Cephalozia ~leniceps (Aust) Lindb 27~ 3746 (66)7987 (139)141 39 46 79 89 141
34
Ceffialoziella 9ivaricata (Sm) Schiffn Cephaloziella ~hista (Jack) Schiffn
46 90 46 (119) 141
Cephaloziella hampeana (Nees) Schiffn
46
Cephaloziella rubella (Nees) Warnst
46 39
Cephaloziella rubella var sullivantii
(Aust) K Mull 46 39
Cephaloziella rubella var bifida (Lindb) Douin
35
46
Chiloscyphus pallescens (Ehrh ~ Hof~ Dum Chiloscyphus pallescens var fragilis
39J 46 88 89J 90 (Roth) K Milll 46
Chiloscyphus E9lyanthos (L) Corda
1 24 39 46
Chiloscyphus polyanthos vax rivularis
(Schrad) Nees 39 46
poundpoundnocepha1um conicum (1) Lindb
12 28 39 46 (49) 87 88 8990 120 121 148
Cololejeunea biddlecomiae (Aust) Evans
39 46 62 87 88 89 90
36
Dipl0]hyllum apicu1atum (Evans) Steph Fossombronia foveo1ata Lindb
(35)41 46 90 140 1 t 38 (39) t 46
Frullania bolanderi Aust
13 46 62 88 90 139 140
Frullanla ari ttoniae Evans
39 46 69 (71) 87 88 89 90
Frullania eboracensis Gott
1 9 28 4 39 46 (57) (59) 69 70 (71) 87 88 89 90 92 93 97 148
Frullania inflata Gott
46 87 88 89 (139)
Frullania oakesiana Aust Frullania plana Sullo
(35) 42 46 87 (139) (140) 92
Frullania ri paria Hampe ~ Lehm
46 69 71 87 89
Frullania ~elwynianna Pears
46 89
Geocalyx SEaveolens (Schrad) Nees
394688 8990 (119) 140
Frullania tamarisci (L) Dum subsp sPagrayana
(Mont) Hatt 28 34 l~6 t (57) 88 89 90
38
Mar pan thus seutatUB ( Web amp Mohr) Spruce Jamesoniella autumnalis (DC) Steph
(35) 39 46 90 1 7 28 39 46 56 (57) 87 8889 90 (119) 137 141
Jungermannia confertissima Nees
137 141
Jungermannia crenuliform1s Aust
79 137 141
Jungermannia ~il1ima Sm
46 90 141 Jungermannia exsert1folia subsp corifo~ (Dum~
Vana (35) 46 79 141
39
Jungermannia hyalina Lyell Jungerrr6nnia 1eiantha Grolle
40 46 79 136 141 39 46 79 88 90 1)7 141
Jungermannia IXlaxis Lindb Jungermannia pumila With
46 67 79 (139) (140) 141 )9 46 90 1)6 141
Jungermannia sphaerocarpa Hook Kurzia setacea (Web) Grolle
4679136140IIH 136
Lejeunea cavifo1ia (Ehrh) Lindb Lepidozia reptans (1) Dum
1 7 27 39 46 88 89 90 137 14010 I ( 27) 46 I 69 (71) 88
lophocolea heterophylla (Schrad) Dum Lophocolea minor Nees
1 )4 39 46 (57) 87 88 89 27 39 46 88 89 90 (119) 90 (93)94 119 136
Lophozia al~stris (Schleich ~ Web) Evans Lophozia ascendens (Warnst) Schust
46 90 137 141 136 140 141
41
loIilozia badensls (Gatt ~ Rabenh)
46 141
lophozia blcrenata (Schmid sectA Hoffm)
Dum 46 141
lophozia collaris (Nees) Dum
39 46 141
lophozla excisa (Dicks) Dum
27 4679 141
lophozia gillmani (Aust) Schust lophozia gIlttulata (Undo tl H Arnell)
36 39 40 46 136 (140) 141 Evans 39 46 136 141
42
141
Lophozia heterocolEDs (Thed) MA Howe IoPhozia ipcisa (Schard) Dum
27 39 46 89 141 8 27 31 46 79 90 140 141
LoQhozia longidens Macoun Lophozia ventricosa
1 46 79 90 141
Lophozia ventricosa var silvicola
(Buch) Jones 141 LoEhozla ventricosa vax longif1ora (Nees)
Macoun 39 46 79 141
43
~~ fragrans (Balbis) Frye et Clark Mannia pi 10sa (Hornem) Frye et Clark
46 (139) (140) 9 46 79 116 139
Mannia triandra (Scop) Gro 11e
46 (140)
Marchantia pound91ymorpha L
1 13 28 3946 (49) 78 87 88 89 90 (Ill) 121 122 147
Mo~rckia hibernica (HookJ Gott
46 89
Metzgeria conjugata Lindb
44
46
Mylia anomala (Hook) S Gray Notothylas orbicularis (Schwein) Sullo
39 46 141 (35)45 46
Nowellia curvifolia (Dicks) Mitt
17 27 3846 88 89 90 141
Odontoschisma denudatuJI1 (Nees ~ Mart)
Dum 46 81 137 141
Pallavicinia lyellii (Hook~ Carruth
90
Odontoschisma macounii (Aust) Under
45 46 (66) (139) 140 141
45
=
Pellia epiphllla (L) CordaPellia endiviifolia (Dicks~ Dum
27 39 46 90 13724 39 46
Pellia neesiana Limp bull Phaeoceros
39 46 89 27 39 45 46 97
Porella pinnata L
13 27 I 46 90 Plagiochila asplenioides (L) Dum
27 I 39 1+6 88 89 I 90 137
46
~e11a platYEhylla (L) Pfeiff Porella platyphylloi~ (Schwein) Lindb
17 27 39 46 69 (71)79 87 88 89 9 34 46 (57) 59 87 88 89 92 90 92 93 77
Preissia quadrata (Seop) Nees
1 36 3946 (71) 88 89 90 121 123
137
Ptllidium ciliare (L) Hampe
1 27 39 46 132 141
Dum
(22) 28 3946 (57) 87 88 89 90 92
Ptilidium ~lcherrium (G Web) Hampe
(9) 13 394656 (57)70 87 88 89 90 93 (119) 136 141
47
Radula obconica Sullo Reboulia hemlsphaerlca (L) Raddi
39 46 88 89 (43) 46 (59) 70 90 t Cf 124 125 (139)
Riccardia ~atifrons Lindb
1 27 39 46 89
Riccardia multifi~ (L) S Gray
39 46 88 145
Riccia beyrichiana Hampe ~ Lehm
46 Riccardia palmata (Hedw) Carruth
39 46 90
48
Riccia bifurca Hoffm Riccia cavernosa Hoffm
48 13739 46 79 137
Riccia fluitans L
242731454679
Riccla frostli Aust
48 79
Riccia sullicantii Aust
79 (137) Riccia sorocarpa Bisch 46 (137)
49
Ricciocarpus natans (1) Corda Sca~ia apiculata Spruce
13 27 46 79 89 110 39 46 (139) 141
ScaJltll1ia carinthica Jack ~ Lindb
137 t 141
Scapania ~ (Mart) Durn
39467990137141
Scapania digenil Schiffn
90
ScaEsectflia cuspiduligera (Nees) K Mull
46 (139) (140) (141)
50
Scapania glaucoeerha1a (Tayl) Aust Seapania gyrrmostomophilia Kaal
39 46 (139) 141 41 44 46 64 (139) (140) 141
ScaP3poundlia irrigua Gott ~ almiddot
7 39 46 79 137 (140L 141
Seapania nemorosa (L) Dum
1 39 46 89 90 137
Scapmi~ ~~ta Smiddotlch
137 141 51
~caPnia poundludicola LDeske amp K Mull Scapania saxicola Schust
46 (66) 141 136 141
Sc~~ scandica (H Arnell ~ Buch) Macv
139 140
Scapania subalpina (Nees) Durn
39 40 46 79 137 141
Scapania undulata (L) Dum
1 42 46 79 141
Scapania umbrosa (Schrad) Dum
137 140 141
52
Trichocolea tomentella (Ehrh) Dum Triomarla exsecta (Schmid) Schiffn
27 (35) 39 46 47 88 137 141 8 28 39 46 90 140 141
Tritomaria exsectiformis (Breidl) Loe~ke Tritomaria Sluinguedentata (Huds) Buch
39 -t6 89 90 136 140 141 7 39 46 141
Trltomaria scitula (Tayl) Joerg
46 136 (139) 141
53
Agn 2 Major Plant Communitiee of Wsconsln aaed on surwvor recorda bull co 18laquo) After Curtismiddot 1959
tit beol Forest e Phillie
Northernmiddot hardwood forest reg Pine 5cMInna
copy Southernmiddot hordwood forest CD Oak Savanna
10
SYSTEMATIC ARRANGEMENT OF WISCONSIN BRYOPHYTES
ANTHOCEROTAE (Hornworts)
1 Anthocerotaceae Anthoceros 1 emnd Prosk
macounii M A Howe punctatum 1
Phaeoceros Prosk laevis (1) Prosk subsp laevis
Notothylas Sul1 orbicularis (Schwein) Sullo
HEPATICAE (liverworts)
2 PseudolepicoJeaceae Blephllrostoma (Dum emend Undb) Dum
trlchophyllum (L) Dum subsp trlchophylluni
3 Ptilidiaceae Ptilidium Nees
ci1iare (L) Hampe pulcherrimum (G Web) Hampe
4 Trichocoleaceae Trichocolea Dum nom cons
tomentella (Ehrh) Dum
5 Lepidoziaceae Kunia von Martens
setacea (Web) GroUe
Lepidozia (Dum) Dum reptans (1) Dum
Bazzania S Gray trilobata (1) S Gray var triZobata
6 Calypogejaceae 011ypogeja Raddi
flSsa (L) Raddi subsp issa integristipula Steph muelleriana (Schiffn) K Mull subsp muelleriana neesiana (Mass et Carest) K Mull sphagnicola (H Arnell et J Perss) Warnst et loeske trichomanis (1) Corda
11
7 Cephaloziaceae Cepluzlozla (Dum emend Schiffn) Dum
biscwpidata (L) Dum subsp biclupidIIta catenulllttl (Hub) Lindb coMivens (Dicks) Lindb var connivens lacinultzta Jack ampX Spruce lunulifolilz (Dum) Dum macounii (Aust) Aust pleniceps (Aust) Lindh var pleniceps
Noweilia Mitt cunifolilz (Dicks) Mitt
8 Adelanthaceae Odontoschimuz (Dwn) Dum
denudatum (Nees ampX Mart) Dum Vir denudatum mtlCount (Aust) Underw
c
9 Cephaloziellaceae CephtlJoziellll (Spruce) Schiffn nom C()II$
divarlctzt4 (Sm) sclurfn val divarlcat(l elachhta (bdc) Schiffn 1uImpetuw (Nees) Schirfn IUbellll (Nees) Warnst vat IUbeIl4rube (Nees) Warnst var biid4 (Lindb) Dquinrube (Nees) WarDSt Var sulliwmtii (Aust) K MuD
10 Geo~c~ larptuttllus Nees
scutatus (Web et Mohr) Spruce
Geocalyx Nees grGVeolens (Schrad) Nees
11 Lophocoleaceae Lophocolea (Dum) Dum
heteropllyll4 (Schrad) Own minorNees
Chiloscypbus Corda om cons pallescens (Ehrh ampX Hoffm) Dum varfragllis (Roth) K Mun polyantitos (L) Corda var poiyanthos pol)llaquo1lthos (L) Corda var rlvulllris (Schrld) Nees
12 PlagiochiJaceae Plagio(hila Dum nom cons
tlBpienioides (L) Dum var asplenioidu
13 Jungermanniaceae Btriilophoril Loeske
attenutlta (Mart) Loeske bIubtlta (Schmid ampX Scltreb) Loeske
12
Lophom (Dum) Dum alpestris (Schleich ~ Web) Evans ascendens (Warnst) Schust badenais (Gott ex Rabenh) Schiffn var bademii bierenata (Schmid ex H)ffm) Dum eollam (Nees) Dum exci8a (Dicks) Dum var excisa gllimanii (Aust) Schust varglllmanii guttulata (Lindb et H Arnell) Evans heterocolpos (Thed) M A Howe var heterocolpos ineistl (Schrad) Dum var tnelsa longidens (Lindb) Macoun subsplongidens ventricosa (Dicks) Dum var ventricosa ventricolfl (Dicks) Dum varlontifIotrl (Nees) Macoun ventricosa (Dicks) Dum var silvlcola (Bueh) Jones
Tritomarill Schiffn ex Loeske exseeta (Schrad) Loeske exsecttformi$ (Breidl) Loeske subsp exsectiormfs quinquedentata (Huds) Buch var quinquedentata scitula (rayl) Joerg
Anastrophyllum (Spruce) Schiffn hellemnum (Noes) Schust michtzuxii (Web) Buch ex Evans minutum (Schrob) Schust var minutum
Mylill S Gray nom cons anomala (Hook) S Gray
Jamesoniella (Spruce) Carring autumnalis (DC) Steph var autumnalis
Jungemumnia L emend Dum eonfertissimtt Nees crenuliformis Aust exsertijolill Steph subsp cordifolia (Dum) Vana graciUima Sm hyalina Lyell leillnthll Grolle polaris Lindb pumila With sphaerocarpa Hook
14 Scapaniaceae Diplophyllum (Dum) Dum nom cons
apiculatum (Evans) Steph var apiculatum
Scapanill (Dum) Dum nom cons apkuata Spruce carinthillca Jack ex Lindb curta (Mart) Dum var curta
13
cuspiduligera (Nees) K Mull var cuspiduiigera degenii Schlffn ex K Mull var degenii glllucocephaJa (Tayt) Aust gymnostomophila Kaal irrigwJ (Nees) Gott et al subsp irrigua lingulata Buch varlingulata mucronata Buch subsp mucronata nemorosa (l) Dum paludicola Toeske et K Mun var paludicola paludosa (K Mull) K Mull saxicola Schust scandiea (H Arnell et Buch) Macv var scandiea 8Ubalp~ (Nees) Dum umbroSll (SchIad) Dum undulata (L) Dum var undulata
15 Radulaceae Raduia Dum nom COllS
compianata (L) Dum obconica Sull
16 Porellaceae Parella L
pinTlllta L patyphylla (L) Pfeiff platyphylloidt(l (Schwein) lindb
17 Jubulaceae FruUanill Raddi
bolanderi Aust brittoniae Evans eboracemis Gott inlllta Gott oakesiana Aust plana Sull rlparia Hampe ex Lehm selwyniana Pears taTnl1iJci (L) Dum subsp asagmyana (Mont) Hatt
18 lejeuneaceae Calolejeunea (Spruce) Schlffn
biddlecomiae (Aust) Evans
Lejeunea Ub nom cons cavifolia (Ehrh) Lindb emend Buch
19 Codoniaceae Fossombronia Raddi
foveolata Lindb
20 Pa11aviciniaceae Paklvieinlil S Gray nom cons
lyeiii (Hook) Carruth
14
Moerckia Gott hibemica (Hook) Gott
21 PeWaceae Pellfa Raddi nom cons propos
endivlifolfa (Dicks) Dwn epiphylllz (L) Corda neeBiana (GottlUmpr
22 Metzgeriaceae Metzgerla Raddi
conjugata Undb
23 Blasiacae B L
pulilm L
24 Aneurareae AneumDum
pinguis (L) Dum
Riccfl1dia S Gray nom cons latifrons Undb multiftda (L) S Gray var multifida palmata (Hedw) Carruth
25 Aytoniaceae Reboulill Raddi nom conI
hemllphaerica (L) Radeli
ManniaOpiz fragrrms (Balbis) Frye el ClarK pilosa (Hornem) Frye et Clark trltmdra (Scop) GroDe
26 Conocephalaceae Conocephalum Wigg nom cons
conicum (L) Lindb
27 Marchantiaceae Preiuia Corda
quadrata (Scop) Nees
Marchantia L polymorpha L
28 Ricciaceae Riccwcarpus Corda
natans (L) Corda
Riccia 1 beyrichiana Hampe ex Lehm bifurca Horfm
15
i
CtlJPemostI Hoffnt fuitu L rostii Aust BOIOCtI1fHI Bi~ sulltvtmtii Aust
MUSCI (Mosses)
29 Sphagnaceae SpIutgnum L
capillaceum (Weiss) Schrank var Cllpil1tlceum caplllllceum (Weiss) Vir tenellum (Schimp) Andr centrille C Jens ex Amen et C Jens compactummiddot OC ex Lam et Dc ClUpidatum Ehrh ex Hoffin var aupidatum jimbrltltum Wits ex J Hookmiddot fulCUm (Scbimp) KUngrr glrgenBOhIlll Russ imbrlctltum Hornsch ex Russ muelllmitil4m BOd 1IIIIU (RUII) C Jens przIutre L papIIIoIum Lindb pulchrum (Lindb ex braithw) Wamst recumun P-Beauv var recumun recum P-Beauv var tenue Klinggr riptlrlum Ansstr ruPQWItWamst squtl1fOlUm Crome subftltens Russ et Wamst ex Wamst IUbIeCfUldum Nees ex Sturm var subsecundum subsecundum Nees ex Sturm varrutem (Nees Hormch et sUUm) Hueb teres (Sdlimp) Anptr e C Hartm
WtlI7IItorjii Russ wuljfanum Gira
30 Andreaeaceae AndNrrerI Hedw
rupeitrlr Oedw var pest
31 Fissident~e Fluid_ lledw
trdiantoid Hedw byoidei Bedw var bryolda cristaM Wils ex Mitt exiguus sun VIr exigUw onttmUi (B-Pyl) Steud jtuberi Lesq et James obtusifolhu Wits var obtwifoliul ollPlUlldQldes Heel ~litSuU lubbtulkril Bed tfDdfoliul lledw llridubu (S~ ) Wahlenb vallirlIU1u
16
32 Ditrichaceae Pleuridium Brid
subulatum (Hedw) Rabenh
Ditrichum Hampe heteromallum (Hedw) Britt linellre (Sw) lindb
~ pallidum (Hedw) Hampe pusiUum (Hedw) Hampe
Saelania Lindb gIlucescens (Hedw) Bomanss et Broth
Ceratodon Brid purpureus (He(lw) Brid var prupurew
Distichium BSG capillaceum (Hedw) BSG inclinatum (Hedw) BSG
33 Bryoxipbiaceae Bryoxlphium Mitt
norvegicum (Brid) Mitt
34 Seligeriaceae Seligeria BSG
cumpylopoda Kindb ex Macoun et Kindb pusillll (Hedw) BSG recurvata (Hedw) BSG
Bindia BSG acuta (Hedw) BSG
35 Dicranaceae BlUehia Schwegr
lexUOSll (Sw ~ Schwaegr) C MueU sullivantii Aust
Trematodon Michx ambiguus (Hedw) Hornsch
Dicranellll (C Muell) Schimp cerviculata (Hedw) Schimp val cillPicuiata
heteromalla (Hedw) Schimp ~ rufescens (With) Schimp
schreberiana (Hedw) Schhnp var schreberiana schreberitJna (Hedw) Schimp var robusta Schimp ex Braithw subulata (Hedw) Schimp varia (Hedw) Schimp
Dicranodontium BSG denudatum (Brid) Britt ex Williams
Panzeucobryum (Lindh) Loeske Iongifolium (Hedw) oelke varlongifolium
17
Tortella (Undb) Limpr fragilis (Drumm) Umpr humilis (Hedw) JeM torlUoSfl (Hedw) Umpr
Hyophila Brid involuta (Hoole) Jaeg et Sauerb
Didymodon Hedw rigidulus Hedw
BryoerythrophyUum Chen recurvirostnon (Hedw) Chen
Btubula Hedw acula (Brid) Brid var acuta convoiuta Hedw fallox Hedw unguicultltll Hedw
Acaulon C MueD muticum (Uedw) C Muell var rufeseens (Iaeg) Crum
PJl~lm Hedw CUSPidtltum Hedw
De8mlltodon Brid cernuu (Hueb) BSG oblUsifoliw (Schwaegr) Schimp
T01IUla Hedw mueronifolia Schwaegr rurall (Hedw) Gaertn Meyer et Scherb var ruralis
39 Grimmiaceae Grlmmill Hedw
alpicola Hedw var alpicola alpicola Hedw var dupretii (Ther) Crum alpicola Hedw var rlvularl8 (Brid) Wahlenb apocarpa Hedw var apocarpa apocarpa Hedw var ambipa (Sull) Jonea apocarpa Hedw var conjerttl (Funck) Spreng apocarpa Hedw var strleta (Tum) Hook et Tayl CiIIyptrrlta Hook ex Drumm donniana Sm var donn_ hemumti Crum ltlevla (Brid) Brid montana BSG oineyi SuD oVllll8 (Hedw) lindb piUfe P-Beauv pltJgiopodill Hedw pulvinata (Hedw) Sm varbullpulvinata teretinervl Limpr
18
Orothera C Muell leana (Sull) C Muell
Rfwbdoweisia crispata (With) Undb
Cynodontium BSG ex Schimp alpestre (Wahlenb) Milde strumiferum (Hedw) Undb
Dichodontium Schimp pellucidum (Hedw) Schimp
Oncophorus (Slid) Brid vuens (Hedw) Brid wahlenbergii (Brid)
Dicranum Hedw bonjeanii DeNot ex Usa drummondii C Muell lagellare Hedw varj1ogellare fulvum Hook fuscescens Turn var fuscescens montanum Hedw muehlenbeckii BSG var muehlenbeckii polysetum Sw sabuletorum Ren et Card scoparium Hedw undulatum Brid viride (Sull et Lesq ex Sull) lindb
36 Leucobryaceae Leucobryum Hampe
glaucum (Hedw) Angstr ex Fr
37 Encalyptaceae Encalypta Hedw
ciliata Hedw var ciliata procera Bruch
38 Pottiaceae Astomum Hampe
muhlenbergianum (Sw) Grout
Weissia Hedw controversa Hedw var controversa
Gymnostomum Nees Homsch et Stunn aeruginosum Sm recurvirostrum Hedw var recurvirostrum
Trichostomum Bruch spirale Prout
19
40 Ephemeraceae Ephemerum Hampe
cohaerens (Hedw) Hampe crassineium (Schwaegr) Hampe val crassinervium spinulosum Bruch et Schimp ex Schimp
41 Funariaceae AphflllolThegma SuD
serratum (1 Hook et Wils ex Drumm) Sullo
Physcomitrium (Brid) Fuernr hookeri Hampe immersum Sullo pyriforme (Hedw) Hampe
Funaria Hedw americana Undb hygrometrica Hedw vaT hygrometrica
42 Splachnaceae Splachnum Hedw
ampullaceum Hedw
43 Schistostegaceae Schistostega Mohr
pennata (Hedw) Web el Mohr
44 Bryaceae Pohlia Hedw
annolina (Hedw) Lindb var annotina camea (Tum) Lindb cruda (Hedw) Undb elongata Hedw iliformis (Dicks) Andr lescurlana (Sull) Grout nutans (Hedw) Lindb obrusifolia (Brid) L Koch proligera (Undb ex Umpr) Undb ex Amell wahlenbergii (Web et Mohr) Andr
Leptobryum (B S G) Wits pyriforme (Hedw) Wils
Bryum Hedw algovicum Sendtnex C Muell argenteum Hedw caespiticium Hedw capillare Hedw var capillare creberrlmum Tayt lonchocaulon C Muell pallens (Brid) Sw ex Roehl pseudotrlquetrum (Hedw) Gaertn Meyer et Scherb varpudotriquetrum
20
pseudotriquetrum (Hedw) GaertnMeyer et Scherb var bimum (Schreb) Ulj stenotrichum C Muell var stenotrichum tortiolium Funck ex Brid uliginosum (Brid) B S G violaceum Crundw et Nyh weigelii Spreng
Rhodobryum (Schimp) Hampe roseum (Hedw) Limpr
45 Mniaceae Mnium Hedw
ciliare (C Muell) Schimp cinclidioides Hueb cuspidarum Hedw drummondii Bruch et Schimp longirostrum Brid lycopodioides Schwaegr marginatum (With) Brid ex P Beauv mediumB~S G var medium onhorrhynchum Brid pseudopunctatum Bruch et Schimp punctalUm Hedw var punctatum punctatum Hedw var elatum Schirnp rugicum Laur spinufosum B S G stellare Hedw
46 Aulacomruaceae Au1acomnium Schwaegr
androgynum (Hedw) Schwaegr heterostichum (Hedw) B S G paJustre (Hedw) Schwaegr var paJustre
47 Meesiaceae Amblyodon B S G
deillbotus (Hedw) B S G
Meesia Hedw triquetra (Richt) Angstr uJiginosa Hedw
48 Bartramiaceae Plagiopus Brid
oederiana (Sw) Umpr
Bartramia Hedw pomiformis Hedw var pomiformis
Philonotis Brid fontana (Hedw) Brid varfontana longseta (Michx) Britt varlongseta marchica (Hedw) Brid
21
~
~
49 Timmiaceae Timmia Hedw
megapolitana Hedw var megapolitana
50 Orthotrichaceae Zygodon Hook et Tayl
viridissimus (Dicks) Brid var rupestris lindb ex C J Hartm
Amphidium Schimp lapponicum (Hedw) Schimp mougeotii (B S G) Schimp
Orthotrichum Hedw anomalum Hedw obtuaifolium Brid ohioense Sullo et Lesq ex Aust pumilumSw sordidum Sullo et Lesq ex Aust speciosum Neesax Sturm var eegam (Schwaegr ex Hook et Grev) W~rnst stellDtwn Brid strrmgulatum P Beauv
Uleta Mohr ex Web cOfIICtata (p Beallv) Hamrn crispa (Hedw) Brid
Drummondill Hook ex Drumm prorepens (Hedw) Brltt
51 Fontinalaceae Fantinalis Hedw
antipyretica Hedw var antipyrettca antipyretica Hedw forma rabusta (Card) Card dalecarlica Schimp ex B S G var dalecarlica flqccidtJ Ren et Card hypnoides C J Hartrn var hypnoides hypnoides C J Hartm var durillei (Schimp) HU$ll
missourlcaCard novae-angliae Sullo var novae-angliae splulgrUlolill (C Moen) Wijket M~S
DichelY1l1ll Myr capilloceum (With) Myr falcatum (Bedw) Myr HIllescens B S G
52 Climaciaceae aimecium Web et Mohr ex Mohr
americllnum Brid dedroides (Hedw) Web et Mohr
22
S3 Hedwigiaceae Hedwigia P Beauv
ciliata (Hedw) P Beauv
54 Cryphaeaceae Forsstroemia lindh
ohioense (Sull) lindh
55 Leucodontaceae Leucodon Schwaegr
brachypus Blid var brachypus brachypus Brid var aniJrewsitlnus Crum et Anderson julaceus (Hedw) SuU
56 Neckeraceae Neckera Hedw
pennata Hedw var pennata
1fomalia (Brid) B S G trichomanoides (Hedw) B S G
57 Theliaceae Thelia Sull
asprellll Sullo hirtella (Hedw) Sullo lescurii SuU
Myurella B SG julacea (Schwaegr) B S G
sibrica (C MueU) Reim var sibrica
58 Fabroniaceae Fabronio Raddi
ciliaris (Brid) Bnd
Anacamptodon Brid splachnoides (Groet ex Brid) Brid
Schwetschkeopsis Broth fabronia (Schwaegr) Broth
59 Leskeaceae Lindbergia Kindb
brachyptera (Mitt) Kindb
Leskea Hedw gracilescens Hedw obscura Hedw poiycarpa Hedw
Leskeella (limpr) Loeske nervosa (Brid) Loeske
23
Pterlgy1lllndrum Hedw filiforme Hedw
60 Thuidiaceae Haplohymenfum Dozy et MoIk
trlste (Ces ex De Not) Kiodb
Anomodon Hook et Tayl attenuatus (Hedw) Hueb minor (Hedw) Fuemr rostratus (Hedw) Schimp viticulosus (Hedw) Hook et Tayl
Haplocladium (C Mucn) C MueU microphyllum (Hedw) Broth virginillnum (Brld) Broth ~bull
Thuidium B S G abietinum (Hedw) B S G delicatulum (Hedw) B S G vardelicatulum delicatulum (Hedw) B S G var radicans (Kindb) Crum Steere et Anderson minutulum (Hedw) B S G pygmaeum B S G recognftum (Hedw) Lindb scitum (p Beauv) Aust
Helodium (Sull) Warnst blandowli (Web et Mohr) Warnst var blandowli blandowli (Web et Mohr) Wamst var helodioides (Ren et Card ex Roell) Crum
Steere et Anderson paludomm (Sull) AuS
61 Amblystegiaceae Cratoneuron (Sull) Spruce
commulatum (Hedw) Roth var commutfltum commutatum (Hedw) Roth var fakatum (Brid) Moenk ilicinum (Hedw) Spruce var filcinum
Campylium (Sun) Mitt chrysophyllum (Brid) 1 Lange var chrysophyllum hispidulum (Brid) Mitt var hllJidulum polygamum (B S G)C lens varpolygtlmUm stellatum (Hedw) C1ens vat stelltum
Leptodictyum (Schimp) Warnst laxirete (Card et Ther) Broth riparium (Hedw) Wamat trichopodium (Schultz) Warns l1acillans (Sull) Broth
Hygroamblystegium Loelke jlul1iatile (Hedw) oeske var luViatile
24
fluviatile (Hedw) Loeske var oWltum Grout noterophilum (Sun et Lesq ex Sull) Warnst tenox (Hedw) Jenn var tenax tenox (Hedw) Jenn var spinilolium Schimp) Jenn
Amblystegium BSG juratzkanum Schimp serpens (Hedw) BSG varlum (Hedw) Undb var varium
Platydictya Berk confervoides (Brid) Crum jungermanniodes (Brid) Crum minutissimum (Sull et Lesq ex Sull) Crum subtile (Hedw) Crum
Drepanocladus (C Muell) Roth aduncus (Hedw) Warnst var aduncus aduncus (Hedw) Warnst var capillilolius (Wamst)Riebm aduncus (Hedw) Warnst var kneiffii (BSG) Moenk aduncus (Hedw) Warnst var polycorpus (Bland ex Voit) Roth exannulatus (BSG) Warnst fluitans (Hedw) Warnst var fluitans revolvens (Sw) Wamst var revolvens sendtneri (Schimp) Warnst var sendtnerl uncinatus (Hedw) Warnst var uncinatus uncinatus ffiedw) Warnst var piwnosus (Schimp) Wamst vernicoms (Lindb ex C Hartm) Warnst
Hygrohypnum Lindb eugyrium (BSG) loeske luridum (Hedw Jenn varluridum ochraecum (Tum ex Wils) Loeske var ochraceunt
Calliergon (Sull) Kindb cordilolium Hedw Kindb glganteum (Schimp) Kindb var giganteum richardsonii (Mitt) Kindb ex Wamst stramineum (Brid) Kindb trilarium (Web et Mohr) Kindb
Calliergonella Loeske cuspidata (Hedw) Loeske
Calliergidium (Ren) Grout bakeri (Ren) Grout
Scorpidium (Schimp) Urnpr scorpiodides (Hedw) limpr
62 Brachytbeciaceae Tomenthypnum loeske
nitens (Hedw) Loeske var mtens
25
Homalotheciella (Card) Broth subcapillata (Hedw) Broth
Brachythecium BSG acuminatum (Hedw) Awt acutum (Mitt) Sullo calcareum IGndb campestTe (C MuelL) BSG cunum (Lindb) Umpr digastrum C Muell et Kindb ex Macoun et Kindb oxycladon (Brid) Jaeg et Sauerb var oxycladon plumosum (Hedw) BSG populeum (Hedw) BSG var populeum reflexum (Starke ex Web et Mohr) DsG var relexum reiexum (Starke ex Web et Mohr) DSG var pacificum Renet Card ex Roell rlvulare BSG var rivulare tabulum (Hedw) BSC var tabuum salebrosum (Web et Mohr) BSG var salebrosum velutinum (Hedw) BSG Vat velutinum
Bryhniil Kaur graminicolor (Brid) Grout var graminicolor novae-angliae (Sull et Lesq ex sun) Grout
Rhynchostegium BSG senulatum (Hedw) Ja4lg et Sauerb
Rhynchostegiella (BSG) Limpr compacta (C MueU) Loeske
EU1hynchium 8SG hians (Hedw) Sande Lac pulchellum (Hedw) Jenn var pulchelium riparloides (Hedw) Rich
Stokeriella (Kindb) Robins pmelonga (Hedw) Robins var stokesii (Tum) Crum
63 Entodontaceae Entodon C Muell
brevisetus (Hook et Wils ex Wils) Lindb cladorrhizans (Hedw) C Muell seductrlx (Hedw) C MueU var wluctrix
Pleurozium Mitt schreberl (8rid) Mitt var schreberi
64 Plagiotheciaceae Pltlglothecium BSG
CflVifolium (Brid) Iwats var CtWifolium denticulatum (Hedw) 8SG laetum BSG ltztebricola BSG
26
6S SernatophyUaceae Brotherella Loeske ex Fleisch
recumins (Michx) Fleisch tenuirostris (Bruch et Schimp ex Sull) Fleisch
Sematophyllum Mitt demissum (Wils) Mitt
66 Hypnaceae Pl(itygyrium BSG
repens (Brid) BSG
PylaisieUa Kindb intricata (Hedw) Grout polyantha (Hedw) Grout selwynii (Kindb) Crum Steere et Anderson
Homomallium (Schimp) Loeske adnatum (Hedw) Broth
Hypnum Hedw cupressiforme Hedw var cupressiforme cupressifonne Hedw varfiliforme Brid curviolium Hedw fertile Sendtn imponens Hedw lindbergii Mitt palescens (Hedw) P-Beauv pratense Koch ex Brid
Calicladium Crum haldanianum (Grev) Crum
sopterygiopsis Iwats muelleriana (Schimp) Iwats
Isopterygium Mitt e1egans (Brid) lindb pulchellum (Hedw) Jaeg et Sauerb tenerum (Sw) Mitt
Taxiphyllum Fleisch depllQtum (Bruch et Schimp ex sun) Fleisch taxirameum (MitL) Fleisch
Herzogiella Broth seligeri (Brid) Iwats striatella (Brid) Iwats ruracea (Lindh) Iwats
hilium De Not crista-castrenna (Hedw) De Not
27
67 Rhytidiaceae Rhytidium (Sull) Kindb
rugosum (Hedw) Kindb
Rhytidfodelphus (Lindb ex Iimpr) Warnst triquetrus (Hedw) Warnst var triquetrus
68 Hylocomiaceae Hylocomium BSG
pyrenaicum (Spruce) lindb splendens (Hedw) BSG varsplendens
69 Buxbaumiaceae Buxbaumia Hedw
aphyllll Hedw
Diphyscium Mohr follosum (Hedw) Mohr
70 Tetraphidaceae Tetraphi3
pellucida Hedw
71 Polytrichaceae Atrichum PmiddotBeauv
angustatum (Brid) BSG cri8pum (James) Sullo oerstedianum (C Muell) Mitt teneUum (Roehl) BSG undulatum (Hedw) P middotBeauv var unduiatum
Pogonatum P-Beauv alpinum (Hedw) Roehl var alpinum umigerum (Hedw) P-Beauv
Poiytrichum Hedw commune Hedw var commune ormosum Hedw juniperinum Hedw longisetum Brid ohioense Ren et Card piliferum Hedw var pilierum strictum Brid
28
MAPS OF BRYOPHYTE TAXA
REPORTED FOR WISCONSIN
HORNWORTS AND LIVERWORTS
29
bull bull WSHeV~N S vE II
0 ~
e~RRON ~USK
(J~ (j -
CHIPPE A DUNN
0 o0 poundAU CLAIRE
~ TREMshyPEAshyLEAU
MICE
ONEIDA
Q LINCOLN
TAYLOR
Q t-~----- MOIRA THON
CLARK
o
10
LA FAVETTE
Hornwor ts amp Liverworts
- None - 1 - 25 taxa
- 26 - 50 tRXshy
- 51 - 75 tax)
- 7c - 100 taxa
FONDOU j LAC ~
DODGE
i O JEfFERSON
WALWORTH
30
Anastrophylltlfl helleranum (Nees) Schust Anastrophyllum michauxil (We~ Buch ~
46 88 136 141 Evans 38 46 141
Anastrophyllum mlnut~ (Schreb) Schuster Aneura plnguls (L) Durn
89 21 384687 88 89 90 (138) (145)
Anthoceros macounil MA Howe ~oceros punctatus L
21 (45) (144) 34
31
Barbilophozia attenuata (Mart) Loeske Barbilophozia barbata (Schmid ~ Schre~
24 141 Loeske 28 39 46 89 90 141
Bazzania trilobata (1) S Gray
1 26 39 46 (57) (59) 79 88 89
90 119 141
BlePbarostoma trichoffiyllum (1) Dumort
1 8 26 39 46 89 90 137 141
Blasia pusilla 1
1 39 46 90
Calypogeja fissa (1) Raddi
90
32
$129geja integristipula Steph C~Ypound2geja muelleriana (Schifm) K Mull
88 89 89
CallpoundO~eja neesiana (Mass amp Carest) K Mull
39 46 89 140 (141)
~pogeja sJlapound2icola (H Arnell tl J bull
Perss) Warnst et Loeske (119) 141
CeIalozia biscuspidata (L) Dum
1 37 I 46 90 t 137 141
CalYpogej~ trichomanis (L) Corda
26 39 (119) 137
33
CeEhalozia catenulata (Hub) Lindb Cephalozia connivens (Dicks) Lindb
39 46 79 141 39 4679 88 (89) 90 119 141
Cephalozia lacinulata Jack ~ Spruce
141 Cephalozia lunulifolia (Dum~ Dum I 28 39 46 79 88 89 90 141
Cepoundhalozia macouni (Aust) Aust Cephalozia ~leniceps (Aust) Lindb 27~ 3746 (66)7987 (139)141 39 46 79 89 141
34
Ceffialoziella 9ivaricata (Sm) Schiffn Cephaloziella ~hista (Jack) Schiffn
46 90 46 (119) 141
Cephaloziella hampeana (Nees) Schiffn
46
Cephaloziella rubella (Nees) Warnst
46 39
Cephaloziella rubella var sullivantii
(Aust) K Mull 46 39
Cephaloziella rubella var bifida (Lindb) Douin
35
46
Chiloscyphus pallescens (Ehrh ~ Hof~ Dum Chiloscyphus pallescens var fragilis
39J 46 88 89J 90 (Roth) K Milll 46
Chiloscyphus E9lyanthos (L) Corda
1 24 39 46
Chiloscyphus polyanthos vax rivularis
(Schrad) Nees 39 46
poundpoundnocepha1um conicum (1) Lindb
12 28 39 46 (49) 87 88 8990 120 121 148
Cololejeunea biddlecomiae (Aust) Evans
39 46 62 87 88 89 90
36
Dipl0]hyllum apicu1atum (Evans) Steph Fossombronia foveo1ata Lindb
(35)41 46 90 140 1 t 38 (39) t 46
Frullania bolanderi Aust
13 46 62 88 90 139 140
Frullanla ari ttoniae Evans
39 46 69 (71) 87 88 89 90
Frullania eboracensis Gott
1 9 28 4 39 46 (57) (59) 69 70 (71) 87 88 89 90 92 93 97 148
Frullania inflata Gott
46 87 88 89 (139)
Frullania oakesiana Aust Frullania plana Sullo
(35) 42 46 87 (139) (140) 92
Frullania ri paria Hampe ~ Lehm
46 69 71 87 89
Frullania ~elwynianna Pears
46 89
Geocalyx SEaveolens (Schrad) Nees
394688 8990 (119) 140
Frullania tamarisci (L) Dum subsp sPagrayana
(Mont) Hatt 28 34 l~6 t (57) 88 89 90
38
Mar pan thus seutatUB ( Web amp Mohr) Spruce Jamesoniella autumnalis (DC) Steph
(35) 39 46 90 1 7 28 39 46 56 (57) 87 8889 90 (119) 137 141
Jungermannia confertissima Nees
137 141
Jungermannia crenuliform1s Aust
79 137 141
Jungermannia ~il1ima Sm
46 90 141 Jungermannia exsert1folia subsp corifo~ (Dum~
Vana (35) 46 79 141
39
Jungermannia hyalina Lyell Jungerrr6nnia 1eiantha Grolle
40 46 79 136 141 39 46 79 88 90 1)7 141
Jungermannia IXlaxis Lindb Jungermannia pumila With
46 67 79 (139) (140) 141 )9 46 90 1)6 141
Jungermannia sphaerocarpa Hook Kurzia setacea (Web) Grolle
4679136140IIH 136
Lejeunea cavifo1ia (Ehrh) Lindb Lepidozia reptans (1) Dum
1 7 27 39 46 88 89 90 137 14010 I ( 27) 46 I 69 (71) 88
lophocolea heterophylla (Schrad) Dum Lophocolea minor Nees
1 )4 39 46 (57) 87 88 89 27 39 46 88 89 90 (119) 90 (93)94 119 136
Lophozia al~stris (Schleich ~ Web) Evans Lophozia ascendens (Warnst) Schust
46 90 137 141 136 140 141
41
loIilozia badensls (Gatt ~ Rabenh)
46 141
lophozia blcrenata (Schmid sectA Hoffm)
Dum 46 141
lophozia collaris (Nees) Dum
39 46 141
lophozla excisa (Dicks) Dum
27 4679 141
lophozia gillmani (Aust) Schust lophozia gIlttulata (Undo tl H Arnell)
36 39 40 46 136 (140) 141 Evans 39 46 136 141
42
141
Lophozia heterocolEDs (Thed) MA Howe IoPhozia ipcisa (Schard) Dum
27 39 46 89 141 8 27 31 46 79 90 140 141
LoQhozia longidens Macoun Lophozia ventricosa
1 46 79 90 141
Lophozia ventricosa var silvicola
(Buch) Jones 141 LoEhozla ventricosa vax longif1ora (Nees)
Macoun 39 46 79 141
43
~~ fragrans (Balbis) Frye et Clark Mannia pi 10sa (Hornem) Frye et Clark
46 (139) (140) 9 46 79 116 139
Mannia triandra (Scop) Gro 11e
46 (140)
Marchantia pound91ymorpha L
1 13 28 3946 (49) 78 87 88 89 90 (Ill) 121 122 147
Mo~rckia hibernica (HookJ Gott
46 89
Metzgeria conjugata Lindb
44
46
Mylia anomala (Hook) S Gray Notothylas orbicularis (Schwein) Sullo
39 46 141 (35)45 46
Nowellia curvifolia (Dicks) Mitt
17 27 3846 88 89 90 141
Odontoschisma denudatuJI1 (Nees ~ Mart)
Dum 46 81 137 141
Pallavicinia lyellii (Hook~ Carruth
90
Odontoschisma macounii (Aust) Under
45 46 (66) (139) 140 141
45
=
Pellia epiphllla (L) CordaPellia endiviifolia (Dicks~ Dum
27 39 46 90 13724 39 46
Pellia neesiana Limp bull Phaeoceros
39 46 89 27 39 45 46 97
Porella pinnata L
13 27 I 46 90 Plagiochila asplenioides (L) Dum
27 I 39 1+6 88 89 I 90 137
46
~e11a platYEhylla (L) Pfeiff Porella platyphylloi~ (Schwein) Lindb
17 27 39 46 69 (71)79 87 88 89 9 34 46 (57) 59 87 88 89 92 90 92 93 77
Preissia quadrata (Seop) Nees
1 36 3946 (71) 88 89 90 121 123
137
Ptllidium ciliare (L) Hampe
1 27 39 46 132 141
Dum
(22) 28 3946 (57) 87 88 89 90 92
Ptilidium ~lcherrium (G Web) Hampe
(9) 13 394656 (57)70 87 88 89 90 93 (119) 136 141
47
Radula obconica Sullo Reboulia hemlsphaerlca (L) Raddi
39 46 88 89 (43) 46 (59) 70 90 t Cf 124 125 (139)
Riccardia ~atifrons Lindb
1 27 39 46 89
Riccardia multifi~ (L) S Gray
39 46 88 145
Riccia beyrichiana Hampe ~ Lehm
46 Riccardia palmata (Hedw) Carruth
39 46 90
48
Riccia bifurca Hoffm Riccia cavernosa Hoffm
48 13739 46 79 137
Riccia fluitans L
242731454679
Riccla frostli Aust
48 79
Riccia sullicantii Aust
79 (137) Riccia sorocarpa Bisch 46 (137)
49
Ricciocarpus natans (1) Corda Sca~ia apiculata Spruce
13 27 46 79 89 110 39 46 (139) 141
ScaJltll1ia carinthica Jack ~ Lindb
137 t 141
Scapania ~ (Mart) Durn
39467990137141
Scapania digenil Schiffn
90
ScaEsectflia cuspiduligera (Nees) K Mull
46 (139) (140) (141)
50
Scapania glaucoeerha1a (Tayl) Aust Seapania gyrrmostomophilia Kaal
39 46 (139) 141 41 44 46 64 (139) (140) 141
ScaP3poundlia irrigua Gott ~ almiddot
7 39 46 79 137 (140L 141
Seapania nemorosa (L) Dum
1 39 46 89 90 137
Scapmi~ ~~ta Smiddotlch
137 141 51
~caPnia poundludicola LDeske amp K Mull Scapania saxicola Schust
46 (66) 141 136 141
Sc~~ scandica (H Arnell ~ Buch) Macv
139 140
Scapania subalpina (Nees) Durn
39 40 46 79 137 141
Scapania undulata (L) Dum
1 42 46 79 141
Scapania umbrosa (Schrad) Dum
137 140 141
52
Trichocolea tomentella (Ehrh) Dum Triomarla exsecta (Schmid) Schiffn
27 (35) 39 46 47 88 137 141 8 28 39 46 90 140 141
Tritomaria exsectiformis (Breidl) Loe~ke Tritomaria Sluinguedentata (Huds) Buch
39 -t6 89 90 136 140 141 7 39 46 141
Trltomaria scitula (Tayl) Joerg
46 136 (139) 141
53
SYSTEMATIC ARRANGEMENT OF WISCONSIN BRYOPHYTES
ANTHOCEROTAE (Hornworts)
1 Anthocerotaceae Anthoceros 1 emnd Prosk
macounii M A Howe punctatum 1
Phaeoceros Prosk laevis (1) Prosk subsp laevis
Notothylas Sul1 orbicularis (Schwein) Sullo
HEPATICAE (liverworts)
2 PseudolepicoJeaceae Blephllrostoma (Dum emend Undb) Dum
trlchophyllum (L) Dum subsp trlchophylluni
3 Ptilidiaceae Ptilidium Nees
ci1iare (L) Hampe pulcherrimum (G Web) Hampe
4 Trichocoleaceae Trichocolea Dum nom cons
tomentella (Ehrh) Dum
5 Lepidoziaceae Kunia von Martens
setacea (Web) GroUe
Lepidozia (Dum) Dum reptans (1) Dum
Bazzania S Gray trilobata (1) S Gray var triZobata
6 Calypogejaceae 011ypogeja Raddi
flSsa (L) Raddi subsp issa integristipula Steph muelleriana (Schiffn) K Mull subsp muelleriana neesiana (Mass et Carest) K Mull sphagnicola (H Arnell et J Perss) Warnst et loeske trichomanis (1) Corda
11
7 Cephaloziaceae Cepluzlozla (Dum emend Schiffn) Dum
biscwpidata (L) Dum subsp biclupidIIta catenulllttl (Hub) Lindb coMivens (Dicks) Lindb var connivens lacinultzta Jack ampX Spruce lunulifolilz (Dum) Dum macounii (Aust) Aust pleniceps (Aust) Lindh var pleniceps
Noweilia Mitt cunifolilz (Dicks) Mitt
8 Adelanthaceae Odontoschimuz (Dwn) Dum
denudatum (Nees ampX Mart) Dum Vir denudatum mtlCount (Aust) Underw
c
9 Cephaloziellaceae CephtlJoziellll (Spruce) Schiffn nom C()II$
divarlctzt4 (Sm) sclurfn val divarlcat(l elachhta (bdc) Schiffn 1uImpetuw (Nees) Schirfn IUbellll (Nees) Warnst vat IUbeIl4rube (Nees) Warnst var biid4 (Lindb) Dquinrube (Nees) WarDSt Var sulliwmtii (Aust) K MuD
10 Geo~c~ larptuttllus Nees
scutatus (Web et Mohr) Spruce
Geocalyx Nees grGVeolens (Schrad) Nees
11 Lophocoleaceae Lophocolea (Dum) Dum
heteropllyll4 (Schrad) Own minorNees
Chiloscypbus Corda om cons pallescens (Ehrh ampX Hoffm) Dum varfragllis (Roth) K Mun polyantitos (L) Corda var poiyanthos pol)llaquo1lthos (L) Corda var rlvulllris (Schrld) Nees
12 PlagiochiJaceae Plagio(hila Dum nom cons
tlBpienioides (L) Dum var asplenioidu
13 Jungermanniaceae Btriilophoril Loeske
attenutlta (Mart) Loeske bIubtlta (Schmid ampX Scltreb) Loeske
12
Lophom (Dum) Dum alpestris (Schleich ~ Web) Evans ascendens (Warnst) Schust badenais (Gott ex Rabenh) Schiffn var bademii bierenata (Schmid ex H)ffm) Dum eollam (Nees) Dum exci8a (Dicks) Dum var excisa gllimanii (Aust) Schust varglllmanii guttulata (Lindb et H Arnell) Evans heterocolpos (Thed) M A Howe var heterocolpos ineistl (Schrad) Dum var tnelsa longidens (Lindb) Macoun subsplongidens ventricosa (Dicks) Dum var ventricosa ventricolfl (Dicks) Dum varlontifIotrl (Nees) Macoun ventricosa (Dicks) Dum var silvlcola (Bueh) Jones
Tritomarill Schiffn ex Loeske exseeta (Schrad) Loeske exsecttformi$ (Breidl) Loeske subsp exsectiormfs quinquedentata (Huds) Buch var quinquedentata scitula (rayl) Joerg
Anastrophyllum (Spruce) Schiffn hellemnum (Noes) Schust michtzuxii (Web) Buch ex Evans minutum (Schrob) Schust var minutum
Mylill S Gray nom cons anomala (Hook) S Gray
Jamesoniella (Spruce) Carring autumnalis (DC) Steph var autumnalis
Jungemumnia L emend Dum eonfertissimtt Nees crenuliformis Aust exsertijolill Steph subsp cordifolia (Dum) Vana graciUima Sm hyalina Lyell leillnthll Grolle polaris Lindb pumila With sphaerocarpa Hook
14 Scapaniaceae Diplophyllum (Dum) Dum nom cons
apiculatum (Evans) Steph var apiculatum
Scapanill (Dum) Dum nom cons apkuata Spruce carinthillca Jack ex Lindb curta (Mart) Dum var curta
13
cuspiduligera (Nees) K Mull var cuspiduiigera degenii Schlffn ex K Mull var degenii glllucocephaJa (Tayt) Aust gymnostomophila Kaal irrigwJ (Nees) Gott et al subsp irrigua lingulata Buch varlingulata mucronata Buch subsp mucronata nemorosa (l) Dum paludicola Toeske et K Mun var paludicola paludosa (K Mull) K Mull saxicola Schust scandiea (H Arnell et Buch) Macv var scandiea 8Ubalp~ (Nees) Dum umbroSll (SchIad) Dum undulata (L) Dum var undulata
15 Radulaceae Raduia Dum nom COllS
compianata (L) Dum obconica Sull
16 Porellaceae Parella L
pinTlllta L patyphylla (L) Pfeiff platyphylloidt(l (Schwein) lindb
17 Jubulaceae FruUanill Raddi
bolanderi Aust brittoniae Evans eboracemis Gott inlllta Gott oakesiana Aust plana Sull rlparia Hampe ex Lehm selwyniana Pears taTnl1iJci (L) Dum subsp asagmyana (Mont) Hatt
18 lejeuneaceae Calolejeunea (Spruce) Schlffn
biddlecomiae (Aust) Evans
Lejeunea Ub nom cons cavifolia (Ehrh) Lindb emend Buch
19 Codoniaceae Fossombronia Raddi
foveolata Lindb
20 Pa11aviciniaceae Paklvieinlil S Gray nom cons
lyeiii (Hook) Carruth
14
Moerckia Gott hibemica (Hook) Gott
21 PeWaceae Pellfa Raddi nom cons propos
endivlifolfa (Dicks) Dwn epiphylllz (L) Corda neeBiana (GottlUmpr
22 Metzgeriaceae Metzgerla Raddi
conjugata Undb
23 Blasiacae B L
pulilm L
24 Aneurareae AneumDum
pinguis (L) Dum
Riccfl1dia S Gray nom cons latifrons Undb multiftda (L) S Gray var multifida palmata (Hedw) Carruth
25 Aytoniaceae Reboulill Raddi nom conI
hemllphaerica (L) Radeli
ManniaOpiz fragrrms (Balbis) Frye el ClarK pilosa (Hornem) Frye et Clark trltmdra (Scop) GroDe
26 Conocephalaceae Conocephalum Wigg nom cons
conicum (L) Lindb
27 Marchantiaceae Preiuia Corda
quadrata (Scop) Nees
Marchantia L polymorpha L
28 Ricciaceae Riccwcarpus Corda
natans (L) Corda
Riccia 1 beyrichiana Hampe ex Lehm bifurca Horfm
15
i
CtlJPemostI Hoffnt fuitu L rostii Aust BOIOCtI1fHI Bi~ sulltvtmtii Aust
MUSCI (Mosses)
29 Sphagnaceae SpIutgnum L
capillaceum (Weiss) Schrank var Cllpil1tlceum caplllllceum (Weiss) Vir tenellum (Schimp) Andr centrille C Jens ex Amen et C Jens compactummiddot OC ex Lam et Dc ClUpidatum Ehrh ex Hoffin var aupidatum jimbrltltum Wits ex J Hookmiddot fulCUm (Scbimp) KUngrr glrgenBOhIlll Russ imbrlctltum Hornsch ex Russ muelllmitil4m BOd 1IIIIU (RUII) C Jens przIutre L papIIIoIum Lindb pulchrum (Lindb ex braithw) Wamst recumun P-Beauv var recumun recum P-Beauv var tenue Klinggr riptlrlum Ansstr ruPQWItWamst squtl1fOlUm Crome subftltens Russ et Wamst ex Wamst IUbIeCfUldum Nees ex Sturm var subsecundum subsecundum Nees ex Sturm varrutem (Nees Hormch et sUUm) Hueb teres (Sdlimp) Anptr e C Hartm
WtlI7IItorjii Russ wuljfanum Gira
30 Andreaeaceae AndNrrerI Hedw
rupeitrlr Oedw var pest
31 Fissident~e Fluid_ lledw
trdiantoid Hedw byoidei Bedw var bryolda cristaM Wils ex Mitt exiguus sun VIr exigUw onttmUi (B-Pyl) Steud jtuberi Lesq et James obtusifolhu Wits var obtwifoliul ollPlUlldQldes Heel ~litSuU lubbtulkril Bed tfDdfoliul lledw llridubu (S~ ) Wahlenb vallirlIU1u
16
32 Ditrichaceae Pleuridium Brid
subulatum (Hedw) Rabenh
Ditrichum Hampe heteromallum (Hedw) Britt linellre (Sw) lindb
~ pallidum (Hedw) Hampe pusiUum (Hedw) Hampe
Saelania Lindb gIlucescens (Hedw) Bomanss et Broth
Ceratodon Brid purpureus (He(lw) Brid var prupurew
Distichium BSG capillaceum (Hedw) BSG inclinatum (Hedw) BSG
33 Bryoxipbiaceae Bryoxlphium Mitt
norvegicum (Brid) Mitt
34 Seligeriaceae Seligeria BSG
cumpylopoda Kindb ex Macoun et Kindb pusillll (Hedw) BSG recurvata (Hedw) BSG
Bindia BSG acuta (Hedw) BSG
35 Dicranaceae BlUehia Schwegr
lexUOSll (Sw ~ Schwaegr) C MueU sullivantii Aust
Trematodon Michx ambiguus (Hedw) Hornsch
Dicranellll (C Muell) Schimp cerviculata (Hedw) Schimp val cillPicuiata
heteromalla (Hedw) Schimp ~ rufescens (With) Schimp
schreberiana (Hedw) Schhnp var schreberiana schreberitJna (Hedw) Schimp var robusta Schimp ex Braithw subulata (Hedw) Schimp varia (Hedw) Schimp
Dicranodontium BSG denudatum (Brid) Britt ex Williams
Panzeucobryum (Lindh) Loeske Iongifolium (Hedw) oelke varlongifolium
17
Tortella (Undb) Limpr fragilis (Drumm) Umpr humilis (Hedw) JeM torlUoSfl (Hedw) Umpr
Hyophila Brid involuta (Hoole) Jaeg et Sauerb
Didymodon Hedw rigidulus Hedw
BryoerythrophyUum Chen recurvirostnon (Hedw) Chen
Btubula Hedw acula (Brid) Brid var acuta convoiuta Hedw fallox Hedw unguicultltll Hedw
Acaulon C MueD muticum (Uedw) C Muell var rufeseens (Iaeg) Crum
PJl~lm Hedw CUSPidtltum Hedw
De8mlltodon Brid cernuu (Hueb) BSG oblUsifoliw (Schwaegr) Schimp
T01IUla Hedw mueronifolia Schwaegr rurall (Hedw) Gaertn Meyer et Scherb var ruralis
39 Grimmiaceae Grlmmill Hedw
alpicola Hedw var alpicola alpicola Hedw var dupretii (Ther) Crum alpicola Hedw var rlvularl8 (Brid) Wahlenb apocarpa Hedw var apocarpa apocarpa Hedw var ambipa (Sull) Jonea apocarpa Hedw var conjerttl (Funck) Spreng apocarpa Hedw var strleta (Tum) Hook et Tayl CiIIyptrrlta Hook ex Drumm donniana Sm var donn_ hemumti Crum ltlevla (Brid) Brid montana BSG oineyi SuD oVllll8 (Hedw) lindb piUfe P-Beauv pltJgiopodill Hedw pulvinata (Hedw) Sm varbullpulvinata teretinervl Limpr
18
Orothera C Muell leana (Sull) C Muell
Rfwbdoweisia crispata (With) Undb
Cynodontium BSG ex Schimp alpestre (Wahlenb) Milde strumiferum (Hedw) Undb
Dichodontium Schimp pellucidum (Hedw) Schimp
Oncophorus (Slid) Brid vuens (Hedw) Brid wahlenbergii (Brid)
Dicranum Hedw bonjeanii DeNot ex Usa drummondii C Muell lagellare Hedw varj1ogellare fulvum Hook fuscescens Turn var fuscescens montanum Hedw muehlenbeckii BSG var muehlenbeckii polysetum Sw sabuletorum Ren et Card scoparium Hedw undulatum Brid viride (Sull et Lesq ex Sull) lindb
36 Leucobryaceae Leucobryum Hampe
glaucum (Hedw) Angstr ex Fr
37 Encalyptaceae Encalypta Hedw
ciliata Hedw var ciliata procera Bruch
38 Pottiaceae Astomum Hampe
muhlenbergianum (Sw) Grout
Weissia Hedw controversa Hedw var controversa
Gymnostomum Nees Homsch et Stunn aeruginosum Sm recurvirostrum Hedw var recurvirostrum
Trichostomum Bruch spirale Prout
19
40 Ephemeraceae Ephemerum Hampe
cohaerens (Hedw) Hampe crassineium (Schwaegr) Hampe val crassinervium spinulosum Bruch et Schimp ex Schimp
41 Funariaceae AphflllolThegma SuD
serratum (1 Hook et Wils ex Drumm) Sullo
Physcomitrium (Brid) Fuernr hookeri Hampe immersum Sullo pyriforme (Hedw) Hampe
Funaria Hedw americana Undb hygrometrica Hedw vaT hygrometrica
42 Splachnaceae Splachnum Hedw
ampullaceum Hedw
43 Schistostegaceae Schistostega Mohr
pennata (Hedw) Web el Mohr
44 Bryaceae Pohlia Hedw
annolina (Hedw) Lindb var annotina camea (Tum) Lindb cruda (Hedw) Undb elongata Hedw iliformis (Dicks) Andr lescurlana (Sull) Grout nutans (Hedw) Lindb obrusifolia (Brid) L Koch proligera (Undb ex Umpr) Undb ex Amell wahlenbergii (Web et Mohr) Andr
Leptobryum (B S G) Wits pyriforme (Hedw) Wils
Bryum Hedw algovicum Sendtnex C Muell argenteum Hedw caespiticium Hedw capillare Hedw var capillare creberrlmum Tayt lonchocaulon C Muell pallens (Brid) Sw ex Roehl pseudotrlquetrum (Hedw) Gaertn Meyer et Scherb varpudotriquetrum
20
pseudotriquetrum (Hedw) GaertnMeyer et Scherb var bimum (Schreb) Ulj stenotrichum C Muell var stenotrichum tortiolium Funck ex Brid uliginosum (Brid) B S G violaceum Crundw et Nyh weigelii Spreng
Rhodobryum (Schimp) Hampe roseum (Hedw) Limpr
45 Mniaceae Mnium Hedw
ciliare (C Muell) Schimp cinclidioides Hueb cuspidarum Hedw drummondii Bruch et Schimp longirostrum Brid lycopodioides Schwaegr marginatum (With) Brid ex P Beauv mediumB~S G var medium onhorrhynchum Brid pseudopunctatum Bruch et Schimp punctalUm Hedw var punctatum punctatum Hedw var elatum Schirnp rugicum Laur spinufosum B S G stellare Hedw
46 Aulacomruaceae Au1acomnium Schwaegr
androgynum (Hedw) Schwaegr heterostichum (Hedw) B S G paJustre (Hedw) Schwaegr var paJustre
47 Meesiaceae Amblyodon B S G
deillbotus (Hedw) B S G
Meesia Hedw triquetra (Richt) Angstr uJiginosa Hedw
48 Bartramiaceae Plagiopus Brid
oederiana (Sw) Umpr
Bartramia Hedw pomiformis Hedw var pomiformis
Philonotis Brid fontana (Hedw) Brid varfontana longseta (Michx) Britt varlongseta marchica (Hedw) Brid
21
~
~
49 Timmiaceae Timmia Hedw
megapolitana Hedw var megapolitana
50 Orthotrichaceae Zygodon Hook et Tayl
viridissimus (Dicks) Brid var rupestris lindb ex C J Hartm
Amphidium Schimp lapponicum (Hedw) Schimp mougeotii (B S G) Schimp
Orthotrichum Hedw anomalum Hedw obtuaifolium Brid ohioense Sullo et Lesq ex Aust pumilumSw sordidum Sullo et Lesq ex Aust speciosum Neesax Sturm var eegam (Schwaegr ex Hook et Grev) W~rnst stellDtwn Brid strrmgulatum P Beauv
Uleta Mohr ex Web cOfIICtata (p Beallv) Hamrn crispa (Hedw) Brid
Drummondill Hook ex Drumm prorepens (Hedw) Brltt
51 Fontinalaceae Fantinalis Hedw
antipyretica Hedw var antipyrettca antipyretica Hedw forma rabusta (Card) Card dalecarlica Schimp ex B S G var dalecarlica flqccidtJ Ren et Card hypnoides C J Hartrn var hypnoides hypnoides C J Hartm var durillei (Schimp) HU$ll
missourlcaCard novae-angliae Sullo var novae-angliae splulgrUlolill (C Moen) Wijket M~S
DichelY1l1ll Myr capilloceum (With) Myr falcatum (Bedw) Myr HIllescens B S G
52 Climaciaceae aimecium Web et Mohr ex Mohr
americllnum Brid dedroides (Hedw) Web et Mohr
22
S3 Hedwigiaceae Hedwigia P Beauv
ciliata (Hedw) P Beauv
54 Cryphaeaceae Forsstroemia lindh
ohioense (Sull) lindh
55 Leucodontaceae Leucodon Schwaegr
brachypus Blid var brachypus brachypus Brid var aniJrewsitlnus Crum et Anderson julaceus (Hedw) SuU
56 Neckeraceae Neckera Hedw
pennata Hedw var pennata
1fomalia (Brid) B S G trichomanoides (Hedw) B S G
57 Theliaceae Thelia Sull
asprellll Sullo hirtella (Hedw) Sullo lescurii SuU
Myurella B SG julacea (Schwaegr) B S G
sibrica (C MueU) Reim var sibrica
58 Fabroniaceae Fabronio Raddi
ciliaris (Brid) Bnd
Anacamptodon Brid splachnoides (Groet ex Brid) Brid
Schwetschkeopsis Broth fabronia (Schwaegr) Broth
59 Leskeaceae Lindbergia Kindb
brachyptera (Mitt) Kindb
Leskea Hedw gracilescens Hedw obscura Hedw poiycarpa Hedw
Leskeella (limpr) Loeske nervosa (Brid) Loeske
23
Pterlgy1lllndrum Hedw filiforme Hedw
60 Thuidiaceae Haplohymenfum Dozy et MoIk
trlste (Ces ex De Not) Kiodb
Anomodon Hook et Tayl attenuatus (Hedw) Hueb minor (Hedw) Fuemr rostratus (Hedw) Schimp viticulosus (Hedw) Hook et Tayl
Haplocladium (C Mucn) C MueU microphyllum (Hedw) Broth virginillnum (Brld) Broth ~bull
Thuidium B S G abietinum (Hedw) B S G delicatulum (Hedw) B S G vardelicatulum delicatulum (Hedw) B S G var radicans (Kindb) Crum Steere et Anderson minutulum (Hedw) B S G pygmaeum B S G recognftum (Hedw) Lindb scitum (p Beauv) Aust
Helodium (Sull) Warnst blandowli (Web et Mohr) Warnst var blandowli blandowli (Web et Mohr) Wamst var helodioides (Ren et Card ex Roell) Crum
Steere et Anderson paludomm (Sull) AuS
61 Amblystegiaceae Cratoneuron (Sull) Spruce
commulatum (Hedw) Roth var commutfltum commutatum (Hedw) Roth var fakatum (Brid) Moenk ilicinum (Hedw) Spruce var filcinum
Campylium (Sun) Mitt chrysophyllum (Brid) 1 Lange var chrysophyllum hispidulum (Brid) Mitt var hllJidulum polygamum (B S G)C lens varpolygtlmUm stellatum (Hedw) C1ens vat stelltum
Leptodictyum (Schimp) Warnst laxirete (Card et Ther) Broth riparium (Hedw) Wamat trichopodium (Schultz) Warns l1acillans (Sull) Broth
Hygroamblystegium Loelke jlul1iatile (Hedw) oeske var luViatile
24
fluviatile (Hedw) Loeske var oWltum Grout noterophilum (Sun et Lesq ex Sull) Warnst tenox (Hedw) Jenn var tenax tenox (Hedw) Jenn var spinilolium Schimp) Jenn
Amblystegium BSG juratzkanum Schimp serpens (Hedw) BSG varlum (Hedw) Undb var varium
Platydictya Berk confervoides (Brid) Crum jungermanniodes (Brid) Crum minutissimum (Sull et Lesq ex Sull) Crum subtile (Hedw) Crum
Drepanocladus (C Muell) Roth aduncus (Hedw) Warnst var aduncus aduncus (Hedw) Warnst var capillilolius (Wamst)Riebm aduncus (Hedw) Warnst var kneiffii (BSG) Moenk aduncus (Hedw) Warnst var polycorpus (Bland ex Voit) Roth exannulatus (BSG) Warnst fluitans (Hedw) Warnst var fluitans revolvens (Sw) Wamst var revolvens sendtneri (Schimp) Warnst var sendtnerl uncinatus (Hedw) Warnst var uncinatus uncinatus ffiedw) Warnst var piwnosus (Schimp) Wamst vernicoms (Lindb ex C Hartm) Warnst
Hygrohypnum Lindb eugyrium (BSG) loeske luridum (Hedw Jenn varluridum ochraecum (Tum ex Wils) Loeske var ochraceunt
Calliergon (Sull) Kindb cordilolium Hedw Kindb glganteum (Schimp) Kindb var giganteum richardsonii (Mitt) Kindb ex Wamst stramineum (Brid) Kindb trilarium (Web et Mohr) Kindb
Calliergonella Loeske cuspidata (Hedw) Loeske
Calliergidium (Ren) Grout bakeri (Ren) Grout
Scorpidium (Schimp) Urnpr scorpiodides (Hedw) limpr
62 Brachytbeciaceae Tomenthypnum loeske
nitens (Hedw) Loeske var mtens
25
Homalotheciella (Card) Broth subcapillata (Hedw) Broth
Brachythecium BSG acuminatum (Hedw) Awt acutum (Mitt) Sullo calcareum IGndb campestTe (C MuelL) BSG cunum (Lindb) Umpr digastrum C Muell et Kindb ex Macoun et Kindb oxycladon (Brid) Jaeg et Sauerb var oxycladon plumosum (Hedw) BSG populeum (Hedw) BSG var populeum reflexum (Starke ex Web et Mohr) DsG var relexum reiexum (Starke ex Web et Mohr) DSG var pacificum Renet Card ex Roell rlvulare BSG var rivulare tabulum (Hedw) BSC var tabuum salebrosum (Web et Mohr) BSG var salebrosum velutinum (Hedw) BSG Vat velutinum
Bryhniil Kaur graminicolor (Brid) Grout var graminicolor novae-angliae (Sull et Lesq ex sun) Grout
Rhynchostegium BSG senulatum (Hedw) Ja4lg et Sauerb
Rhynchostegiella (BSG) Limpr compacta (C MueU) Loeske
EU1hynchium 8SG hians (Hedw) Sande Lac pulchellum (Hedw) Jenn var pulchelium riparloides (Hedw) Rich
Stokeriella (Kindb) Robins pmelonga (Hedw) Robins var stokesii (Tum) Crum
63 Entodontaceae Entodon C Muell
brevisetus (Hook et Wils ex Wils) Lindb cladorrhizans (Hedw) C Muell seductrlx (Hedw) C MueU var wluctrix
Pleurozium Mitt schreberl (8rid) Mitt var schreberi
64 Plagiotheciaceae Pltlglothecium BSG
CflVifolium (Brid) Iwats var CtWifolium denticulatum (Hedw) 8SG laetum BSG ltztebricola BSG
26
6S SernatophyUaceae Brotherella Loeske ex Fleisch
recumins (Michx) Fleisch tenuirostris (Bruch et Schimp ex Sull) Fleisch
Sematophyllum Mitt demissum (Wils) Mitt
66 Hypnaceae Pl(itygyrium BSG
repens (Brid) BSG
PylaisieUa Kindb intricata (Hedw) Grout polyantha (Hedw) Grout selwynii (Kindb) Crum Steere et Anderson
Homomallium (Schimp) Loeske adnatum (Hedw) Broth
Hypnum Hedw cupressiforme Hedw var cupressiforme cupressifonne Hedw varfiliforme Brid curviolium Hedw fertile Sendtn imponens Hedw lindbergii Mitt palescens (Hedw) P-Beauv pratense Koch ex Brid
Calicladium Crum haldanianum (Grev) Crum
sopterygiopsis Iwats muelleriana (Schimp) Iwats
Isopterygium Mitt e1egans (Brid) lindb pulchellum (Hedw) Jaeg et Sauerb tenerum (Sw) Mitt
Taxiphyllum Fleisch depllQtum (Bruch et Schimp ex sun) Fleisch taxirameum (MitL) Fleisch
Herzogiella Broth seligeri (Brid) Iwats striatella (Brid) Iwats ruracea (Lindh) Iwats
hilium De Not crista-castrenna (Hedw) De Not
27
67 Rhytidiaceae Rhytidium (Sull) Kindb
rugosum (Hedw) Kindb
Rhytidfodelphus (Lindb ex Iimpr) Warnst triquetrus (Hedw) Warnst var triquetrus
68 Hylocomiaceae Hylocomium BSG
pyrenaicum (Spruce) lindb splendens (Hedw) BSG varsplendens
69 Buxbaumiaceae Buxbaumia Hedw
aphyllll Hedw
Diphyscium Mohr follosum (Hedw) Mohr
70 Tetraphidaceae Tetraphi3
pellucida Hedw
71 Polytrichaceae Atrichum PmiddotBeauv
angustatum (Brid) BSG cri8pum (James) Sullo oerstedianum (C Muell) Mitt teneUum (Roehl) BSG undulatum (Hedw) P middotBeauv var unduiatum
Pogonatum P-Beauv alpinum (Hedw) Roehl var alpinum umigerum (Hedw) P-Beauv
Poiytrichum Hedw commune Hedw var commune ormosum Hedw juniperinum Hedw longisetum Brid ohioense Ren et Card piliferum Hedw var pilierum strictum Brid
28
MAPS OF BRYOPHYTE TAXA
REPORTED FOR WISCONSIN
HORNWORTS AND LIVERWORTS
29
bull bull WSHeV~N S vE II
0 ~
e~RRON ~USK
(J~ (j -
CHIPPE A DUNN
0 o0 poundAU CLAIRE
~ TREMshyPEAshyLEAU
MICE
ONEIDA
Q LINCOLN
TAYLOR
Q t-~----- MOIRA THON
CLARK
o
10
LA FAVETTE
Hornwor ts amp Liverworts
- None - 1 - 25 taxa
- 26 - 50 tRXshy
- 51 - 75 tax)
- 7c - 100 taxa
FONDOU j LAC ~
DODGE
i O JEfFERSON
WALWORTH
30
Anastrophylltlfl helleranum (Nees) Schust Anastrophyllum michauxil (We~ Buch ~
46 88 136 141 Evans 38 46 141
Anastrophyllum mlnut~ (Schreb) Schuster Aneura plnguls (L) Durn
89 21 384687 88 89 90 (138) (145)
Anthoceros macounil MA Howe ~oceros punctatus L
21 (45) (144) 34
31
Barbilophozia attenuata (Mart) Loeske Barbilophozia barbata (Schmid ~ Schre~
24 141 Loeske 28 39 46 89 90 141
Bazzania trilobata (1) S Gray
1 26 39 46 (57) (59) 79 88 89
90 119 141
BlePbarostoma trichoffiyllum (1) Dumort
1 8 26 39 46 89 90 137 141
Blasia pusilla 1
1 39 46 90
Calypogeja fissa (1) Raddi
90
32
$129geja integristipula Steph C~Ypound2geja muelleriana (Schifm) K Mull
88 89 89
CallpoundO~eja neesiana (Mass amp Carest) K Mull
39 46 89 140 (141)
~pogeja sJlapound2icola (H Arnell tl J bull
Perss) Warnst et Loeske (119) 141
CeIalozia biscuspidata (L) Dum
1 37 I 46 90 t 137 141
CalYpogej~ trichomanis (L) Corda
26 39 (119) 137
33
CeEhalozia catenulata (Hub) Lindb Cephalozia connivens (Dicks) Lindb
39 46 79 141 39 4679 88 (89) 90 119 141
Cephalozia lacinulata Jack ~ Spruce
141 Cephalozia lunulifolia (Dum~ Dum I 28 39 46 79 88 89 90 141
Cepoundhalozia macouni (Aust) Aust Cephalozia ~leniceps (Aust) Lindb 27~ 3746 (66)7987 (139)141 39 46 79 89 141
34
Ceffialoziella 9ivaricata (Sm) Schiffn Cephaloziella ~hista (Jack) Schiffn
46 90 46 (119) 141
Cephaloziella hampeana (Nees) Schiffn
46
Cephaloziella rubella (Nees) Warnst
46 39
Cephaloziella rubella var sullivantii
(Aust) K Mull 46 39
Cephaloziella rubella var bifida (Lindb) Douin
35
46
Chiloscyphus pallescens (Ehrh ~ Hof~ Dum Chiloscyphus pallescens var fragilis
39J 46 88 89J 90 (Roth) K Milll 46
Chiloscyphus E9lyanthos (L) Corda
1 24 39 46
Chiloscyphus polyanthos vax rivularis
(Schrad) Nees 39 46
poundpoundnocepha1um conicum (1) Lindb
12 28 39 46 (49) 87 88 8990 120 121 148
Cololejeunea biddlecomiae (Aust) Evans
39 46 62 87 88 89 90
36
Dipl0]hyllum apicu1atum (Evans) Steph Fossombronia foveo1ata Lindb
(35)41 46 90 140 1 t 38 (39) t 46
Frullania bolanderi Aust
13 46 62 88 90 139 140
Frullanla ari ttoniae Evans
39 46 69 (71) 87 88 89 90
Frullania eboracensis Gott
1 9 28 4 39 46 (57) (59) 69 70 (71) 87 88 89 90 92 93 97 148
Frullania inflata Gott
46 87 88 89 (139)
Frullania oakesiana Aust Frullania plana Sullo
(35) 42 46 87 (139) (140) 92
Frullania ri paria Hampe ~ Lehm
46 69 71 87 89
Frullania ~elwynianna Pears
46 89
Geocalyx SEaveolens (Schrad) Nees
394688 8990 (119) 140
Frullania tamarisci (L) Dum subsp sPagrayana
(Mont) Hatt 28 34 l~6 t (57) 88 89 90
38
Mar pan thus seutatUB ( Web amp Mohr) Spruce Jamesoniella autumnalis (DC) Steph
(35) 39 46 90 1 7 28 39 46 56 (57) 87 8889 90 (119) 137 141
Jungermannia confertissima Nees
137 141
Jungermannia crenuliform1s Aust
79 137 141
Jungermannia ~il1ima Sm
46 90 141 Jungermannia exsert1folia subsp corifo~ (Dum~
Vana (35) 46 79 141
39
Jungermannia hyalina Lyell Jungerrr6nnia 1eiantha Grolle
40 46 79 136 141 39 46 79 88 90 1)7 141
Jungermannia IXlaxis Lindb Jungermannia pumila With
46 67 79 (139) (140) 141 )9 46 90 1)6 141
Jungermannia sphaerocarpa Hook Kurzia setacea (Web) Grolle
4679136140IIH 136
Lejeunea cavifo1ia (Ehrh) Lindb Lepidozia reptans (1) Dum
1 7 27 39 46 88 89 90 137 14010 I ( 27) 46 I 69 (71) 88
lophocolea heterophylla (Schrad) Dum Lophocolea minor Nees
1 )4 39 46 (57) 87 88 89 27 39 46 88 89 90 (119) 90 (93)94 119 136
Lophozia al~stris (Schleich ~ Web) Evans Lophozia ascendens (Warnst) Schust
46 90 137 141 136 140 141
41
loIilozia badensls (Gatt ~ Rabenh)
46 141
lophozia blcrenata (Schmid sectA Hoffm)
Dum 46 141
lophozia collaris (Nees) Dum
39 46 141
lophozla excisa (Dicks) Dum
27 4679 141
lophozia gillmani (Aust) Schust lophozia gIlttulata (Undo tl H Arnell)
36 39 40 46 136 (140) 141 Evans 39 46 136 141
42
141
Lophozia heterocolEDs (Thed) MA Howe IoPhozia ipcisa (Schard) Dum
27 39 46 89 141 8 27 31 46 79 90 140 141
LoQhozia longidens Macoun Lophozia ventricosa
1 46 79 90 141
Lophozia ventricosa var silvicola
(Buch) Jones 141 LoEhozla ventricosa vax longif1ora (Nees)
Macoun 39 46 79 141
43
~~ fragrans (Balbis) Frye et Clark Mannia pi 10sa (Hornem) Frye et Clark
46 (139) (140) 9 46 79 116 139
Mannia triandra (Scop) Gro 11e
46 (140)
Marchantia pound91ymorpha L
1 13 28 3946 (49) 78 87 88 89 90 (Ill) 121 122 147
Mo~rckia hibernica (HookJ Gott
46 89
Metzgeria conjugata Lindb
44
46
Mylia anomala (Hook) S Gray Notothylas orbicularis (Schwein) Sullo
39 46 141 (35)45 46
Nowellia curvifolia (Dicks) Mitt
17 27 3846 88 89 90 141
Odontoschisma denudatuJI1 (Nees ~ Mart)
Dum 46 81 137 141
Pallavicinia lyellii (Hook~ Carruth
90
Odontoschisma macounii (Aust) Under
45 46 (66) (139) 140 141
45
=
Pellia epiphllla (L) CordaPellia endiviifolia (Dicks~ Dum
27 39 46 90 13724 39 46
Pellia neesiana Limp bull Phaeoceros
39 46 89 27 39 45 46 97
Porella pinnata L
13 27 I 46 90 Plagiochila asplenioides (L) Dum
27 I 39 1+6 88 89 I 90 137
46
~e11a platYEhylla (L) Pfeiff Porella platyphylloi~ (Schwein) Lindb
17 27 39 46 69 (71)79 87 88 89 9 34 46 (57) 59 87 88 89 92 90 92 93 77
Preissia quadrata (Seop) Nees
1 36 3946 (71) 88 89 90 121 123
137
Ptllidium ciliare (L) Hampe
1 27 39 46 132 141
Dum
(22) 28 3946 (57) 87 88 89 90 92
Ptilidium ~lcherrium (G Web) Hampe
(9) 13 394656 (57)70 87 88 89 90 93 (119) 136 141
47
Radula obconica Sullo Reboulia hemlsphaerlca (L) Raddi
39 46 88 89 (43) 46 (59) 70 90 t Cf 124 125 (139)
Riccardia ~atifrons Lindb
1 27 39 46 89
Riccardia multifi~ (L) S Gray
39 46 88 145
Riccia beyrichiana Hampe ~ Lehm
46 Riccardia palmata (Hedw) Carruth
39 46 90
48
Riccia bifurca Hoffm Riccia cavernosa Hoffm
48 13739 46 79 137
Riccia fluitans L
242731454679
Riccla frostli Aust
48 79
Riccia sullicantii Aust
79 (137) Riccia sorocarpa Bisch 46 (137)
49
Ricciocarpus natans (1) Corda Sca~ia apiculata Spruce
13 27 46 79 89 110 39 46 (139) 141
ScaJltll1ia carinthica Jack ~ Lindb
137 t 141
Scapania ~ (Mart) Durn
39467990137141
Scapania digenil Schiffn
90
ScaEsectflia cuspiduligera (Nees) K Mull
46 (139) (140) (141)
50
Scapania glaucoeerha1a (Tayl) Aust Seapania gyrrmostomophilia Kaal
39 46 (139) 141 41 44 46 64 (139) (140) 141
ScaP3poundlia irrigua Gott ~ almiddot
7 39 46 79 137 (140L 141
Seapania nemorosa (L) Dum
1 39 46 89 90 137
Scapmi~ ~~ta Smiddotlch
137 141 51
~caPnia poundludicola LDeske amp K Mull Scapania saxicola Schust
46 (66) 141 136 141
Sc~~ scandica (H Arnell ~ Buch) Macv
139 140
Scapania subalpina (Nees) Durn
39 40 46 79 137 141
Scapania undulata (L) Dum
1 42 46 79 141
Scapania umbrosa (Schrad) Dum
137 140 141
52
Trichocolea tomentella (Ehrh) Dum Triomarla exsecta (Schmid) Schiffn
27 (35) 39 46 47 88 137 141 8 28 39 46 90 140 141
Tritomaria exsectiformis (Breidl) Loe~ke Tritomaria Sluinguedentata (Huds) Buch
39 -t6 89 90 136 140 141 7 39 46 141
Trltomaria scitula (Tayl) Joerg
46 136 (139) 141
53
7 Cephaloziaceae Cepluzlozla (Dum emend Schiffn) Dum
biscwpidata (L) Dum subsp biclupidIIta catenulllttl (Hub) Lindb coMivens (Dicks) Lindb var connivens lacinultzta Jack ampX Spruce lunulifolilz (Dum) Dum macounii (Aust) Aust pleniceps (Aust) Lindh var pleniceps
Noweilia Mitt cunifolilz (Dicks) Mitt
8 Adelanthaceae Odontoschimuz (Dwn) Dum
denudatum (Nees ampX Mart) Dum Vir denudatum mtlCount (Aust) Underw
c
9 Cephaloziellaceae CephtlJoziellll (Spruce) Schiffn nom C()II$
divarlctzt4 (Sm) sclurfn val divarlcat(l elachhta (bdc) Schiffn 1uImpetuw (Nees) Schirfn IUbellll (Nees) Warnst vat IUbeIl4rube (Nees) Warnst var biid4 (Lindb) Dquinrube (Nees) WarDSt Var sulliwmtii (Aust) K MuD
10 Geo~c~ larptuttllus Nees
scutatus (Web et Mohr) Spruce
Geocalyx Nees grGVeolens (Schrad) Nees
11 Lophocoleaceae Lophocolea (Dum) Dum
heteropllyll4 (Schrad) Own minorNees
Chiloscypbus Corda om cons pallescens (Ehrh ampX Hoffm) Dum varfragllis (Roth) K Mun polyantitos (L) Corda var poiyanthos pol)llaquo1lthos (L) Corda var rlvulllris (Schrld) Nees
12 PlagiochiJaceae Plagio(hila Dum nom cons
tlBpienioides (L) Dum var asplenioidu
13 Jungermanniaceae Btriilophoril Loeske
attenutlta (Mart) Loeske bIubtlta (Schmid ampX Scltreb) Loeske
12
Lophom (Dum) Dum alpestris (Schleich ~ Web) Evans ascendens (Warnst) Schust badenais (Gott ex Rabenh) Schiffn var bademii bierenata (Schmid ex H)ffm) Dum eollam (Nees) Dum exci8a (Dicks) Dum var excisa gllimanii (Aust) Schust varglllmanii guttulata (Lindb et H Arnell) Evans heterocolpos (Thed) M A Howe var heterocolpos ineistl (Schrad) Dum var tnelsa longidens (Lindb) Macoun subsplongidens ventricosa (Dicks) Dum var ventricosa ventricolfl (Dicks) Dum varlontifIotrl (Nees) Macoun ventricosa (Dicks) Dum var silvlcola (Bueh) Jones
Tritomarill Schiffn ex Loeske exseeta (Schrad) Loeske exsecttformi$ (Breidl) Loeske subsp exsectiormfs quinquedentata (Huds) Buch var quinquedentata scitula (rayl) Joerg
Anastrophyllum (Spruce) Schiffn hellemnum (Noes) Schust michtzuxii (Web) Buch ex Evans minutum (Schrob) Schust var minutum
Mylill S Gray nom cons anomala (Hook) S Gray
Jamesoniella (Spruce) Carring autumnalis (DC) Steph var autumnalis
Jungemumnia L emend Dum eonfertissimtt Nees crenuliformis Aust exsertijolill Steph subsp cordifolia (Dum) Vana graciUima Sm hyalina Lyell leillnthll Grolle polaris Lindb pumila With sphaerocarpa Hook
14 Scapaniaceae Diplophyllum (Dum) Dum nom cons
apiculatum (Evans) Steph var apiculatum
Scapanill (Dum) Dum nom cons apkuata Spruce carinthillca Jack ex Lindb curta (Mart) Dum var curta
13
cuspiduligera (Nees) K Mull var cuspiduiigera degenii Schlffn ex K Mull var degenii glllucocephaJa (Tayt) Aust gymnostomophila Kaal irrigwJ (Nees) Gott et al subsp irrigua lingulata Buch varlingulata mucronata Buch subsp mucronata nemorosa (l) Dum paludicola Toeske et K Mun var paludicola paludosa (K Mull) K Mull saxicola Schust scandiea (H Arnell et Buch) Macv var scandiea 8Ubalp~ (Nees) Dum umbroSll (SchIad) Dum undulata (L) Dum var undulata
15 Radulaceae Raduia Dum nom COllS
compianata (L) Dum obconica Sull
16 Porellaceae Parella L
pinTlllta L patyphylla (L) Pfeiff platyphylloidt(l (Schwein) lindb
17 Jubulaceae FruUanill Raddi
bolanderi Aust brittoniae Evans eboracemis Gott inlllta Gott oakesiana Aust plana Sull rlparia Hampe ex Lehm selwyniana Pears taTnl1iJci (L) Dum subsp asagmyana (Mont) Hatt
18 lejeuneaceae Calolejeunea (Spruce) Schlffn
biddlecomiae (Aust) Evans
Lejeunea Ub nom cons cavifolia (Ehrh) Lindb emend Buch
19 Codoniaceae Fossombronia Raddi
foveolata Lindb
20 Pa11aviciniaceae Paklvieinlil S Gray nom cons
lyeiii (Hook) Carruth
14
Moerckia Gott hibemica (Hook) Gott
21 PeWaceae Pellfa Raddi nom cons propos
endivlifolfa (Dicks) Dwn epiphylllz (L) Corda neeBiana (GottlUmpr
22 Metzgeriaceae Metzgerla Raddi
conjugata Undb
23 Blasiacae B L
pulilm L
24 Aneurareae AneumDum
pinguis (L) Dum
Riccfl1dia S Gray nom cons latifrons Undb multiftda (L) S Gray var multifida palmata (Hedw) Carruth
25 Aytoniaceae Reboulill Raddi nom conI
hemllphaerica (L) Radeli
ManniaOpiz fragrrms (Balbis) Frye el ClarK pilosa (Hornem) Frye et Clark trltmdra (Scop) GroDe
26 Conocephalaceae Conocephalum Wigg nom cons
conicum (L) Lindb
27 Marchantiaceae Preiuia Corda
quadrata (Scop) Nees
Marchantia L polymorpha L
28 Ricciaceae Riccwcarpus Corda
natans (L) Corda
Riccia 1 beyrichiana Hampe ex Lehm bifurca Horfm
15
i
CtlJPemostI Hoffnt fuitu L rostii Aust BOIOCtI1fHI Bi~ sulltvtmtii Aust
MUSCI (Mosses)
29 Sphagnaceae SpIutgnum L
capillaceum (Weiss) Schrank var Cllpil1tlceum caplllllceum (Weiss) Vir tenellum (Schimp) Andr centrille C Jens ex Amen et C Jens compactummiddot OC ex Lam et Dc ClUpidatum Ehrh ex Hoffin var aupidatum jimbrltltum Wits ex J Hookmiddot fulCUm (Scbimp) KUngrr glrgenBOhIlll Russ imbrlctltum Hornsch ex Russ muelllmitil4m BOd 1IIIIU (RUII) C Jens przIutre L papIIIoIum Lindb pulchrum (Lindb ex braithw) Wamst recumun P-Beauv var recumun recum P-Beauv var tenue Klinggr riptlrlum Ansstr ruPQWItWamst squtl1fOlUm Crome subftltens Russ et Wamst ex Wamst IUbIeCfUldum Nees ex Sturm var subsecundum subsecundum Nees ex Sturm varrutem (Nees Hormch et sUUm) Hueb teres (Sdlimp) Anptr e C Hartm
WtlI7IItorjii Russ wuljfanum Gira
30 Andreaeaceae AndNrrerI Hedw
rupeitrlr Oedw var pest
31 Fissident~e Fluid_ lledw
trdiantoid Hedw byoidei Bedw var bryolda cristaM Wils ex Mitt exiguus sun VIr exigUw onttmUi (B-Pyl) Steud jtuberi Lesq et James obtusifolhu Wits var obtwifoliul ollPlUlldQldes Heel ~litSuU lubbtulkril Bed tfDdfoliul lledw llridubu (S~ ) Wahlenb vallirlIU1u
16
32 Ditrichaceae Pleuridium Brid
subulatum (Hedw) Rabenh
Ditrichum Hampe heteromallum (Hedw) Britt linellre (Sw) lindb
~ pallidum (Hedw) Hampe pusiUum (Hedw) Hampe
Saelania Lindb gIlucescens (Hedw) Bomanss et Broth
Ceratodon Brid purpureus (He(lw) Brid var prupurew
Distichium BSG capillaceum (Hedw) BSG inclinatum (Hedw) BSG
33 Bryoxipbiaceae Bryoxlphium Mitt
norvegicum (Brid) Mitt
34 Seligeriaceae Seligeria BSG
cumpylopoda Kindb ex Macoun et Kindb pusillll (Hedw) BSG recurvata (Hedw) BSG
Bindia BSG acuta (Hedw) BSG
35 Dicranaceae BlUehia Schwegr
lexUOSll (Sw ~ Schwaegr) C MueU sullivantii Aust
Trematodon Michx ambiguus (Hedw) Hornsch
Dicranellll (C Muell) Schimp cerviculata (Hedw) Schimp val cillPicuiata
heteromalla (Hedw) Schimp ~ rufescens (With) Schimp
schreberiana (Hedw) Schhnp var schreberiana schreberitJna (Hedw) Schimp var robusta Schimp ex Braithw subulata (Hedw) Schimp varia (Hedw) Schimp
Dicranodontium BSG denudatum (Brid) Britt ex Williams
Panzeucobryum (Lindh) Loeske Iongifolium (Hedw) oelke varlongifolium
17
Tortella (Undb) Limpr fragilis (Drumm) Umpr humilis (Hedw) JeM torlUoSfl (Hedw) Umpr
Hyophila Brid involuta (Hoole) Jaeg et Sauerb
Didymodon Hedw rigidulus Hedw
BryoerythrophyUum Chen recurvirostnon (Hedw) Chen
Btubula Hedw acula (Brid) Brid var acuta convoiuta Hedw fallox Hedw unguicultltll Hedw
Acaulon C MueD muticum (Uedw) C Muell var rufeseens (Iaeg) Crum
PJl~lm Hedw CUSPidtltum Hedw
De8mlltodon Brid cernuu (Hueb) BSG oblUsifoliw (Schwaegr) Schimp
T01IUla Hedw mueronifolia Schwaegr rurall (Hedw) Gaertn Meyer et Scherb var ruralis
39 Grimmiaceae Grlmmill Hedw
alpicola Hedw var alpicola alpicola Hedw var dupretii (Ther) Crum alpicola Hedw var rlvularl8 (Brid) Wahlenb apocarpa Hedw var apocarpa apocarpa Hedw var ambipa (Sull) Jonea apocarpa Hedw var conjerttl (Funck) Spreng apocarpa Hedw var strleta (Tum) Hook et Tayl CiIIyptrrlta Hook ex Drumm donniana Sm var donn_ hemumti Crum ltlevla (Brid) Brid montana BSG oineyi SuD oVllll8 (Hedw) lindb piUfe P-Beauv pltJgiopodill Hedw pulvinata (Hedw) Sm varbullpulvinata teretinervl Limpr
18
Orothera C Muell leana (Sull) C Muell
Rfwbdoweisia crispata (With) Undb
Cynodontium BSG ex Schimp alpestre (Wahlenb) Milde strumiferum (Hedw) Undb
Dichodontium Schimp pellucidum (Hedw) Schimp
Oncophorus (Slid) Brid vuens (Hedw) Brid wahlenbergii (Brid)
Dicranum Hedw bonjeanii DeNot ex Usa drummondii C Muell lagellare Hedw varj1ogellare fulvum Hook fuscescens Turn var fuscescens montanum Hedw muehlenbeckii BSG var muehlenbeckii polysetum Sw sabuletorum Ren et Card scoparium Hedw undulatum Brid viride (Sull et Lesq ex Sull) lindb
36 Leucobryaceae Leucobryum Hampe
glaucum (Hedw) Angstr ex Fr
37 Encalyptaceae Encalypta Hedw
ciliata Hedw var ciliata procera Bruch
38 Pottiaceae Astomum Hampe
muhlenbergianum (Sw) Grout
Weissia Hedw controversa Hedw var controversa
Gymnostomum Nees Homsch et Stunn aeruginosum Sm recurvirostrum Hedw var recurvirostrum
Trichostomum Bruch spirale Prout
19
40 Ephemeraceae Ephemerum Hampe
cohaerens (Hedw) Hampe crassineium (Schwaegr) Hampe val crassinervium spinulosum Bruch et Schimp ex Schimp
41 Funariaceae AphflllolThegma SuD
serratum (1 Hook et Wils ex Drumm) Sullo
Physcomitrium (Brid) Fuernr hookeri Hampe immersum Sullo pyriforme (Hedw) Hampe
Funaria Hedw americana Undb hygrometrica Hedw vaT hygrometrica
42 Splachnaceae Splachnum Hedw
ampullaceum Hedw
43 Schistostegaceae Schistostega Mohr
pennata (Hedw) Web el Mohr
44 Bryaceae Pohlia Hedw
annolina (Hedw) Lindb var annotina camea (Tum) Lindb cruda (Hedw) Undb elongata Hedw iliformis (Dicks) Andr lescurlana (Sull) Grout nutans (Hedw) Lindb obrusifolia (Brid) L Koch proligera (Undb ex Umpr) Undb ex Amell wahlenbergii (Web et Mohr) Andr
Leptobryum (B S G) Wits pyriforme (Hedw) Wils
Bryum Hedw algovicum Sendtnex C Muell argenteum Hedw caespiticium Hedw capillare Hedw var capillare creberrlmum Tayt lonchocaulon C Muell pallens (Brid) Sw ex Roehl pseudotrlquetrum (Hedw) Gaertn Meyer et Scherb varpudotriquetrum
20
pseudotriquetrum (Hedw) GaertnMeyer et Scherb var bimum (Schreb) Ulj stenotrichum C Muell var stenotrichum tortiolium Funck ex Brid uliginosum (Brid) B S G violaceum Crundw et Nyh weigelii Spreng
Rhodobryum (Schimp) Hampe roseum (Hedw) Limpr
45 Mniaceae Mnium Hedw
ciliare (C Muell) Schimp cinclidioides Hueb cuspidarum Hedw drummondii Bruch et Schimp longirostrum Brid lycopodioides Schwaegr marginatum (With) Brid ex P Beauv mediumB~S G var medium onhorrhynchum Brid pseudopunctatum Bruch et Schimp punctalUm Hedw var punctatum punctatum Hedw var elatum Schirnp rugicum Laur spinufosum B S G stellare Hedw
46 Aulacomruaceae Au1acomnium Schwaegr
androgynum (Hedw) Schwaegr heterostichum (Hedw) B S G paJustre (Hedw) Schwaegr var paJustre
47 Meesiaceae Amblyodon B S G
deillbotus (Hedw) B S G
Meesia Hedw triquetra (Richt) Angstr uJiginosa Hedw
48 Bartramiaceae Plagiopus Brid
oederiana (Sw) Umpr
Bartramia Hedw pomiformis Hedw var pomiformis
Philonotis Brid fontana (Hedw) Brid varfontana longseta (Michx) Britt varlongseta marchica (Hedw) Brid
21
~
~
49 Timmiaceae Timmia Hedw
megapolitana Hedw var megapolitana
50 Orthotrichaceae Zygodon Hook et Tayl
viridissimus (Dicks) Brid var rupestris lindb ex C J Hartm
Amphidium Schimp lapponicum (Hedw) Schimp mougeotii (B S G) Schimp
Orthotrichum Hedw anomalum Hedw obtuaifolium Brid ohioense Sullo et Lesq ex Aust pumilumSw sordidum Sullo et Lesq ex Aust speciosum Neesax Sturm var eegam (Schwaegr ex Hook et Grev) W~rnst stellDtwn Brid strrmgulatum P Beauv
Uleta Mohr ex Web cOfIICtata (p Beallv) Hamrn crispa (Hedw) Brid
Drummondill Hook ex Drumm prorepens (Hedw) Brltt
51 Fontinalaceae Fantinalis Hedw
antipyretica Hedw var antipyrettca antipyretica Hedw forma rabusta (Card) Card dalecarlica Schimp ex B S G var dalecarlica flqccidtJ Ren et Card hypnoides C J Hartrn var hypnoides hypnoides C J Hartm var durillei (Schimp) HU$ll
missourlcaCard novae-angliae Sullo var novae-angliae splulgrUlolill (C Moen) Wijket M~S
DichelY1l1ll Myr capilloceum (With) Myr falcatum (Bedw) Myr HIllescens B S G
52 Climaciaceae aimecium Web et Mohr ex Mohr
americllnum Brid dedroides (Hedw) Web et Mohr
22
S3 Hedwigiaceae Hedwigia P Beauv
ciliata (Hedw) P Beauv
54 Cryphaeaceae Forsstroemia lindh
ohioense (Sull) lindh
55 Leucodontaceae Leucodon Schwaegr
brachypus Blid var brachypus brachypus Brid var aniJrewsitlnus Crum et Anderson julaceus (Hedw) SuU
56 Neckeraceae Neckera Hedw
pennata Hedw var pennata
1fomalia (Brid) B S G trichomanoides (Hedw) B S G
57 Theliaceae Thelia Sull
asprellll Sullo hirtella (Hedw) Sullo lescurii SuU
Myurella B SG julacea (Schwaegr) B S G
sibrica (C MueU) Reim var sibrica
58 Fabroniaceae Fabronio Raddi
ciliaris (Brid) Bnd
Anacamptodon Brid splachnoides (Groet ex Brid) Brid
Schwetschkeopsis Broth fabronia (Schwaegr) Broth
59 Leskeaceae Lindbergia Kindb
brachyptera (Mitt) Kindb
Leskea Hedw gracilescens Hedw obscura Hedw poiycarpa Hedw
Leskeella (limpr) Loeske nervosa (Brid) Loeske
23
Pterlgy1lllndrum Hedw filiforme Hedw
60 Thuidiaceae Haplohymenfum Dozy et MoIk
trlste (Ces ex De Not) Kiodb
Anomodon Hook et Tayl attenuatus (Hedw) Hueb minor (Hedw) Fuemr rostratus (Hedw) Schimp viticulosus (Hedw) Hook et Tayl
Haplocladium (C Mucn) C MueU microphyllum (Hedw) Broth virginillnum (Brld) Broth ~bull
Thuidium B S G abietinum (Hedw) B S G delicatulum (Hedw) B S G vardelicatulum delicatulum (Hedw) B S G var radicans (Kindb) Crum Steere et Anderson minutulum (Hedw) B S G pygmaeum B S G recognftum (Hedw) Lindb scitum (p Beauv) Aust
Helodium (Sull) Warnst blandowli (Web et Mohr) Warnst var blandowli blandowli (Web et Mohr) Wamst var helodioides (Ren et Card ex Roell) Crum
Steere et Anderson paludomm (Sull) AuS
61 Amblystegiaceae Cratoneuron (Sull) Spruce
commulatum (Hedw) Roth var commutfltum commutatum (Hedw) Roth var fakatum (Brid) Moenk ilicinum (Hedw) Spruce var filcinum
Campylium (Sun) Mitt chrysophyllum (Brid) 1 Lange var chrysophyllum hispidulum (Brid) Mitt var hllJidulum polygamum (B S G)C lens varpolygtlmUm stellatum (Hedw) C1ens vat stelltum
Leptodictyum (Schimp) Warnst laxirete (Card et Ther) Broth riparium (Hedw) Wamat trichopodium (Schultz) Warns l1acillans (Sull) Broth
Hygroamblystegium Loelke jlul1iatile (Hedw) oeske var luViatile
24
fluviatile (Hedw) Loeske var oWltum Grout noterophilum (Sun et Lesq ex Sull) Warnst tenox (Hedw) Jenn var tenax tenox (Hedw) Jenn var spinilolium Schimp) Jenn
Amblystegium BSG juratzkanum Schimp serpens (Hedw) BSG varlum (Hedw) Undb var varium
Platydictya Berk confervoides (Brid) Crum jungermanniodes (Brid) Crum minutissimum (Sull et Lesq ex Sull) Crum subtile (Hedw) Crum
Drepanocladus (C Muell) Roth aduncus (Hedw) Warnst var aduncus aduncus (Hedw) Warnst var capillilolius (Wamst)Riebm aduncus (Hedw) Warnst var kneiffii (BSG) Moenk aduncus (Hedw) Warnst var polycorpus (Bland ex Voit) Roth exannulatus (BSG) Warnst fluitans (Hedw) Warnst var fluitans revolvens (Sw) Wamst var revolvens sendtneri (Schimp) Warnst var sendtnerl uncinatus (Hedw) Warnst var uncinatus uncinatus ffiedw) Warnst var piwnosus (Schimp) Wamst vernicoms (Lindb ex C Hartm) Warnst
Hygrohypnum Lindb eugyrium (BSG) loeske luridum (Hedw Jenn varluridum ochraecum (Tum ex Wils) Loeske var ochraceunt
Calliergon (Sull) Kindb cordilolium Hedw Kindb glganteum (Schimp) Kindb var giganteum richardsonii (Mitt) Kindb ex Wamst stramineum (Brid) Kindb trilarium (Web et Mohr) Kindb
Calliergonella Loeske cuspidata (Hedw) Loeske
Calliergidium (Ren) Grout bakeri (Ren) Grout
Scorpidium (Schimp) Urnpr scorpiodides (Hedw) limpr
62 Brachytbeciaceae Tomenthypnum loeske
nitens (Hedw) Loeske var mtens
25
Homalotheciella (Card) Broth subcapillata (Hedw) Broth
Brachythecium BSG acuminatum (Hedw) Awt acutum (Mitt) Sullo calcareum IGndb campestTe (C MuelL) BSG cunum (Lindb) Umpr digastrum C Muell et Kindb ex Macoun et Kindb oxycladon (Brid) Jaeg et Sauerb var oxycladon plumosum (Hedw) BSG populeum (Hedw) BSG var populeum reflexum (Starke ex Web et Mohr) DsG var relexum reiexum (Starke ex Web et Mohr) DSG var pacificum Renet Card ex Roell rlvulare BSG var rivulare tabulum (Hedw) BSC var tabuum salebrosum (Web et Mohr) BSG var salebrosum velutinum (Hedw) BSG Vat velutinum
Bryhniil Kaur graminicolor (Brid) Grout var graminicolor novae-angliae (Sull et Lesq ex sun) Grout
Rhynchostegium BSG senulatum (Hedw) Ja4lg et Sauerb
Rhynchostegiella (BSG) Limpr compacta (C MueU) Loeske
EU1hynchium 8SG hians (Hedw) Sande Lac pulchellum (Hedw) Jenn var pulchelium riparloides (Hedw) Rich
Stokeriella (Kindb) Robins pmelonga (Hedw) Robins var stokesii (Tum) Crum
63 Entodontaceae Entodon C Muell
brevisetus (Hook et Wils ex Wils) Lindb cladorrhizans (Hedw) C Muell seductrlx (Hedw) C MueU var wluctrix
Pleurozium Mitt schreberl (8rid) Mitt var schreberi
64 Plagiotheciaceae Pltlglothecium BSG
CflVifolium (Brid) Iwats var CtWifolium denticulatum (Hedw) 8SG laetum BSG ltztebricola BSG
26
6S SernatophyUaceae Brotherella Loeske ex Fleisch
recumins (Michx) Fleisch tenuirostris (Bruch et Schimp ex Sull) Fleisch
Sematophyllum Mitt demissum (Wils) Mitt
66 Hypnaceae Pl(itygyrium BSG
repens (Brid) BSG
PylaisieUa Kindb intricata (Hedw) Grout polyantha (Hedw) Grout selwynii (Kindb) Crum Steere et Anderson
Homomallium (Schimp) Loeske adnatum (Hedw) Broth
Hypnum Hedw cupressiforme Hedw var cupressiforme cupressifonne Hedw varfiliforme Brid curviolium Hedw fertile Sendtn imponens Hedw lindbergii Mitt palescens (Hedw) P-Beauv pratense Koch ex Brid
Calicladium Crum haldanianum (Grev) Crum
sopterygiopsis Iwats muelleriana (Schimp) Iwats
Isopterygium Mitt e1egans (Brid) lindb pulchellum (Hedw) Jaeg et Sauerb tenerum (Sw) Mitt
Taxiphyllum Fleisch depllQtum (Bruch et Schimp ex sun) Fleisch taxirameum (MitL) Fleisch
Herzogiella Broth seligeri (Brid) Iwats striatella (Brid) Iwats ruracea (Lindh) Iwats
hilium De Not crista-castrenna (Hedw) De Not
27
67 Rhytidiaceae Rhytidium (Sull) Kindb
rugosum (Hedw) Kindb
Rhytidfodelphus (Lindb ex Iimpr) Warnst triquetrus (Hedw) Warnst var triquetrus
68 Hylocomiaceae Hylocomium BSG
pyrenaicum (Spruce) lindb splendens (Hedw) BSG varsplendens
69 Buxbaumiaceae Buxbaumia Hedw
aphyllll Hedw
Diphyscium Mohr follosum (Hedw) Mohr
70 Tetraphidaceae Tetraphi3
pellucida Hedw
71 Polytrichaceae Atrichum PmiddotBeauv
angustatum (Brid) BSG cri8pum (James) Sullo oerstedianum (C Muell) Mitt teneUum (Roehl) BSG undulatum (Hedw) P middotBeauv var unduiatum
Pogonatum P-Beauv alpinum (Hedw) Roehl var alpinum umigerum (Hedw) P-Beauv
Poiytrichum Hedw commune Hedw var commune ormosum Hedw juniperinum Hedw longisetum Brid ohioense Ren et Card piliferum Hedw var pilierum strictum Brid
28
MAPS OF BRYOPHYTE TAXA
REPORTED FOR WISCONSIN
HORNWORTS AND LIVERWORTS
29
bull bull WSHeV~N S vE II
0 ~
e~RRON ~USK
(J~ (j -
CHIPPE A DUNN
0 o0 poundAU CLAIRE
~ TREMshyPEAshyLEAU
MICE
ONEIDA
Q LINCOLN
TAYLOR
Q t-~----- MOIRA THON
CLARK
o
10
LA FAVETTE
Hornwor ts amp Liverworts
- None - 1 - 25 taxa
- 26 - 50 tRXshy
- 51 - 75 tax)
- 7c - 100 taxa
FONDOU j LAC ~
DODGE
i O JEfFERSON
WALWORTH
30
Anastrophylltlfl helleranum (Nees) Schust Anastrophyllum michauxil (We~ Buch ~
46 88 136 141 Evans 38 46 141
Anastrophyllum mlnut~ (Schreb) Schuster Aneura plnguls (L) Durn
89 21 384687 88 89 90 (138) (145)
Anthoceros macounil MA Howe ~oceros punctatus L
21 (45) (144) 34
31
Barbilophozia attenuata (Mart) Loeske Barbilophozia barbata (Schmid ~ Schre~
24 141 Loeske 28 39 46 89 90 141
Bazzania trilobata (1) S Gray
1 26 39 46 (57) (59) 79 88 89
90 119 141
BlePbarostoma trichoffiyllum (1) Dumort
1 8 26 39 46 89 90 137 141
Blasia pusilla 1
1 39 46 90
Calypogeja fissa (1) Raddi
90
32
$129geja integristipula Steph C~Ypound2geja muelleriana (Schifm) K Mull
88 89 89
CallpoundO~eja neesiana (Mass amp Carest) K Mull
39 46 89 140 (141)
~pogeja sJlapound2icola (H Arnell tl J bull
Perss) Warnst et Loeske (119) 141
CeIalozia biscuspidata (L) Dum
1 37 I 46 90 t 137 141
CalYpogej~ trichomanis (L) Corda
26 39 (119) 137
33
CeEhalozia catenulata (Hub) Lindb Cephalozia connivens (Dicks) Lindb
39 46 79 141 39 4679 88 (89) 90 119 141
Cephalozia lacinulata Jack ~ Spruce
141 Cephalozia lunulifolia (Dum~ Dum I 28 39 46 79 88 89 90 141
Cepoundhalozia macouni (Aust) Aust Cephalozia ~leniceps (Aust) Lindb 27~ 3746 (66)7987 (139)141 39 46 79 89 141
34
Ceffialoziella 9ivaricata (Sm) Schiffn Cephaloziella ~hista (Jack) Schiffn
46 90 46 (119) 141
Cephaloziella hampeana (Nees) Schiffn
46
Cephaloziella rubella (Nees) Warnst
46 39
Cephaloziella rubella var sullivantii
(Aust) K Mull 46 39
Cephaloziella rubella var bifida (Lindb) Douin
35
46
Chiloscyphus pallescens (Ehrh ~ Hof~ Dum Chiloscyphus pallescens var fragilis
39J 46 88 89J 90 (Roth) K Milll 46
Chiloscyphus E9lyanthos (L) Corda
1 24 39 46
Chiloscyphus polyanthos vax rivularis
(Schrad) Nees 39 46
poundpoundnocepha1um conicum (1) Lindb
12 28 39 46 (49) 87 88 8990 120 121 148
Cololejeunea biddlecomiae (Aust) Evans
39 46 62 87 88 89 90
36
Dipl0]hyllum apicu1atum (Evans) Steph Fossombronia foveo1ata Lindb
(35)41 46 90 140 1 t 38 (39) t 46
Frullania bolanderi Aust
13 46 62 88 90 139 140
Frullanla ari ttoniae Evans
39 46 69 (71) 87 88 89 90
Frullania eboracensis Gott
1 9 28 4 39 46 (57) (59) 69 70 (71) 87 88 89 90 92 93 97 148
Frullania inflata Gott
46 87 88 89 (139)
Frullania oakesiana Aust Frullania plana Sullo
(35) 42 46 87 (139) (140) 92
Frullania ri paria Hampe ~ Lehm
46 69 71 87 89
Frullania ~elwynianna Pears
46 89
Geocalyx SEaveolens (Schrad) Nees
394688 8990 (119) 140
Frullania tamarisci (L) Dum subsp sPagrayana
(Mont) Hatt 28 34 l~6 t (57) 88 89 90
38
Mar pan thus seutatUB ( Web amp Mohr) Spruce Jamesoniella autumnalis (DC) Steph
(35) 39 46 90 1 7 28 39 46 56 (57) 87 8889 90 (119) 137 141
Jungermannia confertissima Nees
137 141
Jungermannia crenuliform1s Aust
79 137 141
Jungermannia ~il1ima Sm
46 90 141 Jungermannia exsert1folia subsp corifo~ (Dum~
Vana (35) 46 79 141
39
Jungermannia hyalina Lyell Jungerrr6nnia 1eiantha Grolle
40 46 79 136 141 39 46 79 88 90 1)7 141
Jungermannia IXlaxis Lindb Jungermannia pumila With
46 67 79 (139) (140) 141 )9 46 90 1)6 141
Jungermannia sphaerocarpa Hook Kurzia setacea (Web) Grolle
4679136140IIH 136
Lejeunea cavifo1ia (Ehrh) Lindb Lepidozia reptans (1) Dum
1 7 27 39 46 88 89 90 137 14010 I ( 27) 46 I 69 (71) 88
lophocolea heterophylla (Schrad) Dum Lophocolea minor Nees
1 )4 39 46 (57) 87 88 89 27 39 46 88 89 90 (119) 90 (93)94 119 136
Lophozia al~stris (Schleich ~ Web) Evans Lophozia ascendens (Warnst) Schust
46 90 137 141 136 140 141
41
loIilozia badensls (Gatt ~ Rabenh)
46 141
lophozia blcrenata (Schmid sectA Hoffm)
Dum 46 141
lophozia collaris (Nees) Dum
39 46 141
lophozla excisa (Dicks) Dum
27 4679 141
lophozia gillmani (Aust) Schust lophozia gIlttulata (Undo tl H Arnell)
36 39 40 46 136 (140) 141 Evans 39 46 136 141
42
141
Lophozia heterocolEDs (Thed) MA Howe IoPhozia ipcisa (Schard) Dum
27 39 46 89 141 8 27 31 46 79 90 140 141
LoQhozia longidens Macoun Lophozia ventricosa
1 46 79 90 141
Lophozia ventricosa var silvicola
(Buch) Jones 141 LoEhozla ventricosa vax longif1ora (Nees)
Macoun 39 46 79 141
43
~~ fragrans (Balbis) Frye et Clark Mannia pi 10sa (Hornem) Frye et Clark
46 (139) (140) 9 46 79 116 139
Mannia triandra (Scop) Gro 11e
46 (140)
Marchantia pound91ymorpha L
1 13 28 3946 (49) 78 87 88 89 90 (Ill) 121 122 147
Mo~rckia hibernica (HookJ Gott
46 89
Metzgeria conjugata Lindb
44
46
Mylia anomala (Hook) S Gray Notothylas orbicularis (Schwein) Sullo
39 46 141 (35)45 46
Nowellia curvifolia (Dicks) Mitt
17 27 3846 88 89 90 141
Odontoschisma denudatuJI1 (Nees ~ Mart)
Dum 46 81 137 141
Pallavicinia lyellii (Hook~ Carruth
90
Odontoschisma macounii (Aust) Under
45 46 (66) (139) 140 141
45
=
Pellia epiphllla (L) CordaPellia endiviifolia (Dicks~ Dum
27 39 46 90 13724 39 46
Pellia neesiana Limp bull Phaeoceros
39 46 89 27 39 45 46 97
Porella pinnata L
13 27 I 46 90 Plagiochila asplenioides (L) Dum
27 I 39 1+6 88 89 I 90 137
46
~e11a platYEhylla (L) Pfeiff Porella platyphylloi~ (Schwein) Lindb
17 27 39 46 69 (71)79 87 88 89 9 34 46 (57) 59 87 88 89 92 90 92 93 77
Preissia quadrata (Seop) Nees
1 36 3946 (71) 88 89 90 121 123
137
Ptllidium ciliare (L) Hampe
1 27 39 46 132 141
Dum
(22) 28 3946 (57) 87 88 89 90 92
Ptilidium ~lcherrium (G Web) Hampe
(9) 13 394656 (57)70 87 88 89 90 93 (119) 136 141
47
Radula obconica Sullo Reboulia hemlsphaerlca (L) Raddi
39 46 88 89 (43) 46 (59) 70 90 t Cf 124 125 (139)
Riccardia ~atifrons Lindb
1 27 39 46 89
Riccardia multifi~ (L) S Gray
39 46 88 145
Riccia beyrichiana Hampe ~ Lehm
46 Riccardia palmata (Hedw) Carruth
39 46 90
48
Riccia bifurca Hoffm Riccia cavernosa Hoffm
48 13739 46 79 137
Riccia fluitans L
242731454679
Riccla frostli Aust
48 79
Riccia sullicantii Aust
79 (137) Riccia sorocarpa Bisch 46 (137)
49
Ricciocarpus natans (1) Corda Sca~ia apiculata Spruce
13 27 46 79 89 110 39 46 (139) 141
ScaJltll1ia carinthica Jack ~ Lindb
137 t 141
Scapania ~ (Mart) Durn
39467990137141
Scapania digenil Schiffn
90
ScaEsectflia cuspiduligera (Nees) K Mull
46 (139) (140) (141)
50
Scapania glaucoeerha1a (Tayl) Aust Seapania gyrrmostomophilia Kaal
39 46 (139) 141 41 44 46 64 (139) (140) 141
ScaP3poundlia irrigua Gott ~ almiddot
7 39 46 79 137 (140L 141
Seapania nemorosa (L) Dum
1 39 46 89 90 137
Scapmi~ ~~ta Smiddotlch
137 141 51
~caPnia poundludicola LDeske amp K Mull Scapania saxicola Schust
46 (66) 141 136 141
Sc~~ scandica (H Arnell ~ Buch) Macv
139 140
Scapania subalpina (Nees) Durn
39 40 46 79 137 141
Scapania undulata (L) Dum
1 42 46 79 141
Scapania umbrosa (Schrad) Dum
137 140 141
52
Trichocolea tomentella (Ehrh) Dum Triomarla exsecta (Schmid) Schiffn
27 (35) 39 46 47 88 137 141 8 28 39 46 90 140 141
Tritomaria exsectiformis (Breidl) Loe~ke Tritomaria Sluinguedentata (Huds) Buch
39 -t6 89 90 136 140 141 7 39 46 141
Trltomaria scitula (Tayl) Joerg
46 136 (139) 141
53
Lophom (Dum) Dum alpestris (Schleich ~ Web) Evans ascendens (Warnst) Schust badenais (Gott ex Rabenh) Schiffn var bademii bierenata (Schmid ex H)ffm) Dum eollam (Nees) Dum exci8a (Dicks) Dum var excisa gllimanii (Aust) Schust varglllmanii guttulata (Lindb et H Arnell) Evans heterocolpos (Thed) M A Howe var heterocolpos ineistl (Schrad) Dum var tnelsa longidens (Lindb) Macoun subsplongidens ventricosa (Dicks) Dum var ventricosa ventricolfl (Dicks) Dum varlontifIotrl (Nees) Macoun ventricosa (Dicks) Dum var silvlcola (Bueh) Jones
Tritomarill Schiffn ex Loeske exseeta (Schrad) Loeske exsecttformi$ (Breidl) Loeske subsp exsectiormfs quinquedentata (Huds) Buch var quinquedentata scitula (rayl) Joerg
Anastrophyllum (Spruce) Schiffn hellemnum (Noes) Schust michtzuxii (Web) Buch ex Evans minutum (Schrob) Schust var minutum
Mylill S Gray nom cons anomala (Hook) S Gray
Jamesoniella (Spruce) Carring autumnalis (DC) Steph var autumnalis
Jungemumnia L emend Dum eonfertissimtt Nees crenuliformis Aust exsertijolill Steph subsp cordifolia (Dum) Vana graciUima Sm hyalina Lyell leillnthll Grolle polaris Lindb pumila With sphaerocarpa Hook
14 Scapaniaceae Diplophyllum (Dum) Dum nom cons
apiculatum (Evans) Steph var apiculatum
Scapanill (Dum) Dum nom cons apkuata Spruce carinthillca Jack ex Lindb curta (Mart) Dum var curta
13
cuspiduligera (Nees) K Mull var cuspiduiigera degenii Schlffn ex K Mull var degenii glllucocephaJa (Tayt) Aust gymnostomophila Kaal irrigwJ (Nees) Gott et al subsp irrigua lingulata Buch varlingulata mucronata Buch subsp mucronata nemorosa (l) Dum paludicola Toeske et K Mun var paludicola paludosa (K Mull) K Mull saxicola Schust scandiea (H Arnell et Buch) Macv var scandiea 8Ubalp~ (Nees) Dum umbroSll (SchIad) Dum undulata (L) Dum var undulata
15 Radulaceae Raduia Dum nom COllS
compianata (L) Dum obconica Sull
16 Porellaceae Parella L
pinTlllta L patyphylla (L) Pfeiff platyphylloidt(l (Schwein) lindb
17 Jubulaceae FruUanill Raddi
bolanderi Aust brittoniae Evans eboracemis Gott inlllta Gott oakesiana Aust plana Sull rlparia Hampe ex Lehm selwyniana Pears taTnl1iJci (L) Dum subsp asagmyana (Mont) Hatt
18 lejeuneaceae Calolejeunea (Spruce) Schlffn
biddlecomiae (Aust) Evans
Lejeunea Ub nom cons cavifolia (Ehrh) Lindb emend Buch
19 Codoniaceae Fossombronia Raddi
foveolata Lindb
20 Pa11aviciniaceae Paklvieinlil S Gray nom cons
lyeiii (Hook) Carruth
14
Moerckia Gott hibemica (Hook) Gott
21 PeWaceae Pellfa Raddi nom cons propos
endivlifolfa (Dicks) Dwn epiphylllz (L) Corda neeBiana (GottlUmpr
22 Metzgeriaceae Metzgerla Raddi
conjugata Undb
23 Blasiacae B L
pulilm L
24 Aneurareae AneumDum
pinguis (L) Dum
Riccfl1dia S Gray nom cons latifrons Undb multiftda (L) S Gray var multifida palmata (Hedw) Carruth
25 Aytoniaceae Reboulill Raddi nom conI
hemllphaerica (L) Radeli
ManniaOpiz fragrrms (Balbis) Frye el ClarK pilosa (Hornem) Frye et Clark trltmdra (Scop) GroDe
26 Conocephalaceae Conocephalum Wigg nom cons
conicum (L) Lindb
27 Marchantiaceae Preiuia Corda
quadrata (Scop) Nees
Marchantia L polymorpha L
28 Ricciaceae Riccwcarpus Corda
natans (L) Corda
Riccia 1 beyrichiana Hampe ex Lehm bifurca Horfm
15
i
CtlJPemostI Hoffnt fuitu L rostii Aust BOIOCtI1fHI Bi~ sulltvtmtii Aust
MUSCI (Mosses)
29 Sphagnaceae SpIutgnum L
capillaceum (Weiss) Schrank var Cllpil1tlceum caplllllceum (Weiss) Vir tenellum (Schimp) Andr centrille C Jens ex Amen et C Jens compactummiddot OC ex Lam et Dc ClUpidatum Ehrh ex Hoffin var aupidatum jimbrltltum Wits ex J Hookmiddot fulCUm (Scbimp) KUngrr glrgenBOhIlll Russ imbrlctltum Hornsch ex Russ muelllmitil4m BOd 1IIIIU (RUII) C Jens przIutre L papIIIoIum Lindb pulchrum (Lindb ex braithw) Wamst recumun P-Beauv var recumun recum P-Beauv var tenue Klinggr riptlrlum Ansstr ruPQWItWamst squtl1fOlUm Crome subftltens Russ et Wamst ex Wamst IUbIeCfUldum Nees ex Sturm var subsecundum subsecundum Nees ex Sturm varrutem (Nees Hormch et sUUm) Hueb teres (Sdlimp) Anptr e C Hartm
WtlI7IItorjii Russ wuljfanum Gira
30 Andreaeaceae AndNrrerI Hedw
rupeitrlr Oedw var pest
31 Fissident~e Fluid_ lledw
trdiantoid Hedw byoidei Bedw var bryolda cristaM Wils ex Mitt exiguus sun VIr exigUw onttmUi (B-Pyl) Steud jtuberi Lesq et James obtusifolhu Wits var obtwifoliul ollPlUlldQldes Heel ~litSuU lubbtulkril Bed tfDdfoliul lledw llridubu (S~ ) Wahlenb vallirlIU1u
16
32 Ditrichaceae Pleuridium Brid
subulatum (Hedw) Rabenh
Ditrichum Hampe heteromallum (Hedw) Britt linellre (Sw) lindb
~ pallidum (Hedw) Hampe pusiUum (Hedw) Hampe
Saelania Lindb gIlucescens (Hedw) Bomanss et Broth
Ceratodon Brid purpureus (He(lw) Brid var prupurew
Distichium BSG capillaceum (Hedw) BSG inclinatum (Hedw) BSG
33 Bryoxipbiaceae Bryoxlphium Mitt
norvegicum (Brid) Mitt
34 Seligeriaceae Seligeria BSG
cumpylopoda Kindb ex Macoun et Kindb pusillll (Hedw) BSG recurvata (Hedw) BSG
Bindia BSG acuta (Hedw) BSG
35 Dicranaceae BlUehia Schwegr
lexUOSll (Sw ~ Schwaegr) C MueU sullivantii Aust
Trematodon Michx ambiguus (Hedw) Hornsch
Dicranellll (C Muell) Schimp cerviculata (Hedw) Schimp val cillPicuiata
heteromalla (Hedw) Schimp ~ rufescens (With) Schimp
schreberiana (Hedw) Schhnp var schreberiana schreberitJna (Hedw) Schimp var robusta Schimp ex Braithw subulata (Hedw) Schimp varia (Hedw) Schimp
Dicranodontium BSG denudatum (Brid) Britt ex Williams
Panzeucobryum (Lindh) Loeske Iongifolium (Hedw) oelke varlongifolium
17
Tortella (Undb) Limpr fragilis (Drumm) Umpr humilis (Hedw) JeM torlUoSfl (Hedw) Umpr
Hyophila Brid involuta (Hoole) Jaeg et Sauerb
Didymodon Hedw rigidulus Hedw
BryoerythrophyUum Chen recurvirostnon (Hedw) Chen
Btubula Hedw acula (Brid) Brid var acuta convoiuta Hedw fallox Hedw unguicultltll Hedw
Acaulon C MueD muticum (Uedw) C Muell var rufeseens (Iaeg) Crum
PJl~lm Hedw CUSPidtltum Hedw
De8mlltodon Brid cernuu (Hueb) BSG oblUsifoliw (Schwaegr) Schimp
T01IUla Hedw mueronifolia Schwaegr rurall (Hedw) Gaertn Meyer et Scherb var ruralis
39 Grimmiaceae Grlmmill Hedw
alpicola Hedw var alpicola alpicola Hedw var dupretii (Ther) Crum alpicola Hedw var rlvularl8 (Brid) Wahlenb apocarpa Hedw var apocarpa apocarpa Hedw var ambipa (Sull) Jonea apocarpa Hedw var conjerttl (Funck) Spreng apocarpa Hedw var strleta (Tum) Hook et Tayl CiIIyptrrlta Hook ex Drumm donniana Sm var donn_ hemumti Crum ltlevla (Brid) Brid montana BSG oineyi SuD oVllll8 (Hedw) lindb piUfe P-Beauv pltJgiopodill Hedw pulvinata (Hedw) Sm varbullpulvinata teretinervl Limpr
18
Orothera C Muell leana (Sull) C Muell
Rfwbdoweisia crispata (With) Undb
Cynodontium BSG ex Schimp alpestre (Wahlenb) Milde strumiferum (Hedw) Undb
Dichodontium Schimp pellucidum (Hedw) Schimp
Oncophorus (Slid) Brid vuens (Hedw) Brid wahlenbergii (Brid)
Dicranum Hedw bonjeanii DeNot ex Usa drummondii C Muell lagellare Hedw varj1ogellare fulvum Hook fuscescens Turn var fuscescens montanum Hedw muehlenbeckii BSG var muehlenbeckii polysetum Sw sabuletorum Ren et Card scoparium Hedw undulatum Brid viride (Sull et Lesq ex Sull) lindb
36 Leucobryaceae Leucobryum Hampe
glaucum (Hedw) Angstr ex Fr
37 Encalyptaceae Encalypta Hedw
ciliata Hedw var ciliata procera Bruch
38 Pottiaceae Astomum Hampe
muhlenbergianum (Sw) Grout
Weissia Hedw controversa Hedw var controversa
Gymnostomum Nees Homsch et Stunn aeruginosum Sm recurvirostrum Hedw var recurvirostrum
Trichostomum Bruch spirale Prout
19
40 Ephemeraceae Ephemerum Hampe
cohaerens (Hedw) Hampe crassineium (Schwaegr) Hampe val crassinervium spinulosum Bruch et Schimp ex Schimp
41 Funariaceae AphflllolThegma SuD
serratum (1 Hook et Wils ex Drumm) Sullo
Physcomitrium (Brid) Fuernr hookeri Hampe immersum Sullo pyriforme (Hedw) Hampe
Funaria Hedw americana Undb hygrometrica Hedw vaT hygrometrica
42 Splachnaceae Splachnum Hedw
ampullaceum Hedw
43 Schistostegaceae Schistostega Mohr
pennata (Hedw) Web el Mohr
44 Bryaceae Pohlia Hedw
annolina (Hedw) Lindb var annotina camea (Tum) Lindb cruda (Hedw) Undb elongata Hedw iliformis (Dicks) Andr lescurlana (Sull) Grout nutans (Hedw) Lindb obrusifolia (Brid) L Koch proligera (Undb ex Umpr) Undb ex Amell wahlenbergii (Web et Mohr) Andr
Leptobryum (B S G) Wits pyriforme (Hedw) Wils
Bryum Hedw algovicum Sendtnex C Muell argenteum Hedw caespiticium Hedw capillare Hedw var capillare creberrlmum Tayt lonchocaulon C Muell pallens (Brid) Sw ex Roehl pseudotrlquetrum (Hedw) Gaertn Meyer et Scherb varpudotriquetrum
20
pseudotriquetrum (Hedw) GaertnMeyer et Scherb var bimum (Schreb) Ulj stenotrichum C Muell var stenotrichum tortiolium Funck ex Brid uliginosum (Brid) B S G violaceum Crundw et Nyh weigelii Spreng
Rhodobryum (Schimp) Hampe roseum (Hedw) Limpr
45 Mniaceae Mnium Hedw
ciliare (C Muell) Schimp cinclidioides Hueb cuspidarum Hedw drummondii Bruch et Schimp longirostrum Brid lycopodioides Schwaegr marginatum (With) Brid ex P Beauv mediumB~S G var medium onhorrhynchum Brid pseudopunctatum Bruch et Schimp punctalUm Hedw var punctatum punctatum Hedw var elatum Schirnp rugicum Laur spinufosum B S G stellare Hedw
46 Aulacomruaceae Au1acomnium Schwaegr
androgynum (Hedw) Schwaegr heterostichum (Hedw) B S G paJustre (Hedw) Schwaegr var paJustre
47 Meesiaceae Amblyodon B S G
deillbotus (Hedw) B S G
Meesia Hedw triquetra (Richt) Angstr uJiginosa Hedw
48 Bartramiaceae Plagiopus Brid
oederiana (Sw) Umpr
Bartramia Hedw pomiformis Hedw var pomiformis
Philonotis Brid fontana (Hedw) Brid varfontana longseta (Michx) Britt varlongseta marchica (Hedw) Brid
21
~
~
49 Timmiaceae Timmia Hedw
megapolitana Hedw var megapolitana
50 Orthotrichaceae Zygodon Hook et Tayl
viridissimus (Dicks) Brid var rupestris lindb ex C J Hartm
Amphidium Schimp lapponicum (Hedw) Schimp mougeotii (B S G) Schimp
Orthotrichum Hedw anomalum Hedw obtuaifolium Brid ohioense Sullo et Lesq ex Aust pumilumSw sordidum Sullo et Lesq ex Aust speciosum Neesax Sturm var eegam (Schwaegr ex Hook et Grev) W~rnst stellDtwn Brid strrmgulatum P Beauv
Uleta Mohr ex Web cOfIICtata (p Beallv) Hamrn crispa (Hedw) Brid
Drummondill Hook ex Drumm prorepens (Hedw) Brltt
51 Fontinalaceae Fantinalis Hedw
antipyretica Hedw var antipyrettca antipyretica Hedw forma rabusta (Card) Card dalecarlica Schimp ex B S G var dalecarlica flqccidtJ Ren et Card hypnoides C J Hartrn var hypnoides hypnoides C J Hartm var durillei (Schimp) HU$ll
missourlcaCard novae-angliae Sullo var novae-angliae splulgrUlolill (C Moen) Wijket M~S
DichelY1l1ll Myr capilloceum (With) Myr falcatum (Bedw) Myr HIllescens B S G
52 Climaciaceae aimecium Web et Mohr ex Mohr
americllnum Brid dedroides (Hedw) Web et Mohr
22
S3 Hedwigiaceae Hedwigia P Beauv
ciliata (Hedw) P Beauv
54 Cryphaeaceae Forsstroemia lindh
ohioense (Sull) lindh
55 Leucodontaceae Leucodon Schwaegr
brachypus Blid var brachypus brachypus Brid var aniJrewsitlnus Crum et Anderson julaceus (Hedw) SuU
56 Neckeraceae Neckera Hedw
pennata Hedw var pennata
1fomalia (Brid) B S G trichomanoides (Hedw) B S G
57 Theliaceae Thelia Sull
asprellll Sullo hirtella (Hedw) Sullo lescurii SuU
Myurella B SG julacea (Schwaegr) B S G
sibrica (C MueU) Reim var sibrica
58 Fabroniaceae Fabronio Raddi
ciliaris (Brid) Bnd
Anacamptodon Brid splachnoides (Groet ex Brid) Brid
Schwetschkeopsis Broth fabronia (Schwaegr) Broth
59 Leskeaceae Lindbergia Kindb
brachyptera (Mitt) Kindb
Leskea Hedw gracilescens Hedw obscura Hedw poiycarpa Hedw
Leskeella (limpr) Loeske nervosa (Brid) Loeske
23
Pterlgy1lllndrum Hedw filiforme Hedw
60 Thuidiaceae Haplohymenfum Dozy et MoIk
trlste (Ces ex De Not) Kiodb
Anomodon Hook et Tayl attenuatus (Hedw) Hueb minor (Hedw) Fuemr rostratus (Hedw) Schimp viticulosus (Hedw) Hook et Tayl
Haplocladium (C Mucn) C MueU microphyllum (Hedw) Broth virginillnum (Brld) Broth ~bull
Thuidium B S G abietinum (Hedw) B S G delicatulum (Hedw) B S G vardelicatulum delicatulum (Hedw) B S G var radicans (Kindb) Crum Steere et Anderson minutulum (Hedw) B S G pygmaeum B S G recognftum (Hedw) Lindb scitum (p Beauv) Aust
Helodium (Sull) Warnst blandowli (Web et Mohr) Warnst var blandowli blandowli (Web et Mohr) Wamst var helodioides (Ren et Card ex Roell) Crum
Steere et Anderson paludomm (Sull) AuS
61 Amblystegiaceae Cratoneuron (Sull) Spruce
commulatum (Hedw) Roth var commutfltum commutatum (Hedw) Roth var fakatum (Brid) Moenk ilicinum (Hedw) Spruce var filcinum
Campylium (Sun) Mitt chrysophyllum (Brid) 1 Lange var chrysophyllum hispidulum (Brid) Mitt var hllJidulum polygamum (B S G)C lens varpolygtlmUm stellatum (Hedw) C1ens vat stelltum
Leptodictyum (Schimp) Warnst laxirete (Card et Ther) Broth riparium (Hedw) Wamat trichopodium (Schultz) Warns l1acillans (Sull) Broth
Hygroamblystegium Loelke jlul1iatile (Hedw) oeske var luViatile
24
fluviatile (Hedw) Loeske var oWltum Grout noterophilum (Sun et Lesq ex Sull) Warnst tenox (Hedw) Jenn var tenax tenox (Hedw) Jenn var spinilolium Schimp) Jenn
Amblystegium BSG juratzkanum Schimp serpens (Hedw) BSG varlum (Hedw) Undb var varium
Platydictya Berk confervoides (Brid) Crum jungermanniodes (Brid) Crum minutissimum (Sull et Lesq ex Sull) Crum subtile (Hedw) Crum
Drepanocladus (C Muell) Roth aduncus (Hedw) Warnst var aduncus aduncus (Hedw) Warnst var capillilolius (Wamst)Riebm aduncus (Hedw) Warnst var kneiffii (BSG) Moenk aduncus (Hedw) Warnst var polycorpus (Bland ex Voit) Roth exannulatus (BSG) Warnst fluitans (Hedw) Warnst var fluitans revolvens (Sw) Wamst var revolvens sendtneri (Schimp) Warnst var sendtnerl uncinatus (Hedw) Warnst var uncinatus uncinatus ffiedw) Warnst var piwnosus (Schimp) Wamst vernicoms (Lindb ex C Hartm) Warnst
Hygrohypnum Lindb eugyrium (BSG) loeske luridum (Hedw Jenn varluridum ochraecum (Tum ex Wils) Loeske var ochraceunt
Calliergon (Sull) Kindb cordilolium Hedw Kindb glganteum (Schimp) Kindb var giganteum richardsonii (Mitt) Kindb ex Wamst stramineum (Brid) Kindb trilarium (Web et Mohr) Kindb
Calliergonella Loeske cuspidata (Hedw) Loeske
Calliergidium (Ren) Grout bakeri (Ren) Grout
Scorpidium (Schimp) Urnpr scorpiodides (Hedw) limpr
62 Brachytbeciaceae Tomenthypnum loeske
nitens (Hedw) Loeske var mtens
25
Homalotheciella (Card) Broth subcapillata (Hedw) Broth
Brachythecium BSG acuminatum (Hedw) Awt acutum (Mitt) Sullo calcareum IGndb campestTe (C MuelL) BSG cunum (Lindb) Umpr digastrum C Muell et Kindb ex Macoun et Kindb oxycladon (Brid) Jaeg et Sauerb var oxycladon plumosum (Hedw) BSG populeum (Hedw) BSG var populeum reflexum (Starke ex Web et Mohr) DsG var relexum reiexum (Starke ex Web et Mohr) DSG var pacificum Renet Card ex Roell rlvulare BSG var rivulare tabulum (Hedw) BSC var tabuum salebrosum (Web et Mohr) BSG var salebrosum velutinum (Hedw) BSG Vat velutinum
Bryhniil Kaur graminicolor (Brid) Grout var graminicolor novae-angliae (Sull et Lesq ex sun) Grout
Rhynchostegium BSG senulatum (Hedw) Ja4lg et Sauerb
Rhynchostegiella (BSG) Limpr compacta (C MueU) Loeske
EU1hynchium 8SG hians (Hedw) Sande Lac pulchellum (Hedw) Jenn var pulchelium riparloides (Hedw) Rich
Stokeriella (Kindb) Robins pmelonga (Hedw) Robins var stokesii (Tum) Crum
63 Entodontaceae Entodon C Muell
brevisetus (Hook et Wils ex Wils) Lindb cladorrhizans (Hedw) C Muell seductrlx (Hedw) C MueU var wluctrix
Pleurozium Mitt schreberl (8rid) Mitt var schreberi
64 Plagiotheciaceae Pltlglothecium BSG
CflVifolium (Brid) Iwats var CtWifolium denticulatum (Hedw) 8SG laetum BSG ltztebricola BSG
26
6S SernatophyUaceae Brotherella Loeske ex Fleisch
recumins (Michx) Fleisch tenuirostris (Bruch et Schimp ex Sull) Fleisch
Sematophyllum Mitt demissum (Wils) Mitt
66 Hypnaceae Pl(itygyrium BSG
repens (Brid) BSG
PylaisieUa Kindb intricata (Hedw) Grout polyantha (Hedw) Grout selwynii (Kindb) Crum Steere et Anderson
Homomallium (Schimp) Loeske adnatum (Hedw) Broth
Hypnum Hedw cupressiforme Hedw var cupressiforme cupressifonne Hedw varfiliforme Brid curviolium Hedw fertile Sendtn imponens Hedw lindbergii Mitt palescens (Hedw) P-Beauv pratense Koch ex Brid
Calicladium Crum haldanianum (Grev) Crum
sopterygiopsis Iwats muelleriana (Schimp) Iwats
Isopterygium Mitt e1egans (Brid) lindb pulchellum (Hedw) Jaeg et Sauerb tenerum (Sw) Mitt
Taxiphyllum Fleisch depllQtum (Bruch et Schimp ex sun) Fleisch taxirameum (MitL) Fleisch
Herzogiella Broth seligeri (Brid) Iwats striatella (Brid) Iwats ruracea (Lindh) Iwats
hilium De Not crista-castrenna (Hedw) De Not
27
67 Rhytidiaceae Rhytidium (Sull) Kindb
rugosum (Hedw) Kindb
Rhytidfodelphus (Lindb ex Iimpr) Warnst triquetrus (Hedw) Warnst var triquetrus
68 Hylocomiaceae Hylocomium BSG
pyrenaicum (Spruce) lindb splendens (Hedw) BSG varsplendens
69 Buxbaumiaceae Buxbaumia Hedw
aphyllll Hedw
Diphyscium Mohr follosum (Hedw) Mohr
70 Tetraphidaceae Tetraphi3
pellucida Hedw
71 Polytrichaceae Atrichum PmiddotBeauv
angustatum (Brid) BSG cri8pum (James) Sullo oerstedianum (C Muell) Mitt teneUum (Roehl) BSG undulatum (Hedw) P middotBeauv var unduiatum
Pogonatum P-Beauv alpinum (Hedw) Roehl var alpinum umigerum (Hedw) P-Beauv
Poiytrichum Hedw commune Hedw var commune ormosum Hedw juniperinum Hedw longisetum Brid ohioense Ren et Card piliferum Hedw var pilierum strictum Brid
28
MAPS OF BRYOPHYTE TAXA
REPORTED FOR WISCONSIN
HORNWORTS AND LIVERWORTS
29
bull bull WSHeV~N S vE II
0 ~
e~RRON ~USK
(J~ (j -
CHIPPE A DUNN
0 o0 poundAU CLAIRE
~ TREMshyPEAshyLEAU
MICE
ONEIDA
Q LINCOLN
TAYLOR
Q t-~----- MOIRA THON
CLARK
o
10
LA FAVETTE
Hornwor ts amp Liverworts
- None - 1 - 25 taxa
- 26 - 50 tRXshy
- 51 - 75 tax)
- 7c - 100 taxa
FONDOU j LAC ~
DODGE
i O JEfFERSON
WALWORTH
30
Anastrophylltlfl helleranum (Nees) Schust Anastrophyllum michauxil (We~ Buch ~
46 88 136 141 Evans 38 46 141
Anastrophyllum mlnut~ (Schreb) Schuster Aneura plnguls (L) Durn
89 21 384687 88 89 90 (138) (145)
Anthoceros macounil MA Howe ~oceros punctatus L
21 (45) (144) 34
31
Barbilophozia attenuata (Mart) Loeske Barbilophozia barbata (Schmid ~ Schre~
24 141 Loeske 28 39 46 89 90 141
Bazzania trilobata (1) S Gray
1 26 39 46 (57) (59) 79 88 89
90 119 141
BlePbarostoma trichoffiyllum (1) Dumort
1 8 26 39 46 89 90 137 141
Blasia pusilla 1
1 39 46 90
Calypogeja fissa (1) Raddi
90
32
$129geja integristipula Steph C~Ypound2geja muelleriana (Schifm) K Mull
88 89 89
CallpoundO~eja neesiana (Mass amp Carest) K Mull
39 46 89 140 (141)
~pogeja sJlapound2icola (H Arnell tl J bull
Perss) Warnst et Loeske (119) 141
CeIalozia biscuspidata (L) Dum
1 37 I 46 90 t 137 141
CalYpogej~ trichomanis (L) Corda
26 39 (119) 137
33
CeEhalozia catenulata (Hub) Lindb Cephalozia connivens (Dicks) Lindb
39 46 79 141 39 4679 88 (89) 90 119 141
Cephalozia lacinulata Jack ~ Spruce
141 Cephalozia lunulifolia (Dum~ Dum I 28 39 46 79 88 89 90 141
Cepoundhalozia macouni (Aust) Aust Cephalozia ~leniceps (Aust) Lindb 27~ 3746 (66)7987 (139)141 39 46 79 89 141
34
Ceffialoziella 9ivaricata (Sm) Schiffn Cephaloziella ~hista (Jack) Schiffn
46 90 46 (119) 141
Cephaloziella hampeana (Nees) Schiffn
46
Cephaloziella rubella (Nees) Warnst
46 39
Cephaloziella rubella var sullivantii
(Aust) K Mull 46 39
Cephaloziella rubella var bifida (Lindb) Douin
35
46
Chiloscyphus pallescens (Ehrh ~ Hof~ Dum Chiloscyphus pallescens var fragilis
39J 46 88 89J 90 (Roth) K Milll 46
Chiloscyphus E9lyanthos (L) Corda
1 24 39 46
Chiloscyphus polyanthos vax rivularis
(Schrad) Nees 39 46
poundpoundnocepha1um conicum (1) Lindb
12 28 39 46 (49) 87 88 8990 120 121 148
Cololejeunea biddlecomiae (Aust) Evans
39 46 62 87 88 89 90
36
Dipl0]hyllum apicu1atum (Evans) Steph Fossombronia foveo1ata Lindb
(35)41 46 90 140 1 t 38 (39) t 46
Frullania bolanderi Aust
13 46 62 88 90 139 140
Frullanla ari ttoniae Evans
39 46 69 (71) 87 88 89 90
Frullania eboracensis Gott
1 9 28 4 39 46 (57) (59) 69 70 (71) 87 88 89 90 92 93 97 148
Frullania inflata Gott
46 87 88 89 (139)
Frullania oakesiana Aust Frullania plana Sullo
(35) 42 46 87 (139) (140) 92
Frullania ri paria Hampe ~ Lehm
46 69 71 87 89
Frullania ~elwynianna Pears
46 89
Geocalyx SEaveolens (Schrad) Nees
394688 8990 (119) 140
Frullania tamarisci (L) Dum subsp sPagrayana
(Mont) Hatt 28 34 l~6 t (57) 88 89 90
38
Mar pan thus seutatUB ( Web amp Mohr) Spruce Jamesoniella autumnalis (DC) Steph
(35) 39 46 90 1 7 28 39 46 56 (57) 87 8889 90 (119) 137 141
Jungermannia confertissima Nees
137 141
Jungermannia crenuliform1s Aust
79 137 141
Jungermannia ~il1ima Sm
46 90 141 Jungermannia exsert1folia subsp corifo~ (Dum~
Vana (35) 46 79 141
39
Jungermannia hyalina Lyell Jungerrr6nnia 1eiantha Grolle
40 46 79 136 141 39 46 79 88 90 1)7 141
Jungermannia IXlaxis Lindb Jungermannia pumila With
46 67 79 (139) (140) 141 )9 46 90 1)6 141
Jungermannia sphaerocarpa Hook Kurzia setacea (Web) Grolle
4679136140IIH 136
Lejeunea cavifo1ia (Ehrh) Lindb Lepidozia reptans (1) Dum
1 7 27 39 46 88 89 90 137 14010 I ( 27) 46 I 69 (71) 88
lophocolea heterophylla (Schrad) Dum Lophocolea minor Nees
1 )4 39 46 (57) 87 88 89 27 39 46 88 89 90 (119) 90 (93)94 119 136
Lophozia al~stris (Schleich ~ Web) Evans Lophozia ascendens (Warnst) Schust
46 90 137 141 136 140 141
41
loIilozia badensls (Gatt ~ Rabenh)
46 141
lophozia blcrenata (Schmid sectA Hoffm)
Dum 46 141
lophozia collaris (Nees) Dum
39 46 141
lophozla excisa (Dicks) Dum
27 4679 141
lophozia gillmani (Aust) Schust lophozia gIlttulata (Undo tl H Arnell)
36 39 40 46 136 (140) 141 Evans 39 46 136 141
42
141
Lophozia heterocolEDs (Thed) MA Howe IoPhozia ipcisa (Schard) Dum
27 39 46 89 141 8 27 31 46 79 90 140 141
LoQhozia longidens Macoun Lophozia ventricosa
1 46 79 90 141
Lophozia ventricosa var silvicola
(Buch) Jones 141 LoEhozla ventricosa vax longif1ora (Nees)
Macoun 39 46 79 141
43
~~ fragrans (Balbis) Frye et Clark Mannia pi 10sa (Hornem) Frye et Clark
46 (139) (140) 9 46 79 116 139
Mannia triandra (Scop) Gro 11e
46 (140)
Marchantia pound91ymorpha L
1 13 28 3946 (49) 78 87 88 89 90 (Ill) 121 122 147
Mo~rckia hibernica (HookJ Gott
46 89
Metzgeria conjugata Lindb
44
46
Mylia anomala (Hook) S Gray Notothylas orbicularis (Schwein) Sullo
39 46 141 (35)45 46
Nowellia curvifolia (Dicks) Mitt
17 27 3846 88 89 90 141
Odontoschisma denudatuJI1 (Nees ~ Mart)
Dum 46 81 137 141
Pallavicinia lyellii (Hook~ Carruth
90
Odontoschisma macounii (Aust) Under
45 46 (66) (139) 140 141
45
=
Pellia epiphllla (L) CordaPellia endiviifolia (Dicks~ Dum
27 39 46 90 13724 39 46
Pellia neesiana Limp bull Phaeoceros
39 46 89 27 39 45 46 97
Porella pinnata L
13 27 I 46 90 Plagiochila asplenioides (L) Dum
27 I 39 1+6 88 89 I 90 137
46
~e11a platYEhylla (L) Pfeiff Porella platyphylloi~ (Schwein) Lindb
17 27 39 46 69 (71)79 87 88 89 9 34 46 (57) 59 87 88 89 92 90 92 93 77
Preissia quadrata (Seop) Nees
1 36 3946 (71) 88 89 90 121 123
137
Ptllidium ciliare (L) Hampe
1 27 39 46 132 141
Dum
(22) 28 3946 (57) 87 88 89 90 92
Ptilidium ~lcherrium (G Web) Hampe
(9) 13 394656 (57)70 87 88 89 90 93 (119) 136 141
47
Radula obconica Sullo Reboulia hemlsphaerlca (L) Raddi
39 46 88 89 (43) 46 (59) 70 90 t Cf 124 125 (139)
Riccardia ~atifrons Lindb
1 27 39 46 89
Riccardia multifi~ (L) S Gray
39 46 88 145
Riccia beyrichiana Hampe ~ Lehm
46 Riccardia palmata (Hedw) Carruth
39 46 90
48
Riccia bifurca Hoffm Riccia cavernosa Hoffm
48 13739 46 79 137
Riccia fluitans L
242731454679
Riccla frostli Aust
48 79
Riccia sullicantii Aust
79 (137) Riccia sorocarpa Bisch 46 (137)
49
Ricciocarpus natans (1) Corda Sca~ia apiculata Spruce
13 27 46 79 89 110 39 46 (139) 141
ScaJltll1ia carinthica Jack ~ Lindb
137 t 141
Scapania ~ (Mart) Durn
39467990137141
Scapania digenil Schiffn
90
ScaEsectflia cuspiduligera (Nees) K Mull
46 (139) (140) (141)
50
Scapania glaucoeerha1a (Tayl) Aust Seapania gyrrmostomophilia Kaal
39 46 (139) 141 41 44 46 64 (139) (140) 141
ScaP3poundlia irrigua Gott ~ almiddot
7 39 46 79 137 (140L 141
Seapania nemorosa (L) Dum
1 39 46 89 90 137
Scapmi~ ~~ta Smiddotlch
137 141 51
~caPnia poundludicola LDeske amp K Mull Scapania saxicola Schust
46 (66) 141 136 141
Sc~~ scandica (H Arnell ~ Buch) Macv
139 140
Scapania subalpina (Nees) Durn
39 40 46 79 137 141
Scapania undulata (L) Dum
1 42 46 79 141
Scapania umbrosa (Schrad) Dum
137 140 141
52
Trichocolea tomentella (Ehrh) Dum Triomarla exsecta (Schmid) Schiffn
27 (35) 39 46 47 88 137 141 8 28 39 46 90 140 141
Tritomaria exsectiformis (Breidl) Loe~ke Tritomaria Sluinguedentata (Huds) Buch
39 -t6 89 90 136 140 141 7 39 46 141
Trltomaria scitula (Tayl) Joerg
46 136 (139) 141
53
cuspiduligera (Nees) K Mull var cuspiduiigera degenii Schlffn ex K Mull var degenii glllucocephaJa (Tayt) Aust gymnostomophila Kaal irrigwJ (Nees) Gott et al subsp irrigua lingulata Buch varlingulata mucronata Buch subsp mucronata nemorosa (l) Dum paludicola Toeske et K Mun var paludicola paludosa (K Mull) K Mull saxicola Schust scandiea (H Arnell et Buch) Macv var scandiea 8Ubalp~ (Nees) Dum umbroSll (SchIad) Dum undulata (L) Dum var undulata
15 Radulaceae Raduia Dum nom COllS
compianata (L) Dum obconica Sull
16 Porellaceae Parella L
pinTlllta L patyphylla (L) Pfeiff platyphylloidt(l (Schwein) lindb
17 Jubulaceae FruUanill Raddi
bolanderi Aust brittoniae Evans eboracemis Gott inlllta Gott oakesiana Aust plana Sull rlparia Hampe ex Lehm selwyniana Pears taTnl1iJci (L) Dum subsp asagmyana (Mont) Hatt
18 lejeuneaceae Calolejeunea (Spruce) Schlffn
biddlecomiae (Aust) Evans
Lejeunea Ub nom cons cavifolia (Ehrh) Lindb emend Buch
19 Codoniaceae Fossombronia Raddi
foveolata Lindb
20 Pa11aviciniaceae Paklvieinlil S Gray nom cons
lyeiii (Hook) Carruth
14
Moerckia Gott hibemica (Hook) Gott
21 PeWaceae Pellfa Raddi nom cons propos
endivlifolfa (Dicks) Dwn epiphylllz (L) Corda neeBiana (GottlUmpr
22 Metzgeriaceae Metzgerla Raddi
conjugata Undb
23 Blasiacae B L
pulilm L
24 Aneurareae AneumDum
pinguis (L) Dum
Riccfl1dia S Gray nom cons latifrons Undb multiftda (L) S Gray var multifida palmata (Hedw) Carruth
25 Aytoniaceae Reboulill Raddi nom conI
hemllphaerica (L) Radeli
ManniaOpiz fragrrms (Balbis) Frye el ClarK pilosa (Hornem) Frye et Clark trltmdra (Scop) GroDe
26 Conocephalaceae Conocephalum Wigg nom cons
conicum (L) Lindb
27 Marchantiaceae Preiuia Corda
quadrata (Scop) Nees
Marchantia L polymorpha L
28 Ricciaceae Riccwcarpus Corda
natans (L) Corda
Riccia 1 beyrichiana Hampe ex Lehm bifurca Horfm
15
i
CtlJPemostI Hoffnt fuitu L rostii Aust BOIOCtI1fHI Bi~ sulltvtmtii Aust
MUSCI (Mosses)
29 Sphagnaceae SpIutgnum L
capillaceum (Weiss) Schrank var Cllpil1tlceum caplllllceum (Weiss) Vir tenellum (Schimp) Andr centrille C Jens ex Amen et C Jens compactummiddot OC ex Lam et Dc ClUpidatum Ehrh ex Hoffin var aupidatum jimbrltltum Wits ex J Hookmiddot fulCUm (Scbimp) KUngrr glrgenBOhIlll Russ imbrlctltum Hornsch ex Russ muelllmitil4m BOd 1IIIIU (RUII) C Jens przIutre L papIIIoIum Lindb pulchrum (Lindb ex braithw) Wamst recumun P-Beauv var recumun recum P-Beauv var tenue Klinggr riptlrlum Ansstr ruPQWItWamst squtl1fOlUm Crome subftltens Russ et Wamst ex Wamst IUbIeCfUldum Nees ex Sturm var subsecundum subsecundum Nees ex Sturm varrutem (Nees Hormch et sUUm) Hueb teres (Sdlimp) Anptr e C Hartm
WtlI7IItorjii Russ wuljfanum Gira
30 Andreaeaceae AndNrrerI Hedw
rupeitrlr Oedw var pest
31 Fissident~e Fluid_ lledw
trdiantoid Hedw byoidei Bedw var bryolda cristaM Wils ex Mitt exiguus sun VIr exigUw onttmUi (B-Pyl) Steud jtuberi Lesq et James obtusifolhu Wits var obtwifoliul ollPlUlldQldes Heel ~litSuU lubbtulkril Bed tfDdfoliul lledw llridubu (S~ ) Wahlenb vallirlIU1u
16
32 Ditrichaceae Pleuridium Brid
subulatum (Hedw) Rabenh
Ditrichum Hampe heteromallum (Hedw) Britt linellre (Sw) lindb
~ pallidum (Hedw) Hampe pusiUum (Hedw) Hampe
Saelania Lindb gIlucescens (Hedw) Bomanss et Broth
Ceratodon Brid purpureus (He(lw) Brid var prupurew
Distichium BSG capillaceum (Hedw) BSG inclinatum (Hedw) BSG
33 Bryoxipbiaceae Bryoxlphium Mitt
norvegicum (Brid) Mitt
34 Seligeriaceae Seligeria BSG
cumpylopoda Kindb ex Macoun et Kindb pusillll (Hedw) BSG recurvata (Hedw) BSG
Bindia BSG acuta (Hedw) BSG
35 Dicranaceae BlUehia Schwegr
lexUOSll (Sw ~ Schwaegr) C MueU sullivantii Aust
Trematodon Michx ambiguus (Hedw) Hornsch
Dicranellll (C Muell) Schimp cerviculata (Hedw) Schimp val cillPicuiata
heteromalla (Hedw) Schimp ~ rufescens (With) Schimp
schreberiana (Hedw) Schhnp var schreberiana schreberitJna (Hedw) Schimp var robusta Schimp ex Braithw subulata (Hedw) Schimp varia (Hedw) Schimp
Dicranodontium BSG denudatum (Brid) Britt ex Williams
Panzeucobryum (Lindh) Loeske Iongifolium (Hedw) oelke varlongifolium
17
Tortella (Undb) Limpr fragilis (Drumm) Umpr humilis (Hedw) JeM torlUoSfl (Hedw) Umpr
Hyophila Brid involuta (Hoole) Jaeg et Sauerb
Didymodon Hedw rigidulus Hedw
BryoerythrophyUum Chen recurvirostnon (Hedw) Chen
Btubula Hedw acula (Brid) Brid var acuta convoiuta Hedw fallox Hedw unguicultltll Hedw
Acaulon C MueD muticum (Uedw) C Muell var rufeseens (Iaeg) Crum
PJl~lm Hedw CUSPidtltum Hedw
De8mlltodon Brid cernuu (Hueb) BSG oblUsifoliw (Schwaegr) Schimp
T01IUla Hedw mueronifolia Schwaegr rurall (Hedw) Gaertn Meyer et Scherb var ruralis
39 Grimmiaceae Grlmmill Hedw
alpicola Hedw var alpicola alpicola Hedw var dupretii (Ther) Crum alpicola Hedw var rlvularl8 (Brid) Wahlenb apocarpa Hedw var apocarpa apocarpa Hedw var ambipa (Sull) Jonea apocarpa Hedw var conjerttl (Funck) Spreng apocarpa Hedw var strleta (Tum) Hook et Tayl CiIIyptrrlta Hook ex Drumm donniana Sm var donn_ hemumti Crum ltlevla (Brid) Brid montana BSG oineyi SuD oVllll8 (Hedw) lindb piUfe P-Beauv pltJgiopodill Hedw pulvinata (Hedw) Sm varbullpulvinata teretinervl Limpr
18
Orothera C Muell leana (Sull) C Muell
Rfwbdoweisia crispata (With) Undb
Cynodontium BSG ex Schimp alpestre (Wahlenb) Milde strumiferum (Hedw) Undb
Dichodontium Schimp pellucidum (Hedw) Schimp
Oncophorus (Slid) Brid vuens (Hedw) Brid wahlenbergii (Brid)
Dicranum Hedw bonjeanii DeNot ex Usa drummondii C Muell lagellare Hedw varj1ogellare fulvum Hook fuscescens Turn var fuscescens montanum Hedw muehlenbeckii BSG var muehlenbeckii polysetum Sw sabuletorum Ren et Card scoparium Hedw undulatum Brid viride (Sull et Lesq ex Sull) lindb
36 Leucobryaceae Leucobryum Hampe
glaucum (Hedw) Angstr ex Fr
37 Encalyptaceae Encalypta Hedw
ciliata Hedw var ciliata procera Bruch
38 Pottiaceae Astomum Hampe
muhlenbergianum (Sw) Grout
Weissia Hedw controversa Hedw var controversa
Gymnostomum Nees Homsch et Stunn aeruginosum Sm recurvirostrum Hedw var recurvirostrum
Trichostomum Bruch spirale Prout
19
40 Ephemeraceae Ephemerum Hampe
cohaerens (Hedw) Hampe crassineium (Schwaegr) Hampe val crassinervium spinulosum Bruch et Schimp ex Schimp
41 Funariaceae AphflllolThegma SuD
serratum (1 Hook et Wils ex Drumm) Sullo
Physcomitrium (Brid) Fuernr hookeri Hampe immersum Sullo pyriforme (Hedw) Hampe
Funaria Hedw americana Undb hygrometrica Hedw vaT hygrometrica
42 Splachnaceae Splachnum Hedw
ampullaceum Hedw
43 Schistostegaceae Schistostega Mohr
pennata (Hedw) Web el Mohr
44 Bryaceae Pohlia Hedw
annolina (Hedw) Lindb var annotina camea (Tum) Lindb cruda (Hedw) Undb elongata Hedw iliformis (Dicks) Andr lescurlana (Sull) Grout nutans (Hedw) Lindb obrusifolia (Brid) L Koch proligera (Undb ex Umpr) Undb ex Amell wahlenbergii (Web et Mohr) Andr
Leptobryum (B S G) Wits pyriforme (Hedw) Wils
Bryum Hedw algovicum Sendtnex C Muell argenteum Hedw caespiticium Hedw capillare Hedw var capillare creberrlmum Tayt lonchocaulon C Muell pallens (Brid) Sw ex Roehl pseudotrlquetrum (Hedw) Gaertn Meyer et Scherb varpudotriquetrum
20
pseudotriquetrum (Hedw) GaertnMeyer et Scherb var bimum (Schreb) Ulj stenotrichum C Muell var stenotrichum tortiolium Funck ex Brid uliginosum (Brid) B S G violaceum Crundw et Nyh weigelii Spreng
Rhodobryum (Schimp) Hampe roseum (Hedw) Limpr
45 Mniaceae Mnium Hedw
ciliare (C Muell) Schimp cinclidioides Hueb cuspidarum Hedw drummondii Bruch et Schimp longirostrum Brid lycopodioides Schwaegr marginatum (With) Brid ex P Beauv mediumB~S G var medium onhorrhynchum Brid pseudopunctatum Bruch et Schimp punctalUm Hedw var punctatum punctatum Hedw var elatum Schirnp rugicum Laur spinufosum B S G stellare Hedw
46 Aulacomruaceae Au1acomnium Schwaegr
androgynum (Hedw) Schwaegr heterostichum (Hedw) B S G paJustre (Hedw) Schwaegr var paJustre
47 Meesiaceae Amblyodon B S G
deillbotus (Hedw) B S G
Meesia Hedw triquetra (Richt) Angstr uJiginosa Hedw
48 Bartramiaceae Plagiopus Brid
oederiana (Sw) Umpr
Bartramia Hedw pomiformis Hedw var pomiformis
Philonotis Brid fontana (Hedw) Brid varfontana longseta (Michx) Britt varlongseta marchica (Hedw) Brid
21
~
~
49 Timmiaceae Timmia Hedw
megapolitana Hedw var megapolitana
50 Orthotrichaceae Zygodon Hook et Tayl
viridissimus (Dicks) Brid var rupestris lindb ex C J Hartm
Amphidium Schimp lapponicum (Hedw) Schimp mougeotii (B S G) Schimp
Orthotrichum Hedw anomalum Hedw obtuaifolium Brid ohioense Sullo et Lesq ex Aust pumilumSw sordidum Sullo et Lesq ex Aust speciosum Neesax Sturm var eegam (Schwaegr ex Hook et Grev) W~rnst stellDtwn Brid strrmgulatum P Beauv
Uleta Mohr ex Web cOfIICtata (p Beallv) Hamrn crispa (Hedw) Brid
Drummondill Hook ex Drumm prorepens (Hedw) Brltt
51 Fontinalaceae Fantinalis Hedw
antipyretica Hedw var antipyrettca antipyretica Hedw forma rabusta (Card) Card dalecarlica Schimp ex B S G var dalecarlica flqccidtJ Ren et Card hypnoides C J Hartrn var hypnoides hypnoides C J Hartm var durillei (Schimp) HU$ll
missourlcaCard novae-angliae Sullo var novae-angliae splulgrUlolill (C Moen) Wijket M~S
DichelY1l1ll Myr capilloceum (With) Myr falcatum (Bedw) Myr HIllescens B S G
52 Climaciaceae aimecium Web et Mohr ex Mohr
americllnum Brid dedroides (Hedw) Web et Mohr
22
S3 Hedwigiaceae Hedwigia P Beauv
ciliata (Hedw) P Beauv
54 Cryphaeaceae Forsstroemia lindh
ohioense (Sull) lindh
55 Leucodontaceae Leucodon Schwaegr
brachypus Blid var brachypus brachypus Brid var aniJrewsitlnus Crum et Anderson julaceus (Hedw) SuU
56 Neckeraceae Neckera Hedw
pennata Hedw var pennata
1fomalia (Brid) B S G trichomanoides (Hedw) B S G
57 Theliaceae Thelia Sull
asprellll Sullo hirtella (Hedw) Sullo lescurii SuU
Myurella B SG julacea (Schwaegr) B S G
sibrica (C MueU) Reim var sibrica
58 Fabroniaceae Fabronio Raddi
ciliaris (Brid) Bnd
Anacamptodon Brid splachnoides (Groet ex Brid) Brid
Schwetschkeopsis Broth fabronia (Schwaegr) Broth
59 Leskeaceae Lindbergia Kindb
brachyptera (Mitt) Kindb
Leskea Hedw gracilescens Hedw obscura Hedw poiycarpa Hedw
Leskeella (limpr) Loeske nervosa (Brid) Loeske
23
Pterlgy1lllndrum Hedw filiforme Hedw
60 Thuidiaceae Haplohymenfum Dozy et MoIk
trlste (Ces ex De Not) Kiodb
Anomodon Hook et Tayl attenuatus (Hedw) Hueb minor (Hedw) Fuemr rostratus (Hedw) Schimp viticulosus (Hedw) Hook et Tayl
Haplocladium (C Mucn) C MueU microphyllum (Hedw) Broth virginillnum (Brld) Broth ~bull
Thuidium B S G abietinum (Hedw) B S G delicatulum (Hedw) B S G vardelicatulum delicatulum (Hedw) B S G var radicans (Kindb) Crum Steere et Anderson minutulum (Hedw) B S G pygmaeum B S G recognftum (Hedw) Lindb scitum (p Beauv) Aust
Helodium (Sull) Warnst blandowli (Web et Mohr) Warnst var blandowli blandowli (Web et Mohr) Wamst var helodioides (Ren et Card ex Roell) Crum
Steere et Anderson paludomm (Sull) AuS
61 Amblystegiaceae Cratoneuron (Sull) Spruce
commulatum (Hedw) Roth var commutfltum commutatum (Hedw) Roth var fakatum (Brid) Moenk ilicinum (Hedw) Spruce var filcinum
Campylium (Sun) Mitt chrysophyllum (Brid) 1 Lange var chrysophyllum hispidulum (Brid) Mitt var hllJidulum polygamum (B S G)C lens varpolygtlmUm stellatum (Hedw) C1ens vat stelltum
Leptodictyum (Schimp) Warnst laxirete (Card et Ther) Broth riparium (Hedw) Wamat trichopodium (Schultz) Warns l1acillans (Sull) Broth
Hygroamblystegium Loelke jlul1iatile (Hedw) oeske var luViatile
24
fluviatile (Hedw) Loeske var oWltum Grout noterophilum (Sun et Lesq ex Sull) Warnst tenox (Hedw) Jenn var tenax tenox (Hedw) Jenn var spinilolium Schimp) Jenn
Amblystegium BSG juratzkanum Schimp serpens (Hedw) BSG varlum (Hedw) Undb var varium
Platydictya Berk confervoides (Brid) Crum jungermanniodes (Brid) Crum minutissimum (Sull et Lesq ex Sull) Crum subtile (Hedw) Crum
Drepanocladus (C Muell) Roth aduncus (Hedw) Warnst var aduncus aduncus (Hedw) Warnst var capillilolius (Wamst)Riebm aduncus (Hedw) Warnst var kneiffii (BSG) Moenk aduncus (Hedw) Warnst var polycorpus (Bland ex Voit) Roth exannulatus (BSG) Warnst fluitans (Hedw) Warnst var fluitans revolvens (Sw) Wamst var revolvens sendtneri (Schimp) Warnst var sendtnerl uncinatus (Hedw) Warnst var uncinatus uncinatus ffiedw) Warnst var piwnosus (Schimp) Wamst vernicoms (Lindb ex C Hartm) Warnst
Hygrohypnum Lindb eugyrium (BSG) loeske luridum (Hedw Jenn varluridum ochraecum (Tum ex Wils) Loeske var ochraceunt
Calliergon (Sull) Kindb cordilolium Hedw Kindb glganteum (Schimp) Kindb var giganteum richardsonii (Mitt) Kindb ex Wamst stramineum (Brid) Kindb trilarium (Web et Mohr) Kindb
Calliergonella Loeske cuspidata (Hedw) Loeske
Calliergidium (Ren) Grout bakeri (Ren) Grout
Scorpidium (Schimp) Urnpr scorpiodides (Hedw) limpr
62 Brachytbeciaceae Tomenthypnum loeske
nitens (Hedw) Loeske var mtens
25
Homalotheciella (Card) Broth subcapillata (Hedw) Broth
Brachythecium BSG acuminatum (Hedw) Awt acutum (Mitt) Sullo calcareum IGndb campestTe (C MuelL) BSG cunum (Lindb) Umpr digastrum C Muell et Kindb ex Macoun et Kindb oxycladon (Brid) Jaeg et Sauerb var oxycladon plumosum (Hedw) BSG populeum (Hedw) BSG var populeum reflexum (Starke ex Web et Mohr) DsG var relexum reiexum (Starke ex Web et Mohr) DSG var pacificum Renet Card ex Roell rlvulare BSG var rivulare tabulum (Hedw) BSC var tabuum salebrosum (Web et Mohr) BSG var salebrosum velutinum (Hedw) BSG Vat velutinum
Bryhniil Kaur graminicolor (Brid) Grout var graminicolor novae-angliae (Sull et Lesq ex sun) Grout
Rhynchostegium BSG senulatum (Hedw) Ja4lg et Sauerb
Rhynchostegiella (BSG) Limpr compacta (C MueU) Loeske
EU1hynchium 8SG hians (Hedw) Sande Lac pulchellum (Hedw) Jenn var pulchelium riparloides (Hedw) Rich
Stokeriella (Kindb) Robins pmelonga (Hedw) Robins var stokesii (Tum) Crum
63 Entodontaceae Entodon C Muell
brevisetus (Hook et Wils ex Wils) Lindb cladorrhizans (Hedw) C Muell seductrlx (Hedw) C MueU var wluctrix
Pleurozium Mitt schreberl (8rid) Mitt var schreberi
64 Plagiotheciaceae Pltlglothecium BSG
CflVifolium (Brid) Iwats var CtWifolium denticulatum (Hedw) 8SG laetum BSG ltztebricola BSG
26
6S SernatophyUaceae Brotherella Loeske ex Fleisch
recumins (Michx) Fleisch tenuirostris (Bruch et Schimp ex Sull) Fleisch
Sematophyllum Mitt demissum (Wils) Mitt
66 Hypnaceae Pl(itygyrium BSG
repens (Brid) BSG
PylaisieUa Kindb intricata (Hedw) Grout polyantha (Hedw) Grout selwynii (Kindb) Crum Steere et Anderson
Homomallium (Schimp) Loeske adnatum (Hedw) Broth
Hypnum Hedw cupressiforme Hedw var cupressiforme cupressifonne Hedw varfiliforme Brid curviolium Hedw fertile Sendtn imponens Hedw lindbergii Mitt palescens (Hedw) P-Beauv pratense Koch ex Brid
Calicladium Crum haldanianum (Grev) Crum
sopterygiopsis Iwats muelleriana (Schimp) Iwats
Isopterygium Mitt e1egans (Brid) lindb pulchellum (Hedw) Jaeg et Sauerb tenerum (Sw) Mitt
Taxiphyllum Fleisch depllQtum (Bruch et Schimp ex sun) Fleisch taxirameum (MitL) Fleisch
Herzogiella Broth seligeri (Brid) Iwats striatella (Brid) Iwats ruracea (Lindh) Iwats
hilium De Not crista-castrenna (Hedw) De Not
27
67 Rhytidiaceae Rhytidium (Sull) Kindb
rugosum (Hedw) Kindb
Rhytidfodelphus (Lindb ex Iimpr) Warnst triquetrus (Hedw) Warnst var triquetrus
68 Hylocomiaceae Hylocomium BSG
pyrenaicum (Spruce) lindb splendens (Hedw) BSG varsplendens
69 Buxbaumiaceae Buxbaumia Hedw
aphyllll Hedw
Diphyscium Mohr follosum (Hedw) Mohr
70 Tetraphidaceae Tetraphi3
pellucida Hedw
71 Polytrichaceae Atrichum PmiddotBeauv
angustatum (Brid) BSG cri8pum (James) Sullo oerstedianum (C Muell) Mitt teneUum (Roehl) BSG undulatum (Hedw) P middotBeauv var unduiatum
Pogonatum P-Beauv alpinum (Hedw) Roehl var alpinum umigerum (Hedw) P-Beauv
Poiytrichum Hedw commune Hedw var commune ormosum Hedw juniperinum Hedw longisetum Brid ohioense Ren et Card piliferum Hedw var pilierum strictum Brid
28
MAPS OF BRYOPHYTE TAXA
REPORTED FOR WISCONSIN
HORNWORTS AND LIVERWORTS
29
bull bull WSHeV~N S vE II
0 ~
e~RRON ~USK
(J~ (j -
CHIPPE A DUNN
0 o0 poundAU CLAIRE
~ TREMshyPEAshyLEAU
MICE
ONEIDA
Q LINCOLN
TAYLOR
Q t-~----- MOIRA THON
CLARK
o
10
LA FAVETTE
Hornwor ts amp Liverworts
- None - 1 - 25 taxa
- 26 - 50 tRXshy
- 51 - 75 tax)
- 7c - 100 taxa
FONDOU j LAC ~
DODGE
i O JEfFERSON
WALWORTH
30
Anastrophylltlfl helleranum (Nees) Schust Anastrophyllum michauxil (We~ Buch ~
46 88 136 141 Evans 38 46 141
Anastrophyllum mlnut~ (Schreb) Schuster Aneura plnguls (L) Durn
89 21 384687 88 89 90 (138) (145)
Anthoceros macounil MA Howe ~oceros punctatus L
21 (45) (144) 34
31
Barbilophozia attenuata (Mart) Loeske Barbilophozia barbata (Schmid ~ Schre~
24 141 Loeske 28 39 46 89 90 141
Bazzania trilobata (1) S Gray
1 26 39 46 (57) (59) 79 88 89
90 119 141
BlePbarostoma trichoffiyllum (1) Dumort
1 8 26 39 46 89 90 137 141
Blasia pusilla 1
1 39 46 90
Calypogeja fissa (1) Raddi
90
32
$129geja integristipula Steph C~Ypound2geja muelleriana (Schifm) K Mull
88 89 89
CallpoundO~eja neesiana (Mass amp Carest) K Mull
39 46 89 140 (141)
~pogeja sJlapound2icola (H Arnell tl J bull
Perss) Warnst et Loeske (119) 141
CeIalozia biscuspidata (L) Dum
1 37 I 46 90 t 137 141
CalYpogej~ trichomanis (L) Corda
26 39 (119) 137
33
CeEhalozia catenulata (Hub) Lindb Cephalozia connivens (Dicks) Lindb
39 46 79 141 39 4679 88 (89) 90 119 141
Cephalozia lacinulata Jack ~ Spruce
141 Cephalozia lunulifolia (Dum~ Dum I 28 39 46 79 88 89 90 141
Cepoundhalozia macouni (Aust) Aust Cephalozia ~leniceps (Aust) Lindb 27~ 3746 (66)7987 (139)141 39 46 79 89 141
34
Ceffialoziella 9ivaricata (Sm) Schiffn Cephaloziella ~hista (Jack) Schiffn
46 90 46 (119) 141
Cephaloziella hampeana (Nees) Schiffn
46
Cephaloziella rubella (Nees) Warnst
46 39
Cephaloziella rubella var sullivantii
(Aust) K Mull 46 39
Cephaloziella rubella var bifida (Lindb) Douin
35
46
Chiloscyphus pallescens (Ehrh ~ Hof~ Dum Chiloscyphus pallescens var fragilis
39J 46 88 89J 90 (Roth) K Milll 46
Chiloscyphus E9lyanthos (L) Corda
1 24 39 46
Chiloscyphus polyanthos vax rivularis
(Schrad) Nees 39 46
poundpoundnocepha1um conicum (1) Lindb
12 28 39 46 (49) 87 88 8990 120 121 148
Cololejeunea biddlecomiae (Aust) Evans
39 46 62 87 88 89 90
36
Dipl0]hyllum apicu1atum (Evans) Steph Fossombronia foveo1ata Lindb
(35)41 46 90 140 1 t 38 (39) t 46
Frullania bolanderi Aust
13 46 62 88 90 139 140
Frullanla ari ttoniae Evans
39 46 69 (71) 87 88 89 90
Frullania eboracensis Gott
1 9 28 4 39 46 (57) (59) 69 70 (71) 87 88 89 90 92 93 97 148
Frullania inflata Gott
46 87 88 89 (139)
Frullania oakesiana Aust Frullania plana Sullo
(35) 42 46 87 (139) (140) 92
Frullania ri paria Hampe ~ Lehm
46 69 71 87 89
Frullania ~elwynianna Pears
46 89
Geocalyx SEaveolens (Schrad) Nees
394688 8990 (119) 140
Frullania tamarisci (L) Dum subsp sPagrayana
(Mont) Hatt 28 34 l~6 t (57) 88 89 90
38
Mar pan thus seutatUB ( Web amp Mohr) Spruce Jamesoniella autumnalis (DC) Steph
(35) 39 46 90 1 7 28 39 46 56 (57) 87 8889 90 (119) 137 141
Jungermannia confertissima Nees
137 141
Jungermannia crenuliform1s Aust
79 137 141
Jungermannia ~il1ima Sm
46 90 141 Jungermannia exsert1folia subsp corifo~ (Dum~
Vana (35) 46 79 141
39
Jungermannia hyalina Lyell Jungerrr6nnia 1eiantha Grolle
40 46 79 136 141 39 46 79 88 90 1)7 141
Jungermannia IXlaxis Lindb Jungermannia pumila With
46 67 79 (139) (140) 141 )9 46 90 1)6 141
Jungermannia sphaerocarpa Hook Kurzia setacea (Web) Grolle
4679136140IIH 136
Lejeunea cavifo1ia (Ehrh) Lindb Lepidozia reptans (1) Dum
1 7 27 39 46 88 89 90 137 14010 I ( 27) 46 I 69 (71) 88
lophocolea heterophylla (Schrad) Dum Lophocolea minor Nees
1 )4 39 46 (57) 87 88 89 27 39 46 88 89 90 (119) 90 (93)94 119 136
Lophozia al~stris (Schleich ~ Web) Evans Lophozia ascendens (Warnst) Schust
46 90 137 141 136 140 141
41
loIilozia badensls (Gatt ~ Rabenh)
46 141
lophozia blcrenata (Schmid sectA Hoffm)
Dum 46 141
lophozia collaris (Nees) Dum
39 46 141
lophozla excisa (Dicks) Dum
27 4679 141
lophozia gillmani (Aust) Schust lophozia gIlttulata (Undo tl H Arnell)
36 39 40 46 136 (140) 141 Evans 39 46 136 141
42
141
Lophozia heterocolEDs (Thed) MA Howe IoPhozia ipcisa (Schard) Dum
27 39 46 89 141 8 27 31 46 79 90 140 141
LoQhozia longidens Macoun Lophozia ventricosa
1 46 79 90 141
Lophozia ventricosa var silvicola
(Buch) Jones 141 LoEhozla ventricosa vax longif1ora (Nees)
Macoun 39 46 79 141
43
~~ fragrans (Balbis) Frye et Clark Mannia pi 10sa (Hornem) Frye et Clark
46 (139) (140) 9 46 79 116 139
Mannia triandra (Scop) Gro 11e
46 (140)
Marchantia pound91ymorpha L
1 13 28 3946 (49) 78 87 88 89 90 (Ill) 121 122 147
Mo~rckia hibernica (HookJ Gott
46 89
Metzgeria conjugata Lindb
44
46
Mylia anomala (Hook) S Gray Notothylas orbicularis (Schwein) Sullo
39 46 141 (35)45 46
Nowellia curvifolia (Dicks) Mitt
17 27 3846 88 89 90 141
Odontoschisma denudatuJI1 (Nees ~ Mart)
Dum 46 81 137 141
Pallavicinia lyellii (Hook~ Carruth
90
Odontoschisma macounii (Aust) Under
45 46 (66) (139) 140 141
45
=
Pellia epiphllla (L) CordaPellia endiviifolia (Dicks~ Dum
27 39 46 90 13724 39 46
Pellia neesiana Limp bull Phaeoceros
39 46 89 27 39 45 46 97
Porella pinnata L
13 27 I 46 90 Plagiochila asplenioides (L) Dum
27 I 39 1+6 88 89 I 90 137
46
~e11a platYEhylla (L) Pfeiff Porella platyphylloi~ (Schwein) Lindb
17 27 39 46 69 (71)79 87 88 89 9 34 46 (57) 59 87 88 89 92 90 92 93 77
Preissia quadrata (Seop) Nees
1 36 3946 (71) 88 89 90 121 123
137
Ptllidium ciliare (L) Hampe
1 27 39 46 132 141
Dum
(22) 28 3946 (57) 87 88 89 90 92
Ptilidium ~lcherrium (G Web) Hampe
(9) 13 394656 (57)70 87 88 89 90 93 (119) 136 141
47
Radula obconica Sullo Reboulia hemlsphaerlca (L) Raddi
39 46 88 89 (43) 46 (59) 70 90 t Cf 124 125 (139)
Riccardia ~atifrons Lindb
1 27 39 46 89
Riccardia multifi~ (L) S Gray
39 46 88 145
Riccia beyrichiana Hampe ~ Lehm
46 Riccardia palmata (Hedw) Carruth
39 46 90
48
Riccia bifurca Hoffm Riccia cavernosa Hoffm
48 13739 46 79 137
Riccia fluitans L
242731454679
Riccla frostli Aust
48 79
Riccia sullicantii Aust
79 (137) Riccia sorocarpa Bisch 46 (137)
49
Ricciocarpus natans (1) Corda Sca~ia apiculata Spruce
13 27 46 79 89 110 39 46 (139) 141
ScaJltll1ia carinthica Jack ~ Lindb
137 t 141
Scapania ~ (Mart) Durn
39467990137141
Scapania digenil Schiffn
90
ScaEsectflia cuspiduligera (Nees) K Mull
46 (139) (140) (141)
50
Scapania glaucoeerha1a (Tayl) Aust Seapania gyrrmostomophilia Kaal
39 46 (139) 141 41 44 46 64 (139) (140) 141
ScaP3poundlia irrigua Gott ~ almiddot
7 39 46 79 137 (140L 141
Seapania nemorosa (L) Dum
1 39 46 89 90 137
Scapmi~ ~~ta Smiddotlch
137 141 51
~caPnia poundludicola LDeske amp K Mull Scapania saxicola Schust
46 (66) 141 136 141
Sc~~ scandica (H Arnell ~ Buch) Macv
139 140
Scapania subalpina (Nees) Durn
39 40 46 79 137 141
Scapania undulata (L) Dum
1 42 46 79 141
Scapania umbrosa (Schrad) Dum
137 140 141
52
Trichocolea tomentella (Ehrh) Dum Triomarla exsecta (Schmid) Schiffn
27 (35) 39 46 47 88 137 141 8 28 39 46 90 140 141
Tritomaria exsectiformis (Breidl) Loe~ke Tritomaria Sluinguedentata (Huds) Buch
39 -t6 89 90 136 140 141 7 39 46 141
Trltomaria scitula (Tayl) Joerg
46 136 (139) 141
53
Moerckia Gott hibemica (Hook) Gott
21 PeWaceae Pellfa Raddi nom cons propos
endivlifolfa (Dicks) Dwn epiphylllz (L) Corda neeBiana (GottlUmpr
22 Metzgeriaceae Metzgerla Raddi
conjugata Undb
23 Blasiacae B L
pulilm L
24 Aneurareae AneumDum
pinguis (L) Dum
Riccfl1dia S Gray nom cons latifrons Undb multiftda (L) S Gray var multifida palmata (Hedw) Carruth
25 Aytoniaceae Reboulill Raddi nom conI
hemllphaerica (L) Radeli
ManniaOpiz fragrrms (Balbis) Frye el ClarK pilosa (Hornem) Frye et Clark trltmdra (Scop) GroDe
26 Conocephalaceae Conocephalum Wigg nom cons
conicum (L) Lindb
27 Marchantiaceae Preiuia Corda
quadrata (Scop) Nees
Marchantia L polymorpha L
28 Ricciaceae Riccwcarpus Corda
natans (L) Corda
Riccia 1 beyrichiana Hampe ex Lehm bifurca Horfm
15
i
CtlJPemostI Hoffnt fuitu L rostii Aust BOIOCtI1fHI Bi~ sulltvtmtii Aust
MUSCI (Mosses)
29 Sphagnaceae SpIutgnum L
capillaceum (Weiss) Schrank var Cllpil1tlceum caplllllceum (Weiss) Vir tenellum (Schimp) Andr centrille C Jens ex Amen et C Jens compactummiddot OC ex Lam et Dc ClUpidatum Ehrh ex Hoffin var aupidatum jimbrltltum Wits ex J Hookmiddot fulCUm (Scbimp) KUngrr glrgenBOhIlll Russ imbrlctltum Hornsch ex Russ muelllmitil4m BOd 1IIIIU (RUII) C Jens przIutre L papIIIoIum Lindb pulchrum (Lindb ex braithw) Wamst recumun P-Beauv var recumun recum P-Beauv var tenue Klinggr riptlrlum Ansstr ruPQWItWamst squtl1fOlUm Crome subftltens Russ et Wamst ex Wamst IUbIeCfUldum Nees ex Sturm var subsecundum subsecundum Nees ex Sturm varrutem (Nees Hormch et sUUm) Hueb teres (Sdlimp) Anptr e C Hartm
WtlI7IItorjii Russ wuljfanum Gira
30 Andreaeaceae AndNrrerI Hedw
rupeitrlr Oedw var pest
31 Fissident~e Fluid_ lledw
trdiantoid Hedw byoidei Bedw var bryolda cristaM Wils ex Mitt exiguus sun VIr exigUw onttmUi (B-Pyl) Steud jtuberi Lesq et James obtusifolhu Wits var obtwifoliul ollPlUlldQldes Heel ~litSuU lubbtulkril Bed tfDdfoliul lledw llridubu (S~ ) Wahlenb vallirlIU1u
16
32 Ditrichaceae Pleuridium Brid
subulatum (Hedw) Rabenh
Ditrichum Hampe heteromallum (Hedw) Britt linellre (Sw) lindb
~ pallidum (Hedw) Hampe pusiUum (Hedw) Hampe
Saelania Lindb gIlucescens (Hedw) Bomanss et Broth
Ceratodon Brid purpureus (He(lw) Brid var prupurew
Distichium BSG capillaceum (Hedw) BSG inclinatum (Hedw) BSG
33 Bryoxipbiaceae Bryoxlphium Mitt
norvegicum (Brid) Mitt
34 Seligeriaceae Seligeria BSG
cumpylopoda Kindb ex Macoun et Kindb pusillll (Hedw) BSG recurvata (Hedw) BSG
Bindia BSG acuta (Hedw) BSG
35 Dicranaceae BlUehia Schwegr
lexUOSll (Sw ~ Schwaegr) C MueU sullivantii Aust
Trematodon Michx ambiguus (Hedw) Hornsch
Dicranellll (C Muell) Schimp cerviculata (Hedw) Schimp val cillPicuiata
heteromalla (Hedw) Schimp ~ rufescens (With) Schimp
schreberiana (Hedw) Schhnp var schreberiana schreberitJna (Hedw) Schimp var robusta Schimp ex Braithw subulata (Hedw) Schimp varia (Hedw) Schimp
Dicranodontium BSG denudatum (Brid) Britt ex Williams
Panzeucobryum (Lindh) Loeske Iongifolium (Hedw) oelke varlongifolium
17
Tortella (Undb) Limpr fragilis (Drumm) Umpr humilis (Hedw) JeM torlUoSfl (Hedw) Umpr
Hyophila Brid involuta (Hoole) Jaeg et Sauerb
Didymodon Hedw rigidulus Hedw
BryoerythrophyUum Chen recurvirostnon (Hedw) Chen
Btubula Hedw acula (Brid) Brid var acuta convoiuta Hedw fallox Hedw unguicultltll Hedw
Acaulon C MueD muticum (Uedw) C Muell var rufeseens (Iaeg) Crum
PJl~lm Hedw CUSPidtltum Hedw
De8mlltodon Brid cernuu (Hueb) BSG oblUsifoliw (Schwaegr) Schimp
T01IUla Hedw mueronifolia Schwaegr rurall (Hedw) Gaertn Meyer et Scherb var ruralis
39 Grimmiaceae Grlmmill Hedw
alpicola Hedw var alpicola alpicola Hedw var dupretii (Ther) Crum alpicola Hedw var rlvularl8 (Brid) Wahlenb apocarpa Hedw var apocarpa apocarpa Hedw var ambipa (Sull) Jonea apocarpa Hedw var conjerttl (Funck) Spreng apocarpa Hedw var strleta (Tum) Hook et Tayl CiIIyptrrlta Hook ex Drumm donniana Sm var donn_ hemumti Crum ltlevla (Brid) Brid montana BSG oineyi SuD oVllll8 (Hedw) lindb piUfe P-Beauv pltJgiopodill Hedw pulvinata (Hedw) Sm varbullpulvinata teretinervl Limpr
18
Orothera C Muell leana (Sull) C Muell
Rfwbdoweisia crispata (With) Undb
Cynodontium BSG ex Schimp alpestre (Wahlenb) Milde strumiferum (Hedw) Undb
Dichodontium Schimp pellucidum (Hedw) Schimp
Oncophorus (Slid) Brid vuens (Hedw) Brid wahlenbergii (Brid)
Dicranum Hedw bonjeanii DeNot ex Usa drummondii C Muell lagellare Hedw varj1ogellare fulvum Hook fuscescens Turn var fuscescens montanum Hedw muehlenbeckii BSG var muehlenbeckii polysetum Sw sabuletorum Ren et Card scoparium Hedw undulatum Brid viride (Sull et Lesq ex Sull) lindb
36 Leucobryaceae Leucobryum Hampe
glaucum (Hedw) Angstr ex Fr
37 Encalyptaceae Encalypta Hedw
ciliata Hedw var ciliata procera Bruch
38 Pottiaceae Astomum Hampe
muhlenbergianum (Sw) Grout
Weissia Hedw controversa Hedw var controversa
Gymnostomum Nees Homsch et Stunn aeruginosum Sm recurvirostrum Hedw var recurvirostrum
Trichostomum Bruch spirale Prout
19
40 Ephemeraceae Ephemerum Hampe
cohaerens (Hedw) Hampe crassineium (Schwaegr) Hampe val crassinervium spinulosum Bruch et Schimp ex Schimp
41 Funariaceae AphflllolThegma SuD
serratum (1 Hook et Wils ex Drumm) Sullo
Physcomitrium (Brid) Fuernr hookeri Hampe immersum Sullo pyriforme (Hedw) Hampe
Funaria Hedw americana Undb hygrometrica Hedw vaT hygrometrica
42 Splachnaceae Splachnum Hedw
ampullaceum Hedw
43 Schistostegaceae Schistostega Mohr
pennata (Hedw) Web el Mohr
44 Bryaceae Pohlia Hedw
annolina (Hedw) Lindb var annotina camea (Tum) Lindb cruda (Hedw) Undb elongata Hedw iliformis (Dicks) Andr lescurlana (Sull) Grout nutans (Hedw) Lindb obrusifolia (Brid) L Koch proligera (Undb ex Umpr) Undb ex Amell wahlenbergii (Web et Mohr) Andr
Leptobryum (B S G) Wits pyriforme (Hedw) Wils
Bryum Hedw algovicum Sendtnex C Muell argenteum Hedw caespiticium Hedw capillare Hedw var capillare creberrlmum Tayt lonchocaulon C Muell pallens (Brid) Sw ex Roehl pseudotrlquetrum (Hedw) Gaertn Meyer et Scherb varpudotriquetrum
20
pseudotriquetrum (Hedw) GaertnMeyer et Scherb var bimum (Schreb) Ulj stenotrichum C Muell var stenotrichum tortiolium Funck ex Brid uliginosum (Brid) B S G violaceum Crundw et Nyh weigelii Spreng
Rhodobryum (Schimp) Hampe roseum (Hedw) Limpr
45 Mniaceae Mnium Hedw
ciliare (C Muell) Schimp cinclidioides Hueb cuspidarum Hedw drummondii Bruch et Schimp longirostrum Brid lycopodioides Schwaegr marginatum (With) Brid ex P Beauv mediumB~S G var medium onhorrhynchum Brid pseudopunctatum Bruch et Schimp punctalUm Hedw var punctatum punctatum Hedw var elatum Schirnp rugicum Laur spinufosum B S G stellare Hedw
46 Aulacomruaceae Au1acomnium Schwaegr
androgynum (Hedw) Schwaegr heterostichum (Hedw) B S G paJustre (Hedw) Schwaegr var paJustre
47 Meesiaceae Amblyodon B S G
deillbotus (Hedw) B S G
Meesia Hedw triquetra (Richt) Angstr uJiginosa Hedw
48 Bartramiaceae Plagiopus Brid
oederiana (Sw) Umpr
Bartramia Hedw pomiformis Hedw var pomiformis
Philonotis Brid fontana (Hedw) Brid varfontana longseta (Michx) Britt varlongseta marchica (Hedw) Brid
21
~
~
49 Timmiaceae Timmia Hedw
megapolitana Hedw var megapolitana
50 Orthotrichaceae Zygodon Hook et Tayl
viridissimus (Dicks) Brid var rupestris lindb ex C J Hartm
Amphidium Schimp lapponicum (Hedw) Schimp mougeotii (B S G) Schimp
Orthotrichum Hedw anomalum Hedw obtuaifolium Brid ohioense Sullo et Lesq ex Aust pumilumSw sordidum Sullo et Lesq ex Aust speciosum Neesax Sturm var eegam (Schwaegr ex Hook et Grev) W~rnst stellDtwn Brid strrmgulatum P Beauv
Uleta Mohr ex Web cOfIICtata (p Beallv) Hamrn crispa (Hedw) Brid
Drummondill Hook ex Drumm prorepens (Hedw) Brltt
51 Fontinalaceae Fantinalis Hedw
antipyretica Hedw var antipyrettca antipyretica Hedw forma rabusta (Card) Card dalecarlica Schimp ex B S G var dalecarlica flqccidtJ Ren et Card hypnoides C J Hartrn var hypnoides hypnoides C J Hartm var durillei (Schimp) HU$ll
missourlcaCard novae-angliae Sullo var novae-angliae splulgrUlolill (C Moen) Wijket M~S
DichelY1l1ll Myr capilloceum (With) Myr falcatum (Bedw) Myr HIllescens B S G
52 Climaciaceae aimecium Web et Mohr ex Mohr
americllnum Brid dedroides (Hedw) Web et Mohr
22
S3 Hedwigiaceae Hedwigia P Beauv
ciliata (Hedw) P Beauv
54 Cryphaeaceae Forsstroemia lindh
ohioense (Sull) lindh
55 Leucodontaceae Leucodon Schwaegr
brachypus Blid var brachypus brachypus Brid var aniJrewsitlnus Crum et Anderson julaceus (Hedw) SuU
56 Neckeraceae Neckera Hedw
pennata Hedw var pennata
1fomalia (Brid) B S G trichomanoides (Hedw) B S G
57 Theliaceae Thelia Sull
asprellll Sullo hirtella (Hedw) Sullo lescurii SuU
Myurella B SG julacea (Schwaegr) B S G
sibrica (C MueU) Reim var sibrica
58 Fabroniaceae Fabronio Raddi
ciliaris (Brid) Bnd
Anacamptodon Brid splachnoides (Groet ex Brid) Brid
Schwetschkeopsis Broth fabronia (Schwaegr) Broth
59 Leskeaceae Lindbergia Kindb
brachyptera (Mitt) Kindb
Leskea Hedw gracilescens Hedw obscura Hedw poiycarpa Hedw
Leskeella (limpr) Loeske nervosa (Brid) Loeske
23
Pterlgy1lllndrum Hedw filiforme Hedw
60 Thuidiaceae Haplohymenfum Dozy et MoIk
trlste (Ces ex De Not) Kiodb
Anomodon Hook et Tayl attenuatus (Hedw) Hueb minor (Hedw) Fuemr rostratus (Hedw) Schimp viticulosus (Hedw) Hook et Tayl
Haplocladium (C Mucn) C MueU microphyllum (Hedw) Broth virginillnum (Brld) Broth ~bull
Thuidium B S G abietinum (Hedw) B S G delicatulum (Hedw) B S G vardelicatulum delicatulum (Hedw) B S G var radicans (Kindb) Crum Steere et Anderson minutulum (Hedw) B S G pygmaeum B S G recognftum (Hedw) Lindb scitum (p Beauv) Aust
Helodium (Sull) Warnst blandowli (Web et Mohr) Warnst var blandowli blandowli (Web et Mohr) Wamst var helodioides (Ren et Card ex Roell) Crum
Steere et Anderson paludomm (Sull) AuS
61 Amblystegiaceae Cratoneuron (Sull) Spruce
commulatum (Hedw) Roth var commutfltum commutatum (Hedw) Roth var fakatum (Brid) Moenk ilicinum (Hedw) Spruce var filcinum
Campylium (Sun) Mitt chrysophyllum (Brid) 1 Lange var chrysophyllum hispidulum (Brid) Mitt var hllJidulum polygamum (B S G)C lens varpolygtlmUm stellatum (Hedw) C1ens vat stelltum
Leptodictyum (Schimp) Warnst laxirete (Card et Ther) Broth riparium (Hedw) Wamat trichopodium (Schultz) Warns l1acillans (Sull) Broth
Hygroamblystegium Loelke jlul1iatile (Hedw) oeske var luViatile
24
fluviatile (Hedw) Loeske var oWltum Grout noterophilum (Sun et Lesq ex Sull) Warnst tenox (Hedw) Jenn var tenax tenox (Hedw) Jenn var spinilolium Schimp) Jenn
Amblystegium BSG juratzkanum Schimp serpens (Hedw) BSG varlum (Hedw) Undb var varium
Platydictya Berk confervoides (Brid) Crum jungermanniodes (Brid) Crum minutissimum (Sull et Lesq ex Sull) Crum subtile (Hedw) Crum
Drepanocladus (C Muell) Roth aduncus (Hedw) Warnst var aduncus aduncus (Hedw) Warnst var capillilolius (Wamst)Riebm aduncus (Hedw) Warnst var kneiffii (BSG) Moenk aduncus (Hedw) Warnst var polycorpus (Bland ex Voit) Roth exannulatus (BSG) Warnst fluitans (Hedw) Warnst var fluitans revolvens (Sw) Wamst var revolvens sendtneri (Schimp) Warnst var sendtnerl uncinatus (Hedw) Warnst var uncinatus uncinatus ffiedw) Warnst var piwnosus (Schimp) Wamst vernicoms (Lindb ex C Hartm) Warnst
Hygrohypnum Lindb eugyrium (BSG) loeske luridum (Hedw Jenn varluridum ochraecum (Tum ex Wils) Loeske var ochraceunt
Calliergon (Sull) Kindb cordilolium Hedw Kindb glganteum (Schimp) Kindb var giganteum richardsonii (Mitt) Kindb ex Wamst stramineum (Brid) Kindb trilarium (Web et Mohr) Kindb
Calliergonella Loeske cuspidata (Hedw) Loeske
Calliergidium (Ren) Grout bakeri (Ren) Grout
Scorpidium (Schimp) Urnpr scorpiodides (Hedw) limpr
62 Brachytbeciaceae Tomenthypnum loeske
nitens (Hedw) Loeske var mtens
25
Homalotheciella (Card) Broth subcapillata (Hedw) Broth
Brachythecium BSG acuminatum (Hedw) Awt acutum (Mitt) Sullo calcareum IGndb campestTe (C MuelL) BSG cunum (Lindb) Umpr digastrum C Muell et Kindb ex Macoun et Kindb oxycladon (Brid) Jaeg et Sauerb var oxycladon plumosum (Hedw) BSG populeum (Hedw) BSG var populeum reflexum (Starke ex Web et Mohr) DsG var relexum reiexum (Starke ex Web et Mohr) DSG var pacificum Renet Card ex Roell rlvulare BSG var rivulare tabulum (Hedw) BSC var tabuum salebrosum (Web et Mohr) BSG var salebrosum velutinum (Hedw) BSG Vat velutinum
Bryhniil Kaur graminicolor (Brid) Grout var graminicolor novae-angliae (Sull et Lesq ex sun) Grout
Rhynchostegium BSG senulatum (Hedw) Ja4lg et Sauerb
Rhynchostegiella (BSG) Limpr compacta (C MueU) Loeske
EU1hynchium 8SG hians (Hedw) Sande Lac pulchellum (Hedw) Jenn var pulchelium riparloides (Hedw) Rich
Stokeriella (Kindb) Robins pmelonga (Hedw) Robins var stokesii (Tum) Crum
63 Entodontaceae Entodon C Muell
brevisetus (Hook et Wils ex Wils) Lindb cladorrhizans (Hedw) C Muell seductrlx (Hedw) C MueU var wluctrix
Pleurozium Mitt schreberl (8rid) Mitt var schreberi
64 Plagiotheciaceae Pltlglothecium BSG
CflVifolium (Brid) Iwats var CtWifolium denticulatum (Hedw) 8SG laetum BSG ltztebricola BSG
26
6S SernatophyUaceae Brotherella Loeske ex Fleisch
recumins (Michx) Fleisch tenuirostris (Bruch et Schimp ex Sull) Fleisch
Sematophyllum Mitt demissum (Wils) Mitt
66 Hypnaceae Pl(itygyrium BSG
repens (Brid) BSG
PylaisieUa Kindb intricata (Hedw) Grout polyantha (Hedw) Grout selwynii (Kindb) Crum Steere et Anderson
Homomallium (Schimp) Loeske adnatum (Hedw) Broth
Hypnum Hedw cupressiforme Hedw var cupressiforme cupressifonne Hedw varfiliforme Brid curviolium Hedw fertile Sendtn imponens Hedw lindbergii Mitt palescens (Hedw) P-Beauv pratense Koch ex Brid
Calicladium Crum haldanianum (Grev) Crum
sopterygiopsis Iwats muelleriana (Schimp) Iwats
Isopterygium Mitt e1egans (Brid) lindb pulchellum (Hedw) Jaeg et Sauerb tenerum (Sw) Mitt
Taxiphyllum Fleisch depllQtum (Bruch et Schimp ex sun) Fleisch taxirameum (MitL) Fleisch
Herzogiella Broth seligeri (Brid) Iwats striatella (Brid) Iwats ruracea (Lindh) Iwats
hilium De Not crista-castrenna (Hedw) De Not
27
67 Rhytidiaceae Rhytidium (Sull) Kindb
rugosum (Hedw) Kindb
Rhytidfodelphus (Lindb ex Iimpr) Warnst triquetrus (Hedw) Warnst var triquetrus
68 Hylocomiaceae Hylocomium BSG
pyrenaicum (Spruce) lindb splendens (Hedw) BSG varsplendens
69 Buxbaumiaceae Buxbaumia Hedw
aphyllll Hedw
Diphyscium Mohr follosum (Hedw) Mohr
70 Tetraphidaceae Tetraphi3
pellucida Hedw
71 Polytrichaceae Atrichum PmiddotBeauv
angustatum (Brid) BSG cri8pum (James) Sullo oerstedianum (C Muell) Mitt teneUum (Roehl) BSG undulatum (Hedw) P middotBeauv var unduiatum
Pogonatum P-Beauv alpinum (Hedw) Roehl var alpinum umigerum (Hedw) P-Beauv
Poiytrichum Hedw commune Hedw var commune ormosum Hedw juniperinum Hedw longisetum Brid ohioense Ren et Card piliferum Hedw var pilierum strictum Brid
28
MAPS OF BRYOPHYTE TAXA
REPORTED FOR WISCONSIN
HORNWORTS AND LIVERWORTS
29
bull bull WSHeV~N S vE II
0 ~
e~RRON ~USK
(J~ (j -
CHIPPE A DUNN
0 o0 poundAU CLAIRE
~ TREMshyPEAshyLEAU
MICE
ONEIDA
Q LINCOLN
TAYLOR
Q t-~----- MOIRA THON
CLARK
o
10
LA FAVETTE
Hornwor ts amp Liverworts
- None - 1 - 25 taxa
- 26 - 50 tRXshy
- 51 - 75 tax)
- 7c - 100 taxa
FONDOU j LAC ~
DODGE
i O JEfFERSON
WALWORTH
30
Anastrophylltlfl helleranum (Nees) Schust Anastrophyllum michauxil (We~ Buch ~
46 88 136 141 Evans 38 46 141
Anastrophyllum mlnut~ (Schreb) Schuster Aneura plnguls (L) Durn
89 21 384687 88 89 90 (138) (145)
Anthoceros macounil MA Howe ~oceros punctatus L
21 (45) (144) 34
31
Barbilophozia attenuata (Mart) Loeske Barbilophozia barbata (Schmid ~ Schre~
24 141 Loeske 28 39 46 89 90 141
Bazzania trilobata (1) S Gray
1 26 39 46 (57) (59) 79 88 89
90 119 141
BlePbarostoma trichoffiyllum (1) Dumort
1 8 26 39 46 89 90 137 141
Blasia pusilla 1
1 39 46 90
Calypogeja fissa (1) Raddi
90
32
$129geja integristipula Steph C~Ypound2geja muelleriana (Schifm) K Mull
88 89 89
CallpoundO~eja neesiana (Mass amp Carest) K Mull
39 46 89 140 (141)
~pogeja sJlapound2icola (H Arnell tl J bull
Perss) Warnst et Loeske (119) 141
CeIalozia biscuspidata (L) Dum
1 37 I 46 90 t 137 141
CalYpogej~ trichomanis (L) Corda
26 39 (119) 137
33
CeEhalozia catenulata (Hub) Lindb Cephalozia connivens (Dicks) Lindb
39 46 79 141 39 4679 88 (89) 90 119 141
Cephalozia lacinulata Jack ~ Spruce
141 Cephalozia lunulifolia (Dum~ Dum I 28 39 46 79 88 89 90 141
Cepoundhalozia macouni (Aust) Aust Cephalozia ~leniceps (Aust) Lindb 27~ 3746 (66)7987 (139)141 39 46 79 89 141
34
Ceffialoziella 9ivaricata (Sm) Schiffn Cephaloziella ~hista (Jack) Schiffn
46 90 46 (119) 141
Cephaloziella hampeana (Nees) Schiffn
46
Cephaloziella rubella (Nees) Warnst
46 39
Cephaloziella rubella var sullivantii
(Aust) K Mull 46 39
Cephaloziella rubella var bifida (Lindb) Douin
35
46
Chiloscyphus pallescens (Ehrh ~ Hof~ Dum Chiloscyphus pallescens var fragilis
39J 46 88 89J 90 (Roth) K Milll 46
Chiloscyphus E9lyanthos (L) Corda
1 24 39 46
Chiloscyphus polyanthos vax rivularis
(Schrad) Nees 39 46
poundpoundnocepha1um conicum (1) Lindb
12 28 39 46 (49) 87 88 8990 120 121 148
Cololejeunea biddlecomiae (Aust) Evans
39 46 62 87 88 89 90
36
Dipl0]hyllum apicu1atum (Evans) Steph Fossombronia foveo1ata Lindb
(35)41 46 90 140 1 t 38 (39) t 46
Frullania bolanderi Aust
13 46 62 88 90 139 140
Frullanla ari ttoniae Evans
39 46 69 (71) 87 88 89 90
Frullania eboracensis Gott
1 9 28 4 39 46 (57) (59) 69 70 (71) 87 88 89 90 92 93 97 148
Frullania inflata Gott
46 87 88 89 (139)
Frullania oakesiana Aust Frullania plana Sullo
(35) 42 46 87 (139) (140) 92
Frullania ri paria Hampe ~ Lehm
46 69 71 87 89
Frullania ~elwynianna Pears
46 89
Geocalyx SEaveolens (Schrad) Nees
394688 8990 (119) 140
Frullania tamarisci (L) Dum subsp sPagrayana
(Mont) Hatt 28 34 l~6 t (57) 88 89 90
38
Mar pan thus seutatUB ( Web amp Mohr) Spruce Jamesoniella autumnalis (DC) Steph
(35) 39 46 90 1 7 28 39 46 56 (57) 87 8889 90 (119) 137 141
Jungermannia confertissima Nees
137 141
Jungermannia crenuliform1s Aust
79 137 141
Jungermannia ~il1ima Sm
46 90 141 Jungermannia exsert1folia subsp corifo~ (Dum~
Vana (35) 46 79 141
39
Jungermannia hyalina Lyell Jungerrr6nnia 1eiantha Grolle
40 46 79 136 141 39 46 79 88 90 1)7 141
Jungermannia IXlaxis Lindb Jungermannia pumila With
46 67 79 (139) (140) 141 )9 46 90 1)6 141
Jungermannia sphaerocarpa Hook Kurzia setacea (Web) Grolle
4679136140IIH 136
Lejeunea cavifo1ia (Ehrh) Lindb Lepidozia reptans (1) Dum
1 7 27 39 46 88 89 90 137 14010 I ( 27) 46 I 69 (71) 88
lophocolea heterophylla (Schrad) Dum Lophocolea minor Nees
1 )4 39 46 (57) 87 88 89 27 39 46 88 89 90 (119) 90 (93)94 119 136
Lophozia al~stris (Schleich ~ Web) Evans Lophozia ascendens (Warnst) Schust
46 90 137 141 136 140 141
41
loIilozia badensls (Gatt ~ Rabenh)
46 141
lophozia blcrenata (Schmid sectA Hoffm)
Dum 46 141
lophozia collaris (Nees) Dum
39 46 141
lophozla excisa (Dicks) Dum
27 4679 141
lophozia gillmani (Aust) Schust lophozia gIlttulata (Undo tl H Arnell)
36 39 40 46 136 (140) 141 Evans 39 46 136 141
42
141
Lophozia heterocolEDs (Thed) MA Howe IoPhozia ipcisa (Schard) Dum
27 39 46 89 141 8 27 31 46 79 90 140 141
LoQhozia longidens Macoun Lophozia ventricosa
1 46 79 90 141
Lophozia ventricosa var silvicola
(Buch) Jones 141 LoEhozla ventricosa vax longif1ora (Nees)
Macoun 39 46 79 141
43
~~ fragrans (Balbis) Frye et Clark Mannia pi 10sa (Hornem) Frye et Clark
46 (139) (140) 9 46 79 116 139
Mannia triandra (Scop) Gro 11e
46 (140)
Marchantia pound91ymorpha L
1 13 28 3946 (49) 78 87 88 89 90 (Ill) 121 122 147
Mo~rckia hibernica (HookJ Gott
46 89
Metzgeria conjugata Lindb
44
46
Mylia anomala (Hook) S Gray Notothylas orbicularis (Schwein) Sullo
39 46 141 (35)45 46
Nowellia curvifolia (Dicks) Mitt
17 27 3846 88 89 90 141
Odontoschisma denudatuJI1 (Nees ~ Mart)
Dum 46 81 137 141
Pallavicinia lyellii (Hook~ Carruth
90
Odontoschisma macounii (Aust) Under
45 46 (66) (139) 140 141
45
=
Pellia epiphllla (L) CordaPellia endiviifolia (Dicks~ Dum
27 39 46 90 13724 39 46
Pellia neesiana Limp bull Phaeoceros
39 46 89 27 39 45 46 97
Porella pinnata L
13 27 I 46 90 Plagiochila asplenioides (L) Dum
27 I 39 1+6 88 89 I 90 137
46
~e11a platYEhylla (L) Pfeiff Porella platyphylloi~ (Schwein) Lindb
17 27 39 46 69 (71)79 87 88 89 9 34 46 (57) 59 87 88 89 92 90 92 93 77
Preissia quadrata (Seop) Nees
1 36 3946 (71) 88 89 90 121 123
137
Ptllidium ciliare (L) Hampe
1 27 39 46 132 141
Dum
(22) 28 3946 (57) 87 88 89 90 92
Ptilidium ~lcherrium (G Web) Hampe
(9) 13 394656 (57)70 87 88 89 90 93 (119) 136 141
47
Radula obconica Sullo Reboulia hemlsphaerlca (L) Raddi
39 46 88 89 (43) 46 (59) 70 90 t Cf 124 125 (139)
Riccardia ~atifrons Lindb
1 27 39 46 89
Riccardia multifi~ (L) S Gray
39 46 88 145
Riccia beyrichiana Hampe ~ Lehm
46 Riccardia palmata (Hedw) Carruth
39 46 90
48
Riccia bifurca Hoffm Riccia cavernosa Hoffm
48 13739 46 79 137
Riccia fluitans L
242731454679
Riccla frostli Aust
48 79
Riccia sullicantii Aust
79 (137) Riccia sorocarpa Bisch 46 (137)
49
Ricciocarpus natans (1) Corda Sca~ia apiculata Spruce
13 27 46 79 89 110 39 46 (139) 141
ScaJltll1ia carinthica Jack ~ Lindb
137 t 141
Scapania ~ (Mart) Durn
39467990137141
Scapania digenil Schiffn
90
ScaEsectflia cuspiduligera (Nees) K Mull
46 (139) (140) (141)
50
Scapania glaucoeerha1a (Tayl) Aust Seapania gyrrmostomophilia Kaal
39 46 (139) 141 41 44 46 64 (139) (140) 141
ScaP3poundlia irrigua Gott ~ almiddot
7 39 46 79 137 (140L 141
Seapania nemorosa (L) Dum
1 39 46 89 90 137
Scapmi~ ~~ta Smiddotlch
137 141 51
~caPnia poundludicola LDeske amp K Mull Scapania saxicola Schust
46 (66) 141 136 141
Sc~~ scandica (H Arnell ~ Buch) Macv
139 140
Scapania subalpina (Nees) Durn
39 40 46 79 137 141
Scapania undulata (L) Dum
1 42 46 79 141
Scapania umbrosa (Schrad) Dum
137 140 141
52
Trichocolea tomentella (Ehrh) Dum Triomarla exsecta (Schmid) Schiffn
27 (35) 39 46 47 88 137 141 8 28 39 46 90 140 141
Tritomaria exsectiformis (Breidl) Loe~ke Tritomaria Sluinguedentata (Huds) Buch
39 -t6 89 90 136 140 141 7 39 46 141
Trltomaria scitula (Tayl) Joerg
46 136 (139) 141
53
CtlJPemostI Hoffnt fuitu L rostii Aust BOIOCtI1fHI Bi~ sulltvtmtii Aust
MUSCI (Mosses)
29 Sphagnaceae SpIutgnum L
capillaceum (Weiss) Schrank var Cllpil1tlceum caplllllceum (Weiss) Vir tenellum (Schimp) Andr centrille C Jens ex Amen et C Jens compactummiddot OC ex Lam et Dc ClUpidatum Ehrh ex Hoffin var aupidatum jimbrltltum Wits ex J Hookmiddot fulCUm (Scbimp) KUngrr glrgenBOhIlll Russ imbrlctltum Hornsch ex Russ muelllmitil4m BOd 1IIIIU (RUII) C Jens przIutre L papIIIoIum Lindb pulchrum (Lindb ex braithw) Wamst recumun P-Beauv var recumun recum P-Beauv var tenue Klinggr riptlrlum Ansstr ruPQWItWamst squtl1fOlUm Crome subftltens Russ et Wamst ex Wamst IUbIeCfUldum Nees ex Sturm var subsecundum subsecundum Nees ex Sturm varrutem (Nees Hormch et sUUm) Hueb teres (Sdlimp) Anptr e C Hartm
WtlI7IItorjii Russ wuljfanum Gira
30 Andreaeaceae AndNrrerI Hedw
rupeitrlr Oedw var pest
31 Fissident~e Fluid_ lledw
trdiantoid Hedw byoidei Bedw var bryolda cristaM Wils ex Mitt exiguus sun VIr exigUw onttmUi (B-Pyl) Steud jtuberi Lesq et James obtusifolhu Wits var obtwifoliul ollPlUlldQldes Heel ~litSuU lubbtulkril Bed tfDdfoliul lledw llridubu (S~ ) Wahlenb vallirlIU1u
16
32 Ditrichaceae Pleuridium Brid
subulatum (Hedw) Rabenh
Ditrichum Hampe heteromallum (Hedw) Britt linellre (Sw) lindb
~ pallidum (Hedw) Hampe pusiUum (Hedw) Hampe
Saelania Lindb gIlucescens (Hedw) Bomanss et Broth
Ceratodon Brid purpureus (He(lw) Brid var prupurew
Distichium BSG capillaceum (Hedw) BSG inclinatum (Hedw) BSG
33 Bryoxipbiaceae Bryoxlphium Mitt
norvegicum (Brid) Mitt
34 Seligeriaceae Seligeria BSG
cumpylopoda Kindb ex Macoun et Kindb pusillll (Hedw) BSG recurvata (Hedw) BSG
Bindia BSG acuta (Hedw) BSG
35 Dicranaceae BlUehia Schwegr
lexUOSll (Sw ~ Schwaegr) C MueU sullivantii Aust
Trematodon Michx ambiguus (Hedw) Hornsch
Dicranellll (C Muell) Schimp cerviculata (Hedw) Schimp val cillPicuiata
heteromalla (Hedw) Schimp ~ rufescens (With) Schimp
schreberiana (Hedw) Schhnp var schreberiana schreberitJna (Hedw) Schimp var robusta Schimp ex Braithw subulata (Hedw) Schimp varia (Hedw) Schimp
Dicranodontium BSG denudatum (Brid) Britt ex Williams
Panzeucobryum (Lindh) Loeske Iongifolium (Hedw) oelke varlongifolium
17
Tortella (Undb) Limpr fragilis (Drumm) Umpr humilis (Hedw) JeM torlUoSfl (Hedw) Umpr
Hyophila Brid involuta (Hoole) Jaeg et Sauerb
Didymodon Hedw rigidulus Hedw
BryoerythrophyUum Chen recurvirostnon (Hedw) Chen
Btubula Hedw acula (Brid) Brid var acuta convoiuta Hedw fallox Hedw unguicultltll Hedw
Acaulon C MueD muticum (Uedw) C Muell var rufeseens (Iaeg) Crum
PJl~lm Hedw CUSPidtltum Hedw
De8mlltodon Brid cernuu (Hueb) BSG oblUsifoliw (Schwaegr) Schimp
T01IUla Hedw mueronifolia Schwaegr rurall (Hedw) Gaertn Meyer et Scherb var ruralis
39 Grimmiaceae Grlmmill Hedw
alpicola Hedw var alpicola alpicola Hedw var dupretii (Ther) Crum alpicola Hedw var rlvularl8 (Brid) Wahlenb apocarpa Hedw var apocarpa apocarpa Hedw var ambipa (Sull) Jonea apocarpa Hedw var conjerttl (Funck) Spreng apocarpa Hedw var strleta (Tum) Hook et Tayl CiIIyptrrlta Hook ex Drumm donniana Sm var donn_ hemumti Crum ltlevla (Brid) Brid montana BSG oineyi SuD oVllll8 (Hedw) lindb piUfe P-Beauv pltJgiopodill Hedw pulvinata (Hedw) Sm varbullpulvinata teretinervl Limpr
18
Orothera C Muell leana (Sull) C Muell
Rfwbdoweisia crispata (With) Undb
Cynodontium BSG ex Schimp alpestre (Wahlenb) Milde strumiferum (Hedw) Undb
Dichodontium Schimp pellucidum (Hedw) Schimp
Oncophorus (Slid) Brid vuens (Hedw) Brid wahlenbergii (Brid)
Dicranum Hedw bonjeanii DeNot ex Usa drummondii C Muell lagellare Hedw varj1ogellare fulvum Hook fuscescens Turn var fuscescens montanum Hedw muehlenbeckii BSG var muehlenbeckii polysetum Sw sabuletorum Ren et Card scoparium Hedw undulatum Brid viride (Sull et Lesq ex Sull) lindb
36 Leucobryaceae Leucobryum Hampe
glaucum (Hedw) Angstr ex Fr
37 Encalyptaceae Encalypta Hedw
ciliata Hedw var ciliata procera Bruch
38 Pottiaceae Astomum Hampe
muhlenbergianum (Sw) Grout
Weissia Hedw controversa Hedw var controversa
Gymnostomum Nees Homsch et Stunn aeruginosum Sm recurvirostrum Hedw var recurvirostrum
Trichostomum Bruch spirale Prout
19
40 Ephemeraceae Ephemerum Hampe
cohaerens (Hedw) Hampe crassineium (Schwaegr) Hampe val crassinervium spinulosum Bruch et Schimp ex Schimp
41 Funariaceae AphflllolThegma SuD
serratum (1 Hook et Wils ex Drumm) Sullo
Physcomitrium (Brid) Fuernr hookeri Hampe immersum Sullo pyriforme (Hedw) Hampe
Funaria Hedw americana Undb hygrometrica Hedw vaT hygrometrica
42 Splachnaceae Splachnum Hedw
ampullaceum Hedw
43 Schistostegaceae Schistostega Mohr
pennata (Hedw) Web el Mohr
44 Bryaceae Pohlia Hedw
annolina (Hedw) Lindb var annotina camea (Tum) Lindb cruda (Hedw) Undb elongata Hedw iliformis (Dicks) Andr lescurlana (Sull) Grout nutans (Hedw) Lindb obrusifolia (Brid) L Koch proligera (Undb ex Umpr) Undb ex Amell wahlenbergii (Web et Mohr) Andr
Leptobryum (B S G) Wits pyriforme (Hedw) Wils
Bryum Hedw algovicum Sendtnex C Muell argenteum Hedw caespiticium Hedw capillare Hedw var capillare creberrlmum Tayt lonchocaulon C Muell pallens (Brid) Sw ex Roehl pseudotrlquetrum (Hedw) Gaertn Meyer et Scherb varpudotriquetrum
20
pseudotriquetrum (Hedw) GaertnMeyer et Scherb var bimum (Schreb) Ulj stenotrichum C Muell var stenotrichum tortiolium Funck ex Brid uliginosum (Brid) B S G violaceum Crundw et Nyh weigelii Spreng
Rhodobryum (Schimp) Hampe roseum (Hedw) Limpr
45 Mniaceae Mnium Hedw
ciliare (C Muell) Schimp cinclidioides Hueb cuspidarum Hedw drummondii Bruch et Schimp longirostrum Brid lycopodioides Schwaegr marginatum (With) Brid ex P Beauv mediumB~S G var medium onhorrhynchum Brid pseudopunctatum Bruch et Schimp punctalUm Hedw var punctatum punctatum Hedw var elatum Schirnp rugicum Laur spinufosum B S G stellare Hedw
46 Aulacomruaceae Au1acomnium Schwaegr
androgynum (Hedw) Schwaegr heterostichum (Hedw) B S G paJustre (Hedw) Schwaegr var paJustre
47 Meesiaceae Amblyodon B S G
deillbotus (Hedw) B S G
Meesia Hedw triquetra (Richt) Angstr uJiginosa Hedw
48 Bartramiaceae Plagiopus Brid
oederiana (Sw) Umpr
Bartramia Hedw pomiformis Hedw var pomiformis
Philonotis Brid fontana (Hedw) Brid varfontana longseta (Michx) Britt varlongseta marchica (Hedw) Brid
21
~
~
49 Timmiaceae Timmia Hedw
megapolitana Hedw var megapolitana
50 Orthotrichaceae Zygodon Hook et Tayl
viridissimus (Dicks) Brid var rupestris lindb ex C J Hartm
Amphidium Schimp lapponicum (Hedw) Schimp mougeotii (B S G) Schimp
Orthotrichum Hedw anomalum Hedw obtuaifolium Brid ohioense Sullo et Lesq ex Aust pumilumSw sordidum Sullo et Lesq ex Aust speciosum Neesax Sturm var eegam (Schwaegr ex Hook et Grev) W~rnst stellDtwn Brid strrmgulatum P Beauv
Uleta Mohr ex Web cOfIICtata (p Beallv) Hamrn crispa (Hedw) Brid
Drummondill Hook ex Drumm prorepens (Hedw) Brltt
51 Fontinalaceae Fantinalis Hedw
antipyretica Hedw var antipyrettca antipyretica Hedw forma rabusta (Card) Card dalecarlica Schimp ex B S G var dalecarlica flqccidtJ Ren et Card hypnoides C J Hartrn var hypnoides hypnoides C J Hartm var durillei (Schimp) HU$ll
missourlcaCard novae-angliae Sullo var novae-angliae splulgrUlolill (C Moen) Wijket M~S
DichelY1l1ll Myr capilloceum (With) Myr falcatum (Bedw) Myr HIllescens B S G
52 Climaciaceae aimecium Web et Mohr ex Mohr
americllnum Brid dedroides (Hedw) Web et Mohr
22
S3 Hedwigiaceae Hedwigia P Beauv
ciliata (Hedw) P Beauv
54 Cryphaeaceae Forsstroemia lindh
ohioense (Sull) lindh
55 Leucodontaceae Leucodon Schwaegr
brachypus Blid var brachypus brachypus Brid var aniJrewsitlnus Crum et Anderson julaceus (Hedw) SuU
56 Neckeraceae Neckera Hedw
pennata Hedw var pennata
1fomalia (Brid) B S G trichomanoides (Hedw) B S G
57 Theliaceae Thelia Sull
asprellll Sullo hirtella (Hedw) Sullo lescurii SuU
Myurella B SG julacea (Schwaegr) B S G
sibrica (C MueU) Reim var sibrica
58 Fabroniaceae Fabronio Raddi
ciliaris (Brid) Bnd
Anacamptodon Brid splachnoides (Groet ex Brid) Brid
Schwetschkeopsis Broth fabronia (Schwaegr) Broth
59 Leskeaceae Lindbergia Kindb
brachyptera (Mitt) Kindb
Leskea Hedw gracilescens Hedw obscura Hedw poiycarpa Hedw
Leskeella (limpr) Loeske nervosa (Brid) Loeske
23
Pterlgy1lllndrum Hedw filiforme Hedw
60 Thuidiaceae Haplohymenfum Dozy et MoIk
trlste (Ces ex De Not) Kiodb
Anomodon Hook et Tayl attenuatus (Hedw) Hueb minor (Hedw) Fuemr rostratus (Hedw) Schimp viticulosus (Hedw) Hook et Tayl
Haplocladium (C Mucn) C MueU microphyllum (Hedw) Broth virginillnum (Brld) Broth ~bull
Thuidium B S G abietinum (Hedw) B S G delicatulum (Hedw) B S G vardelicatulum delicatulum (Hedw) B S G var radicans (Kindb) Crum Steere et Anderson minutulum (Hedw) B S G pygmaeum B S G recognftum (Hedw) Lindb scitum (p Beauv) Aust
Helodium (Sull) Warnst blandowli (Web et Mohr) Warnst var blandowli blandowli (Web et Mohr) Wamst var helodioides (Ren et Card ex Roell) Crum
Steere et Anderson paludomm (Sull) AuS
61 Amblystegiaceae Cratoneuron (Sull) Spruce
commulatum (Hedw) Roth var commutfltum commutatum (Hedw) Roth var fakatum (Brid) Moenk ilicinum (Hedw) Spruce var filcinum
Campylium (Sun) Mitt chrysophyllum (Brid) 1 Lange var chrysophyllum hispidulum (Brid) Mitt var hllJidulum polygamum (B S G)C lens varpolygtlmUm stellatum (Hedw) C1ens vat stelltum
Leptodictyum (Schimp) Warnst laxirete (Card et Ther) Broth riparium (Hedw) Wamat trichopodium (Schultz) Warns l1acillans (Sull) Broth
Hygroamblystegium Loelke jlul1iatile (Hedw) oeske var luViatile
24
fluviatile (Hedw) Loeske var oWltum Grout noterophilum (Sun et Lesq ex Sull) Warnst tenox (Hedw) Jenn var tenax tenox (Hedw) Jenn var spinilolium Schimp) Jenn
Amblystegium BSG juratzkanum Schimp serpens (Hedw) BSG varlum (Hedw) Undb var varium
Platydictya Berk confervoides (Brid) Crum jungermanniodes (Brid) Crum minutissimum (Sull et Lesq ex Sull) Crum subtile (Hedw) Crum
Drepanocladus (C Muell) Roth aduncus (Hedw) Warnst var aduncus aduncus (Hedw) Warnst var capillilolius (Wamst)Riebm aduncus (Hedw) Warnst var kneiffii (BSG) Moenk aduncus (Hedw) Warnst var polycorpus (Bland ex Voit) Roth exannulatus (BSG) Warnst fluitans (Hedw) Warnst var fluitans revolvens (Sw) Wamst var revolvens sendtneri (Schimp) Warnst var sendtnerl uncinatus (Hedw) Warnst var uncinatus uncinatus ffiedw) Warnst var piwnosus (Schimp) Wamst vernicoms (Lindb ex C Hartm) Warnst
Hygrohypnum Lindb eugyrium (BSG) loeske luridum (Hedw Jenn varluridum ochraecum (Tum ex Wils) Loeske var ochraceunt
Calliergon (Sull) Kindb cordilolium Hedw Kindb glganteum (Schimp) Kindb var giganteum richardsonii (Mitt) Kindb ex Wamst stramineum (Brid) Kindb trilarium (Web et Mohr) Kindb
Calliergonella Loeske cuspidata (Hedw) Loeske
Calliergidium (Ren) Grout bakeri (Ren) Grout
Scorpidium (Schimp) Urnpr scorpiodides (Hedw) limpr
62 Brachytbeciaceae Tomenthypnum loeske
nitens (Hedw) Loeske var mtens
25
Homalotheciella (Card) Broth subcapillata (Hedw) Broth
Brachythecium BSG acuminatum (Hedw) Awt acutum (Mitt) Sullo calcareum IGndb campestTe (C MuelL) BSG cunum (Lindb) Umpr digastrum C Muell et Kindb ex Macoun et Kindb oxycladon (Brid) Jaeg et Sauerb var oxycladon plumosum (Hedw) BSG populeum (Hedw) BSG var populeum reflexum (Starke ex Web et Mohr) DsG var relexum reiexum (Starke ex Web et Mohr) DSG var pacificum Renet Card ex Roell rlvulare BSG var rivulare tabulum (Hedw) BSC var tabuum salebrosum (Web et Mohr) BSG var salebrosum velutinum (Hedw) BSG Vat velutinum
Bryhniil Kaur graminicolor (Brid) Grout var graminicolor novae-angliae (Sull et Lesq ex sun) Grout
Rhynchostegium BSG senulatum (Hedw) Ja4lg et Sauerb
Rhynchostegiella (BSG) Limpr compacta (C MueU) Loeske
EU1hynchium 8SG hians (Hedw) Sande Lac pulchellum (Hedw) Jenn var pulchelium riparloides (Hedw) Rich
Stokeriella (Kindb) Robins pmelonga (Hedw) Robins var stokesii (Tum) Crum
63 Entodontaceae Entodon C Muell
brevisetus (Hook et Wils ex Wils) Lindb cladorrhizans (Hedw) C Muell seductrlx (Hedw) C MueU var wluctrix
Pleurozium Mitt schreberl (8rid) Mitt var schreberi
64 Plagiotheciaceae Pltlglothecium BSG
CflVifolium (Brid) Iwats var CtWifolium denticulatum (Hedw) 8SG laetum BSG ltztebricola BSG
26
6S SernatophyUaceae Brotherella Loeske ex Fleisch
recumins (Michx) Fleisch tenuirostris (Bruch et Schimp ex Sull) Fleisch
Sematophyllum Mitt demissum (Wils) Mitt
66 Hypnaceae Pl(itygyrium BSG
repens (Brid) BSG
PylaisieUa Kindb intricata (Hedw) Grout polyantha (Hedw) Grout selwynii (Kindb) Crum Steere et Anderson
Homomallium (Schimp) Loeske adnatum (Hedw) Broth
Hypnum Hedw cupressiforme Hedw var cupressiforme cupressifonne Hedw varfiliforme Brid curviolium Hedw fertile Sendtn imponens Hedw lindbergii Mitt palescens (Hedw) P-Beauv pratense Koch ex Brid
Calicladium Crum haldanianum (Grev) Crum
sopterygiopsis Iwats muelleriana (Schimp) Iwats
Isopterygium Mitt e1egans (Brid) lindb pulchellum (Hedw) Jaeg et Sauerb tenerum (Sw) Mitt
Taxiphyllum Fleisch depllQtum (Bruch et Schimp ex sun) Fleisch taxirameum (MitL) Fleisch
Herzogiella Broth seligeri (Brid) Iwats striatella (Brid) Iwats ruracea (Lindh) Iwats
hilium De Not crista-castrenna (Hedw) De Not
27
67 Rhytidiaceae Rhytidium (Sull) Kindb
rugosum (Hedw) Kindb
Rhytidfodelphus (Lindb ex Iimpr) Warnst triquetrus (Hedw) Warnst var triquetrus
68 Hylocomiaceae Hylocomium BSG
pyrenaicum (Spruce) lindb splendens (Hedw) BSG varsplendens
69 Buxbaumiaceae Buxbaumia Hedw
aphyllll Hedw
Diphyscium Mohr follosum (Hedw) Mohr
70 Tetraphidaceae Tetraphi3
pellucida Hedw
71 Polytrichaceae Atrichum PmiddotBeauv
angustatum (Brid) BSG cri8pum (James) Sullo oerstedianum (C Muell) Mitt teneUum (Roehl) BSG undulatum (Hedw) P middotBeauv var unduiatum
Pogonatum P-Beauv alpinum (Hedw) Roehl var alpinum umigerum (Hedw) P-Beauv
Poiytrichum Hedw commune Hedw var commune ormosum Hedw juniperinum Hedw longisetum Brid ohioense Ren et Card piliferum Hedw var pilierum strictum Brid
28
MAPS OF BRYOPHYTE TAXA
REPORTED FOR WISCONSIN
HORNWORTS AND LIVERWORTS
29
bull bull WSHeV~N S vE II
0 ~
e~RRON ~USK
(J~ (j -
CHIPPE A DUNN
0 o0 poundAU CLAIRE
~ TREMshyPEAshyLEAU
MICE
ONEIDA
Q LINCOLN
TAYLOR
Q t-~----- MOIRA THON
CLARK
o
10
LA FAVETTE
Hornwor ts amp Liverworts
- None - 1 - 25 taxa
- 26 - 50 tRXshy
- 51 - 75 tax)
- 7c - 100 taxa
FONDOU j LAC ~
DODGE
i O JEfFERSON
WALWORTH
30
Anastrophylltlfl helleranum (Nees) Schust Anastrophyllum michauxil (We~ Buch ~
46 88 136 141 Evans 38 46 141
Anastrophyllum mlnut~ (Schreb) Schuster Aneura plnguls (L) Durn
89 21 384687 88 89 90 (138) (145)
Anthoceros macounil MA Howe ~oceros punctatus L
21 (45) (144) 34
31
Barbilophozia attenuata (Mart) Loeske Barbilophozia barbata (Schmid ~ Schre~
24 141 Loeske 28 39 46 89 90 141
Bazzania trilobata (1) S Gray
1 26 39 46 (57) (59) 79 88 89
90 119 141
BlePbarostoma trichoffiyllum (1) Dumort
1 8 26 39 46 89 90 137 141
Blasia pusilla 1
1 39 46 90
Calypogeja fissa (1) Raddi
90
32
$129geja integristipula Steph C~Ypound2geja muelleriana (Schifm) K Mull
88 89 89
CallpoundO~eja neesiana (Mass amp Carest) K Mull
39 46 89 140 (141)
~pogeja sJlapound2icola (H Arnell tl J bull
Perss) Warnst et Loeske (119) 141
CeIalozia biscuspidata (L) Dum
1 37 I 46 90 t 137 141
CalYpogej~ trichomanis (L) Corda
26 39 (119) 137
33
CeEhalozia catenulata (Hub) Lindb Cephalozia connivens (Dicks) Lindb
39 46 79 141 39 4679 88 (89) 90 119 141
Cephalozia lacinulata Jack ~ Spruce
141 Cephalozia lunulifolia (Dum~ Dum I 28 39 46 79 88 89 90 141
Cepoundhalozia macouni (Aust) Aust Cephalozia ~leniceps (Aust) Lindb 27~ 3746 (66)7987 (139)141 39 46 79 89 141
34
Ceffialoziella 9ivaricata (Sm) Schiffn Cephaloziella ~hista (Jack) Schiffn
46 90 46 (119) 141
Cephaloziella hampeana (Nees) Schiffn
46
Cephaloziella rubella (Nees) Warnst
46 39
Cephaloziella rubella var sullivantii
(Aust) K Mull 46 39
Cephaloziella rubella var bifida (Lindb) Douin
35
46
Chiloscyphus pallescens (Ehrh ~ Hof~ Dum Chiloscyphus pallescens var fragilis
39J 46 88 89J 90 (Roth) K Milll 46
Chiloscyphus E9lyanthos (L) Corda
1 24 39 46
Chiloscyphus polyanthos vax rivularis
(Schrad) Nees 39 46
poundpoundnocepha1um conicum (1) Lindb
12 28 39 46 (49) 87 88 8990 120 121 148
Cololejeunea biddlecomiae (Aust) Evans
39 46 62 87 88 89 90
36
Dipl0]hyllum apicu1atum (Evans) Steph Fossombronia foveo1ata Lindb
(35)41 46 90 140 1 t 38 (39) t 46
Frullania bolanderi Aust
13 46 62 88 90 139 140
Frullanla ari ttoniae Evans
39 46 69 (71) 87 88 89 90
Frullania eboracensis Gott
1 9 28 4 39 46 (57) (59) 69 70 (71) 87 88 89 90 92 93 97 148
Frullania inflata Gott
46 87 88 89 (139)
Frullania oakesiana Aust Frullania plana Sullo
(35) 42 46 87 (139) (140) 92
Frullania ri paria Hampe ~ Lehm
46 69 71 87 89
Frullania ~elwynianna Pears
46 89
Geocalyx SEaveolens (Schrad) Nees
394688 8990 (119) 140
Frullania tamarisci (L) Dum subsp sPagrayana
(Mont) Hatt 28 34 l~6 t (57) 88 89 90
38
Mar pan thus seutatUB ( Web amp Mohr) Spruce Jamesoniella autumnalis (DC) Steph
(35) 39 46 90 1 7 28 39 46 56 (57) 87 8889 90 (119) 137 141
Jungermannia confertissima Nees
137 141
Jungermannia crenuliform1s Aust
79 137 141
Jungermannia ~il1ima Sm
46 90 141 Jungermannia exsert1folia subsp corifo~ (Dum~
Vana (35) 46 79 141
39
Jungermannia hyalina Lyell Jungerrr6nnia 1eiantha Grolle
40 46 79 136 141 39 46 79 88 90 1)7 141
Jungermannia IXlaxis Lindb Jungermannia pumila With
46 67 79 (139) (140) 141 )9 46 90 1)6 141
Jungermannia sphaerocarpa Hook Kurzia setacea (Web) Grolle
4679136140IIH 136
Lejeunea cavifo1ia (Ehrh) Lindb Lepidozia reptans (1) Dum
1 7 27 39 46 88 89 90 137 14010 I ( 27) 46 I 69 (71) 88
lophocolea heterophylla (Schrad) Dum Lophocolea minor Nees
1 )4 39 46 (57) 87 88 89 27 39 46 88 89 90 (119) 90 (93)94 119 136
Lophozia al~stris (Schleich ~ Web) Evans Lophozia ascendens (Warnst) Schust
46 90 137 141 136 140 141
41
loIilozia badensls (Gatt ~ Rabenh)
46 141
lophozia blcrenata (Schmid sectA Hoffm)
Dum 46 141
lophozia collaris (Nees) Dum
39 46 141
lophozla excisa (Dicks) Dum
27 4679 141
lophozia gillmani (Aust) Schust lophozia gIlttulata (Undo tl H Arnell)
36 39 40 46 136 (140) 141 Evans 39 46 136 141
42
141
Lophozia heterocolEDs (Thed) MA Howe IoPhozia ipcisa (Schard) Dum
27 39 46 89 141 8 27 31 46 79 90 140 141
LoQhozia longidens Macoun Lophozia ventricosa
1 46 79 90 141
Lophozia ventricosa var silvicola
(Buch) Jones 141 LoEhozla ventricosa vax longif1ora (Nees)
Macoun 39 46 79 141
43
~~ fragrans (Balbis) Frye et Clark Mannia pi 10sa (Hornem) Frye et Clark
46 (139) (140) 9 46 79 116 139
Mannia triandra (Scop) Gro 11e
46 (140)
Marchantia pound91ymorpha L
1 13 28 3946 (49) 78 87 88 89 90 (Ill) 121 122 147
Mo~rckia hibernica (HookJ Gott
46 89
Metzgeria conjugata Lindb
44
46
Mylia anomala (Hook) S Gray Notothylas orbicularis (Schwein) Sullo
39 46 141 (35)45 46
Nowellia curvifolia (Dicks) Mitt
17 27 3846 88 89 90 141
Odontoschisma denudatuJI1 (Nees ~ Mart)
Dum 46 81 137 141
Pallavicinia lyellii (Hook~ Carruth
90
Odontoschisma macounii (Aust) Under
45 46 (66) (139) 140 141
45
=
Pellia epiphllla (L) CordaPellia endiviifolia (Dicks~ Dum
27 39 46 90 13724 39 46
Pellia neesiana Limp bull Phaeoceros
39 46 89 27 39 45 46 97
Porella pinnata L
13 27 I 46 90 Plagiochila asplenioides (L) Dum
27 I 39 1+6 88 89 I 90 137
46
~e11a platYEhylla (L) Pfeiff Porella platyphylloi~ (Schwein) Lindb
17 27 39 46 69 (71)79 87 88 89 9 34 46 (57) 59 87 88 89 92 90 92 93 77
Preissia quadrata (Seop) Nees
1 36 3946 (71) 88 89 90 121 123
137
Ptllidium ciliare (L) Hampe
1 27 39 46 132 141
Dum
(22) 28 3946 (57) 87 88 89 90 92
Ptilidium ~lcherrium (G Web) Hampe
(9) 13 394656 (57)70 87 88 89 90 93 (119) 136 141
47
Radula obconica Sullo Reboulia hemlsphaerlca (L) Raddi
39 46 88 89 (43) 46 (59) 70 90 t Cf 124 125 (139)
Riccardia ~atifrons Lindb
1 27 39 46 89
Riccardia multifi~ (L) S Gray
39 46 88 145
Riccia beyrichiana Hampe ~ Lehm
46 Riccardia palmata (Hedw) Carruth
39 46 90
48
Riccia bifurca Hoffm Riccia cavernosa Hoffm
48 13739 46 79 137
Riccia fluitans L
242731454679
Riccla frostli Aust
48 79
Riccia sullicantii Aust
79 (137) Riccia sorocarpa Bisch 46 (137)
49
Ricciocarpus natans (1) Corda Sca~ia apiculata Spruce
13 27 46 79 89 110 39 46 (139) 141
ScaJltll1ia carinthica Jack ~ Lindb
137 t 141
Scapania ~ (Mart) Durn
39467990137141
Scapania digenil Schiffn
90
ScaEsectflia cuspiduligera (Nees) K Mull
46 (139) (140) (141)
50
Scapania glaucoeerha1a (Tayl) Aust Seapania gyrrmostomophilia Kaal
39 46 (139) 141 41 44 46 64 (139) (140) 141
ScaP3poundlia irrigua Gott ~ almiddot
7 39 46 79 137 (140L 141
Seapania nemorosa (L) Dum
1 39 46 89 90 137
Scapmi~ ~~ta Smiddotlch
137 141 51
~caPnia poundludicola LDeske amp K Mull Scapania saxicola Schust
46 (66) 141 136 141
Sc~~ scandica (H Arnell ~ Buch) Macv
139 140
Scapania subalpina (Nees) Durn
39 40 46 79 137 141
Scapania undulata (L) Dum
1 42 46 79 141
Scapania umbrosa (Schrad) Dum
137 140 141
52
Trichocolea tomentella (Ehrh) Dum Triomarla exsecta (Schmid) Schiffn
27 (35) 39 46 47 88 137 141 8 28 39 46 90 140 141
Tritomaria exsectiformis (Breidl) Loe~ke Tritomaria Sluinguedentata (Huds) Buch
39 -t6 89 90 136 140 141 7 39 46 141
Trltomaria scitula (Tayl) Joerg
46 136 (139) 141
53
32 Ditrichaceae Pleuridium Brid
subulatum (Hedw) Rabenh
Ditrichum Hampe heteromallum (Hedw) Britt linellre (Sw) lindb
~ pallidum (Hedw) Hampe pusiUum (Hedw) Hampe
Saelania Lindb gIlucescens (Hedw) Bomanss et Broth
Ceratodon Brid purpureus (He(lw) Brid var prupurew
Distichium BSG capillaceum (Hedw) BSG inclinatum (Hedw) BSG
33 Bryoxipbiaceae Bryoxlphium Mitt
norvegicum (Brid) Mitt
34 Seligeriaceae Seligeria BSG
cumpylopoda Kindb ex Macoun et Kindb pusillll (Hedw) BSG recurvata (Hedw) BSG
Bindia BSG acuta (Hedw) BSG
35 Dicranaceae BlUehia Schwegr
lexUOSll (Sw ~ Schwaegr) C MueU sullivantii Aust
Trematodon Michx ambiguus (Hedw) Hornsch
Dicranellll (C Muell) Schimp cerviculata (Hedw) Schimp val cillPicuiata
heteromalla (Hedw) Schimp ~ rufescens (With) Schimp
schreberiana (Hedw) Schhnp var schreberiana schreberitJna (Hedw) Schimp var robusta Schimp ex Braithw subulata (Hedw) Schimp varia (Hedw) Schimp
Dicranodontium BSG denudatum (Brid) Britt ex Williams
Panzeucobryum (Lindh) Loeske Iongifolium (Hedw) oelke varlongifolium
17
Tortella (Undb) Limpr fragilis (Drumm) Umpr humilis (Hedw) JeM torlUoSfl (Hedw) Umpr
Hyophila Brid involuta (Hoole) Jaeg et Sauerb
Didymodon Hedw rigidulus Hedw
BryoerythrophyUum Chen recurvirostnon (Hedw) Chen
Btubula Hedw acula (Brid) Brid var acuta convoiuta Hedw fallox Hedw unguicultltll Hedw
Acaulon C MueD muticum (Uedw) C Muell var rufeseens (Iaeg) Crum
PJl~lm Hedw CUSPidtltum Hedw
De8mlltodon Brid cernuu (Hueb) BSG oblUsifoliw (Schwaegr) Schimp
T01IUla Hedw mueronifolia Schwaegr rurall (Hedw) Gaertn Meyer et Scherb var ruralis
39 Grimmiaceae Grlmmill Hedw
alpicola Hedw var alpicola alpicola Hedw var dupretii (Ther) Crum alpicola Hedw var rlvularl8 (Brid) Wahlenb apocarpa Hedw var apocarpa apocarpa Hedw var ambipa (Sull) Jonea apocarpa Hedw var conjerttl (Funck) Spreng apocarpa Hedw var strleta (Tum) Hook et Tayl CiIIyptrrlta Hook ex Drumm donniana Sm var donn_ hemumti Crum ltlevla (Brid) Brid montana BSG oineyi SuD oVllll8 (Hedw) lindb piUfe P-Beauv pltJgiopodill Hedw pulvinata (Hedw) Sm varbullpulvinata teretinervl Limpr
18
Orothera C Muell leana (Sull) C Muell
Rfwbdoweisia crispata (With) Undb
Cynodontium BSG ex Schimp alpestre (Wahlenb) Milde strumiferum (Hedw) Undb
Dichodontium Schimp pellucidum (Hedw) Schimp
Oncophorus (Slid) Brid vuens (Hedw) Brid wahlenbergii (Brid)
Dicranum Hedw bonjeanii DeNot ex Usa drummondii C Muell lagellare Hedw varj1ogellare fulvum Hook fuscescens Turn var fuscescens montanum Hedw muehlenbeckii BSG var muehlenbeckii polysetum Sw sabuletorum Ren et Card scoparium Hedw undulatum Brid viride (Sull et Lesq ex Sull) lindb
36 Leucobryaceae Leucobryum Hampe
glaucum (Hedw) Angstr ex Fr
37 Encalyptaceae Encalypta Hedw
ciliata Hedw var ciliata procera Bruch
38 Pottiaceae Astomum Hampe
muhlenbergianum (Sw) Grout
Weissia Hedw controversa Hedw var controversa
Gymnostomum Nees Homsch et Stunn aeruginosum Sm recurvirostrum Hedw var recurvirostrum
Trichostomum Bruch spirale Prout
19
40 Ephemeraceae Ephemerum Hampe
cohaerens (Hedw) Hampe crassineium (Schwaegr) Hampe val crassinervium spinulosum Bruch et Schimp ex Schimp
41 Funariaceae AphflllolThegma SuD
serratum (1 Hook et Wils ex Drumm) Sullo
Physcomitrium (Brid) Fuernr hookeri Hampe immersum Sullo pyriforme (Hedw) Hampe
Funaria Hedw americana Undb hygrometrica Hedw vaT hygrometrica
42 Splachnaceae Splachnum Hedw
ampullaceum Hedw
43 Schistostegaceae Schistostega Mohr
pennata (Hedw) Web el Mohr
44 Bryaceae Pohlia Hedw
annolina (Hedw) Lindb var annotina camea (Tum) Lindb cruda (Hedw) Undb elongata Hedw iliformis (Dicks) Andr lescurlana (Sull) Grout nutans (Hedw) Lindb obrusifolia (Brid) L Koch proligera (Undb ex Umpr) Undb ex Amell wahlenbergii (Web et Mohr) Andr
Leptobryum (B S G) Wits pyriforme (Hedw) Wils
Bryum Hedw algovicum Sendtnex C Muell argenteum Hedw caespiticium Hedw capillare Hedw var capillare creberrlmum Tayt lonchocaulon C Muell pallens (Brid) Sw ex Roehl pseudotrlquetrum (Hedw) Gaertn Meyer et Scherb varpudotriquetrum
20
pseudotriquetrum (Hedw) GaertnMeyer et Scherb var bimum (Schreb) Ulj stenotrichum C Muell var stenotrichum tortiolium Funck ex Brid uliginosum (Brid) B S G violaceum Crundw et Nyh weigelii Spreng
Rhodobryum (Schimp) Hampe roseum (Hedw) Limpr
45 Mniaceae Mnium Hedw
ciliare (C Muell) Schimp cinclidioides Hueb cuspidarum Hedw drummondii Bruch et Schimp longirostrum Brid lycopodioides Schwaegr marginatum (With) Brid ex P Beauv mediumB~S G var medium onhorrhynchum Brid pseudopunctatum Bruch et Schimp punctalUm Hedw var punctatum punctatum Hedw var elatum Schirnp rugicum Laur spinufosum B S G stellare Hedw
46 Aulacomruaceae Au1acomnium Schwaegr
androgynum (Hedw) Schwaegr heterostichum (Hedw) B S G paJustre (Hedw) Schwaegr var paJustre
47 Meesiaceae Amblyodon B S G
deillbotus (Hedw) B S G
Meesia Hedw triquetra (Richt) Angstr uJiginosa Hedw
48 Bartramiaceae Plagiopus Brid
oederiana (Sw) Umpr
Bartramia Hedw pomiformis Hedw var pomiformis
Philonotis Brid fontana (Hedw) Brid varfontana longseta (Michx) Britt varlongseta marchica (Hedw) Brid
21
~
~
49 Timmiaceae Timmia Hedw
megapolitana Hedw var megapolitana
50 Orthotrichaceae Zygodon Hook et Tayl
viridissimus (Dicks) Brid var rupestris lindb ex C J Hartm
Amphidium Schimp lapponicum (Hedw) Schimp mougeotii (B S G) Schimp
Orthotrichum Hedw anomalum Hedw obtuaifolium Brid ohioense Sullo et Lesq ex Aust pumilumSw sordidum Sullo et Lesq ex Aust speciosum Neesax Sturm var eegam (Schwaegr ex Hook et Grev) W~rnst stellDtwn Brid strrmgulatum P Beauv
Uleta Mohr ex Web cOfIICtata (p Beallv) Hamrn crispa (Hedw) Brid
Drummondill Hook ex Drumm prorepens (Hedw) Brltt
51 Fontinalaceae Fantinalis Hedw
antipyretica Hedw var antipyrettca antipyretica Hedw forma rabusta (Card) Card dalecarlica Schimp ex B S G var dalecarlica flqccidtJ Ren et Card hypnoides C J Hartrn var hypnoides hypnoides C J Hartm var durillei (Schimp) HU$ll
missourlcaCard novae-angliae Sullo var novae-angliae splulgrUlolill (C Moen) Wijket M~S
DichelY1l1ll Myr capilloceum (With) Myr falcatum (Bedw) Myr HIllescens B S G
52 Climaciaceae aimecium Web et Mohr ex Mohr
americllnum Brid dedroides (Hedw) Web et Mohr
22
S3 Hedwigiaceae Hedwigia P Beauv
ciliata (Hedw) P Beauv
54 Cryphaeaceae Forsstroemia lindh
ohioense (Sull) lindh
55 Leucodontaceae Leucodon Schwaegr
brachypus Blid var brachypus brachypus Brid var aniJrewsitlnus Crum et Anderson julaceus (Hedw) SuU
56 Neckeraceae Neckera Hedw
pennata Hedw var pennata
1fomalia (Brid) B S G trichomanoides (Hedw) B S G
57 Theliaceae Thelia Sull
asprellll Sullo hirtella (Hedw) Sullo lescurii SuU
Myurella B SG julacea (Schwaegr) B S G
sibrica (C MueU) Reim var sibrica
58 Fabroniaceae Fabronio Raddi
ciliaris (Brid) Bnd
Anacamptodon Brid splachnoides (Groet ex Brid) Brid
Schwetschkeopsis Broth fabronia (Schwaegr) Broth
59 Leskeaceae Lindbergia Kindb
brachyptera (Mitt) Kindb
Leskea Hedw gracilescens Hedw obscura Hedw poiycarpa Hedw
Leskeella (limpr) Loeske nervosa (Brid) Loeske
23
Pterlgy1lllndrum Hedw filiforme Hedw
60 Thuidiaceae Haplohymenfum Dozy et MoIk
trlste (Ces ex De Not) Kiodb
Anomodon Hook et Tayl attenuatus (Hedw) Hueb minor (Hedw) Fuemr rostratus (Hedw) Schimp viticulosus (Hedw) Hook et Tayl
Haplocladium (C Mucn) C MueU microphyllum (Hedw) Broth virginillnum (Brld) Broth ~bull
Thuidium B S G abietinum (Hedw) B S G delicatulum (Hedw) B S G vardelicatulum delicatulum (Hedw) B S G var radicans (Kindb) Crum Steere et Anderson minutulum (Hedw) B S G pygmaeum B S G recognftum (Hedw) Lindb scitum (p Beauv) Aust
Helodium (Sull) Warnst blandowli (Web et Mohr) Warnst var blandowli blandowli (Web et Mohr) Wamst var helodioides (Ren et Card ex Roell) Crum
Steere et Anderson paludomm (Sull) AuS
61 Amblystegiaceae Cratoneuron (Sull) Spruce
commulatum (Hedw) Roth var commutfltum commutatum (Hedw) Roth var fakatum (Brid) Moenk ilicinum (Hedw) Spruce var filcinum
Campylium (Sun) Mitt chrysophyllum (Brid) 1 Lange var chrysophyllum hispidulum (Brid) Mitt var hllJidulum polygamum (B S G)C lens varpolygtlmUm stellatum (Hedw) C1ens vat stelltum
Leptodictyum (Schimp) Warnst laxirete (Card et Ther) Broth riparium (Hedw) Wamat trichopodium (Schultz) Warns l1acillans (Sull) Broth
Hygroamblystegium Loelke jlul1iatile (Hedw) oeske var luViatile
24
fluviatile (Hedw) Loeske var oWltum Grout noterophilum (Sun et Lesq ex Sull) Warnst tenox (Hedw) Jenn var tenax tenox (Hedw) Jenn var spinilolium Schimp) Jenn
Amblystegium BSG juratzkanum Schimp serpens (Hedw) BSG varlum (Hedw) Undb var varium
Platydictya Berk confervoides (Brid) Crum jungermanniodes (Brid) Crum minutissimum (Sull et Lesq ex Sull) Crum subtile (Hedw) Crum
Drepanocladus (C Muell) Roth aduncus (Hedw) Warnst var aduncus aduncus (Hedw) Warnst var capillilolius (Wamst)Riebm aduncus (Hedw) Warnst var kneiffii (BSG) Moenk aduncus (Hedw) Warnst var polycorpus (Bland ex Voit) Roth exannulatus (BSG) Warnst fluitans (Hedw) Warnst var fluitans revolvens (Sw) Wamst var revolvens sendtneri (Schimp) Warnst var sendtnerl uncinatus (Hedw) Warnst var uncinatus uncinatus ffiedw) Warnst var piwnosus (Schimp) Wamst vernicoms (Lindb ex C Hartm) Warnst
Hygrohypnum Lindb eugyrium (BSG) loeske luridum (Hedw Jenn varluridum ochraecum (Tum ex Wils) Loeske var ochraceunt
Calliergon (Sull) Kindb cordilolium Hedw Kindb glganteum (Schimp) Kindb var giganteum richardsonii (Mitt) Kindb ex Wamst stramineum (Brid) Kindb trilarium (Web et Mohr) Kindb
Calliergonella Loeske cuspidata (Hedw) Loeske
Calliergidium (Ren) Grout bakeri (Ren) Grout
Scorpidium (Schimp) Urnpr scorpiodides (Hedw) limpr
62 Brachytbeciaceae Tomenthypnum loeske
nitens (Hedw) Loeske var mtens
25
Homalotheciella (Card) Broth subcapillata (Hedw) Broth
Brachythecium BSG acuminatum (Hedw) Awt acutum (Mitt) Sullo calcareum IGndb campestTe (C MuelL) BSG cunum (Lindb) Umpr digastrum C Muell et Kindb ex Macoun et Kindb oxycladon (Brid) Jaeg et Sauerb var oxycladon plumosum (Hedw) BSG populeum (Hedw) BSG var populeum reflexum (Starke ex Web et Mohr) DsG var relexum reiexum (Starke ex Web et Mohr) DSG var pacificum Renet Card ex Roell rlvulare BSG var rivulare tabulum (Hedw) BSC var tabuum salebrosum (Web et Mohr) BSG var salebrosum velutinum (Hedw) BSG Vat velutinum
Bryhniil Kaur graminicolor (Brid) Grout var graminicolor novae-angliae (Sull et Lesq ex sun) Grout
Rhynchostegium BSG senulatum (Hedw) Ja4lg et Sauerb
Rhynchostegiella (BSG) Limpr compacta (C MueU) Loeske
EU1hynchium 8SG hians (Hedw) Sande Lac pulchellum (Hedw) Jenn var pulchelium riparloides (Hedw) Rich
Stokeriella (Kindb) Robins pmelonga (Hedw) Robins var stokesii (Tum) Crum
63 Entodontaceae Entodon C Muell
brevisetus (Hook et Wils ex Wils) Lindb cladorrhizans (Hedw) C Muell seductrlx (Hedw) C MueU var wluctrix
Pleurozium Mitt schreberl (8rid) Mitt var schreberi
64 Plagiotheciaceae Pltlglothecium BSG
CflVifolium (Brid) Iwats var CtWifolium denticulatum (Hedw) 8SG laetum BSG ltztebricola BSG
26
6S SernatophyUaceae Brotherella Loeske ex Fleisch
recumins (Michx) Fleisch tenuirostris (Bruch et Schimp ex Sull) Fleisch
Sematophyllum Mitt demissum (Wils) Mitt
66 Hypnaceae Pl(itygyrium BSG
repens (Brid) BSG
PylaisieUa Kindb intricata (Hedw) Grout polyantha (Hedw) Grout selwynii (Kindb) Crum Steere et Anderson
Homomallium (Schimp) Loeske adnatum (Hedw) Broth
Hypnum Hedw cupressiforme Hedw var cupressiforme cupressifonne Hedw varfiliforme Brid curviolium Hedw fertile Sendtn imponens Hedw lindbergii Mitt palescens (Hedw) P-Beauv pratense Koch ex Brid
Calicladium Crum haldanianum (Grev) Crum
sopterygiopsis Iwats muelleriana (Schimp) Iwats
Isopterygium Mitt e1egans (Brid) lindb pulchellum (Hedw) Jaeg et Sauerb tenerum (Sw) Mitt
Taxiphyllum Fleisch depllQtum (Bruch et Schimp ex sun) Fleisch taxirameum (MitL) Fleisch
Herzogiella Broth seligeri (Brid) Iwats striatella (Brid) Iwats ruracea (Lindh) Iwats
hilium De Not crista-castrenna (Hedw) De Not
27
67 Rhytidiaceae Rhytidium (Sull) Kindb
rugosum (Hedw) Kindb
Rhytidfodelphus (Lindb ex Iimpr) Warnst triquetrus (Hedw) Warnst var triquetrus
68 Hylocomiaceae Hylocomium BSG
pyrenaicum (Spruce) lindb splendens (Hedw) BSG varsplendens
69 Buxbaumiaceae Buxbaumia Hedw
aphyllll Hedw
Diphyscium Mohr follosum (Hedw) Mohr
70 Tetraphidaceae Tetraphi3
pellucida Hedw
71 Polytrichaceae Atrichum PmiddotBeauv
angustatum (Brid) BSG cri8pum (James) Sullo oerstedianum (C Muell) Mitt teneUum (Roehl) BSG undulatum (Hedw) P middotBeauv var unduiatum
Pogonatum P-Beauv alpinum (Hedw) Roehl var alpinum umigerum (Hedw) P-Beauv
Poiytrichum Hedw commune Hedw var commune ormosum Hedw juniperinum Hedw longisetum Brid ohioense Ren et Card piliferum Hedw var pilierum strictum Brid
28
MAPS OF BRYOPHYTE TAXA
REPORTED FOR WISCONSIN
HORNWORTS AND LIVERWORTS
29
bull bull WSHeV~N S vE II
0 ~
e~RRON ~USK
(J~ (j -
CHIPPE A DUNN
0 o0 poundAU CLAIRE
~ TREMshyPEAshyLEAU
MICE
ONEIDA
Q LINCOLN
TAYLOR
Q t-~----- MOIRA THON
CLARK
o
10
LA FAVETTE
Hornwor ts amp Liverworts
- None - 1 - 25 taxa
- 26 - 50 tRXshy
- 51 - 75 tax)
- 7c - 100 taxa
FONDOU j LAC ~
DODGE
i O JEfFERSON
WALWORTH
30
Anastrophylltlfl helleranum (Nees) Schust Anastrophyllum michauxil (We~ Buch ~
46 88 136 141 Evans 38 46 141
Anastrophyllum mlnut~ (Schreb) Schuster Aneura plnguls (L) Durn
89 21 384687 88 89 90 (138) (145)
Anthoceros macounil MA Howe ~oceros punctatus L
21 (45) (144) 34
31
Barbilophozia attenuata (Mart) Loeske Barbilophozia barbata (Schmid ~ Schre~
24 141 Loeske 28 39 46 89 90 141
Bazzania trilobata (1) S Gray
1 26 39 46 (57) (59) 79 88 89
90 119 141
BlePbarostoma trichoffiyllum (1) Dumort
1 8 26 39 46 89 90 137 141
Blasia pusilla 1
1 39 46 90
Calypogeja fissa (1) Raddi
90
32
$129geja integristipula Steph C~Ypound2geja muelleriana (Schifm) K Mull
88 89 89
CallpoundO~eja neesiana (Mass amp Carest) K Mull
39 46 89 140 (141)
~pogeja sJlapound2icola (H Arnell tl J bull
Perss) Warnst et Loeske (119) 141
CeIalozia biscuspidata (L) Dum
1 37 I 46 90 t 137 141
CalYpogej~ trichomanis (L) Corda
26 39 (119) 137
33
CeEhalozia catenulata (Hub) Lindb Cephalozia connivens (Dicks) Lindb
39 46 79 141 39 4679 88 (89) 90 119 141
Cephalozia lacinulata Jack ~ Spruce
141 Cephalozia lunulifolia (Dum~ Dum I 28 39 46 79 88 89 90 141
Cepoundhalozia macouni (Aust) Aust Cephalozia ~leniceps (Aust) Lindb 27~ 3746 (66)7987 (139)141 39 46 79 89 141
34
Ceffialoziella 9ivaricata (Sm) Schiffn Cephaloziella ~hista (Jack) Schiffn
46 90 46 (119) 141
Cephaloziella hampeana (Nees) Schiffn
46
Cephaloziella rubella (Nees) Warnst
46 39
Cephaloziella rubella var sullivantii
(Aust) K Mull 46 39
Cephaloziella rubella var bifida (Lindb) Douin
35
46
Chiloscyphus pallescens (Ehrh ~ Hof~ Dum Chiloscyphus pallescens var fragilis
39J 46 88 89J 90 (Roth) K Milll 46
Chiloscyphus E9lyanthos (L) Corda
1 24 39 46
Chiloscyphus polyanthos vax rivularis
(Schrad) Nees 39 46
poundpoundnocepha1um conicum (1) Lindb
12 28 39 46 (49) 87 88 8990 120 121 148
Cololejeunea biddlecomiae (Aust) Evans
39 46 62 87 88 89 90
36
Dipl0]hyllum apicu1atum (Evans) Steph Fossombronia foveo1ata Lindb
(35)41 46 90 140 1 t 38 (39) t 46
Frullania bolanderi Aust
13 46 62 88 90 139 140
Frullanla ari ttoniae Evans
39 46 69 (71) 87 88 89 90
Frullania eboracensis Gott
1 9 28 4 39 46 (57) (59) 69 70 (71) 87 88 89 90 92 93 97 148
Frullania inflata Gott
46 87 88 89 (139)
Frullania oakesiana Aust Frullania plana Sullo
(35) 42 46 87 (139) (140) 92
Frullania ri paria Hampe ~ Lehm
46 69 71 87 89
Frullania ~elwynianna Pears
46 89
Geocalyx SEaveolens (Schrad) Nees
394688 8990 (119) 140
Frullania tamarisci (L) Dum subsp sPagrayana
(Mont) Hatt 28 34 l~6 t (57) 88 89 90
38
Mar pan thus seutatUB ( Web amp Mohr) Spruce Jamesoniella autumnalis (DC) Steph
(35) 39 46 90 1 7 28 39 46 56 (57) 87 8889 90 (119) 137 141
Jungermannia confertissima Nees
137 141
Jungermannia crenuliform1s Aust
79 137 141
Jungermannia ~il1ima Sm
46 90 141 Jungermannia exsert1folia subsp corifo~ (Dum~
Vana (35) 46 79 141
39
Jungermannia hyalina Lyell Jungerrr6nnia 1eiantha Grolle
40 46 79 136 141 39 46 79 88 90 1)7 141
Jungermannia IXlaxis Lindb Jungermannia pumila With
46 67 79 (139) (140) 141 )9 46 90 1)6 141
Jungermannia sphaerocarpa Hook Kurzia setacea (Web) Grolle
4679136140IIH 136
Lejeunea cavifo1ia (Ehrh) Lindb Lepidozia reptans (1) Dum
1 7 27 39 46 88 89 90 137 14010 I ( 27) 46 I 69 (71) 88
lophocolea heterophylla (Schrad) Dum Lophocolea minor Nees
1 )4 39 46 (57) 87 88 89 27 39 46 88 89 90 (119) 90 (93)94 119 136
Lophozia al~stris (Schleich ~ Web) Evans Lophozia ascendens (Warnst) Schust
46 90 137 141 136 140 141
41
loIilozia badensls (Gatt ~ Rabenh)
46 141
lophozia blcrenata (Schmid sectA Hoffm)
Dum 46 141
lophozia collaris (Nees) Dum
39 46 141
lophozla excisa (Dicks) Dum
27 4679 141
lophozia gillmani (Aust) Schust lophozia gIlttulata (Undo tl H Arnell)
36 39 40 46 136 (140) 141 Evans 39 46 136 141
42
141
Lophozia heterocolEDs (Thed) MA Howe IoPhozia ipcisa (Schard) Dum
27 39 46 89 141 8 27 31 46 79 90 140 141
LoQhozia longidens Macoun Lophozia ventricosa
1 46 79 90 141
Lophozia ventricosa var silvicola
(Buch) Jones 141 LoEhozla ventricosa vax longif1ora (Nees)
Macoun 39 46 79 141
43
~~ fragrans (Balbis) Frye et Clark Mannia pi 10sa (Hornem) Frye et Clark
46 (139) (140) 9 46 79 116 139
Mannia triandra (Scop) Gro 11e
46 (140)
Marchantia pound91ymorpha L
1 13 28 3946 (49) 78 87 88 89 90 (Ill) 121 122 147
Mo~rckia hibernica (HookJ Gott
46 89
Metzgeria conjugata Lindb
44
46
Mylia anomala (Hook) S Gray Notothylas orbicularis (Schwein) Sullo
39 46 141 (35)45 46
Nowellia curvifolia (Dicks) Mitt
17 27 3846 88 89 90 141
Odontoschisma denudatuJI1 (Nees ~ Mart)
Dum 46 81 137 141
Pallavicinia lyellii (Hook~ Carruth
90
Odontoschisma macounii (Aust) Under
45 46 (66) (139) 140 141
45
=
Pellia epiphllla (L) CordaPellia endiviifolia (Dicks~ Dum
27 39 46 90 13724 39 46
Pellia neesiana Limp bull Phaeoceros
39 46 89 27 39 45 46 97
Porella pinnata L
13 27 I 46 90 Plagiochila asplenioides (L) Dum
27 I 39 1+6 88 89 I 90 137
46
~e11a platYEhylla (L) Pfeiff Porella platyphylloi~ (Schwein) Lindb
17 27 39 46 69 (71)79 87 88 89 9 34 46 (57) 59 87 88 89 92 90 92 93 77
Preissia quadrata (Seop) Nees
1 36 3946 (71) 88 89 90 121 123
137
Ptllidium ciliare (L) Hampe
1 27 39 46 132 141
Dum
(22) 28 3946 (57) 87 88 89 90 92
Ptilidium ~lcherrium (G Web) Hampe
(9) 13 394656 (57)70 87 88 89 90 93 (119) 136 141
47
Radula obconica Sullo Reboulia hemlsphaerlca (L) Raddi
39 46 88 89 (43) 46 (59) 70 90 t Cf 124 125 (139)
Riccardia ~atifrons Lindb
1 27 39 46 89
Riccardia multifi~ (L) S Gray
39 46 88 145
Riccia beyrichiana Hampe ~ Lehm
46 Riccardia palmata (Hedw) Carruth
39 46 90
48
Riccia bifurca Hoffm Riccia cavernosa Hoffm
48 13739 46 79 137
Riccia fluitans L
242731454679
Riccla frostli Aust
48 79
Riccia sullicantii Aust
79 (137) Riccia sorocarpa Bisch 46 (137)
49
Ricciocarpus natans (1) Corda Sca~ia apiculata Spruce
13 27 46 79 89 110 39 46 (139) 141
ScaJltll1ia carinthica Jack ~ Lindb
137 t 141
Scapania ~ (Mart) Durn
39467990137141
Scapania digenil Schiffn
90
ScaEsectflia cuspiduligera (Nees) K Mull
46 (139) (140) (141)
50
Scapania glaucoeerha1a (Tayl) Aust Seapania gyrrmostomophilia Kaal
39 46 (139) 141 41 44 46 64 (139) (140) 141
ScaP3poundlia irrigua Gott ~ almiddot
7 39 46 79 137 (140L 141
Seapania nemorosa (L) Dum
1 39 46 89 90 137
Scapmi~ ~~ta Smiddotlch
137 141 51
~caPnia poundludicola LDeske amp K Mull Scapania saxicola Schust
46 (66) 141 136 141
Sc~~ scandica (H Arnell ~ Buch) Macv
139 140
Scapania subalpina (Nees) Durn
39 40 46 79 137 141
Scapania undulata (L) Dum
1 42 46 79 141
Scapania umbrosa (Schrad) Dum
137 140 141
52
Trichocolea tomentella (Ehrh) Dum Triomarla exsecta (Schmid) Schiffn
27 (35) 39 46 47 88 137 141 8 28 39 46 90 140 141
Tritomaria exsectiformis (Breidl) Loe~ke Tritomaria Sluinguedentata (Huds) Buch
39 -t6 89 90 136 140 141 7 39 46 141
Trltomaria scitula (Tayl) Joerg
46 136 (139) 141
53
Tortella (Undb) Limpr fragilis (Drumm) Umpr humilis (Hedw) JeM torlUoSfl (Hedw) Umpr
Hyophila Brid involuta (Hoole) Jaeg et Sauerb
Didymodon Hedw rigidulus Hedw
BryoerythrophyUum Chen recurvirostnon (Hedw) Chen
Btubula Hedw acula (Brid) Brid var acuta convoiuta Hedw fallox Hedw unguicultltll Hedw
Acaulon C MueD muticum (Uedw) C Muell var rufeseens (Iaeg) Crum
PJl~lm Hedw CUSPidtltum Hedw
De8mlltodon Brid cernuu (Hueb) BSG oblUsifoliw (Schwaegr) Schimp
T01IUla Hedw mueronifolia Schwaegr rurall (Hedw) Gaertn Meyer et Scherb var ruralis
39 Grimmiaceae Grlmmill Hedw
alpicola Hedw var alpicola alpicola Hedw var dupretii (Ther) Crum alpicola Hedw var rlvularl8 (Brid) Wahlenb apocarpa Hedw var apocarpa apocarpa Hedw var ambipa (Sull) Jonea apocarpa Hedw var conjerttl (Funck) Spreng apocarpa Hedw var strleta (Tum) Hook et Tayl CiIIyptrrlta Hook ex Drumm donniana Sm var donn_ hemumti Crum ltlevla (Brid) Brid montana BSG oineyi SuD oVllll8 (Hedw) lindb piUfe P-Beauv pltJgiopodill Hedw pulvinata (Hedw) Sm varbullpulvinata teretinervl Limpr
18
Orothera C Muell leana (Sull) C Muell
Rfwbdoweisia crispata (With) Undb
Cynodontium BSG ex Schimp alpestre (Wahlenb) Milde strumiferum (Hedw) Undb
Dichodontium Schimp pellucidum (Hedw) Schimp
Oncophorus (Slid) Brid vuens (Hedw) Brid wahlenbergii (Brid)
Dicranum Hedw bonjeanii DeNot ex Usa drummondii C Muell lagellare Hedw varj1ogellare fulvum Hook fuscescens Turn var fuscescens montanum Hedw muehlenbeckii BSG var muehlenbeckii polysetum Sw sabuletorum Ren et Card scoparium Hedw undulatum Brid viride (Sull et Lesq ex Sull) lindb
36 Leucobryaceae Leucobryum Hampe
glaucum (Hedw) Angstr ex Fr
37 Encalyptaceae Encalypta Hedw
ciliata Hedw var ciliata procera Bruch
38 Pottiaceae Astomum Hampe
muhlenbergianum (Sw) Grout
Weissia Hedw controversa Hedw var controversa
Gymnostomum Nees Homsch et Stunn aeruginosum Sm recurvirostrum Hedw var recurvirostrum
Trichostomum Bruch spirale Prout
19
40 Ephemeraceae Ephemerum Hampe
cohaerens (Hedw) Hampe crassineium (Schwaegr) Hampe val crassinervium spinulosum Bruch et Schimp ex Schimp
41 Funariaceae AphflllolThegma SuD
serratum (1 Hook et Wils ex Drumm) Sullo
Physcomitrium (Brid) Fuernr hookeri Hampe immersum Sullo pyriforme (Hedw) Hampe
Funaria Hedw americana Undb hygrometrica Hedw vaT hygrometrica
42 Splachnaceae Splachnum Hedw
ampullaceum Hedw
43 Schistostegaceae Schistostega Mohr
pennata (Hedw) Web el Mohr
44 Bryaceae Pohlia Hedw
annolina (Hedw) Lindb var annotina camea (Tum) Lindb cruda (Hedw) Undb elongata Hedw iliformis (Dicks) Andr lescurlana (Sull) Grout nutans (Hedw) Lindb obrusifolia (Brid) L Koch proligera (Undb ex Umpr) Undb ex Amell wahlenbergii (Web et Mohr) Andr
Leptobryum (B S G) Wits pyriforme (Hedw) Wils
Bryum Hedw algovicum Sendtnex C Muell argenteum Hedw caespiticium Hedw capillare Hedw var capillare creberrlmum Tayt lonchocaulon C Muell pallens (Brid) Sw ex Roehl pseudotrlquetrum (Hedw) Gaertn Meyer et Scherb varpudotriquetrum
20
pseudotriquetrum (Hedw) GaertnMeyer et Scherb var bimum (Schreb) Ulj stenotrichum C Muell var stenotrichum tortiolium Funck ex Brid uliginosum (Brid) B S G violaceum Crundw et Nyh weigelii Spreng
Rhodobryum (Schimp) Hampe roseum (Hedw) Limpr
45 Mniaceae Mnium Hedw
ciliare (C Muell) Schimp cinclidioides Hueb cuspidarum Hedw drummondii Bruch et Schimp longirostrum Brid lycopodioides Schwaegr marginatum (With) Brid ex P Beauv mediumB~S G var medium onhorrhynchum Brid pseudopunctatum Bruch et Schimp punctalUm Hedw var punctatum punctatum Hedw var elatum Schirnp rugicum Laur spinufosum B S G stellare Hedw
46 Aulacomruaceae Au1acomnium Schwaegr
androgynum (Hedw) Schwaegr heterostichum (Hedw) B S G paJustre (Hedw) Schwaegr var paJustre
47 Meesiaceae Amblyodon B S G
deillbotus (Hedw) B S G
Meesia Hedw triquetra (Richt) Angstr uJiginosa Hedw
48 Bartramiaceae Plagiopus Brid
oederiana (Sw) Umpr
Bartramia Hedw pomiformis Hedw var pomiformis
Philonotis Brid fontana (Hedw) Brid varfontana longseta (Michx) Britt varlongseta marchica (Hedw) Brid
21
~
~
49 Timmiaceae Timmia Hedw
megapolitana Hedw var megapolitana
50 Orthotrichaceae Zygodon Hook et Tayl
viridissimus (Dicks) Brid var rupestris lindb ex C J Hartm
Amphidium Schimp lapponicum (Hedw) Schimp mougeotii (B S G) Schimp
Orthotrichum Hedw anomalum Hedw obtuaifolium Brid ohioense Sullo et Lesq ex Aust pumilumSw sordidum Sullo et Lesq ex Aust speciosum Neesax Sturm var eegam (Schwaegr ex Hook et Grev) W~rnst stellDtwn Brid strrmgulatum P Beauv
Uleta Mohr ex Web cOfIICtata (p Beallv) Hamrn crispa (Hedw) Brid
Drummondill Hook ex Drumm prorepens (Hedw) Brltt
51 Fontinalaceae Fantinalis Hedw
antipyretica Hedw var antipyrettca antipyretica Hedw forma rabusta (Card) Card dalecarlica Schimp ex B S G var dalecarlica flqccidtJ Ren et Card hypnoides C J Hartrn var hypnoides hypnoides C J Hartm var durillei (Schimp) HU$ll
missourlcaCard novae-angliae Sullo var novae-angliae splulgrUlolill (C Moen) Wijket M~S
DichelY1l1ll Myr capilloceum (With) Myr falcatum (Bedw) Myr HIllescens B S G
52 Climaciaceae aimecium Web et Mohr ex Mohr
americllnum Brid dedroides (Hedw) Web et Mohr
22
S3 Hedwigiaceae Hedwigia P Beauv
ciliata (Hedw) P Beauv
54 Cryphaeaceae Forsstroemia lindh
ohioense (Sull) lindh
55 Leucodontaceae Leucodon Schwaegr
brachypus Blid var brachypus brachypus Brid var aniJrewsitlnus Crum et Anderson julaceus (Hedw) SuU
56 Neckeraceae Neckera Hedw
pennata Hedw var pennata
1fomalia (Brid) B S G trichomanoides (Hedw) B S G
57 Theliaceae Thelia Sull
asprellll Sullo hirtella (Hedw) Sullo lescurii SuU
Myurella B SG julacea (Schwaegr) B S G
sibrica (C MueU) Reim var sibrica
58 Fabroniaceae Fabronio Raddi
ciliaris (Brid) Bnd
Anacamptodon Brid splachnoides (Groet ex Brid) Brid
Schwetschkeopsis Broth fabronia (Schwaegr) Broth
59 Leskeaceae Lindbergia Kindb
brachyptera (Mitt) Kindb
Leskea Hedw gracilescens Hedw obscura Hedw poiycarpa Hedw
Leskeella (limpr) Loeske nervosa (Brid) Loeske
23
Pterlgy1lllndrum Hedw filiforme Hedw
60 Thuidiaceae Haplohymenfum Dozy et MoIk
trlste (Ces ex De Not) Kiodb
Anomodon Hook et Tayl attenuatus (Hedw) Hueb minor (Hedw) Fuemr rostratus (Hedw) Schimp viticulosus (Hedw) Hook et Tayl
Haplocladium (C Mucn) C MueU microphyllum (Hedw) Broth virginillnum (Brld) Broth ~bull
Thuidium B S G abietinum (Hedw) B S G delicatulum (Hedw) B S G vardelicatulum delicatulum (Hedw) B S G var radicans (Kindb) Crum Steere et Anderson minutulum (Hedw) B S G pygmaeum B S G recognftum (Hedw) Lindb scitum (p Beauv) Aust
Helodium (Sull) Warnst blandowli (Web et Mohr) Warnst var blandowli blandowli (Web et Mohr) Wamst var helodioides (Ren et Card ex Roell) Crum
Steere et Anderson paludomm (Sull) AuS
61 Amblystegiaceae Cratoneuron (Sull) Spruce
commulatum (Hedw) Roth var commutfltum commutatum (Hedw) Roth var fakatum (Brid) Moenk ilicinum (Hedw) Spruce var filcinum
Campylium (Sun) Mitt chrysophyllum (Brid) 1 Lange var chrysophyllum hispidulum (Brid) Mitt var hllJidulum polygamum (B S G)C lens varpolygtlmUm stellatum (Hedw) C1ens vat stelltum
Leptodictyum (Schimp) Warnst laxirete (Card et Ther) Broth riparium (Hedw) Wamat trichopodium (Schultz) Warns l1acillans (Sull) Broth
Hygroamblystegium Loelke jlul1iatile (Hedw) oeske var luViatile
24
fluviatile (Hedw) Loeske var oWltum Grout noterophilum (Sun et Lesq ex Sull) Warnst tenox (Hedw) Jenn var tenax tenox (Hedw) Jenn var spinilolium Schimp) Jenn
Amblystegium BSG juratzkanum Schimp serpens (Hedw) BSG varlum (Hedw) Undb var varium
Platydictya Berk confervoides (Brid) Crum jungermanniodes (Brid) Crum minutissimum (Sull et Lesq ex Sull) Crum subtile (Hedw) Crum
Drepanocladus (C Muell) Roth aduncus (Hedw) Warnst var aduncus aduncus (Hedw) Warnst var capillilolius (Wamst)Riebm aduncus (Hedw) Warnst var kneiffii (BSG) Moenk aduncus (Hedw) Warnst var polycorpus (Bland ex Voit) Roth exannulatus (BSG) Warnst fluitans (Hedw) Warnst var fluitans revolvens (Sw) Wamst var revolvens sendtneri (Schimp) Warnst var sendtnerl uncinatus (Hedw) Warnst var uncinatus uncinatus ffiedw) Warnst var piwnosus (Schimp) Wamst vernicoms (Lindb ex C Hartm) Warnst
Hygrohypnum Lindb eugyrium (BSG) loeske luridum (Hedw Jenn varluridum ochraecum (Tum ex Wils) Loeske var ochraceunt
Calliergon (Sull) Kindb cordilolium Hedw Kindb glganteum (Schimp) Kindb var giganteum richardsonii (Mitt) Kindb ex Wamst stramineum (Brid) Kindb trilarium (Web et Mohr) Kindb
Calliergonella Loeske cuspidata (Hedw) Loeske
Calliergidium (Ren) Grout bakeri (Ren) Grout
Scorpidium (Schimp) Urnpr scorpiodides (Hedw) limpr
62 Brachytbeciaceae Tomenthypnum loeske
nitens (Hedw) Loeske var mtens
25
Homalotheciella (Card) Broth subcapillata (Hedw) Broth
Brachythecium BSG acuminatum (Hedw) Awt acutum (Mitt) Sullo calcareum IGndb campestTe (C MuelL) BSG cunum (Lindb) Umpr digastrum C Muell et Kindb ex Macoun et Kindb oxycladon (Brid) Jaeg et Sauerb var oxycladon plumosum (Hedw) BSG populeum (Hedw) BSG var populeum reflexum (Starke ex Web et Mohr) DsG var relexum reiexum (Starke ex Web et Mohr) DSG var pacificum Renet Card ex Roell rlvulare BSG var rivulare tabulum (Hedw) BSC var tabuum salebrosum (Web et Mohr) BSG var salebrosum velutinum (Hedw) BSG Vat velutinum
Bryhniil Kaur graminicolor (Brid) Grout var graminicolor novae-angliae (Sull et Lesq ex sun) Grout
Rhynchostegium BSG senulatum (Hedw) Ja4lg et Sauerb
Rhynchostegiella (BSG) Limpr compacta (C MueU) Loeske
EU1hynchium 8SG hians (Hedw) Sande Lac pulchellum (Hedw) Jenn var pulchelium riparloides (Hedw) Rich
Stokeriella (Kindb) Robins pmelonga (Hedw) Robins var stokesii (Tum) Crum
63 Entodontaceae Entodon C Muell
brevisetus (Hook et Wils ex Wils) Lindb cladorrhizans (Hedw) C Muell seductrlx (Hedw) C MueU var wluctrix
Pleurozium Mitt schreberl (8rid) Mitt var schreberi
64 Plagiotheciaceae Pltlglothecium BSG
CflVifolium (Brid) Iwats var CtWifolium denticulatum (Hedw) 8SG laetum BSG ltztebricola BSG
26
6S SernatophyUaceae Brotherella Loeske ex Fleisch
recumins (Michx) Fleisch tenuirostris (Bruch et Schimp ex Sull) Fleisch
Sematophyllum Mitt demissum (Wils) Mitt
66 Hypnaceae Pl(itygyrium BSG
repens (Brid) BSG
PylaisieUa Kindb intricata (Hedw) Grout polyantha (Hedw) Grout selwynii (Kindb) Crum Steere et Anderson
Homomallium (Schimp) Loeske adnatum (Hedw) Broth
Hypnum Hedw cupressiforme Hedw var cupressiforme cupressifonne Hedw varfiliforme Brid curviolium Hedw fertile Sendtn imponens Hedw lindbergii Mitt palescens (Hedw) P-Beauv pratense Koch ex Brid
Calicladium Crum haldanianum (Grev) Crum
sopterygiopsis Iwats muelleriana (Schimp) Iwats
Isopterygium Mitt e1egans (Brid) lindb pulchellum (Hedw) Jaeg et Sauerb tenerum (Sw) Mitt
Taxiphyllum Fleisch depllQtum (Bruch et Schimp ex sun) Fleisch taxirameum (MitL) Fleisch
Herzogiella Broth seligeri (Brid) Iwats striatella (Brid) Iwats ruracea (Lindh) Iwats
hilium De Not crista-castrenna (Hedw) De Not
27
67 Rhytidiaceae Rhytidium (Sull) Kindb
rugosum (Hedw) Kindb
Rhytidfodelphus (Lindb ex Iimpr) Warnst triquetrus (Hedw) Warnst var triquetrus
68 Hylocomiaceae Hylocomium BSG
pyrenaicum (Spruce) lindb splendens (Hedw) BSG varsplendens
69 Buxbaumiaceae Buxbaumia Hedw
aphyllll Hedw
Diphyscium Mohr follosum (Hedw) Mohr
70 Tetraphidaceae Tetraphi3
pellucida Hedw
71 Polytrichaceae Atrichum PmiddotBeauv
angustatum (Brid) BSG cri8pum (James) Sullo oerstedianum (C Muell) Mitt teneUum (Roehl) BSG undulatum (Hedw) P middotBeauv var unduiatum
Pogonatum P-Beauv alpinum (Hedw) Roehl var alpinum umigerum (Hedw) P-Beauv
Poiytrichum Hedw commune Hedw var commune ormosum Hedw juniperinum Hedw longisetum Brid ohioense Ren et Card piliferum Hedw var pilierum strictum Brid
28
MAPS OF BRYOPHYTE TAXA
REPORTED FOR WISCONSIN
HORNWORTS AND LIVERWORTS
29
bull bull WSHeV~N S vE II
0 ~
e~RRON ~USK
(J~ (j -
CHIPPE A DUNN
0 o0 poundAU CLAIRE
~ TREMshyPEAshyLEAU
MICE
ONEIDA
Q LINCOLN
TAYLOR
Q t-~----- MOIRA THON
CLARK
o
10
LA FAVETTE
Hornwor ts amp Liverworts
- None - 1 - 25 taxa
- 26 - 50 tRXshy
- 51 - 75 tax)
- 7c - 100 taxa
FONDOU j LAC ~
DODGE
i O JEfFERSON
WALWORTH
30
Anastrophylltlfl helleranum (Nees) Schust Anastrophyllum michauxil (We~ Buch ~
46 88 136 141 Evans 38 46 141
Anastrophyllum mlnut~ (Schreb) Schuster Aneura plnguls (L) Durn
89 21 384687 88 89 90 (138) (145)
Anthoceros macounil MA Howe ~oceros punctatus L
21 (45) (144) 34
31
Barbilophozia attenuata (Mart) Loeske Barbilophozia barbata (Schmid ~ Schre~
24 141 Loeske 28 39 46 89 90 141
Bazzania trilobata (1) S Gray
1 26 39 46 (57) (59) 79 88 89
90 119 141
BlePbarostoma trichoffiyllum (1) Dumort
1 8 26 39 46 89 90 137 141
Blasia pusilla 1
1 39 46 90
Calypogeja fissa (1) Raddi
90
32
$129geja integristipula Steph C~Ypound2geja muelleriana (Schifm) K Mull
88 89 89
CallpoundO~eja neesiana (Mass amp Carest) K Mull
39 46 89 140 (141)
~pogeja sJlapound2icola (H Arnell tl J bull
Perss) Warnst et Loeske (119) 141
CeIalozia biscuspidata (L) Dum
1 37 I 46 90 t 137 141
CalYpogej~ trichomanis (L) Corda
26 39 (119) 137
33
CeEhalozia catenulata (Hub) Lindb Cephalozia connivens (Dicks) Lindb
39 46 79 141 39 4679 88 (89) 90 119 141
Cephalozia lacinulata Jack ~ Spruce
141 Cephalozia lunulifolia (Dum~ Dum I 28 39 46 79 88 89 90 141
Cepoundhalozia macouni (Aust) Aust Cephalozia ~leniceps (Aust) Lindb 27~ 3746 (66)7987 (139)141 39 46 79 89 141
34
Ceffialoziella 9ivaricata (Sm) Schiffn Cephaloziella ~hista (Jack) Schiffn
46 90 46 (119) 141
Cephaloziella hampeana (Nees) Schiffn
46
Cephaloziella rubella (Nees) Warnst
46 39
Cephaloziella rubella var sullivantii
(Aust) K Mull 46 39
Cephaloziella rubella var bifida (Lindb) Douin
35
46
Chiloscyphus pallescens (Ehrh ~ Hof~ Dum Chiloscyphus pallescens var fragilis
39J 46 88 89J 90 (Roth) K Milll 46
Chiloscyphus E9lyanthos (L) Corda
1 24 39 46
Chiloscyphus polyanthos vax rivularis
(Schrad) Nees 39 46
poundpoundnocepha1um conicum (1) Lindb
12 28 39 46 (49) 87 88 8990 120 121 148
Cololejeunea biddlecomiae (Aust) Evans
39 46 62 87 88 89 90
36
Dipl0]hyllum apicu1atum (Evans) Steph Fossombronia foveo1ata Lindb
(35)41 46 90 140 1 t 38 (39) t 46
Frullania bolanderi Aust
13 46 62 88 90 139 140
Frullanla ari ttoniae Evans
39 46 69 (71) 87 88 89 90
Frullania eboracensis Gott
1 9 28 4 39 46 (57) (59) 69 70 (71) 87 88 89 90 92 93 97 148
Frullania inflata Gott
46 87 88 89 (139)
Frullania oakesiana Aust Frullania plana Sullo
(35) 42 46 87 (139) (140) 92
Frullania ri paria Hampe ~ Lehm
46 69 71 87 89
Frullania ~elwynianna Pears
46 89
Geocalyx SEaveolens (Schrad) Nees
394688 8990 (119) 140
Frullania tamarisci (L) Dum subsp sPagrayana
(Mont) Hatt 28 34 l~6 t (57) 88 89 90
38
Mar pan thus seutatUB ( Web amp Mohr) Spruce Jamesoniella autumnalis (DC) Steph
(35) 39 46 90 1 7 28 39 46 56 (57) 87 8889 90 (119) 137 141
Jungermannia confertissima Nees
137 141
Jungermannia crenuliform1s Aust
79 137 141
Jungermannia ~il1ima Sm
46 90 141 Jungermannia exsert1folia subsp corifo~ (Dum~
Vana (35) 46 79 141
39
Jungermannia hyalina Lyell Jungerrr6nnia 1eiantha Grolle
40 46 79 136 141 39 46 79 88 90 1)7 141
Jungermannia IXlaxis Lindb Jungermannia pumila With
46 67 79 (139) (140) 141 )9 46 90 1)6 141
Jungermannia sphaerocarpa Hook Kurzia setacea (Web) Grolle
4679136140IIH 136
Lejeunea cavifo1ia (Ehrh) Lindb Lepidozia reptans (1) Dum
1 7 27 39 46 88 89 90 137 14010 I ( 27) 46 I 69 (71) 88
lophocolea heterophylla (Schrad) Dum Lophocolea minor Nees
1 )4 39 46 (57) 87 88 89 27 39 46 88 89 90 (119) 90 (93)94 119 136
Lophozia al~stris (Schleich ~ Web) Evans Lophozia ascendens (Warnst) Schust
46 90 137 141 136 140 141
41
loIilozia badensls (Gatt ~ Rabenh)
46 141
lophozia blcrenata (Schmid sectA Hoffm)
Dum 46 141
lophozia collaris (Nees) Dum
39 46 141
lophozla excisa (Dicks) Dum
27 4679 141
lophozia gillmani (Aust) Schust lophozia gIlttulata (Undo tl H Arnell)
36 39 40 46 136 (140) 141 Evans 39 46 136 141
42
141
Lophozia heterocolEDs (Thed) MA Howe IoPhozia ipcisa (Schard) Dum
27 39 46 89 141 8 27 31 46 79 90 140 141
LoQhozia longidens Macoun Lophozia ventricosa
1 46 79 90 141
Lophozia ventricosa var silvicola
(Buch) Jones 141 LoEhozla ventricosa vax longif1ora (Nees)
Macoun 39 46 79 141
43
~~ fragrans (Balbis) Frye et Clark Mannia pi 10sa (Hornem) Frye et Clark
46 (139) (140) 9 46 79 116 139
Mannia triandra (Scop) Gro 11e
46 (140)
Marchantia pound91ymorpha L
1 13 28 3946 (49) 78 87 88 89 90 (Ill) 121 122 147
Mo~rckia hibernica (HookJ Gott
46 89
Metzgeria conjugata Lindb
44
46
Mylia anomala (Hook) S Gray Notothylas orbicularis (Schwein) Sullo
39 46 141 (35)45 46
Nowellia curvifolia (Dicks) Mitt
17 27 3846 88 89 90 141
Odontoschisma denudatuJI1 (Nees ~ Mart)
Dum 46 81 137 141
Pallavicinia lyellii (Hook~ Carruth
90
Odontoschisma macounii (Aust) Under
45 46 (66) (139) 140 141
45
=
Pellia epiphllla (L) CordaPellia endiviifolia (Dicks~ Dum
27 39 46 90 13724 39 46
Pellia neesiana Limp bull Phaeoceros
39 46 89 27 39 45 46 97
Porella pinnata L
13 27 I 46 90 Plagiochila asplenioides (L) Dum
27 I 39 1+6 88 89 I 90 137
46
~e11a platYEhylla (L) Pfeiff Porella platyphylloi~ (Schwein) Lindb
17 27 39 46 69 (71)79 87 88 89 9 34 46 (57) 59 87 88 89 92 90 92 93 77
Preissia quadrata (Seop) Nees
1 36 3946 (71) 88 89 90 121 123
137
Ptllidium ciliare (L) Hampe
1 27 39 46 132 141
Dum
(22) 28 3946 (57) 87 88 89 90 92
Ptilidium ~lcherrium (G Web) Hampe
(9) 13 394656 (57)70 87 88 89 90 93 (119) 136 141
47
Radula obconica Sullo Reboulia hemlsphaerlca (L) Raddi
39 46 88 89 (43) 46 (59) 70 90 t Cf 124 125 (139)
Riccardia ~atifrons Lindb
1 27 39 46 89
Riccardia multifi~ (L) S Gray
39 46 88 145
Riccia beyrichiana Hampe ~ Lehm
46 Riccardia palmata (Hedw) Carruth
39 46 90
48
Riccia bifurca Hoffm Riccia cavernosa Hoffm
48 13739 46 79 137
Riccia fluitans L
242731454679
Riccla frostli Aust
48 79
Riccia sullicantii Aust
79 (137) Riccia sorocarpa Bisch 46 (137)
49
Ricciocarpus natans (1) Corda Sca~ia apiculata Spruce
13 27 46 79 89 110 39 46 (139) 141
ScaJltll1ia carinthica Jack ~ Lindb
137 t 141
Scapania ~ (Mart) Durn
39467990137141
Scapania digenil Schiffn
90
ScaEsectflia cuspiduligera (Nees) K Mull
46 (139) (140) (141)
50
Scapania glaucoeerha1a (Tayl) Aust Seapania gyrrmostomophilia Kaal
39 46 (139) 141 41 44 46 64 (139) (140) 141
ScaP3poundlia irrigua Gott ~ almiddot
7 39 46 79 137 (140L 141
Seapania nemorosa (L) Dum
1 39 46 89 90 137
Scapmi~ ~~ta Smiddotlch
137 141 51
~caPnia poundludicola LDeske amp K Mull Scapania saxicola Schust
46 (66) 141 136 141
Sc~~ scandica (H Arnell ~ Buch) Macv
139 140
Scapania subalpina (Nees) Durn
39 40 46 79 137 141
Scapania undulata (L) Dum
1 42 46 79 141
Scapania umbrosa (Schrad) Dum
137 140 141
52
Trichocolea tomentella (Ehrh) Dum Triomarla exsecta (Schmid) Schiffn
27 (35) 39 46 47 88 137 141 8 28 39 46 90 140 141
Tritomaria exsectiformis (Breidl) Loe~ke Tritomaria Sluinguedentata (Huds) Buch
39 -t6 89 90 136 140 141 7 39 46 141
Trltomaria scitula (Tayl) Joerg
46 136 (139) 141
53
Orothera C Muell leana (Sull) C Muell
Rfwbdoweisia crispata (With) Undb
Cynodontium BSG ex Schimp alpestre (Wahlenb) Milde strumiferum (Hedw) Undb
Dichodontium Schimp pellucidum (Hedw) Schimp
Oncophorus (Slid) Brid vuens (Hedw) Brid wahlenbergii (Brid)
Dicranum Hedw bonjeanii DeNot ex Usa drummondii C Muell lagellare Hedw varj1ogellare fulvum Hook fuscescens Turn var fuscescens montanum Hedw muehlenbeckii BSG var muehlenbeckii polysetum Sw sabuletorum Ren et Card scoparium Hedw undulatum Brid viride (Sull et Lesq ex Sull) lindb
36 Leucobryaceae Leucobryum Hampe
glaucum (Hedw) Angstr ex Fr
37 Encalyptaceae Encalypta Hedw
ciliata Hedw var ciliata procera Bruch
38 Pottiaceae Astomum Hampe
muhlenbergianum (Sw) Grout
Weissia Hedw controversa Hedw var controversa
Gymnostomum Nees Homsch et Stunn aeruginosum Sm recurvirostrum Hedw var recurvirostrum
Trichostomum Bruch spirale Prout
19
40 Ephemeraceae Ephemerum Hampe
cohaerens (Hedw) Hampe crassineium (Schwaegr) Hampe val crassinervium spinulosum Bruch et Schimp ex Schimp
41 Funariaceae AphflllolThegma SuD
serratum (1 Hook et Wils ex Drumm) Sullo
Physcomitrium (Brid) Fuernr hookeri Hampe immersum Sullo pyriforme (Hedw) Hampe
Funaria Hedw americana Undb hygrometrica Hedw vaT hygrometrica
42 Splachnaceae Splachnum Hedw
ampullaceum Hedw
43 Schistostegaceae Schistostega Mohr
pennata (Hedw) Web el Mohr
44 Bryaceae Pohlia Hedw
annolina (Hedw) Lindb var annotina camea (Tum) Lindb cruda (Hedw) Undb elongata Hedw iliformis (Dicks) Andr lescurlana (Sull) Grout nutans (Hedw) Lindb obrusifolia (Brid) L Koch proligera (Undb ex Umpr) Undb ex Amell wahlenbergii (Web et Mohr) Andr
Leptobryum (B S G) Wits pyriforme (Hedw) Wils
Bryum Hedw algovicum Sendtnex C Muell argenteum Hedw caespiticium Hedw capillare Hedw var capillare creberrlmum Tayt lonchocaulon C Muell pallens (Brid) Sw ex Roehl pseudotrlquetrum (Hedw) Gaertn Meyer et Scherb varpudotriquetrum
20
pseudotriquetrum (Hedw) GaertnMeyer et Scherb var bimum (Schreb) Ulj stenotrichum C Muell var stenotrichum tortiolium Funck ex Brid uliginosum (Brid) B S G violaceum Crundw et Nyh weigelii Spreng
Rhodobryum (Schimp) Hampe roseum (Hedw) Limpr
45 Mniaceae Mnium Hedw
ciliare (C Muell) Schimp cinclidioides Hueb cuspidarum Hedw drummondii Bruch et Schimp longirostrum Brid lycopodioides Schwaegr marginatum (With) Brid ex P Beauv mediumB~S G var medium onhorrhynchum Brid pseudopunctatum Bruch et Schimp punctalUm Hedw var punctatum punctatum Hedw var elatum Schirnp rugicum Laur spinufosum B S G stellare Hedw
46 Aulacomruaceae Au1acomnium Schwaegr
androgynum (Hedw) Schwaegr heterostichum (Hedw) B S G paJustre (Hedw) Schwaegr var paJustre
47 Meesiaceae Amblyodon B S G
deillbotus (Hedw) B S G
Meesia Hedw triquetra (Richt) Angstr uJiginosa Hedw
48 Bartramiaceae Plagiopus Brid
oederiana (Sw) Umpr
Bartramia Hedw pomiformis Hedw var pomiformis
Philonotis Brid fontana (Hedw) Brid varfontana longseta (Michx) Britt varlongseta marchica (Hedw) Brid
21
~
~
49 Timmiaceae Timmia Hedw
megapolitana Hedw var megapolitana
50 Orthotrichaceae Zygodon Hook et Tayl
viridissimus (Dicks) Brid var rupestris lindb ex C J Hartm
Amphidium Schimp lapponicum (Hedw) Schimp mougeotii (B S G) Schimp
Orthotrichum Hedw anomalum Hedw obtuaifolium Brid ohioense Sullo et Lesq ex Aust pumilumSw sordidum Sullo et Lesq ex Aust speciosum Neesax Sturm var eegam (Schwaegr ex Hook et Grev) W~rnst stellDtwn Brid strrmgulatum P Beauv
Uleta Mohr ex Web cOfIICtata (p Beallv) Hamrn crispa (Hedw) Brid
Drummondill Hook ex Drumm prorepens (Hedw) Brltt
51 Fontinalaceae Fantinalis Hedw
antipyretica Hedw var antipyrettca antipyretica Hedw forma rabusta (Card) Card dalecarlica Schimp ex B S G var dalecarlica flqccidtJ Ren et Card hypnoides C J Hartrn var hypnoides hypnoides C J Hartm var durillei (Schimp) HU$ll
missourlcaCard novae-angliae Sullo var novae-angliae splulgrUlolill (C Moen) Wijket M~S
DichelY1l1ll Myr capilloceum (With) Myr falcatum (Bedw) Myr HIllescens B S G
52 Climaciaceae aimecium Web et Mohr ex Mohr
americllnum Brid dedroides (Hedw) Web et Mohr
22
S3 Hedwigiaceae Hedwigia P Beauv
ciliata (Hedw) P Beauv
54 Cryphaeaceae Forsstroemia lindh
ohioense (Sull) lindh
55 Leucodontaceae Leucodon Schwaegr
brachypus Blid var brachypus brachypus Brid var aniJrewsitlnus Crum et Anderson julaceus (Hedw) SuU
56 Neckeraceae Neckera Hedw
pennata Hedw var pennata
1fomalia (Brid) B S G trichomanoides (Hedw) B S G
57 Theliaceae Thelia Sull
asprellll Sullo hirtella (Hedw) Sullo lescurii SuU
Myurella B SG julacea (Schwaegr) B S G
sibrica (C MueU) Reim var sibrica
58 Fabroniaceae Fabronio Raddi
ciliaris (Brid) Bnd
Anacamptodon Brid splachnoides (Groet ex Brid) Brid
Schwetschkeopsis Broth fabronia (Schwaegr) Broth
59 Leskeaceae Lindbergia Kindb
brachyptera (Mitt) Kindb
Leskea Hedw gracilescens Hedw obscura Hedw poiycarpa Hedw
Leskeella (limpr) Loeske nervosa (Brid) Loeske
23
Pterlgy1lllndrum Hedw filiforme Hedw
60 Thuidiaceae Haplohymenfum Dozy et MoIk
trlste (Ces ex De Not) Kiodb
Anomodon Hook et Tayl attenuatus (Hedw) Hueb minor (Hedw) Fuemr rostratus (Hedw) Schimp viticulosus (Hedw) Hook et Tayl
Haplocladium (C Mucn) C MueU microphyllum (Hedw) Broth virginillnum (Brld) Broth ~bull
Thuidium B S G abietinum (Hedw) B S G delicatulum (Hedw) B S G vardelicatulum delicatulum (Hedw) B S G var radicans (Kindb) Crum Steere et Anderson minutulum (Hedw) B S G pygmaeum B S G recognftum (Hedw) Lindb scitum (p Beauv) Aust
Helodium (Sull) Warnst blandowli (Web et Mohr) Warnst var blandowli blandowli (Web et Mohr) Wamst var helodioides (Ren et Card ex Roell) Crum
Steere et Anderson paludomm (Sull) AuS
61 Amblystegiaceae Cratoneuron (Sull) Spruce
commulatum (Hedw) Roth var commutfltum commutatum (Hedw) Roth var fakatum (Brid) Moenk ilicinum (Hedw) Spruce var filcinum
Campylium (Sun) Mitt chrysophyllum (Brid) 1 Lange var chrysophyllum hispidulum (Brid) Mitt var hllJidulum polygamum (B S G)C lens varpolygtlmUm stellatum (Hedw) C1ens vat stelltum
Leptodictyum (Schimp) Warnst laxirete (Card et Ther) Broth riparium (Hedw) Wamat trichopodium (Schultz) Warns l1acillans (Sull) Broth
Hygroamblystegium Loelke jlul1iatile (Hedw) oeske var luViatile
24
fluviatile (Hedw) Loeske var oWltum Grout noterophilum (Sun et Lesq ex Sull) Warnst tenox (Hedw) Jenn var tenax tenox (Hedw) Jenn var spinilolium Schimp) Jenn
Amblystegium BSG juratzkanum Schimp serpens (Hedw) BSG varlum (Hedw) Undb var varium
Platydictya Berk confervoides (Brid) Crum jungermanniodes (Brid) Crum minutissimum (Sull et Lesq ex Sull) Crum subtile (Hedw) Crum
Drepanocladus (C Muell) Roth aduncus (Hedw) Warnst var aduncus aduncus (Hedw) Warnst var capillilolius (Wamst)Riebm aduncus (Hedw) Warnst var kneiffii (BSG) Moenk aduncus (Hedw) Warnst var polycorpus (Bland ex Voit) Roth exannulatus (BSG) Warnst fluitans (Hedw) Warnst var fluitans revolvens (Sw) Wamst var revolvens sendtneri (Schimp) Warnst var sendtnerl uncinatus (Hedw) Warnst var uncinatus uncinatus ffiedw) Warnst var piwnosus (Schimp) Wamst vernicoms (Lindb ex C Hartm) Warnst
Hygrohypnum Lindb eugyrium (BSG) loeske luridum (Hedw Jenn varluridum ochraecum (Tum ex Wils) Loeske var ochraceunt
Calliergon (Sull) Kindb cordilolium Hedw Kindb glganteum (Schimp) Kindb var giganteum richardsonii (Mitt) Kindb ex Wamst stramineum (Brid) Kindb trilarium (Web et Mohr) Kindb
Calliergonella Loeske cuspidata (Hedw) Loeske
Calliergidium (Ren) Grout bakeri (Ren) Grout
Scorpidium (Schimp) Urnpr scorpiodides (Hedw) limpr
62 Brachytbeciaceae Tomenthypnum loeske
nitens (Hedw) Loeske var mtens
25
Homalotheciella (Card) Broth subcapillata (Hedw) Broth
Brachythecium BSG acuminatum (Hedw) Awt acutum (Mitt) Sullo calcareum IGndb campestTe (C MuelL) BSG cunum (Lindb) Umpr digastrum C Muell et Kindb ex Macoun et Kindb oxycladon (Brid) Jaeg et Sauerb var oxycladon plumosum (Hedw) BSG populeum (Hedw) BSG var populeum reflexum (Starke ex Web et Mohr) DsG var relexum reiexum (Starke ex Web et Mohr) DSG var pacificum Renet Card ex Roell rlvulare BSG var rivulare tabulum (Hedw) BSC var tabuum salebrosum (Web et Mohr) BSG var salebrosum velutinum (Hedw) BSG Vat velutinum
Bryhniil Kaur graminicolor (Brid) Grout var graminicolor novae-angliae (Sull et Lesq ex sun) Grout
Rhynchostegium BSG senulatum (Hedw) Ja4lg et Sauerb
Rhynchostegiella (BSG) Limpr compacta (C MueU) Loeske
EU1hynchium 8SG hians (Hedw) Sande Lac pulchellum (Hedw) Jenn var pulchelium riparloides (Hedw) Rich
Stokeriella (Kindb) Robins pmelonga (Hedw) Robins var stokesii (Tum) Crum
63 Entodontaceae Entodon C Muell
brevisetus (Hook et Wils ex Wils) Lindb cladorrhizans (Hedw) C Muell seductrlx (Hedw) C MueU var wluctrix
Pleurozium Mitt schreberl (8rid) Mitt var schreberi
64 Plagiotheciaceae Pltlglothecium BSG
CflVifolium (Brid) Iwats var CtWifolium denticulatum (Hedw) 8SG laetum BSG ltztebricola BSG
26
6S SernatophyUaceae Brotherella Loeske ex Fleisch
recumins (Michx) Fleisch tenuirostris (Bruch et Schimp ex Sull) Fleisch
Sematophyllum Mitt demissum (Wils) Mitt
66 Hypnaceae Pl(itygyrium BSG
repens (Brid) BSG
PylaisieUa Kindb intricata (Hedw) Grout polyantha (Hedw) Grout selwynii (Kindb) Crum Steere et Anderson
Homomallium (Schimp) Loeske adnatum (Hedw) Broth
Hypnum Hedw cupressiforme Hedw var cupressiforme cupressifonne Hedw varfiliforme Brid curviolium Hedw fertile Sendtn imponens Hedw lindbergii Mitt palescens (Hedw) P-Beauv pratense Koch ex Brid
Calicladium Crum haldanianum (Grev) Crum
sopterygiopsis Iwats muelleriana (Schimp) Iwats
Isopterygium Mitt e1egans (Brid) lindb pulchellum (Hedw) Jaeg et Sauerb tenerum (Sw) Mitt
Taxiphyllum Fleisch depllQtum (Bruch et Schimp ex sun) Fleisch taxirameum (MitL) Fleisch
Herzogiella Broth seligeri (Brid) Iwats striatella (Brid) Iwats ruracea (Lindh) Iwats
hilium De Not crista-castrenna (Hedw) De Not
27
67 Rhytidiaceae Rhytidium (Sull) Kindb
rugosum (Hedw) Kindb
Rhytidfodelphus (Lindb ex Iimpr) Warnst triquetrus (Hedw) Warnst var triquetrus
68 Hylocomiaceae Hylocomium BSG
pyrenaicum (Spruce) lindb splendens (Hedw) BSG varsplendens
69 Buxbaumiaceae Buxbaumia Hedw
aphyllll Hedw
Diphyscium Mohr follosum (Hedw) Mohr
70 Tetraphidaceae Tetraphi3
pellucida Hedw
71 Polytrichaceae Atrichum PmiddotBeauv
angustatum (Brid) BSG cri8pum (James) Sullo oerstedianum (C Muell) Mitt teneUum (Roehl) BSG undulatum (Hedw) P middotBeauv var unduiatum
Pogonatum P-Beauv alpinum (Hedw) Roehl var alpinum umigerum (Hedw) P-Beauv
Poiytrichum Hedw commune Hedw var commune ormosum Hedw juniperinum Hedw longisetum Brid ohioense Ren et Card piliferum Hedw var pilierum strictum Brid
28
MAPS OF BRYOPHYTE TAXA
REPORTED FOR WISCONSIN
HORNWORTS AND LIVERWORTS
29
bull bull WSHeV~N S vE II
0 ~
e~RRON ~USK
(J~ (j -
CHIPPE A DUNN
0 o0 poundAU CLAIRE
~ TREMshyPEAshyLEAU
MICE
ONEIDA
Q LINCOLN
TAYLOR
Q t-~----- MOIRA THON
CLARK
o
10
LA FAVETTE
Hornwor ts amp Liverworts
- None - 1 - 25 taxa
- 26 - 50 tRXshy
- 51 - 75 tax)
- 7c - 100 taxa
FONDOU j LAC ~
DODGE
i O JEfFERSON
WALWORTH
30
Anastrophylltlfl helleranum (Nees) Schust Anastrophyllum michauxil (We~ Buch ~
46 88 136 141 Evans 38 46 141
Anastrophyllum mlnut~ (Schreb) Schuster Aneura plnguls (L) Durn
89 21 384687 88 89 90 (138) (145)
Anthoceros macounil MA Howe ~oceros punctatus L
21 (45) (144) 34
31
Barbilophozia attenuata (Mart) Loeske Barbilophozia barbata (Schmid ~ Schre~
24 141 Loeske 28 39 46 89 90 141
Bazzania trilobata (1) S Gray
1 26 39 46 (57) (59) 79 88 89
90 119 141
BlePbarostoma trichoffiyllum (1) Dumort
1 8 26 39 46 89 90 137 141
Blasia pusilla 1
1 39 46 90
Calypogeja fissa (1) Raddi
90
32
$129geja integristipula Steph C~Ypound2geja muelleriana (Schifm) K Mull
88 89 89
CallpoundO~eja neesiana (Mass amp Carest) K Mull
39 46 89 140 (141)
~pogeja sJlapound2icola (H Arnell tl J bull
Perss) Warnst et Loeske (119) 141
CeIalozia biscuspidata (L) Dum
1 37 I 46 90 t 137 141
CalYpogej~ trichomanis (L) Corda
26 39 (119) 137
33
CeEhalozia catenulata (Hub) Lindb Cephalozia connivens (Dicks) Lindb
39 46 79 141 39 4679 88 (89) 90 119 141
Cephalozia lacinulata Jack ~ Spruce
141 Cephalozia lunulifolia (Dum~ Dum I 28 39 46 79 88 89 90 141
Cepoundhalozia macouni (Aust) Aust Cephalozia ~leniceps (Aust) Lindb 27~ 3746 (66)7987 (139)141 39 46 79 89 141
34
Ceffialoziella 9ivaricata (Sm) Schiffn Cephaloziella ~hista (Jack) Schiffn
46 90 46 (119) 141
Cephaloziella hampeana (Nees) Schiffn
46
Cephaloziella rubella (Nees) Warnst
46 39
Cephaloziella rubella var sullivantii
(Aust) K Mull 46 39
Cephaloziella rubella var bifida (Lindb) Douin
35
46
Chiloscyphus pallescens (Ehrh ~ Hof~ Dum Chiloscyphus pallescens var fragilis
39J 46 88 89J 90 (Roth) K Milll 46
Chiloscyphus E9lyanthos (L) Corda
1 24 39 46
Chiloscyphus polyanthos vax rivularis
(Schrad) Nees 39 46
poundpoundnocepha1um conicum (1) Lindb
12 28 39 46 (49) 87 88 8990 120 121 148
Cololejeunea biddlecomiae (Aust) Evans
39 46 62 87 88 89 90
36
Dipl0]hyllum apicu1atum (Evans) Steph Fossombronia foveo1ata Lindb
(35)41 46 90 140 1 t 38 (39) t 46
Frullania bolanderi Aust
13 46 62 88 90 139 140
Frullanla ari ttoniae Evans
39 46 69 (71) 87 88 89 90
Frullania eboracensis Gott
1 9 28 4 39 46 (57) (59) 69 70 (71) 87 88 89 90 92 93 97 148
Frullania inflata Gott
46 87 88 89 (139)
Frullania oakesiana Aust Frullania plana Sullo
(35) 42 46 87 (139) (140) 92
Frullania ri paria Hampe ~ Lehm
46 69 71 87 89
Frullania ~elwynianna Pears
46 89
Geocalyx SEaveolens (Schrad) Nees
394688 8990 (119) 140
Frullania tamarisci (L) Dum subsp sPagrayana
(Mont) Hatt 28 34 l~6 t (57) 88 89 90
38
Mar pan thus seutatUB ( Web amp Mohr) Spruce Jamesoniella autumnalis (DC) Steph
(35) 39 46 90 1 7 28 39 46 56 (57) 87 8889 90 (119) 137 141
Jungermannia confertissima Nees
137 141
Jungermannia crenuliform1s Aust
79 137 141
Jungermannia ~il1ima Sm
46 90 141 Jungermannia exsert1folia subsp corifo~ (Dum~
Vana (35) 46 79 141
39
Jungermannia hyalina Lyell Jungerrr6nnia 1eiantha Grolle
40 46 79 136 141 39 46 79 88 90 1)7 141
Jungermannia IXlaxis Lindb Jungermannia pumila With
46 67 79 (139) (140) 141 )9 46 90 1)6 141
Jungermannia sphaerocarpa Hook Kurzia setacea (Web) Grolle
4679136140IIH 136
Lejeunea cavifo1ia (Ehrh) Lindb Lepidozia reptans (1) Dum
1 7 27 39 46 88 89 90 137 14010 I ( 27) 46 I 69 (71) 88
lophocolea heterophylla (Schrad) Dum Lophocolea minor Nees
1 )4 39 46 (57) 87 88 89 27 39 46 88 89 90 (119) 90 (93)94 119 136
Lophozia al~stris (Schleich ~ Web) Evans Lophozia ascendens (Warnst) Schust
46 90 137 141 136 140 141
41
loIilozia badensls (Gatt ~ Rabenh)
46 141
lophozia blcrenata (Schmid sectA Hoffm)
Dum 46 141
lophozia collaris (Nees) Dum
39 46 141
lophozla excisa (Dicks) Dum
27 4679 141
lophozia gillmani (Aust) Schust lophozia gIlttulata (Undo tl H Arnell)
36 39 40 46 136 (140) 141 Evans 39 46 136 141
42
141
Lophozia heterocolEDs (Thed) MA Howe IoPhozia ipcisa (Schard) Dum
27 39 46 89 141 8 27 31 46 79 90 140 141
LoQhozia longidens Macoun Lophozia ventricosa
1 46 79 90 141
Lophozia ventricosa var silvicola
(Buch) Jones 141 LoEhozla ventricosa vax longif1ora (Nees)
Macoun 39 46 79 141
43
~~ fragrans (Balbis) Frye et Clark Mannia pi 10sa (Hornem) Frye et Clark
46 (139) (140) 9 46 79 116 139
Mannia triandra (Scop) Gro 11e
46 (140)
Marchantia pound91ymorpha L
1 13 28 3946 (49) 78 87 88 89 90 (Ill) 121 122 147
Mo~rckia hibernica (HookJ Gott
46 89
Metzgeria conjugata Lindb
44
46
Mylia anomala (Hook) S Gray Notothylas orbicularis (Schwein) Sullo
39 46 141 (35)45 46
Nowellia curvifolia (Dicks) Mitt
17 27 3846 88 89 90 141
Odontoschisma denudatuJI1 (Nees ~ Mart)
Dum 46 81 137 141
Pallavicinia lyellii (Hook~ Carruth
90
Odontoschisma macounii (Aust) Under
45 46 (66) (139) 140 141
45
=
Pellia epiphllla (L) CordaPellia endiviifolia (Dicks~ Dum
27 39 46 90 13724 39 46
Pellia neesiana Limp bull Phaeoceros
39 46 89 27 39 45 46 97
Porella pinnata L
13 27 I 46 90 Plagiochila asplenioides (L) Dum
27 I 39 1+6 88 89 I 90 137
46
~e11a platYEhylla (L) Pfeiff Porella platyphylloi~ (Schwein) Lindb
17 27 39 46 69 (71)79 87 88 89 9 34 46 (57) 59 87 88 89 92 90 92 93 77
Preissia quadrata (Seop) Nees
1 36 3946 (71) 88 89 90 121 123
137
Ptllidium ciliare (L) Hampe
1 27 39 46 132 141
Dum
(22) 28 3946 (57) 87 88 89 90 92
Ptilidium ~lcherrium (G Web) Hampe
(9) 13 394656 (57)70 87 88 89 90 93 (119) 136 141
47
Radula obconica Sullo Reboulia hemlsphaerlca (L) Raddi
39 46 88 89 (43) 46 (59) 70 90 t Cf 124 125 (139)
Riccardia ~atifrons Lindb
1 27 39 46 89
Riccardia multifi~ (L) S Gray
39 46 88 145
Riccia beyrichiana Hampe ~ Lehm
46 Riccardia palmata (Hedw) Carruth
39 46 90
48
Riccia bifurca Hoffm Riccia cavernosa Hoffm
48 13739 46 79 137
Riccia fluitans L
242731454679
Riccla frostli Aust
48 79
Riccia sullicantii Aust
79 (137) Riccia sorocarpa Bisch 46 (137)
49
Ricciocarpus natans (1) Corda Sca~ia apiculata Spruce
13 27 46 79 89 110 39 46 (139) 141
ScaJltll1ia carinthica Jack ~ Lindb
137 t 141
Scapania ~ (Mart) Durn
39467990137141
Scapania digenil Schiffn
90
ScaEsectflia cuspiduligera (Nees) K Mull
46 (139) (140) (141)
50
Scapania glaucoeerha1a (Tayl) Aust Seapania gyrrmostomophilia Kaal
39 46 (139) 141 41 44 46 64 (139) (140) 141
ScaP3poundlia irrigua Gott ~ almiddot
7 39 46 79 137 (140L 141
Seapania nemorosa (L) Dum
1 39 46 89 90 137
Scapmi~ ~~ta Smiddotlch
137 141 51
~caPnia poundludicola LDeske amp K Mull Scapania saxicola Schust
46 (66) 141 136 141
Sc~~ scandica (H Arnell ~ Buch) Macv
139 140
Scapania subalpina (Nees) Durn
39 40 46 79 137 141
Scapania undulata (L) Dum
1 42 46 79 141
Scapania umbrosa (Schrad) Dum
137 140 141
52
Trichocolea tomentella (Ehrh) Dum Triomarla exsecta (Schmid) Schiffn
27 (35) 39 46 47 88 137 141 8 28 39 46 90 140 141
Tritomaria exsectiformis (Breidl) Loe~ke Tritomaria Sluinguedentata (Huds) Buch
39 -t6 89 90 136 140 141 7 39 46 141
Trltomaria scitula (Tayl) Joerg
46 136 (139) 141
53
40 Ephemeraceae Ephemerum Hampe
cohaerens (Hedw) Hampe crassineium (Schwaegr) Hampe val crassinervium spinulosum Bruch et Schimp ex Schimp
41 Funariaceae AphflllolThegma SuD
serratum (1 Hook et Wils ex Drumm) Sullo
Physcomitrium (Brid) Fuernr hookeri Hampe immersum Sullo pyriforme (Hedw) Hampe
Funaria Hedw americana Undb hygrometrica Hedw vaT hygrometrica
42 Splachnaceae Splachnum Hedw
ampullaceum Hedw
43 Schistostegaceae Schistostega Mohr
pennata (Hedw) Web el Mohr
44 Bryaceae Pohlia Hedw
annolina (Hedw) Lindb var annotina camea (Tum) Lindb cruda (Hedw) Undb elongata Hedw iliformis (Dicks) Andr lescurlana (Sull) Grout nutans (Hedw) Lindb obrusifolia (Brid) L Koch proligera (Undb ex Umpr) Undb ex Amell wahlenbergii (Web et Mohr) Andr
Leptobryum (B S G) Wits pyriforme (Hedw) Wils
Bryum Hedw algovicum Sendtnex C Muell argenteum Hedw caespiticium Hedw capillare Hedw var capillare creberrlmum Tayt lonchocaulon C Muell pallens (Brid) Sw ex Roehl pseudotrlquetrum (Hedw) Gaertn Meyer et Scherb varpudotriquetrum
20
pseudotriquetrum (Hedw) GaertnMeyer et Scherb var bimum (Schreb) Ulj stenotrichum C Muell var stenotrichum tortiolium Funck ex Brid uliginosum (Brid) B S G violaceum Crundw et Nyh weigelii Spreng
Rhodobryum (Schimp) Hampe roseum (Hedw) Limpr
45 Mniaceae Mnium Hedw
ciliare (C Muell) Schimp cinclidioides Hueb cuspidarum Hedw drummondii Bruch et Schimp longirostrum Brid lycopodioides Schwaegr marginatum (With) Brid ex P Beauv mediumB~S G var medium onhorrhynchum Brid pseudopunctatum Bruch et Schimp punctalUm Hedw var punctatum punctatum Hedw var elatum Schirnp rugicum Laur spinufosum B S G stellare Hedw
46 Aulacomruaceae Au1acomnium Schwaegr
androgynum (Hedw) Schwaegr heterostichum (Hedw) B S G paJustre (Hedw) Schwaegr var paJustre
47 Meesiaceae Amblyodon B S G
deillbotus (Hedw) B S G
Meesia Hedw triquetra (Richt) Angstr uJiginosa Hedw
48 Bartramiaceae Plagiopus Brid
oederiana (Sw) Umpr
Bartramia Hedw pomiformis Hedw var pomiformis
Philonotis Brid fontana (Hedw) Brid varfontana longseta (Michx) Britt varlongseta marchica (Hedw) Brid
21
~
~
49 Timmiaceae Timmia Hedw
megapolitana Hedw var megapolitana
50 Orthotrichaceae Zygodon Hook et Tayl
viridissimus (Dicks) Brid var rupestris lindb ex C J Hartm
Amphidium Schimp lapponicum (Hedw) Schimp mougeotii (B S G) Schimp
Orthotrichum Hedw anomalum Hedw obtuaifolium Brid ohioense Sullo et Lesq ex Aust pumilumSw sordidum Sullo et Lesq ex Aust speciosum Neesax Sturm var eegam (Schwaegr ex Hook et Grev) W~rnst stellDtwn Brid strrmgulatum P Beauv
Uleta Mohr ex Web cOfIICtata (p Beallv) Hamrn crispa (Hedw) Brid
Drummondill Hook ex Drumm prorepens (Hedw) Brltt
51 Fontinalaceae Fantinalis Hedw
antipyretica Hedw var antipyrettca antipyretica Hedw forma rabusta (Card) Card dalecarlica Schimp ex B S G var dalecarlica flqccidtJ Ren et Card hypnoides C J Hartrn var hypnoides hypnoides C J Hartm var durillei (Schimp) HU$ll
missourlcaCard novae-angliae Sullo var novae-angliae splulgrUlolill (C Moen) Wijket M~S
DichelY1l1ll Myr capilloceum (With) Myr falcatum (Bedw) Myr HIllescens B S G
52 Climaciaceae aimecium Web et Mohr ex Mohr
americllnum Brid dedroides (Hedw) Web et Mohr
22
S3 Hedwigiaceae Hedwigia P Beauv
ciliata (Hedw) P Beauv
54 Cryphaeaceae Forsstroemia lindh
ohioense (Sull) lindh
55 Leucodontaceae Leucodon Schwaegr
brachypus Blid var brachypus brachypus Brid var aniJrewsitlnus Crum et Anderson julaceus (Hedw) SuU
56 Neckeraceae Neckera Hedw
pennata Hedw var pennata
1fomalia (Brid) B S G trichomanoides (Hedw) B S G
57 Theliaceae Thelia Sull
asprellll Sullo hirtella (Hedw) Sullo lescurii SuU
Myurella B SG julacea (Schwaegr) B S G
sibrica (C MueU) Reim var sibrica
58 Fabroniaceae Fabronio Raddi
ciliaris (Brid) Bnd
Anacamptodon Brid splachnoides (Groet ex Brid) Brid
Schwetschkeopsis Broth fabronia (Schwaegr) Broth
59 Leskeaceae Lindbergia Kindb
brachyptera (Mitt) Kindb
Leskea Hedw gracilescens Hedw obscura Hedw poiycarpa Hedw
Leskeella (limpr) Loeske nervosa (Brid) Loeske
23
Pterlgy1lllndrum Hedw filiforme Hedw
60 Thuidiaceae Haplohymenfum Dozy et MoIk
trlste (Ces ex De Not) Kiodb
Anomodon Hook et Tayl attenuatus (Hedw) Hueb minor (Hedw) Fuemr rostratus (Hedw) Schimp viticulosus (Hedw) Hook et Tayl
Haplocladium (C Mucn) C MueU microphyllum (Hedw) Broth virginillnum (Brld) Broth ~bull
Thuidium B S G abietinum (Hedw) B S G delicatulum (Hedw) B S G vardelicatulum delicatulum (Hedw) B S G var radicans (Kindb) Crum Steere et Anderson minutulum (Hedw) B S G pygmaeum B S G recognftum (Hedw) Lindb scitum (p Beauv) Aust
Helodium (Sull) Warnst blandowli (Web et Mohr) Warnst var blandowli blandowli (Web et Mohr) Wamst var helodioides (Ren et Card ex Roell) Crum
Steere et Anderson paludomm (Sull) AuS
61 Amblystegiaceae Cratoneuron (Sull) Spruce
commulatum (Hedw) Roth var commutfltum commutatum (Hedw) Roth var fakatum (Brid) Moenk ilicinum (Hedw) Spruce var filcinum
Campylium (Sun) Mitt chrysophyllum (Brid) 1 Lange var chrysophyllum hispidulum (Brid) Mitt var hllJidulum polygamum (B S G)C lens varpolygtlmUm stellatum (Hedw) C1ens vat stelltum
Leptodictyum (Schimp) Warnst laxirete (Card et Ther) Broth riparium (Hedw) Wamat trichopodium (Schultz) Warns l1acillans (Sull) Broth
Hygroamblystegium Loelke jlul1iatile (Hedw) oeske var luViatile
24
fluviatile (Hedw) Loeske var oWltum Grout noterophilum (Sun et Lesq ex Sull) Warnst tenox (Hedw) Jenn var tenax tenox (Hedw) Jenn var spinilolium Schimp) Jenn
Amblystegium BSG juratzkanum Schimp serpens (Hedw) BSG varlum (Hedw) Undb var varium
Platydictya Berk confervoides (Brid) Crum jungermanniodes (Brid) Crum minutissimum (Sull et Lesq ex Sull) Crum subtile (Hedw) Crum
Drepanocladus (C Muell) Roth aduncus (Hedw) Warnst var aduncus aduncus (Hedw) Warnst var capillilolius (Wamst)Riebm aduncus (Hedw) Warnst var kneiffii (BSG) Moenk aduncus (Hedw) Warnst var polycorpus (Bland ex Voit) Roth exannulatus (BSG) Warnst fluitans (Hedw) Warnst var fluitans revolvens (Sw) Wamst var revolvens sendtneri (Schimp) Warnst var sendtnerl uncinatus (Hedw) Warnst var uncinatus uncinatus ffiedw) Warnst var piwnosus (Schimp) Wamst vernicoms (Lindb ex C Hartm) Warnst
Hygrohypnum Lindb eugyrium (BSG) loeske luridum (Hedw Jenn varluridum ochraecum (Tum ex Wils) Loeske var ochraceunt
Calliergon (Sull) Kindb cordilolium Hedw Kindb glganteum (Schimp) Kindb var giganteum richardsonii (Mitt) Kindb ex Wamst stramineum (Brid) Kindb trilarium (Web et Mohr) Kindb
Calliergonella Loeske cuspidata (Hedw) Loeske
Calliergidium (Ren) Grout bakeri (Ren) Grout
Scorpidium (Schimp) Urnpr scorpiodides (Hedw) limpr
62 Brachytbeciaceae Tomenthypnum loeske
nitens (Hedw) Loeske var mtens
25
Homalotheciella (Card) Broth subcapillata (Hedw) Broth
Brachythecium BSG acuminatum (Hedw) Awt acutum (Mitt) Sullo calcareum IGndb campestTe (C MuelL) BSG cunum (Lindb) Umpr digastrum C Muell et Kindb ex Macoun et Kindb oxycladon (Brid) Jaeg et Sauerb var oxycladon plumosum (Hedw) BSG populeum (Hedw) BSG var populeum reflexum (Starke ex Web et Mohr) DsG var relexum reiexum (Starke ex Web et Mohr) DSG var pacificum Renet Card ex Roell rlvulare BSG var rivulare tabulum (Hedw) BSC var tabuum salebrosum (Web et Mohr) BSG var salebrosum velutinum (Hedw) BSG Vat velutinum
Bryhniil Kaur graminicolor (Brid) Grout var graminicolor novae-angliae (Sull et Lesq ex sun) Grout
Rhynchostegium BSG senulatum (Hedw) Ja4lg et Sauerb
Rhynchostegiella (BSG) Limpr compacta (C MueU) Loeske
EU1hynchium 8SG hians (Hedw) Sande Lac pulchellum (Hedw) Jenn var pulchelium riparloides (Hedw) Rich
Stokeriella (Kindb) Robins pmelonga (Hedw) Robins var stokesii (Tum) Crum
63 Entodontaceae Entodon C Muell
brevisetus (Hook et Wils ex Wils) Lindb cladorrhizans (Hedw) C Muell seductrlx (Hedw) C MueU var wluctrix
Pleurozium Mitt schreberl (8rid) Mitt var schreberi
64 Plagiotheciaceae Pltlglothecium BSG
CflVifolium (Brid) Iwats var CtWifolium denticulatum (Hedw) 8SG laetum BSG ltztebricola BSG
26
6S SernatophyUaceae Brotherella Loeske ex Fleisch
recumins (Michx) Fleisch tenuirostris (Bruch et Schimp ex Sull) Fleisch
Sematophyllum Mitt demissum (Wils) Mitt
66 Hypnaceae Pl(itygyrium BSG
repens (Brid) BSG
PylaisieUa Kindb intricata (Hedw) Grout polyantha (Hedw) Grout selwynii (Kindb) Crum Steere et Anderson
Homomallium (Schimp) Loeske adnatum (Hedw) Broth
Hypnum Hedw cupressiforme Hedw var cupressiforme cupressifonne Hedw varfiliforme Brid curviolium Hedw fertile Sendtn imponens Hedw lindbergii Mitt palescens (Hedw) P-Beauv pratense Koch ex Brid
Calicladium Crum haldanianum (Grev) Crum
sopterygiopsis Iwats muelleriana (Schimp) Iwats
Isopterygium Mitt e1egans (Brid) lindb pulchellum (Hedw) Jaeg et Sauerb tenerum (Sw) Mitt
Taxiphyllum Fleisch depllQtum (Bruch et Schimp ex sun) Fleisch taxirameum (MitL) Fleisch
Herzogiella Broth seligeri (Brid) Iwats striatella (Brid) Iwats ruracea (Lindh) Iwats
hilium De Not crista-castrenna (Hedw) De Not
27
67 Rhytidiaceae Rhytidium (Sull) Kindb
rugosum (Hedw) Kindb
Rhytidfodelphus (Lindb ex Iimpr) Warnst triquetrus (Hedw) Warnst var triquetrus
68 Hylocomiaceae Hylocomium BSG
pyrenaicum (Spruce) lindb splendens (Hedw) BSG varsplendens
69 Buxbaumiaceae Buxbaumia Hedw
aphyllll Hedw
Diphyscium Mohr follosum (Hedw) Mohr
70 Tetraphidaceae Tetraphi3
pellucida Hedw
71 Polytrichaceae Atrichum PmiddotBeauv
angustatum (Brid) BSG cri8pum (James) Sullo oerstedianum (C Muell) Mitt teneUum (Roehl) BSG undulatum (Hedw) P middotBeauv var unduiatum
Pogonatum P-Beauv alpinum (Hedw) Roehl var alpinum umigerum (Hedw) P-Beauv
Poiytrichum Hedw commune Hedw var commune ormosum Hedw juniperinum Hedw longisetum Brid ohioense Ren et Card piliferum Hedw var pilierum strictum Brid
28
MAPS OF BRYOPHYTE TAXA
REPORTED FOR WISCONSIN
HORNWORTS AND LIVERWORTS
29
bull bull WSHeV~N S vE II
0 ~
e~RRON ~USK
(J~ (j -
CHIPPE A DUNN
0 o0 poundAU CLAIRE
~ TREMshyPEAshyLEAU
MICE
ONEIDA
Q LINCOLN
TAYLOR
Q t-~----- MOIRA THON
CLARK
o
10
LA FAVETTE
Hornwor ts amp Liverworts
- None - 1 - 25 taxa
- 26 - 50 tRXshy
- 51 - 75 tax)
- 7c - 100 taxa
FONDOU j LAC ~
DODGE
i O JEfFERSON
WALWORTH
30
Anastrophylltlfl helleranum (Nees) Schust Anastrophyllum michauxil (We~ Buch ~
46 88 136 141 Evans 38 46 141
Anastrophyllum mlnut~ (Schreb) Schuster Aneura plnguls (L) Durn
89 21 384687 88 89 90 (138) (145)
Anthoceros macounil MA Howe ~oceros punctatus L
21 (45) (144) 34
31
Barbilophozia attenuata (Mart) Loeske Barbilophozia barbata (Schmid ~ Schre~
24 141 Loeske 28 39 46 89 90 141
Bazzania trilobata (1) S Gray
1 26 39 46 (57) (59) 79 88 89
90 119 141
BlePbarostoma trichoffiyllum (1) Dumort
1 8 26 39 46 89 90 137 141
Blasia pusilla 1
1 39 46 90
Calypogeja fissa (1) Raddi
90
32
$129geja integristipula Steph C~Ypound2geja muelleriana (Schifm) K Mull
88 89 89
CallpoundO~eja neesiana (Mass amp Carest) K Mull
39 46 89 140 (141)
~pogeja sJlapound2icola (H Arnell tl J bull
Perss) Warnst et Loeske (119) 141
CeIalozia biscuspidata (L) Dum
1 37 I 46 90 t 137 141
CalYpogej~ trichomanis (L) Corda
26 39 (119) 137
33
CeEhalozia catenulata (Hub) Lindb Cephalozia connivens (Dicks) Lindb
39 46 79 141 39 4679 88 (89) 90 119 141
Cephalozia lacinulata Jack ~ Spruce
141 Cephalozia lunulifolia (Dum~ Dum I 28 39 46 79 88 89 90 141
Cepoundhalozia macouni (Aust) Aust Cephalozia ~leniceps (Aust) Lindb 27~ 3746 (66)7987 (139)141 39 46 79 89 141
34
Ceffialoziella 9ivaricata (Sm) Schiffn Cephaloziella ~hista (Jack) Schiffn
46 90 46 (119) 141
Cephaloziella hampeana (Nees) Schiffn
46
Cephaloziella rubella (Nees) Warnst
46 39
Cephaloziella rubella var sullivantii
(Aust) K Mull 46 39
Cephaloziella rubella var bifida (Lindb) Douin
35
46
Chiloscyphus pallescens (Ehrh ~ Hof~ Dum Chiloscyphus pallescens var fragilis
39J 46 88 89J 90 (Roth) K Milll 46
Chiloscyphus E9lyanthos (L) Corda
1 24 39 46
Chiloscyphus polyanthos vax rivularis
(Schrad) Nees 39 46
poundpoundnocepha1um conicum (1) Lindb
12 28 39 46 (49) 87 88 8990 120 121 148
Cololejeunea biddlecomiae (Aust) Evans
39 46 62 87 88 89 90
36
Dipl0]hyllum apicu1atum (Evans) Steph Fossombronia foveo1ata Lindb
(35)41 46 90 140 1 t 38 (39) t 46
Frullania bolanderi Aust
13 46 62 88 90 139 140
Frullanla ari ttoniae Evans
39 46 69 (71) 87 88 89 90
Frullania eboracensis Gott
1 9 28 4 39 46 (57) (59) 69 70 (71) 87 88 89 90 92 93 97 148
Frullania inflata Gott
46 87 88 89 (139)
Frullania oakesiana Aust Frullania plana Sullo
(35) 42 46 87 (139) (140) 92
Frullania ri paria Hampe ~ Lehm
46 69 71 87 89
Frullania ~elwynianna Pears
46 89
Geocalyx SEaveolens (Schrad) Nees
394688 8990 (119) 140
Frullania tamarisci (L) Dum subsp sPagrayana
(Mont) Hatt 28 34 l~6 t (57) 88 89 90
38
Mar pan thus seutatUB ( Web amp Mohr) Spruce Jamesoniella autumnalis (DC) Steph
(35) 39 46 90 1 7 28 39 46 56 (57) 87 8889 90 (119) 137 141
Jungermannia confertissima Nees
137 141
Jungermannia crenuliform1s Aust
79 137 141
Jungermannia ~il1ima Sm
46 90 141 Jungermannia exsert1folia subsp corifo~ (Dum~
Vana (35) 46 79 141
39
Jungermannia hyalina Lyell Jungerrr6nnia 1eiantha Grolle
40 46 79 136 141 39 46 79 88 90 1)7 141
Jungermannia IXlaxis Lindb Jungermannia pumila With
46 67 79 (139) (140) 141 )9 46 90 1)6 141
Jungermannia sphaerocarpa Hook Kurzia setacea (Web) Grolle
4679136140IIH 136
Lejeunea cavifo1ia (Ehrh) Lindb Lepidozia reptans (1) Dum
1 7 27 39 46 88 89 90 137 14010 I ( 27) 46 I 69 (71) 88
lophocolea heterophylla (Schrad) Dum Lophocolea minor Nees
1 )4 39 46 (57) 87 88 89 27 39 46 88 89 90 (119) 90 (93)94 119 136
Lophozia al~stris (Schleich ~ Web) Evans Lophozia ascendens (Warnst) Schust
46 90 137 141 136 140 141
41
loIilozia badensls (Gatt ~ Rabenh)
46 141
lophozia blcrenata (Schmid sectA Hoffm)
Dum 46 141
lophozia collaris (Nees) Dum
39 46 141
lophozla excisa (Dicks) Dum
27 4679 141
lophozia gillmani (Aust) Schust lophozia gIlttulata (Undo tl H Arnell)
36 39 40 46 136 (140) 141 Evans 39 46 136 141
42
141
Lophozia heterocolEDs (Thed) MA Howe IoPhozia ipcisa (Schard) Dum
27 39 46 89 141 8 27 31 46 79 90 140 141
LoQhozia longidens Macoun Lophozia ventricosa
1 46 79 90 141
Lophozia ventricosa var silvicola
(Buch) Jones 141 LoEhozla ventricosa vax longif1ora (Nees)
Macoun 39 46 79 141
43
~~ fragrans (Balbis) Frye et Clark Mannia pi 10sa (Hornem) Frye et Clark
46 (139) (140) 9 46 79 116 139
Mannia triandra (Scop) Gro 11e
46 (140)
Marchantia pound91ymorpha L
1 13 28 3946 (49) 78 87 88 89 90 (Ill) 121 122 147
Mo~rckia hibernica (HookJ Gott
46 89
Metzgeria conjugata Lindb
44
46
Mylia anomala (Hook) S Gray Notothylas orbicularis (Schwein) Sullo
39 46 141 (35)45 46
Nowellia curvifolia (Dicks) Mitt
17 27 3846 88 89 90 141
Odontoschisma denudatuJI1 (Nees ~ Mart)
Dum 46 81 137 141
Pallavicinia lyellii (Hook~ Carruth
90
Odontoschisma macounii (Aust) Under
45 46 (66) (139) 140 141
45
=
Pellia epiphllla (L) CordaPellia endiviifolia (Dicks~ Dum
27 39 46 90 13724 39 46
Pellia neesiana Limp bull Phaeoceros
39 46 89 27 39 45 46 97
Porella pinnata L
13 27 I 46 90 Plagiochila asplenioides (L) Dum
27 I 39 1+6 88 89 I 90 137
46
~e11a platYEhylla (L) Pfeiff Porella platyphylloi~ (Schwein) Lindb
17 27 39 46 69 (71)79 87 88 89 9 34 46 (57) 59 87 88 89 92 90 92 93 77
Preissia quadrata (Seop) Nees
1 36 3946 (71) 88 89 90 121 123
137
Ptllidium ciliare (L) Hampe
1 27 39 46 132 141
Dum
(22) 28 3946 (57) 87 88 89 90 92
Ptilidium ~lcherrium (G Web) Hampe
(9) 13 394656 (57)70 87 88 89 90 93 (119) 136 141
47
Radula obconica Sullo Reboulia hemlsphaerlca (L) Raddi
39 46 88 89 (43) 46 (59) 70 90 t Cf 124 125 (139)
Riccardia ~atifrons Lindb
1 27 39 46 89
Riccardia multifi~ (L) S Gray
39 46 88 145
Riccia beyrichiana Hampe ~ Lehm
46 Riccardia palmata (Hedw) Carruth
39 46 90
48
Riccia bifurca Hoffm Riccia cavernosa Hoffm
48 13739 46 79 137
Riccia fluitans L
242731454679
Riccla frostli Aust
48 79
Riccia sullicantii Aust
79 (137) Riccia sorocarpa Bisch 46 (137)
49
Ricciocarpus natans (1) Corda Sca~ia apiculata Spruce
13 27 46 79 89 110 39 46 (139) 141
ScaJltll1ia carinthica Jack ~ Lindb
137 t 141
Scapania ~ (Mart) Durn
39467990137141
Scapania digenil Schiffn
90
ScaEsectflia cuspiduligera (Nees) K Mull
46 (139) (140) (141)
50
Scapania glaucoeerha1a (Tayl) Aust Seapania gyrrmostomophilia Kaal
39 46 (139) 141 41 44 46 64 (139) (140) 141
ScaP3poundlia irrigua Gott ~ almiddot
7 39 46 79 137 (140L 141
Seapania nemorosa (L) Dum
1 39 46 89 90 137
Scapmi~ ~~ta Smiddotlch
137 141 51
~caPnia poundludicola LDeske amp K Mull Scapania saxicola Schust
46 (66) 141 136 141
Sc~~ scandica (H Arnell ~ Buch) Macv
139 140
Scapania subalpina (Nees) Durn
39 40 46 79 137 141
Scapania undulata (L) Dum
1 42 46 79 141
Scapania umbrosa (Schrad) Dum
137 140 141
52
Trichocolea tomentella (Ehrh) Dum Triomarla exsecta (Schmid) Schiffn
27 (35) 39 46 47 88 137 141 8 28 39 46 90 140 141
Tritomaria exsectiformis (Breidl) Loe~ke Tritomaria Sluinguedentata (Huds) Buch
39 -t6 89 90 136 140 141 7 39 46 141
Trltomaria scitula (Tayl) Joerg
46 136 (139) 141
53
pseudotriquetrum (Hedw) GaertnMeyer et Scherb var bimum (Schreb) Ulj stenotrichum C Muell var stenotrichum tortiolium Funck ex Brid uliginosum (Brid) B S G violaceum Crundw et Nyh weigelii Spreng
Rhodobryum (Schimp) Hampe roseum (Hedw) Limpr
45 Mniaceae Mnium Hedw
ciliare (C Muell) Schimp cinclidioides Hueb cuspidarum Hedw drummondii Bruch et Schimp longirostrum Brid lycopodioides Schwaegr marginatum (With) Brid ex P Beauv mediumB~S G var medium onhorrhynchum Brid pseudopunctatum Bruch et Schimp punctalUm Hedw var punctatum punctatum Hedw var elatum Schirnp rugicum Laur spinufosum B S G stellare Hedw
46 Aulacomruaceae Au1acomnium Schwaegr
androgynum (Hedw) Schwaegr heterostichum (Hedw) B S G paJustre (Hedw) Schwaegr var paJustre
47 Meesiaceae Amblyodon B S G
deillbotus (Hedw) B S G
Meesia Hedw triquetra (Richt) Angstr uJiginosa Hedw
48 Bartramiaceae Plagiopus Brid
oederiana (Sw) Umpr
Bartramia Hedw pomiformis Hedw var pomiformis
Philonotis Brid fontana (Hedw) Brid varfontana longseta (Michx) Britt varlongseta marchica (Hedw) Brid
21
~
~
49 Timmiaceae Timmia Hedw
megapolitana Hedw var megapolitana
50 Orthotrichaceae Zygodon Hook et Tayl
viridissimus (Dicks) Brid var rupestris lindb ex C J Hartm
Amphidium Schimp lapponicum (Hedw) Schimp mougeotii (B S G) Schimp
Orthotrichum Hedw anomalum Hedw obtuaifolium Brid ohioense Sullo et Lesq ex Aust pumilumSw sordidum Sullo et Lesq ex Aust speciosum Neesax Sturm var eegam (Schwaegr ex Hook et Grev) W~rnst stellDtwn Brid strrmgulatum P Beauv
Uleta Mohr ex Web cOfIICtata (p Beallv) Hamrn crispa (Hedw) Brid
Drummondill Hook ex Drumm prorepens (Hedw) Brltt
51 Fontinalaceae Fantinalis Hedw
antipyretica Hedw var antipyrettca antipyretica Hedw forma rabusta (Card) Card dalecarlica Schimp ex B S G var dalecarlica flqccidtJ Ren et Card hypnoides C J Hartrn var hypnoides hypnoides C J Hartm var durillei (Schimp) HU$ll
missourlcaCard novae-angliae Sullo var novae-angliae splulgrUlolill (C Moen) Wijket M~S
DichelY1l1ll Myr capilloceum (With) Myr falcatum (Bedw) Myr HIllescens B S G
52 Climaciaceae aimecium Web et Mohr ex Mohr
americllnum Brid dedroides (Hedw) Web et Mohr
22
S3 Hedwigiaceae Hedwigia P Beauv
ciliata (Hedw) P Beauv
54 Cryphaeaceae Forsstroemia lindh
ohioense (Sull) lindh
55 Leucodontaceae Leucodon Schwaegr
brachypus Blid var brachypus brachypus Brid var aniJrewsitlnus Crum et Anderson julaceus (Hedw) SuU
56 Neckeraceae Neckera Hedw
pennata Hedw var pennata
1fomalia (Brid) B S G trichomanoides (Hedw) B S G
57 Theliaceae Thelia Sull
asprellll Sullo hirtella (Hedw) Sullo lescurii SuU
Myurella B SG julacea (Schwaegr) B S G
sibrica (C MueU) Reim var sibrica
58 Fabroniaceae Fabronio Raddi
ciliaris (Brid) Bnd
Anacamptodon Brid splachnoides (Groet ex Brid) Brid
Schwetschkeopsis Broth fabronia (Schwaegr) Broth
59 Leskeaceae Lindbergia Kindb
brachyptera (Mitt) Kindb
Leskea Hedw gracilescens Hedw obscura Hedw poiycarpa Hedw
Leskeella (limpr) Loeske nervosa (Brid) Loeske
23
Pterlgy1lllndrum Hedw filiforme Hedw
60 Thuidiaceae Haplohymenfum Dozy et MoIk
trlste (Ces ex De Not) Kiodb
Anomodon Hook et Tayl attenuatus (Hedw) Hueb minor (Hedw) Fuemr rostratus (Hedw) Schimp viticulosus (Hedw) Hook et Tayl
Haplocladium (C Mucn) C MueU microphyllum (Hedw) Broth virginillnum (Brld) Broth ~bull
Thuidium B S G abietinum (Hedw) B S G delicatulum (Hedw) B S G vardelicatulum delicatulum (Hedw) B S G var radicans (Kindb) Crum Steere et Anderson minutulum (Hedw) B S G pygmaeum B S G recognftum (Hedw) Lindb scitum (p Beauv) Aust
Helodium (Sull) Warnst blandowli (Web et Mohr) Warnst var blandowli blandowli (Web et Mohr) Wamst var helodioides (Ren et Card ex Roell) Crum
Steere et Anderson paludomm (Sull) AuS
61 Amblystegiaceae Cratoneuron (Sull) Spruce
commulatum (Hedw) Roth var commutfltum commutatum (Hedw) Roth var fakatum (Brid) Moenk ilicinum (Hedw) Spruce var filcinum
Campylium (Sun) Mitt chrysophyllum (Brid) 1 Lange var chrysophyllum hispidulum (Brid) Mitt var hllJidulum polygamum (B S G)C lens varpolygtlmUm stellatum (Hedw) C1ens vat stelltum
Leptodictyum (Schimp) Warnst laxirete (Card et Ther) Broth riparium (Hedw) Wamat trichopodium (Schultz) Warns l1acillans (Sull) Broth
Hygroamblystegium Loelke jlul1iatile (Hedw) oeske var luViatile
24
fluviatile (Hedw) Loeske var oWltum Grout noterophilum (Sun et Lesq ex Sull) Warnst tenox (Hedw) Jenn var tenax tenox (Hedw) Jenn var spinilolium Schimp) Jenn
Amblystegium BSG juratzkanum Schimp serpens (Hedw) BSG varlum (Hedw) Undb var varium
Platydictya Berk confervoides (Brid) Crum jungermanniodes (Brid) Crum minutissimum (Sull et Lesq ex Sull) Crum subtile (Hedw) Crum
Drepanocladus (C Muell) Roth aduncus (Hedw) Warnst var aduncus aduncus (Hedw) Warnst var capillilolius (Wamst)Riebm aduncus (Hedw) Warnst var kneiffii (BSG) Moenk aduncus (Hedw) Warnst var polycorpus (Bland ex Voit) Roth exannulatus (BSG) Warnst fluitans (Hedw) Warnst var fluitans revolvens (Sw) Wamst var revolvens sendtneri (Schimp) Warnst var sendtnerl uncinatus (Hedw) Warnst var uncinatus uncinatus ffiedw) Warnst var piwnosus (Schimp) Wamst vernicoms (Lindb ex C Hartm) Warnst
Hygrohypnum Lindb eugyrium (BSG) loeske luridum (Hedw Jenn varluridum ochraecum (Tum ex Wils) Loeske var ochraceunt
Calliergon (Sull) Kindb cordilolium Hedw Kindb glganteum (Schimp) Kindb var giganteum richardsonii (Mitt) Kindb ex Wamst stramineum (Brid) Kindb trilarium (Web et Mohr) Kindb
Calliergonella Loeske cuspidata (Hedw) Loeske
Calliergidium (Ren) Grout bakeri (Ren) Grout
Scorpidium (Schimp) Urnpr scorpiodides (Hedw) limpr
62 Brachytbeciaceae Tomenthypnum loeske
nitens (Hedw) Loeske var mtens
25
Homalotheciella (Card) Broth subcapillata (Hedw) Broth
Brachythecium BSG acuminatum (Hedw) Awt acutum (Mitt) Sullo calcareum IGndb campestTe (C MuelL) BSG cunum (Lindb) Umpr digastrum C Muell et Kindb ex Macoun et Kindb oxycladon (Brid) Jaeg et Sauerb var oxycladon plumosum (Hedw) BSG populeum (Hedw) BSG var populeum reflexum (Starke ex Web et Mohr) DsG var relexum reiexum (Starke ex Web et Mohr) DSG var pacificum Renet Card ex Roell rlvulare BSG var rivulare tabulum (Hedw) BSC var tabuum salebrosum (Web et Mohr) BSG var salebrosum velutinum (Hedw) BSG Vat velutinum
Bryhniil Kaur graminicolor (Brid) Grout var graminicolor novae-angliae (Sull et Lesq ex sun) Grout
Rhynchostegium BSG senulatum (Hedw) Ja4lg et Sauerb
Rhynchostegiella (BSG) Limpr compacta (C MueU) Loeske
EU1hynchium 8SG hians (Hedw) Sande Lac pulchellum (Hedw) Jenn var pulchelium riparloides (Hedw) Rich
Stokeriella (Kindb) Robins pmelonga (Hedw) Robins var stokesii (Tum) Crum
63 Entodontaceae Entodon C Muell
brevisetus (Hook et Wils ex Wils) Lindb cladorrhizans (Hedw) C Muell seductrlx (Hedw) C MueU var wluctrix
Pleurozium Mitt schreberl (8rid) Mitt var schreberi
64 Plagiotheciaceae Pltlglothecium BSG
CflVifolium (Brid) Iwats var CtWifolium denticulatum (Hedw) 8SG laetum BSG ltztebricola BSG
26
6S SernatophyUaceae Brotherella Loeske ex Fleisch
recumins (Michx) Fleisch tenuirostris (Bruch et Schimp ex Sull) Fleisch
Sematophyllum Mitt demissum (Wils) Mitt
66 Hypnaceae Pl(itygyrium BSG
repens (Brid) BSG
PylaisieUa Kindb intricata (Hedw) Grout polyantha (Hedw) Grout selwynii (Kindb) Crum Steere et Anderson
Homomallium (Schimp) Loeske adnatum (Hedw) Broth
Hypnum Hedw cupressiforme Hedw var cupressiforme cupressifonne Hedw varfiliforme Brid curviolium Hedw fertile Sendtn imponens Hedw lindbergii Mitt palescens (Hedw) P-Beauv pratense Koch ex Brid
Calicladium Crum haldanianum (Grev) Crum
sopterygiopsis Iwats muelleriana (Schimp) Iwats
Isopterygium Mitt e1egans (Brid) lindb pulchellum (Hedw) Jaeg et Sauerb tenerum (Sw) Mitt
Taxiphyllum Fleisch depllQtum (Bruch et Schimp ex sun) Fleisch taxirameum (MitL) Fleisch
Herzogiella Broth seligeri (Brid) Iwats striatella (Brid) Iwats ruracea (Lindh) Iwats
hilium De Not crista-castrenna (Hedw) De Not
27
67 Rhytidiaceae Rhytidium (Sull) Kindb
rugosum (Hedw) Kindb
Rhytidfodelphus (Lindb ex Iimpr) Warnst triquetrus (Hedw) Warnst var triquetrus
68 Hylocomiaceae Hylocomium BSG
pyrenaicum (Spruce) lindb splendens (Hedw) BSG varsplendens
69 Buxbaumiaceae Buxbaumia Hedw
aphyllll Hedw
Diphyscium Mohr follosum (Hedw) Mohr
70 Tetraphidaceae Tetraphi3
pellucida Hedw
71 Polytrichaceae Atrichum PmiddotBeauv
angustatum (Brid) BSG cri8pum (James) Sullo oerstedianum (C Muell) Mitt teneUum (Roehl) BSG undulatum (Hedw) P middotBeauv var unduiatum
Pogonatum P-Beauv alpinum (Hedw) Roehl var alpinum umigerum (Hedw) P-Beauv
Poiytrichum Hedw commune Hedw var commune ormosum Hedw juniperinum Hedw longisetum Brid ohioense Ren et Card piliferum Hedw var pilierum strictum Brid
28
MAPS OF BRYOPHYTE TAXA
REPORTED FOR WISCONSIN
HORNWORTS AND LIVERWORTS
29
bull bull WSHeV~N S vE II
0 ~
e~RRON ~USK
(J~ (j -
CHIPPE A DUNN
0 o0 poundAU CLAIRE
~ TREMshyPEAshyLEAU
MICE
ONEIDA
Q LINCOLN
TAYLOR
Q t-~----- MOIRA THON
CLARK
o
10
LA FAVETTE
Hornwor ts amp Liverworts
- None - 1 - 25 taxa
- 26 - 50 tRXshy
- 51 - 75 tax)
- 7c - 100 taxa
FONDOU j LAC ~
DODGE
i O JEfFERSON
WALWORTH
30
Anastrophylltlfl helleranum (Nees) Schust Anastrophyllum michauxil (We~ Buch ~
46 88 136 141 Evans 38 46 141
Anastrophyllum mlnut~ (Schreb) Schuster Aneura plnguls (L) Durn
89 21 384687 88 89 90 (138) (145)
Anthoceros macounil MA Howe ~oceros punctatus L
21 (45) (144) 34
31
Barbilophozia attenuata (Mart) Loeske Barbilophozia barbata (Schmid ~ Schre~
24 141 Loeske 28 39 46 89 90 141
Bazzania trilobata (1) S Gray
1 26 39 46 (57) (59) 79 88 89
90 119 141
BlePbarostoma trichoffiyllum (1) Dumort
1 8 26 39 46 89 90 137 141
Blasia pusilla 1
1 39 46 90
Calypogeja fissa (1) Raddi
90
32
$129geja integristipula Steph C~Ypound2geja muelleriana (Schifm) K Mull
88 89 89
CallpoundO~eja neesiana (Mass amp Carest) K Mull
39 46 89 140 (141)
~pogeja sJlapound2icola (H Arnell tl J bull
Perss) Warnst et Loeske (119) 141
CeIalozia biscuspidata (L) Dum
1 37 I 46 90 t 137 141
CalYpogej~ trichomanis (L) Corda
26 39 (119) 137
33
CeEhalozia catenulata (Hub) Lindb Cephalozia connivens (Dicks) Lindb
39 46 79 141 39 4679 88 (89) 90 119 141
Cephalozia lacinulata Jack ~ Spruce
141 Cephalozia lunulifolia (Dum~ Dum I 28 39 46 79 88 89 90 141
Cepoundhalozia macouni (Aust) Aust Cephalozia ~leniceps (Aust) Lindb 27~ 3746 (66)7987 (139)141 39 46 79 89 141
34
Ceffialoziella 9ivaricata (Sm) Schiffn Cephaloziella ~hista (Jack) Schiffn
46 90 46 (119) 141
Cephaloziella hampeana (Nees) Schiffn
46
Cephaloziella rubella (Nees) Warnst
46 39
Cephaloziella rubella var sullivantii
(Aust) K Mull 46 39
Cephaloziella rubella var bifida (Lindb) Douin
35
46
Chiloscyphus pallescens (Ehrh ~ Hof~ Dum Chiloscyphus pallescens var fragilis
39J 46 88 89J 90 (Roth) K Milll 46
Chiloscyphus E9lyanthos (L) Corda
1 24 39 46
Chiloscyphus polyanthos vax rivularis
(Schrad) Nees 39 46
poundpoundnocepha1um conicum (1) Lindb
12 28 39 46 (49) 87 88 8990 120 121 148
Cololejeunea biddlecomiae (Aust) Evans
39 46 62 87 88 89 90
36
Dipl0]hyllum apicu1atum (Evans) Steph Fossombronia foveo1ata Lindb
(35)41 46 90 140 1 t 38 (39) t 46
Frullania bolanderi Aust
13 46 62 88 90 139 140
Frullanla ari ttoniae Evans
39 46 69 (71) 87 88 89 90
Frullania eboracensis Gott
1 9 28 4 39 46 (57) (59) 69 70 (71) 87 88 89 90 92 93 97 148
Frullania inflata Gott
46 87 88 89 (139)
Frullania oakesiana Aust Frullania plana Sullo
(35) 42 46 87 (139) (140) 92
Frullania ri paria Hampe ~ Lehm
46 69 71 87 89
Frullania ~elwynianna Pears
46 89
Geocalyx SEaveolens (Schrad) Nees
394688 8990 (119) 140
Frullania tamarisci (L) Dum subsp sPagrayana
(Mont) Hatt 28 34 l~6 t (57) 88 89 90
38
Mar pan thus seutatUB ( Web amp Mohr) Spruce Jamesoniella autumnalis (DC) Steph
(35) 39 46 90 1 7 28 39 46 56 (57) 87 8889 90 (119) 137 141
Jungermannia confertissima Nees
137 141
Jungermannia crenuliform1s Aust
79 137 141
Jungermannia ~il1ima Sm
46 90 141 Jungermannia exsert1folia subsp corifo~ (Dum~
Vana (35) 46 79 141
39
Jungermannia hyalina Lyell Jungerrr6nnia 1eiantha Grolle
40 46 79 136 141 39 46 79 88 90 1)7 141
Jungermannia IXlaxis Lindb Jungermannia pumila With
46 67 79 (139) (140) 141 )9 46 90 1)6 141
Jungermannia sphaerocarpa Hook Kurzia setacea (Web) Grolle
4679136140IIH 136
Lejeunea cavifo1ia (Ehrh) Lindb Lepidozia reptans (1) Dum
1 7 27 39 46 88 89 90 137 14010 I ( 27) 46 I 69 (71) 88
lophocolea heterophylla (Schrad) Dum Lophocolea minor Nees
1 )4 39 46 (57) 87 88 89 27 39 46 88 89 90 (119) 90 (93)94 119 136
Lophozia al~stris (Schleich ~ Web) Evans Lophozia ascendens (Warnst) Schust
46 90 137 141 136 140 141
41
loIilozia badensls (Gatt ~ Rabenh)
46 141
lophozia blcrenata (Schmid sectA Hoffm)
Dum 46 141
lophozia collaris (Nees) Dum
39 46 141
lophozla excisa (Dicks) Dum
27 4679 141
lophozia gillmani (Aust) Schust lophozia gIlttulata (Undo tl H Arnell)
36 39 40 46 136 (140) 141 Evans 39 46 136 141
42
141
Lophozia heterocolEDs (Thed) MA Howe IoPhozia ipcisa (Schard) Dum
27 39 46 89 141 8 27 31 46 79 90 140 141
LoQhozia longidens Macoun Lophozia ventricosa
1 46 79 90 141
Lophozia ventricosa var silvicola
(Buch) Jones 141 LoEhozla ventricosa vax longif1ora (Nees)
Macoun 39 46 79 141
43
~~ fragrans (Balbis) Frye et Clark Mannia pi 10sa (Hornem) Frye et Clark
46 (139) (140) 9 46 79 116 139
Mannia triandra (Scop) Gro 11e
46 (140)
Marchantia pound91ymorpha L
1 13 28 3946 (49) 78 87 88 89 90 (Ill) 121 122 147
Mo~rckia hibernica (HookJ Gott
46 89
Metzgeria conjugata Lindb
44
46
Mylia anomala (Hook) S Gray Notothylas orbicularis (Schwein) Sullo
39 46 141 (35)45 46
Nowellia curvifolia (Dicks) Mitt
17 27 3846 88 89 90 141
Odontoschisma denudatuJI1 (Nees ~ Mart)
Dum 46 81 137 141
Pallavicinia lyellii (Hook~ Carruth
90
Odontoschisma macounii (Aust) Under
45 46 (66) (139) 140 141
45
=
Pellia epiphllla (L) CordaPellia endiviifolia (Dicks~ Dum
27 39 46 90 13724 39 46
Pellia neesiana Limp bull Phaeoceros
39 46 89 27 39 45 46 97
Porella pinnata L
13 27 I 46 90 Plagiochila asplenioides (L) Dum
27 I 39 1+6 88 89 I 90 137
46
~e11a platYEhylla (L) Pfeiff Porella platyphylloi~ (Schwein) Lindb
17 27 39 46 69 (71)79 87 88 89 9 34 46 (57) 59 87 88 89 92 90 92 93 77
Preissia quadrata (Seop) Nees
1 36 3946 (71) 88 89 90 121 123
137
Ptllidium ciliare (L) Hampe
1 27 39 46 132 141
Dum
(22) 28 3946 (57) 87 88 89 90 92
Ptilidium ~lcherrium (G Web) Hampe
(9) 13 394656 (57)70 87 88 89 90 93 (119) 136 141
47
Radula obconica Sullo Reboulia hemlsphaerlca (L) Raddi
39 46 88 89 (43) 46 (59) 70 90 t Cf 124 125 (139)
Riccardia ~atifrons Lindb
1 27 39 46 89
Riccardia multifi~ (L) S Gray
39 46 88 145
Riccia beyrichiana Hampe ~ Lehm
46 Riccardia palmata (Hedw) Carruth
39 46 90
48
Riccia bifurca Hoffm Riccia cavernosa Hoffm
48 13739 46 79 137
Riccia fluitans L
242731454679
Riccla frostli Aust
48 79
Riccia sullicantii Aust
79 (137) Riccia sorocarpa Bisch 46 (137)
49
Ricciocarpus natans (1) Corda Sca~ia apiculata Spruce
13 27 46 79 89 110 39 46 (139) 141
ScaJltll1ia carinthica Jack ~ Lindb
137 t 141
Scapania ~ (Mart) Durn
39467990137141
Scapania digenil Schiffn
90
ScaEsectflia cuspiduligera (Nees) K Mull
46 (139) (140) (141)
50
Scapania glaucoeerha1a (Tayl) Aust Seapania gyrrmostomophilia Kaal
39 46 (139) 141 41 44 46 64 (139) (140) 141
ScaP3poundlia irrigua Gott ~ almiddot
7 39 46 79 137 (140L 141
Seapania nemorosa (L) Dum
1 39 46 89 90 137
Scapmi~ ~~ta Smiddotlch
137 141 51
~caPnia poundludicola LDeske amp K Mull Scapania saxicola Schust
46 (66) 141 136 141
Sc~~ scandica (H Arnell ~ Buch) Macv
139 140
Scapania subalpina (Nees) Durn
39 40 46 79 137 141
Scapania undulata (L) Dum
1 42 46 79 141
Scapania umbrosa (Schrad) Dum
137 140 141
52
Trichocolea tomentella (Ehrh) Dum Triomarla exsecta (Schmid) Schiffn
27 (35) 39 46 47 88 137 141 8 28 39 46 90 140 141
Tritomaria exsectiformis (Breidl) Loe~ke Tritomaria Sluinguedentata (Huds) Buch
39 -t6 89 90 136 140 141 7 39 46 141
Trltomaria scitula (Tayl) Joerg
46 136 (139) 141
53
49 Timmiaceae Timmia Hedw
megapolitana Hedw var megapolitana
50 Orthotrichaceae Zygodon Hook et Tayl
viridissimus (Dicks) Brid var rupestris lindb ex C J Hartm
Amphidium Schimp lapponicum (Hedw) Schimp mougeotii (B S G) Schimp
Orthotrichum Hedw anomalum Hedw obtuaifolium Brid ohioense Sullo et Lesq ex Aust pumilumSw sordidum Sullo et Lesq ex Aust speciosum Neesax Sturm var eegam (Schwaegr ex Hook et Grev) W~rnst stellDtwn Brid strrmgulatum P Beauv
Uleta Mohr ex Web cOfIICtata (p Beallv) Hamrn crispa (Hedw) Brid
Drummondill Hook ex Drumm prorepens (Hedw) Brltt
51 Fontinalaceae Fantinalis Hedw
antipyretica Hedw var antipyrettca antipyretica Hedw forma rabusta (Card) Card dalecarlica Schimp ex B S G var dalecarlica flqccidtJ Ren et Card hypnoides C J Hartrn var hypnoides hypnoides C J Hartm var durillei (Schimp) HU$ll
missourlcaCard novae-angliae Sullo var novae-angliae splulgrUlolill (C Moen) Wijket M~S
DichelY1l1ll Myr capilloceum (With) Myr falcatum (Bedw) Myr HIllescens B S G
52 Climaciaceae aimecium Web et Mohr ex Mohr
americllnum Brid dedroides (Hedw) Web et Mohr
22
S3 Hedwigiaceae Hedwigia P Beauv
ciliata (Hedw) P Beauv
54 Cryphaeaceae Forsstroemia lindh
ohioense (Sull) lindh
55 Leucodontaceae Leucodon Schwaegr
brachypus Blid var brachypus brachypus Brid var aniJrewsitlnus Crum et Anderson julaceus (Hedw) SuU
56 Neckeraceae Neckera Hedw
pennata Hedw var pennata
1fomalia (Brid) B S G trichomanoides (Hedw) B S G
57 Theliaceae Thelia Sull
asprellll Sullo hirtella (Hedw) Sullo lescurii SuU
Myurella B SG julacea (Schwaegr) B S G
sibrica (C MueU) Reim var sibrica
58 Fabroniaceae Fabronio Raddi
ciliaris (Brid) Bnd
Anacamptodon Brid splachnoides (Groet ex Brid) Brid
Schwetschkeopsis Broth fabronia (Schwaegr) Broth
59 Leskeaceae Lindbergia Kindb
brachyptera (Mitt) Kindb
Leskea Hedw gracilescens Hedw obscura Hedw poiycarpa Hedw
Leskeella (limpr) Loeske nervosa (Brid) Loeske
23
Pterlgy1lllndrum Hedw filiforme Hedw
60 Thuidiaceae Haplohymenfum Dozy et MoIk
trlste (Ces ex De Not) Kiodb
Anomodon Hook et Tayl attenuatus (Hedw) Hueb minor (Hedw) Fuemr rostratus (Hedw) Schimp viticulosus (Hedw) Hook et Tayl
Haplocladium (C Mucn) C MueU microphyllum (Hedw) Broth virginillnum (Brld) Broth ~bull
Thuidium B S G abietinum (Hedw) B S G delicatulum (Hedw) B S G vardelicatulum delicatulum (Hedw) B S G var radicans (Kindb) Crum Steere et Anderson minutulum (Hedw) B S G pygmaeum B S G recognftum (Hedw) Lindb scitum (p Beauv) Aust
Helodium (Sull) Warnst blandowli (Web et Mohr) Warnst var blandowli blandowli (Web et Mohr) Wamst var helodioides (Ren et Card ex Roell) Crum
Steere et Anderson paludomm (Sull) AuS
61 Amblystegiaceae Cratoneuron (Sull) Spruce
commulatum (Hedw) Roth var commutfltum commutatum (Hedw) Roth var fakatum (Brid) Moenk ilicinum (Hedw) Spruce var filcinum
Campylium (Sun) Mitt chrysophyllum (Brid) 1 Lange var chrysophyllum hispidulum (Brid) Mitt var hllJidulum polygamum (B S G)C lens varpolygtlmUm stellatum (Hedw) C1ens vat stelltum
Leptodictyum (Schimp) Warnst laxirete (Card et Ther) Broth riparium (Hedw) Wamat trichopodium (Schultz) Warns l1acillans (Sull) Broth
Hygroamblystegium Loelke jlul1iatile (Hedw) oeske var luViatile
24
fluviatile (Hedw) Loeske var oWltum Grout noterophilum (Sun et Lesq ex Sull) Warnst tenox (Hedw) Jenn var tenax tenox (Hedw) Jenn var spinilolium Schimp) Jenn
Amblystegium BSG juratzkanum Schimp serpens (Hedw) BSG varlum (Hedw) Undb var varium
Platydictya Berk confervoides (Brid) Crum jungermanniodes (Brid) Crum minutissimum (Sull et Lesq ex Sull) Crum subtile (Hedw) Crum
Drepanocladus (C Muell) Roth aduncus (Hedw) Warnst var aduncus aduncus (Hedw) Warnst var capillilolius (Wamst)Riebm aduncus (Hedw) Warnst var kneiffii (BSG) Moenk aduncus (Hedw) Warnst var polycorpus (Bland ex Voit) Roth exannulatus (BSG) Warnst fluitans (Hedw) Warnst var fluitans revolvens (Sw) Wamst var revolvens sendtneri (Schimp) Warnst var sendtnerl uncinatus (Hedw) Warnst var uncinatus uncinatus ffiedw) Warnst var piwnosus (Schimp) Wamst vernicoms (Lindb ex C Hartm) Warnst
Hygrohypnum Lindb eugyrium (BSG) loeske luridum (Hedw Jenn varluridum ochraecum (Tum ex Wils) Loeske var ochraceunt
Calliergon (Sull) Kindb cordilolium Hedw Kindb glganteum (Schimp) Kindb var giganteum richardsonii (Mitt) Kindb ex Wamst stramineum (Brid) Kindb trilarium (Web et Mohr) Kindb
Calliergonella Loeske cuspidata (Hedw) Loeske
Calliergidium (Ren) Grout bakeri (Ren) Grout
Scorpidium (Schimp) Urnpr scorpiodides (Hedw) limpr
62 Brachytbeciaceae Tomenthypnum loeske
nitens (Hedw) Loeske var mtens
25
Homalotheciella (Card) Broth subcapillata (Hedw) Broth
Brachythecium BSG acuminatum (Hedw) Awt acutum (Mitt) Sullo calcareum IGndb campestTe (C MuelL) BSG cunum (Lindb) Umpr digastrum C Muell et Kindb ex Macoun et Kindb oxycladon (Brid) Jaeg et Sauerb var oxycladon plumosum (Hedw) BSG populeum (Hedw) BSG var populeum reflexum (Starke ex Web et Mohr) DsG var relexum reiexum (Starke ex Web et Mohr) DSG var pacificum Renet Card ex Roell rlvulare BSG var rivulare tabulum (Hedw) BSC var tabuum salebrosum (Web et Mohr) BSG var salebrosum velutinum (Hedw) BSG Vat velutinum
Bryhniil Kaur graminicolor (Brid) Grout var graminicolor novae-angliae (Sull et Lesq ex sun) Grout
Rhynchostegium BSG senulatum (Hedw) Ja4lg et Sauerb
Rhynchostegiella (BSG) Limpr compacta (C MueU) Loeske
EU1hynchium 8SG hians (Hedw) Sande Lac pulchellum (Hedw) Jenn var pulchelium riparloides (Hedw) Rich
Stokeriella (Kindb) Robins pmelonga (Hedw) Robins var stokesii (Tum) Crum
63 Entodontaceae Entodon C Muell
brevisetus (Hook et Wils ex Wils) Lindb cladorrhizans (Hedw) C Muell seductrlx (Hedw) C MueU var wluctrix
Pleurozium Mitt schreberl (8rid) Mitt var schreberi
64 Plagiotheciaceae Pltlglothecium BSG
CflVifolium (Brid) Iwats var CtWifolium denticulatum (Hedw) 8SG laetum BSG ltztebricola BSG
26
6S SernatophyUaceae Brotherella Loeske ex Fleisch
recumins (Michx) Fleisch tenuirostris (Bruch et Schimp ex Sull) Fleisch
Sematophyllum Mitt demissum (Wils) Mitt
66 Hypnaceae Pl(itygyrium BSG
repens (Brid) BSG
PylaisieUa Kindb intricata (Hedw) Grout polyantha (Hedw) Grout selwynii (Kindb) Crum Steere et Anderson
Homomallium (Schimp) Loeske adnatum (Hedw) Broth
Hypnum Hedw cupressiforme Hedw var cupressiforme cupressifonne Hedw varfiliforme Brid curviolium Hedw fertile Sendtn imponens Hedw lindbergii Mitt palescens (Hedw) P-Beauv pratense Koch ex Brid
Calicladium Crum haldanianum (Grev) Crum
sopterygiopsis Iwats muelleriana (Schimp) Iwats
Isopterygium Mitt e1egans (Brid) lindb pulchellum (Hedw) Jaeg et Sauerb tenerum (Sw) Mitt
Taxiphyllum Fleisch depllQtum (Bruch et Schimp ex sun) Fleisch taxirameum (MitL) Fleisch
Herzogiella Broth seligeri (Brid) Iwats striatella (Brid) Iwats ruracea (Lindh) Iwats
hilium De Not crista-castrenna (Hedw) De Not
27
67 Rhytidiaceae Rhytidium (Sull) Kindb
rugosum (Hedw) Kindb
Rhytidfodelphus (Lindb ex Iimpr) Warnst triquetrus (Hedw) Warnst var triquetrus
68 Hylocomiaceae Hylocomium BSG
pyrenaicum (Spruce) lindb splendens (Hedw) BSG varsplendens
69 Buxbaumiaceae Buxbaumia Hedw
aphyllll Hedw
Diphyscium Mohr follosum (Hedw) Mohr
70 Tetraphidaceae Tetraphi3
pellucida Hedw
71 Polytrichaceae Atrichum PmiddotBeauv
angustatum (Brid) BSG cri8pum (James) Sullo oerstedianum (C Muell) Mitt teneUum (Roehl) BSG undulatum (Hedw) P middotBeauv var unduiatum
Pogonatum P-Beauv alpinum (Hedw) Roehl var alpinum umigerum (Hedw) P-Beauv
Poiytrichum Hedw commune Hedw var commune ormosum Hedw juniperinum Hedw longisetum Brid ohioense Ren et Card piliferum Hedw var pilierum strictum Brid
28
MAPS OF BRYOPHYTE TAXA
REPORTED FOR WISCONSIN
HORNWORTS AND LIVERWORTS
29
bull bull WSHeV~N S vE II
0 ~
e~RRON ~USK
(J~ (j -
CHIPPE A DUNN
0 o0 poundAU CLAIRE
~ TREMshyPEAshyLEAU
MICE
ONEIDA
Q LINCOLN
TAYLOR
Q t-~----- MOIRA THON
CLARK
o
10
LA FAVETTE
Hornwor ts amp Liverworts
- None - 1 - 25 taxa
- 26 - 50 tRXshy
- 51 - 75 tax)
- 7c - 100 taxa
FONDOU j LAC ~
DODGE
i O JEfFERSON
WALWORTH
30
Anastrophylltlfl helleranum (Nees) Schust Anastrophyllum michauxil (We~ Buch ~
46 88 136 141 Evans 38 46 141
Anastrophyllum mlnut~ (Schreb) Schuster Aneura plnguls (L) Durn
89 21 384687 88 89 90 (138) (145)
Anthoceros macounil MA Howe ~oceros punctatus L
21 (45) (144) 34
31
Barbilophozia attenuata (Mart) Loeske Barbilophozia barbata (Schmid ~ Schre~
24 141 Loeske 28 39 46 89 90 141
Bazzania trilobata (1) S Gray
1 26 39 46 (57) (59) 79 88 89
90 119 141
BlePbarostoma trichoffiyllum (1) Dumort
1 8 26 39 46 89 90 137 141
Blasia pusilla 1
1 39 46 90
Calypogeja fissa (1) Raddi
90
32
$129geja integristipula Steph C~Ypound2geja muelleriana (Schifm) K Mull
88 89 89
CallpoundO~eja neesiana (Mass amp Carest) K Mull
39 46 89 140 (141)
~pogeja sJlapound2icola (H Arnell tl J bull
Perss) Warnst et Loeske (119) 141
CeIalozia biscuspidata (L) Dum
1 37 I 46 90 t 137 141
CalYpogej~ trichomanis (L) Corda
26 39 (119) 137
33
CeEhalozia catenulata (Hub) Lindb Cephalozia connivens (Dicks) Lindb
39 46 79 141 39 4679 88 (89) 90 119 141
Cephalozia lacinulata Jack ~ Spruce
141 Cephalozia lunulifolia (Dum~ Dum I 28 39 46 79 88 89 90 141
Cepoundhalozia macouni (Aust) Aust Cephalozia ~leniceps (Aust) Lindb 27~ 3746 (66)7987 (139)141 39 46 79 89 141
34
Ceffialoziella 9ivaricata (Sm) Schiffn Cephaloziella ~hista (Jack) Schiffn
46 90 46 (119) 141
Cephaloziella hampeana (Nees) Schiffn
46
Cephaloziella rubella (Nees) Warnst
46 39
Cephaloziella rubella var sullivantii
(Aust) K Mull 46 39
Cephaloziella rubella var bifida (Lindb) Douin
35
46
Chiloscyphus pallescens (Ehrh ~ Hof~ Dum Chiloscyphus pallescens var fragilis
39J 46 88 89J 90 (Roth) K Milll 46
Chiloscyphus E9lyanthos (L) Corda
1 24 39 46
Chiloscyphus polyanthos vax rivularis
(Schrad) Nees 39 46
poundpoundnocepha1um conicum (1) Lindb
12 28 39 46 (49) 87 88 8990 120 121 148
Cololejeunea biddlecomiae (Aust) Evans
39 46 62 87 88 89 90
36
Dipl0]hyllum apicu1atum (Evans) Steph Fossombronia foveo1ata Lindb
(35)41 46 90 140 1 t 38 (39) t 46
Frullania bolanderi Aust
13 46 62 88 90 139 140
Frullanla ari ttoniae Evans
39 46 69 (71) 87 88 89 90
Frullania eboracensis Gott
1 9 28 4 39 46 (57) (59) 69 70 (71) 87 88 89 90 92 93 97 148
Frullania inflata Gott
46 87 88 89 (139)
Frullania oakesiana Aust Frullania plana Sullo
(35) 42 46 87 (139) (140) 92
Frullania ri paria Hampe ~ Lehm
46 69 71 87 89
Frullania ~elwynianna Pears
46 89
Geocalyx SEaveolens (Schrad) Nees
394688 8990 (119) 140
Frullania tamarisci (L) Dum subsp sPagrayana
(Mont) Hatt 28 34 l~6 t (57) 88 89 90
38
Mar pan thus seutatUB ( Web amp Mohr) Spruce Jamesoniella autumnalis (DC) Steph
(35) 39 46 90 1 7 28 39 46 56 (57) 87 8889 90 (119) 137 141
Jungermannia confertissima Nees
137 141
Jungermannia crenuliform1s Aust
79 137 141
Jungermannia ~il1ima Sm
46 90 141 Jungermannia exsert1folia subsp corifo~ (Dum~
Vana (35) 46 79 141
39
Jungermannia hyalina Lyell Jungerrr6nnia 1eiantha Grolle
40 46 79 136 141 39 46 79 88 90 1)7 141
Jungermannia IXlaxis Lindb Jungermannia pumila With
46 67 79 (139) (140) 141 )9 46 90 1)6 141
Jungermannia sphaerocarpa Hook Kurzia setacea (Web) Grolle
4679136140IIH 136
Lejeunea cavifo1ia (Ehrh) Lindb Lepidozia reptans (1) Dum
1 7 27 39 46 88 89 90 137 14010 I ( 27) 46 I 69 (71) 88
lophocolea heterophylla (Schrad) Dum Lophocolea minor Nees
1 )4 39 46 (57) 87 88 89 27 39 46 88 89 90 (119) 90 (93)94 119 136
Lophozia al~stris (Schleich ~ Web) Evans Lophozia ascendens (Warnst) Schust
46 90 137 141 136 140 141
41
loIilozia badensls (Gatt ~ Rabenh)
46 141
lophozia blcrenata (Schmid sectA Hoffm)
Dum 46 141
lophozia collaris (Nees) Dum
39 46 141
lophozla excisa (Dicks) Dum
27 4679 141
lophozia gillmani (Aust) Schust lophozia gIlttulata (Undo tl H Arnell)
36 39 40 46 136 (140) 141 Evans 39 46 136 141
42
141
Lophozia heterocolEDs (Thed) MA Howe IoPhozia ipcisa (Schard) Dum
27 39 46 89 141 8 27 31 46 79 90 140 141
LoQhozia longidens Macoun Lophozia ventricosa
1 46 79 90 141
Lophozia ventricosa var silvicola
(Buch) Jones 141 LoEhozla ventricosa vax longif1ora (Nees)
Macoun 39 46 79 141
43
~~ fragrans (Balbis) Frye et Clark Mannia pi 10sa (Hornem) Frye et Clark
46 (139) (140) 9 46 79 116 139
Mannia triandra (Scop) Gro 11e
46 (140)
Marchantia pound91ymorpha L
1 13 28 3946 (49) 78 87 88 89 90 (Ill) 121 122 147
Mo~rckia hibernica (HookJ Gott
46 89
Metzgeria conjugata Lindb
44
46
Mylia anomala (Hook) S Gray Notothylas orbicularis (Schwein) Sullo
39 46 141 (35)45 46
Nowellia curvifolia (Dicks) Mitt
17 27 3846 88 89 90 141
Odontoschisma denudatuJI1 (Nees ~ Mart)
Dum 46 81 137 141
Pallavicinia lyellii (Hook~ Carruth
90
Odontoschisma macounii (Aust) Under
45 46 (66) (139) 140 141
45
=
Pellia epiphllla (L) CordaPellia endiviifolia (Dicks~ Dum
27 39 46 90 13724 39 46
Pellia neesiana Limp bull Phaeoceros
39 46 89 27 39 45 46 97
Porella pinnata L
13 27 I 46 90 Plagiochila asplenioides (L) Dum
27 I 39 1+6 88 89 I 90 137
46
~e11a platYEhylla (L) Pfeiff Porella platyphylloi~ (Schwein) Lindb
17 27 39 46 69 (71)79 87 88 89 9 34 46 (57) 59 87 88 89 92 90 92 93 77
Preissia quadrata (Seop) Nees
1 36 3946 (71) 88 89 90 121 123
137
Ptllidium ciliare (L) Hampe
1 27 39 46 132 141
Dum
(22) 28 3946 (57) 87 88 89 90 92
Ptilidium ~lcherrium (G Web) Hampe
(9) 13 394656 (57)70 87 88 89 90 93 (119) 136 141
47
Radula obconica Sullo Reboulia hemlsphaerlca (L) Raddi
39 46 88 89 (43) 46 (59) 70 90 t Cf 124 125 (139)
Riccardia ~atifrons Lindb
1 27 39 46 89
Riccardia multifi~ (L) S Gray
39 46 88 145
Riccia beyrichiana Hampe ~ Lehm
46 Riccardia palmata (Hedw) Carruth
39 46 90
48
Riccia bifurca Hoffm Riccia cavernosa Hoffm
48 13739 46 79 137
Riccia fluitans L
242731454679
Riccla frostli Aust
48 79
Riccia sullicantii Aust
79 (137) Riccia sorocarpa Bisch 46 (137)
49
Ricciocarpus natans (1) Corda Sca~ia apiculata Spruce
13 27 46 79 89 110 39 46 (139) 141
ScaJltll1ia carinthica Jack ~ Lindb
137 t 141
Scapania ~ (Mart) Durn
39467990137141
Scapania digenil Schiffn
90
ScaEsectflia cuspiduligera (Nees) K Mull
46 (139) (140) (141)
50
Scapania glaucoeerha1a (Tayl) Aust Seapania gyrrmostomophilia Kaal
39 46 (139) 141 41 44 46 64 (139) (140) 141
ScaP3poundlia irrigua Gott ~ almiddot
7 39 46 79 137 (140L 141
Seapania nemorosa (L) Dum
1 39 46 89 90 137
Scapmi~ ~~ta Smiddotlch
137 141 51
~caPnia poundludicola LDeske amp K Mull Scapania saxicola Schust
46 (66) 141 136 141
Sc~~ scandica (H Arnell ~ Buch) Macv
139 140
Scapania subalpina (Nees) Durn
39 40 46 79 137 141
Scapania undulata (L) Dum
1 42 46 79 141
Scapania umbrosa (Schrad) Dum
137 140 141
52
Trichocolea tomentella (Ehrh) Dum Triomarla exsecta (Schmid) Schiffn
27 (35) 39 46 47 88 137 141 8 28 39 46 90 140 141
Tritomaria exsectiformis (Breidl) Loe~ke Tritomaria Sluinguedentata (Huds) Buch
39 -t6 89 90 136 140 141 7 39 46 141
Trltomaria scitula (Tayl) Joerg
46 136 (139) 141
53
S3 Hedwigiaceae Hedwigia P Beauv
ciliata (Hedw) P Beauv
54 Cryphaeaceae Forsstroemia lindh
ohioense (Sull) lindh
55 Leucodontaceae Leucodon Schwaegr
brachypus Blid var brachypus brachypus Brid var aniJrewsitlnus Crum et Anderson julaceus (Hedw) SuU
56 Neckeraceae Neckera Hedw
pennata Hedw var pennata
1fomalia (Brid) B S G trichomanoides (Hedw) B S G
57 Theliaceae Thelia Sull
asprellll Sullo hirtella (Hedw) Sullo lescurii SuU
Myurella B SG julacea (Schwaegr) B S G
sibrica (C MueU) Reim var sibrica
58 Fabroniaceae Fabronio Raddi
ciliaris (Brid) Bnd
Anacamptodon Brid splachnoides (Groet ex Brid) Brid
Schwetschkeopsis Broth fabronia (Schwaegr) Broth
59 Leskeaceae Lindbergia Kindb
brachyptera (Mitt) Kindb
Leskea Hedw gracilescens Hedw obscura Hedw poiycarpa Hedw
Leskeella (limpr) Loeske nervosa (Brid) Loeske
23
Pterlgy1lllndrum Hedw filiforme Hedw
60 Thuidiaceae Haplohymenfum Dozy et MoIk
trlste (Ces ex De Not) Kiodb
Anomodon Hook et Tayl attenuatus (Hedw) Hueb minor (Hedw) Fuemr rostratus (Hedw) Schimp viticulosus (Hedw) Hook et Tayl
Haplocladium (C Mucn) C MueU microphyllum (Hedw) Broth virginillnum (Brld) Broth ~bull
Thuidium B S G abietinum (Hedw) B S G delicatulum (Hedw) B S G vardelicatulum delicatulum (Hedw) B S G var radicans (Kindb) Crum Steere et Anderson minutulum (Hedw) B S G pygmaeum B S G recognftum (Hedw) Lindb scitum (p Beauv) Aust
Helodium (Sull) Warnst blandowli (Web et Mohr) Warnst var blandowli blandowli (Web et Mohr) Wamst var helodioides (Ren et Card ex Roell) Crum
Steere et Anderson paludomm (Sull) AuS
61 Amblystegiaceae Cratoneuron (Sull) Spruce
commulatum (Hedw) Roth var commutfltum commutatum (Hedw) Roth var fakatum (Brid) Moenk ilicinum (Hedw) Spruce var filcinum
Campylium (Sun) Mitt chrysophyllum (Brid) 1 Lange var chrysophyllum hispidulum (Brid) Mitt var hllJidulum polygamum (B S G)C lens varpolygtlmUm stellatum (Hedw) C1ens vat stelltum
Leptodictyum (Schimp) Warnst laxirete (Card et Ther) Broth riparium (Hedw) Wamat trichopodium (Schultz) Warns l1acillans (Sull) Broth
Hygroamblystegium Loelke jlul1iatile (Hedw) oeske var luViatile
24
fluviatile (Hedw) Loeske var oWltum Grout noterophilum (Sun et Lesq ex Sull) Warnst tenox (Hedw) Jenn var tenax tenox (Hedw) Jenn var spinilolium Schimp) Jenn
Amblystegium BSG juratzkanum Schimp serpens (Hedw) BSG varlum (Hedw) Undb var varium
Platydictya Berk confervoides (Brid) Crum jungermanniodes (Brid) Crum minutissimum (Sull et Lesq ex Sull) Crum subtile (Hedw) Crum
Drepanocladus (C Muell) Roth aduncus (Hedw) Warnst var aduncus aduncus (Hedw) Warnst var capillilolius (Wamst)Riebm aduncus (Hedw) Warnst var kneiffii (BSG) Moenk aduncus (Hedw) Warnst var polycorpus (Bland ex Voit) Roth exannulatus (BSG) Warnst fluitans (Hedw) Warnst var fluitans revolvens (Sw) Wamst var revolvens sendtneri (Schimp) Warnst var sendtnerl uncinatus (Hedw) Warnst var uncinatus uncinatus ffiedw) Warnst var piwnosus (Schimp) Wamst vernicoms (Lindb ex C Hartm) Warnst
Hygrohypnum Lindb eugyrium (BSG) loeske luridum (Hedw Jenn varluridum ochraecum (Tum ex Wils) Loeske var ochraceunt
Calliergon (Sull) Kindb cordilolium Hedw Kindb glganteum (Schimp) Kindb var giganteum richardsonii (Mitt) Kindb ex Wamst stramineum (Brid) Kindb trilarium (Web et Mohr) Kindb
Calliergonella Loeske cuspidata (Hedw) Loeske
Calliergidium (Ren) Grout bakeri (Ren) Grout
Scorpidium (Schimp) Urnpr scorpiodides (Hedw) limpr
62 Brachytbeciaceae Tomenthypnum loeske
nitens (Hedw) Loeske var mtens
25
Homalotheciella (Card) Broth subcapillata (Hedw) Broth
Brachythecium BSG acuminatum (Hedw) Awt acutum (Mitt) Sullo calcareum IGndb campestTe (C MuelL) BSG cunum (Lindb) Umpr digastrum C Muell et Kindb ex Macoun et Kindb oxycladon (Brid) Jaeg et Sauerb var oxycladon plumosum (Hedw) BSG populeum (Hedw) BSG var populeum reflexum (Starke ex Web et Mohr) DsG var relexum reiexum (Starke ex Web et Mohr) DSG var pacificum Renet Card ex Roell rlvulare BSG var rivulare tabulum (Hedw) BSC var tabuum salebrosum (Web et Mohr) BSG var salebrosum velutinum (Hedw) BSG Vat velutinum
Bryhniil Kaur graminicolor (Brid) Grout var graminicolor novae-angliae (Sull et Lesq ex sun) Grout
Rhynchostegium BSG senulatum (Hedw) Ja4lg et Sauerb
Rhynchostegiella (BSG) Limpr compacta (C MueU) Loeske
EU1hynchium 8SG hians (Hedw) Sande Lac pulchellum (Hedw) Jenn var pulchelium riparloides (Hedw) Rich
Stokeriella (Kindb) Robins pmelonga (Hedw) Robins var stokesii (Tum) Crum
63 Entodontaceae Entodon C Muell
brevisetus (Hook et Wils ex Wils) Lindb cladorrhizans (Hedw) C Muell seductrlx (Hedw) C MueU var wluctrix
Pleurozium Mitt schreberl (8rid) Mitt var schreberi
64 Plagiotheciaceae Pltlglothecium BSG
CflVifolium (Brid) Iwats var CtWifolium denticulatum (Hedw) 8SG laetum BSG ltztebricola BSG
26
6S SernatophyUaceae Brotherella Loeske ex Fleisch
recumins (Michx) Fleisch tenuirostris (Bruch et Schimp ex Sull) Fleisch
Sematophyllum Mitt demissum (Wils) Mitt
66 Hypnaceae Pl(itygyrium BSG
repens (Brid) BSG
PylaisieUa Kindb intricata (Hedw) Grout polyantha (Hedw) Grout selwynii (Kindb) Crum Steere et Anderson
Homomallium (Schimp) Loeske adnatum (Hedw) Broth
Hypnum Hedw cupressiforme Hedw var cupressiforme cupressifonne Hedw varfiliforme Brid curviolium Hedw fertile Sendtn imponens Hedw lindbergii Mitt palescens (Hedw) P-Beauv pratense Koch ex Brid
Calicladium Crum haldanianum (Grev) Crum
sopterygiopsis Iwats muelleriana (Schimp) Iwats
Isopterygium Mitt e1egans (Brid) lindb pulchellum (Hedw) Jaeg et Sauerb tenerum (Sw) Mitt
Taxiphyllum Fleisch depllQtum (Bruch et Schimp ex sun) Fleisch taxirameum (MitL) Fleisch
Herzogiella Broth seligeri (Brid) Iwats striatella (Brid) Iwats ruracea (Lindh) Iwats
hilium De Not crista-castrenna (Hedw) De Not
27
67 Rhytidiaceae Rhytidium (Sull) Kindb
rugosum (Hedw) Kindb
Rhytidfodelphus (Lindb ex Iimpr) Warnst triquetrus (Hedw) Warnst var triquetrus
68 Hylocomiaceae Hylocomium BSG
pyrenaicum (Spruce) lindb splendens (Hedw) BSG varsplendens
69 Buxbaumiaceae Buxbaumia Hedw
aphyllll Hedw
Diphyscium Mohr follosum (Hedw) Mohr
70 Tetraphidaceae Tetraphi3
pellucida Hedw
71 Polytrichaceae Atrichum PmiddotBeauv
angustatum (Brid) BSG cri8pum (James) Sullo oerstedianum (C Muell) Mitt teneUum (Roehl) BSG undulatum (Hedw) P middotBeauv var unduiatum
Pogonatum P-Beauv alpinum (Hedw) Roehl var alpinum umigerum (Hedw) P-Beauv
Poiytrichum Hedw commune Hedw var commune ormosum Hedw juniperinum Hedw longisetum Brid ohioense Ren et Card piliferum Hedw var pilierum strictum Brid
28
MAPS OF BRYOPHYTE TAXA
REPORTED FOR WISCONSIN
HORNWORTS AND LIVERWORTS
29
bull bull WSHeV~N S vE II
0 ~
e~RRON ~USK
(J~ (j -
CHIPPE A DUNN
0 o0 poundAU CLAIRE
~ TREMshyPEAshyLEAU
MICE
ONEIDA
Q LINCOLN
TAYLOR
Q t-~----- MOIRA THON
CLARK
o
10
LA FAVETTE
Hornwor ts amp Liverworts
- None - 1 - 25 taxa
- 26 - 50 tRXshy
- 51 - 75 tax)
- 7c - 100 taxa
FONDOU j LAC ~
DODGE
i O JEfFERSON
WALWORTH
30
Anastrophylltlfl helleranum (Nees) Schust Anastrophyllum michauxil (We~ Buch ~
46 88 136 141 Evans 38 46 141
Anastrophyllum mlnut~ (Schreb) Schuster Aneura plnguls (L) Durn
89 21 384687 88 89 90 (138) (145)
Anthoceros macounil MA Howe ~oceros punctatus L
21 (45) (144) 34
31
Barbilophozia attenuata (Mart) Loeske Barbilophozia barbata (Schmid ~ Schre~
24 141 Loeske 28 39 46 89 90 141
Bazzania trilobata (1) S Gray
1 26 39 46 (57) (59) 79 88 89
90 119 141
BlePbarostoma trichoffiyllum (1) Dumort
1 8 26 39 46 89 90 137 141
Blasia pusilla 1
1 39 46 90
Calypogeja fissa (1) Raddi
90
32
$129geja integristipula Steph C~Ypound2geja muelleriana (Schifm) K Mull
88 89 89
CallpoundO~eja neesiana (Mass amp Carest) K Mull
39 46 89 140 (141)
~pogeja sJlapound2icola (H Arnell tl J bull
Perss) Warnst et Loeske (119) 141
CeIalozia biscuspidata (L) Dum
1 37 I 46 90 t 137 141
CalYpogej~ trichomanis (L) Corda
26 39 (119) 137
33
CeEhalozia catenulata (Hub) Lindb Cephalozia connivens (Dicks) Lindb
39 46 79 141 39 4679 88 (89) 90 119 141
Cephalozia lacinulata Jack ~ Spruce
141 Cephalozia lunulifolia (Dum~ Dum I 28 39 46 79 88 89 90 141
Cepoundhalozia macouni (Aust) Aust Cephalozia ~leniceps (Aust) Lindb 27~ 3746 (66)7987 (139)141 39 46 79 89 141
34
Ceffialoziella 9ivaricata (Sm) Schiffn Cephaloziella ~hista (Jack) Schiffn
46 90 46 (119) 141
Cephaloziella hampeana (Nees) Schiffn
46
Cephaloziella rubella (Nees) Warnst
46 39
Cephaloziella rubella var sullivantii
(Aust) K Mull 46 39
Cephaloziella rubella var bifida (Lindb) Douin
35
46
Chiloscyphus pallescens (Ehrh ~ Hof~ Dum Chiloscyphus pallescens var fragilis
39J 46 88 89J 90 (Roth) K Milll 46
Chiloscyphus E9lyanthos (L) Corda
1 24 39 46
Chiloscyphus polyanthos vax rivularis
(Schrad) Nees 39 46
poundpoundnocepha1um conicum (1) Lindb
12 28 39 46 (49) 87 88 8990 120 121 148
Cololejeunea biddlecomiae (Aust) Evans
39 46 62 87 88 89 90
36
Dipl0]hyllum apicu1atum (Evans) Steph Fossombronia foveo1ata Lindb
(35)41 46 90 140 1 t 38 (39) t 46
Frullania bolanderi Aust
13 46 62 88 90 139 140
Frullanla ari ttoniae Evans
39 46 69 (71) 87 88 89 90
Frullania eboracensis Gott
1 9 28 4 39 46 (57) (59) 69 70 (71) 87 88 89 90 92 93 97 148
Frullania inflata Gott
46 87 88 89 (139)
Frullania oakesiana Aust Frullania plana Sullo
(35) 42 46 87 (139) (140) 92
Frullania ri paria Hampe ~ Lehm
46 69 71 87 89
Frullania ~elwynianna Pears
46 89
Geocalyx SEaveolens (Schrad) Nees
394688 8990 (119) 140
Frullania tamarisci (L) Dum subsp sPagrayana
(Mont) Hatt 28 34 l~6 t (57) 88 89 90
38
Mar pan thus seutatUB ( Web amp Mohr) Spruce Jamesoniella autumnalis (DC) Steph
(35) 39 46 90 1 7 28 39 46 56 (57) 87 8889 90 (119) 137 141
Jungermannia confertissima Nees
137 141
Jungermannia crenuliform1s Aust
79 137 141
Jungermannia ~il1ima Sm
46 90 141 Jungermannia exsert1folia subsp corifo~ (Dum~
Vana (35) 46 79 141
39
Jungermannia hyalina Lyell Jungerrr6nnia 1eiantha Grolle
40 46 79 136 141 39 46 79 88 90 1)7 141
Jungermannia IXlaxis Lindb Jungermannia pumila With
46 67 79 (139) (140) 141 )9 46 90 1)6 141
Jungermannia sphaerocarpa Hook Kurzia setacea (Web) Grolle
4679136140IIH 136
Lejeunea cavifo1ia (Ehrh) Lindb Lepidozia reptans (1) Dum
1 7 27 39 46 88 89 90 137 14010 I ( 27) 46 I 69 (71) 88
lophocolea heterophylla (Schrad) Dum Lophocolea minor Nees
1 )4 39 46 (57) 87 88 89 27 39 46 88 89 90 (119) 90 (93)94 119 136
Lophozia al~stris (Schleich ~ Web) Evans Lophozia ascendens (Warnst) Schust
46 90 137 141 136 140 141
41
loIilozia badensls (Gatt ~ Rabenh)
46 141
lophozia blcrenata (Schmid sectA Hoffm)
Dum 46 141
lophozia collaris (Nees) Dum
39 46 141
lophozla excisa (Dicks) Dum
27 4679 141
lophozia gillmani (Aust) Schust lophozia gIlttulata (Undo tl H Arnell)
36 39 40 46 136 (140) 141 Evans 39 46 136 141
42
141
Lophozia heterocolEDs (Thed) MA Howe IoPhozia ipcisa (Schard) Dum
27 39 46 89 141 8 27 31 46 79 90 140 141
LoQhozia longidens Macoun Lophozia ventricosa
1 46 79 90 141
Lophozia ventricosa var silvicola
(Buch) Jones 141 LoEhozla ventricosa vax longif1ora (Nees)
Macoun 39 46 79 141
43
~~ fragrans (Balbis) Frye et Clark Mannia pi 10sa (Hornem) Frye et Clark
46 (139) (140) 9 46 79 116 139
Mannia triandra (Scop) Gro 11e
46 (140)
Marchantia pound91ymorpha L
1 13 28 3946 (49) 78 87 88 89 90 (Ill) 121 122 147
Mo~rckia hibernica (HookJ Gott
46 89
Metzgeria conjugata Lindb
44
46
Mylia anomala (Hook) S Gray Notothylas orbicularis (Schwein) Sullo
39 46 141 (35)45 46
Nowellia curvifolia (Dicks) Mitt
17 27 3846 88 89 90 141
Odontoschisma denudatuJI1 (Nees ~ Mart)
Dum 46 81 137 141
Pallavicinia lyellii (Hook~ Carruth
90
Odontoschisma macounii (Aust) Under
45 46 (66) (139) 140 141
45
=
Pellia epiphllla (L) CordaPellia endiviifolia (Dicks~ Dum
27 39 46 90 13724 39 46
Pellia neesiana Limp bull Phaeoceros
39 46 89 27 39 45 46 97
Porella pinnata L
13 27 I 46 90 Plagiochila asplenioides (L) Dum
27 I 39 1+6 88 89 I 90 137
46
~e11a platYEhylla (L) Pfeiff Porella platyphylloi~ (Schwein) Lindb
17 27 39 46 69 (71)79 87 88 89 9 34 46 (57) 59 87 88 89 92 90 92 93 77
Preissia quadrata (Seop) Nees
1 36 3946 (71) 88 89 90 121 123
137
Ptllidium ciliare (L) Hampe
1 27 39 46 132 141
Dum
(22) 28 3946 (57) 87 88 89 90 92
Ptilidium ~lcherrium (G Web) Hampe
(9) 13 394656 (57)70 87 88 89 90 93 (119) 136 141
47
Radula obconica Sullo Reboulia hemlsphaerlca (L) Raddi
39 46 88 89 (43) 46 (59) 70 90 t Cf 124 125 (139)
Riccardia ~atifrons Lindb
1 27 39 46 89
Riccardia multifi~ (L) S Gray
39 46 88 145
Riccia beyrichiana Hampe ~ Lehm
46 Riccardia palmata (Hedw) Carruth
39 46 90
48
Riccia bifurca Hoffm Riccia cavernosa Hoffm
48 13739 46 79 137
Riccia fluitans L
242731454679
Riccla frostli Aust
48 79
Riccia sullicantii Aust
79 (137) Riccia sorocarpa Bisch 46 (137)
49
Ricciocarpus natans (1) Corda Sca~ia apiculata Spruce
13 27 46 79 89 110 39 46 (139) 141
ScaJltll1ia carinthica Jack ~ Lindb
137 t 141
Scapania ~ (Mart) Durn
39467990137141
Scapania digenil Schiffn
90
ScaEsectflia cuspiduligera (Nees) K Mull
46 (139) (140) (141)
50
Scapania glaucoeerha1a (Tayl) Aust Seapania gyrrmostomophilia Kaal
39 46 (139) 141 41 44 46 64 (139) (140) 141
ScaP3poundlia irrigua Gott ~ almiddot
7 39 46 79 137 (140L 141
Seapania nemorosa (L) Dum
1 39 46 89 90 137
Scapmi~ ~~ta Smiddotlch
137 141 51
~caPnia poundludicola LDeske amp K Mull Scapania saxicola Schust
46 (66) 141 136 141
Sc~~ scandica (H Arnell ~ Buch) Macv
139 140
Scapania subalpina (Nees) Durn
39 40 46 79 137 141
Scapania undulata (L) Dum
1 42 46 79 141
Scapania umbrosa (Schrad) Dum
137 140 141
52
Trichocolea tomentella (Ehrh) Dum Triomarla exsecta (Schmid) Schiffn
27 (35) 39 46 47 88 137 141 8 28 39 46 90 140 141
Tritomaria exsectiformis (Breidl) Loe~ke Tritomaria Sluinguedentata (Huds) Buch
39 -t6 89 90 136 140 141 7 39 46 141
Trltomaria scitula (Tayl) Joerg
46 136 (139) 141
53
Pterlgy1lllndrum Hedw filiforme Hedw
60 Thuidiaceae Haplohymenfum Dozy et MoIk
trlste (Ces ex De Not) Kiodb
Anomodon Hook et Tayl attenuatus (Hedw) Hueb minor (Hedw) Fuemr rostratus (Hedw) Schimp viticulosus (Hedw) Hook et Tayl
Haplocladium (C Mucn) C MueU microphyllum (Hedw) Broth virginillnum (Brld) Broth ~bull
Thuidium B S G abietinum (Hedw) B S G delicatulum (Hedw) B S G vardelicatulum delicatulum (Hedw) B S G var radicans (Kindb) Crum Steere et Anderson minutulum (Hedw) B S G pygmaeum B S G recognftum (Hedw) Lindb scitum (p Beauv) Aust
Helodium (Sull) Warnst blandowli (Web et Mohr) Warnst var blandowli blandowli (Web et Mohr) Wamst var helodioides (Ren et Card ex Roell) Crum
Steere et Anderson paludomm (Sull) AuS
61 Amblystegiaceae Cratoneuron (Sull) Spruce
commulatum (Hedw) Roth var commutfltum commutatum (Hedw) Roth var fakatum (Brid) Moenk ilicinum (Hedw) Spruce var filcinum
Campylium (Sun) Mitt chrysophyllum (Brid) 1 Lange var chrysophyllum hispidulum (Brid) Mitt var hllJidulum polygamum (B S G)C lens varpolygtlmUm stellatum (Hedw) C1ens vat stelltum
Leptodictyum (Schimp) Warnst laxirete (Card et Ther) Broth riparium (Hedw) Wamat trichopodium (Schultz) Warns l1acillans (Sull) Broth
Hygroamblystegium Loelke jlul1iatile (Hedw) oeske var luViatile
24
fluviatile (Hedw) Loeske var oWltum Grout noterophilum (Sun et Lesq ex Sull) Warnst tenox (Hedw) Jenn var tenax tenox (Hedw) Jenn var spinilolium Schimp) Jenn
Amblystegium BSG juratzkanum Schimp serpens (Hedw) BSG varlum (Hedw) Undb var varium
Platydictya Berk confervoides (Brid) Crum jungermanniodes (Brid) Crum minutissimum (Sull et Lesq ex Sull) Crum subtile (Hedw) Crum
Drepanocladus (C Muell) Roth aduncus (Hedw) Warnst var aduncus aduncus (Hedw) Warnst var capillilolius (Wamst)Riebm aduncus (Hedw) Warnst var kneiffii (BSG) Moenk aduncus (Hedw) Warnst var polycorpus (Bland ex Voit) Roth exannulatus (BSG) Warnst fluitans (Hedw) Warnst var fluitans revolvens (Sw) Wamst var revolvens sendtneri (Schimp) Warnst var sendtnerl uncinatus (Hedw) Warnst var uncinatus uncinatus ffiedw) Warnst var piwnosus (Schimp) Wamst vernicoms (Lindb ex C Hartm) Warnst
Hygrohypnum Lindb eugyrium (BSG) loeske luridum (Hedw Jenn varluridum ochraecum (Tum ex Wils) Loeske var ochraceunt
Calliergon (Sull) Kindb cordilolium Hedw Kindb glganteum (Schimp) Kindb var giganteum richardsonii (Mitt) Kindb ex Wamst stramineum (Brid) Kindb trilarium (Web et Mohr) Kindb
Calliergonella Loeske cuspidata (Hedw) Loeske
Calliergidium (Ren) Grout bakeri (Ren) Grout
Scorpidium (Schimp) Urnpr scorpiodides (Hedw) limpr
62 Brachytbeciaceae Tomenthypnum loeske
nitens (Hedw) Loeske var mtens
25
Homalotheciella (Card) Broth subcapillata (Hedw) Broth
Brachythecium BSG acuminatum (Hedw) Awt acutum (Mitt) Sullo calcareum IGndb campestTe (C MuelL) BSG cunum (Lindb) Umpr digastrum C Muell et Kindb ex Macoun et Kindb oxycladon (Brid) Jaeg et Sauerb var oxycladon plumosum (Hedw) BSG populeum (Hedw) BSG var populeum reflexum (Starke ex Web et Mohr) DsG var relexum reiexum (Starke ex Web et Mohr) DSG var pacificum Renet Card ex Roell rlvulare BSG var rivulare tabulum (Hedw) BSC var tabuum salebrosum (Web et Mohr) BSG var salebrosum velutinum (Hedw) BSG Vat velutinum
Bryhniil Kaur graminicolor (Brid) Grout var graminicolor novae-angliae (Sull et Lesq ex sun) Grout
Rhynchostegium BSG senulatum (Hedw) Ja4lg et Sauerb
Rhynchostegiella (BSG) Limpr compacta (C MueU) Loeske
EU1hynchium 8SG hians (Hedw) Sande Lac pulchellum (Hedw) Jenn var pulchelium riparloides (Hedw) Rich
Stokeriella (Kindb) Robins pmelonga (Hedw) Robins var stokesii (Tum) Crum
63 Entodontaceae Entodon C Muell
brevisetus (Hook et Wils ex Wils) Lindb cladorrhizans (Hedw) C Muell seductrlx (Hedw) C MueU var wluctrix
Pleurozium Mitt schreberl (8rid) Mitt var schreberi
64 Plagiotheciaceae Pltlglothecium BSG
CflVifolium (Brid) Iwats var CtWifolium denticulatum (Hedw) 8SG laetum BSG ltztebricola BSG
26
6S SernatophyUaceae Brotherella Loeske ex Fleisch
recumins (Michx) Fleisch tenuirostris (Bruch et Schimp ex Sull) Fleisch
Sematophyllum Mitt demissum (Wils) Mitt
66 Hypnaceae Pl(itygyrium BSG
repens (Brid) BSG
PylaisieUa Kindb intricata (Hedw) Grout polyantha (Hedw) Grout selwynii (Kindb) Crum Steere et Anderson
Homomallium (Schimp) Loeske adnatum (Hedw) Broth
Hypnum Hedw cupressiforme Hedw var cupressiforme cupressifonne Hedw varfiliforme Brid curviolium Hedw fertile Sendtn imponens Hedw lindbergii Mitt palescens (Hedw) P-Beauv pratense Koch ex Brid
Calicladium Crum haldanianum (Grev) Crum
sopterygiopsis Iwats muelleriana (Schimp) Iwats
Isopterygium Mitt e1egans (Brid) lindb pulchellum (Hedw) Jaeg et Sauerb tenerum (Sw) Mitt
Taxiphyllum Fleisch depllQtum (Bruch et Schimp ex sun) Fleisch taxirameum (MitL) Fleisch
Herzogiella Broth seligeri (Brid) Iwats striatella (Brid) Iwats ruracea (Lindh) Iwats
hilium De Not crista-castrenna (Hedw) De Not
27
67 Rhytidiaceae Rhytidium (Sull) Kindb
rugosum (Hedw) Kindb
Rhytidfodelphus (Lindb ex Iimpr) Warnst triquetrus (Hedw) Warnst var triquetrus
68 Hylocomiaceae Hylocomium BSG
pyrenaicum (Spruce) lindb splendens (Hedw) BSG varsplendens
69 Buxbaumiaceae Buxbaumia Hedw
aphyllll Hedw
Diphyscium Mohr follosum (Hedw) Mohr
70 Tetraphidaceae Tetraphi3
pellucida Hedw
71 Polytrichaceae Atrichum PmiddotBeauv
angustatum (Brid) BSG cri8pum (James) Sullo oerstedianum (C Muell) Mitt teneUum (Roehl) BSG undulatum (Hedw) P middotBeauv var unduiatum
Pogonatum P-Beauv alpinum (Hedw) Roehl var alpinum umigerum (Hedw) P-Beauv
Poiytrichum Hedw commune Hedw var commune ormosum Hedw juniperinum Hedw longisetum Brid ohioense Ren et Card piliferum Hedw var pilierum strictum Brid
28
MAPS OF BRYOPHYTE TAXA
REPORTED FOR WISCONSIN
HORNWORTS AND LIVERWORTS
29
bull bull WSHeV~N S vE II
0 ~
e~RRON ~USK
(J~ (j -
CHIPPE A DUNN
0 o0 poundAU CLAIRE
~ TREMshyPEAshyLEAU
MICE
ONEIDA
Q LINCOLN
TAYLOR
Q t-~----- MOIRA THON
CLARK
o
10
LA FAVETTE
Hornwor ts amp Liverworts
- None - 1 - 25 taxa
- 26 - 50 tRXshy
- 51 - 75 tax)
- 7c - 100 taxa
FONDOU j LAC ~
DODGE
i O JEfFERSON
WALWORTH
30
Anastrophylltlfl helleranum (Nees) Schust Anastrophyllum michauxil (We~ Buch ~
46 88 136 141 Evans 38 46 141
Anastrophyllum mlnut~ (Schreb) Schuster Aneura plnguls (L) Durn
89 21 384687 88 89 90 (138) (145)
Anthoceros macounil MA Howe ~oceros punctatus L
21 (45) (144) 34
31
Barbilophozia attenuata (Mart) Loeske Barbilophozia barbata (Schmid ~ Schre~
24 141 Loeske 28 39 46 89 90 141
Bazzania trilobata (1) S Gray
1 26 39 46 (57) (59) 79 88 89
90 119 141
BlePbarostoma trichoffiyllum (1) Dumort
1 8 26 39 46 89 90 137 141
Blasia pusilla 1
1 39 46 90
Calypogeja fissa (1) Raddi
90
32
$129geja integristipula Steph C~Ypound2geja muelleriana (Schifm) K Mull
88 89 89
CallpoundO~eja neesiana (Mass amp Carest) K Mull
39 46 89 140 (141)
~pogeja sJlapound2icola (H Arnell tl J bull
Perss) Warnst et Loeske (119) 141
CeIalozia biscuspidata (L) Dum
1 37 I 46 90 t 137 141
CalYpogej~ trichomanis (L) Corda
26 39 (119) 137
33
CeEhalozia catenulata (Hub) Lindb Cephalozia connivens (Dicks) Lindb
39 46 79 141 39 4679 88 (89) 90 119 141
Cephalozia lacinulata Jack ~ Spruce
141 Cephalozia lunulifolia (Dum~ Dum I 28 39 46 79 88 89 90 141
Cepoundhalozia macouni (Aust) Aust Cephalozia ~leniceps (Aust) Lindb 27~ 3746 (66)7987 (139)141 39 46 79 89 141
34
Ceffialoziella 9ivaricata (Sm) Schiffn Cephaloziella ~hista (Jack) Schiffn
46 90 46 (119) 141
Cephaloziella hampeana (Nees) Schiffn
46
Cephaloziella rubella (Nees) Warnst
46 39
Cephaloziella rubella var sullivantii
(Aust) K Mull 46 39
Cephaloziella rubella var bifida (Lindb) Douin
35
46
Chiloscyphus pallescens (Ehrh ~ Hof~ Dum Chiloscyphus pallescens var fragilis
39J 46 88 89J 90 (Roth) K Milll 46
Chiloscyphus E9lyanthos (L) Corda
1 24 39 46
Chiloscyphus polyanthos vax rivularis
(Schrad) Nees 39 46
poundpoundnocepha1um conicum (1) Lindb
12 28 39 46 (49) 87 88 8990 120 121 148
Cololejeunea biddlecomiae (Aust) Evans
39 46 62 87 88 89 90
36
Dipl0]hyllum apicu1atum (Evans) Steph Fossombronia foveo1ata Lindb
(35)41 46 90 140 1 t 38 (39) t 46
Frullania bolanderi Aust
13 46 62 88 90 139 140
Frullanla ari ttoniae Evans
39 46 69 (71) 87 88 89 90
Frullania eboracensis Gott
1 9 28 4 39 46 (57) (59) 69 70 (71) 87 88 89 90 92 93 97 148
Frullania inflata Gott
46 87 88 89 (139)
Frullania oakesiana Aust Frullania plana Sullo
(35) 42 46 87 (139) (140) 92
Frullania ri paria Hampe ~ Lehm
46 69 71 87 89
Frullania ~elwynianna Pears
46 89
Geocalyx SEaveolens (Schrad) Nees
394688 8990 (119) 140
Frullania tamarisci (L) Dum subsp sPagrayana
(Mont) Hatt 28 34 l~6 t (57) 88 89 90
38
Mar pan thus seutatUB ( Web amp Mohr) Spruce Jamesoniella autumnalis (DC) Steph
(35) 39 46 90 1 7 28 39 46 56 (57) 87 8889 90 (119) 137 141
Jungermannia confertissima Nees
137 141
Jungermannia crenuliform1s Aust
79 137 141
Jungermannia ~il1ima Sm
46 90 141 Jungermannia exsert1folia subsp corifo~ (Dum~
Vana (35) 46 79 141
39
Jungermannia hyalina Lyell Jungerrr6nnia 1eiantha Grolle
40 46 79 136 141 39 46 79 88 90 1)7 141
Jungermannia IXlaxis Lindb Jungermannia pumila With
46 67 79 (139) (140) 141 )9 46 90 1)6 141
Jungermannia sphaerocarpa Hook Kurzia setacea (Web) Grolle
4679136140IIH 136
Lejeunea cavifo1ia (Ehrh) Lindb Lepidozia reptans (1) Dum
1 7 27 39 46 88 89 90 137 14010 I ( 27) 46 I 69 (71) 88
lophocolea heterophylla (Schrad) Dum Lophocolea minor Nees
1 )4 39 46 (57) 87 88 89 27 39 46 88 89 90 (119) 90 (93)94 119 136
Lophozia al~stris (Schleich ~ Web) Evans Lophozia ascendens (Warnst) Schust
46 90 137 141 136 140 141
41
loIilozia badensls (Gatt ~ Rabenh)
46 141
lophozia blcrenata (Schmid sectA Hoffm)
Dum 46 141
lophozia collaris (Nees) Dum
39 46 141
lophozla excisa (Dicks) Dum
27 4679 141
lophozia gillmani (Aust) Schust lophozia gIlttulata (Undo tl H Arnell)
36 39 40 46 136 (140) 141 Evans 39 46 136 141
42
141
Lophozia heterocolEDs (Thed) MA Howe IoPhozia ipcisa (Schard) Dum
27 39 46 89 141 8 27 31 46 79 90 140 141
LoQhozia longidens Macoun Lophozia ventricosa
1 46 79 90 141
Lophozia ventricosa var silvicola
(Buch) Jones 141 LoEhozla ventricosa vax longif1ora (Nees)
Macoun 39 46 79 141
43
~~ fragrans (Balbis) Frye et Clark Mannia pi 10sa (Hornem) Frye et Clark
46 (139) (140) 9 46 79 116 139
Mannia triandra (Scop) Gro 11e
46 (140)
Marchantia pound91ymorpha L
1 13 28 3946 (49) 78 87 88 89 90 (Ill) 121 122 147
Mo~rckia hibernica (HookJ Gott
46 89
Metzgeria conjugata Lindb
44
46
Mylia anomala (Hook) S Gray Notothylas orbicularis (Schwein) Sullo
39 46 141 (35)45 46
Nowellia curvifolia (Dicks) Mitt
17 27 3846 88 89 90 141
Odontoschisma denudatuJI1 (Nees ~ Mart)
Dum 46 81 137 141
Pallavicinia lyellii (Hook~ Carruth
90
Odontoschisma macounii (Aust) Under
45 46 (66) (139) 140 141
45
=
Pellia epiphllla (L) CordaPellia endiviifolia (Dicks~ Dum
27 39 46 90 13724 39 46
Pellia neesiana Limp bull Phaeoceros
39 46 89 27 39 45 46 97
Porella pinnata L
13 27 I 46 90 Plagiochila asplenioides (L) Dum
27 I 39 1+6 88 89 I 90 137
46
~e11a platYEhylla (L) Pfeiff Porella platyphylloi~ (Schwein) Lindb
17 27 39 46 69 (71)79 87 88 89 9 34 46 (57) 59 87 88 89 92 90 92 93 77
Preissia quadrata (Seop) Nees
1 36 3946 (71) 88 89 90 121 123
137
Ptllidium ciliare (L) Hampe
1 27 39 46 132 141
Dum
(22) 28 3946 (57) 87 88 89 90 92
Ptilidium ~lcherrium (G Web) Hampe
(9) 13 394656 (57)70 87 88 89 90 93 (119) 136 141
47
Radula obconica Sullo Reboulia hemlsphaerlca (L) Raddi
39 46 88 89 (43) 46 (59) 70 90 t Cf 124 125 (139)
Riccardia ~atifrons Lindb
1 27 39 46 89
Riccardia multifi~ (L) S Gray
39 46 88 145
Riccia beyrichiana Hampe ~ Lehm
46 Riccardia palmata (Hedw) Carruth
39 46 90
48
Riccia bifurca Hoffm Riccia cavernosa Hoffm
48 13739 46 79 137
Riccia fluitans L
242731454679
Riccla frostli Aust
48 79
Riccia sullicantii Aust
79 (137) Riccia sorocarpa Bisch 46 (137)
49
Ricciocarpus natans (1) Corda Sca~ia apiculata Spruce
13 27 46 79 89 110 39 46 (139) 141
ScaJltll1ia carinthica Jack ~ Lindb
137 t 141
Scapania ~ (Mart) Durn
39467990137141
Scapania digenil Schiffn
90
ScaEsectflia cuspiduligera (Nees) K Mull
46 (139) (140) (141)
50
Scapania glaucoeerha1a (Tayl) Aust Seapania gyrrmostomophilia Kaal
39 46 (139) 141 41 44 46 64 (139) (140) 141
ScaP3poundlia irrigua Gott ~ almiddot
7 39 46 79 137 (140L 141
Seapania nemorosa (L) Dum
1 39 46 89 90 137
Scapmi~ ~~ta Smiddotlch
137 141 51
~caPnia poundludicola LDeske amp K Mull Scapania saxicola Schust
46 (66) 141 136 141
Sc~~ scandica (H Arnell ~ Buch) Macv
139 140
Scapania subalpina (Nees) Durn
39 40 46 79 137 141
Scapania undulata (L) Dum
1 42 46 79 141
Scapania umbrosa (Schrad) Dum
137 140 141
52
Trichocolea tomentella (Ehrh) Dum Triomarla exsecta (Schmid) Schiffn
27 (35) 39 46 47 88 137 141 8 28 39 46 90 140 141
Tritomaria exsectiformis (Breidl) Loe~ke Tritomaria Sluinguedentata (Huds) Buch
39 -t6 89 90 136 140 141 7 39 46 141
Trltomaria scitula (Tayl) Joerg
46 136 (139) 141
53
fluviatile (Hedw) Loeske var oWltum Grout noterophilum (Sun et Lesq ex Sull) Warnst tenox (Hedw) Jenn var tenax tenox (Hedw) Jenn var spinilolium Schimp) Jenn
Amblystegium BSG juratzkanum Schimp serpens (Hedw) BSG varlum (Hedw) Undb var varium
Platydictya Berk confervoides (Brid) Crum jungermanniodes (Brid) Crum minutissimum (Sull et Lesq ex Sull) Crum subtile (Hedw) Crum
Drepanocladus (C Muell) Roth aduncus (Hedw) Warnst var aduncus aduncus (Hedw) Warnst var capillilolius (Wamst)Riebm aduncus (Hedw) Warnst var kneiffii (BSG) Moenk aduncus (Hedw) Warnst var polycorpus (Bland ex Voit) Roth exannulatus (BSG) Warnst fluitans (Hedw) Warnst var fluitans revolvens (Sw) Wamst var revolvens sendtneri (Schimp) Warnst var sendtnerl uncinatus (Hedw) Warnst var uncinatus uncinatus ffiedw) Warnst var piwnosus (Schimp) Wamst vernicoms (Lindb ex C Hartm) Warnst
Hygrohypnum Lindb eugyrium (BSG) loeske luridum (Hedw Jenn varluridum ochraecum (Tum ex Wils) Loeske var ochraceunt
Calliergon (Sull) Kindb cordilolium Hedw Kindb glganteum (Schimp) Kindb var giganteum richardsonii (Mitt) Kindb ex Wamst stramineum (Brid) Kindb trilarium (Web et Mohr) Kindb
Calliergonella Loeske cuspidata (Hedw) Loeske
Calliergidium (Ren) Grout bakeri (Ren) Grout
Scorpidium (Schimp) Urnpr scorpiodides (Hedw) limpr
62 Brachytbeciaceae Tomenthypnum loeske
nitens (Hedw) Loeske var mtens
25
Homalotheciella (Card) Broth subcapillata (Hedw) Broth
Brachythecium BSG acuminatum (Hedw) Awt acutum (Mitt) Sullo calcareum IGndb campestTe (C MuelL) BSG cunum (Lindb) Umpr digastrum C Muell et Kindb ex Macoun et Kindb oxycladon (Brid) Jaeg et Sauerb var oxycladon plumosum (Hedw) BSG populeum (Hedw) BSG var populeum reflexum (Starke ex Web et Mohr) DsG var relexum reiexum (Starke ex Web et Mohr) DSG var pacificum Renet Card ex Roell rlvulare BSG var rivulare tabulum (Hedw) BSC var tabuum salebrosum (Web et Mohr) BSG var salebrosum velutinum (Hedw) BSG Vat velutinum
Bryhniil Kaur graminicolor (Brid) Grout var graminicolor novae-angliae (Sull et Lesq ex sun) Grout
Rhynchostegium BSG senulatum (Hedw) Ja4lg et Sauerb
Rhynchostegiella (BSG) Limpr compacta (C MueU) Loeske
EU1hynchium 8SG hians (Hedw) Sande Lac pulchellum (Hedw) Jenn var pulchelium riparloides (Hedw) Rich
Stokeriella (Kindb) Robins pmelonga (Hedw) Robins var stokesii (Tum) Crum
63 Entodontaceae Entodon C Muell
brevisetus (Hook et Wils ex Wils) Lindb cladorrhizans (Hedw) C Muell seductrlx (Hedw) C MueU var wluctrix
Pleurozium Mitt schreberl (8rid) Mitt var schreberi
64 Plagiotheciaceae Pltlglothecium BSG
CflVifolium (Brid) Iwats var CtWifolium denticulatum (Hedw) 8SG laetum BSG ltztebricola BSG
26
6S SernatophyUaceae Brotherella Loeske ex Fleisch
recumins (Michx) Fleisch tenuirostris (Bruch et Schimp ex Sull) Fleisch
Sematophyllum Mitt demissum (Wils) Mitt
66 Hypnaceae Pl(itygyrium BSG
repens (Brid) BSG
PylaisieUa Kindb intricata (Hedw) Grout polyantha (Hedw) Grout selwynii (Kindb) Crum Steere et Anderson
Homomallium (Schimp) Loeske adnatum (Hedw) Broth
Hypnum Hedw cupressiforme Hedw var cupressiforme cupressifonne Hedw varfiliforme Brid curviolium Hedw fertile Sendtn imponens Hedw lindbergii Mitt palescens (Hedw) P-Beauv pratense Koch ex Brid
Calicladium Crum haldanianum (Grev) Crum
sopterygiopsis Iwats muelleriana (Schimp) Iwats
Isopterygium Mitt e1egans (Brid) lindb pulchellum (Hedw) Jaeg et Sauerb tenerum (Sw) Mitt
Taxiphyllum Fleisch depllQtum (Bruch et Schimp ex sun) Fleisch taxirameum (MitL) Fleisch
Herzogiella Broth seligeri (Brid) Iwats striatella (Brid) Iwats ruracea (Lindh) Iwats
hilium De Not crista-castrenna (Hedw) De Not
27
67 Rhytidiaceae Rhytidium (Sull) Kindb
rugosum (Hedw) Kindb
Rhytidfodelphus (Lindb ex Iimpr) Warnst triquetrus (Hedw) Warnst var triquetrus
68 Hylocomiaceae Hylocomium BSG
pyrenaicum (Spruce) lindb splendens (Hedw) BSG varsplendens
69 Buxbaumiaceae Buxbaumia Hedw
aphyllll Hedw
Diphyscium Mohr follosum (Hedw) Mohr
70 Tetraphidaceae Tetraphi3
pellucida Hedw
71 Polytrichaceae Atrichum PmiddotBeauv
angustatum (Brid) BSG cri8pum (James) Sullo oerstedianum (C Muell) Mitt teneUum (Roehl) BSG undulatum (Hedw) P middotBeauv var unduiatum
Pogonatum P-Beauv alpinum (Hedw) Roehl var alpinum umigerum (Hedw) P-Beauv
Poiytrichum Hedw commune Hedw var commune ormosum Hedw juniperinum Hedw longisetum Brid ohioense Ren et Card piliferum Hedw var pilierum strictum Brid
28
MAPS OF BRYOPHYTE TAXA
REPORTED FOR WISCONSIN
HORNWORTS AND LIVERWORTS
29
bull bull WSHeV~N S vE II
0 ~
e~RRON ~USK
(J~ (j -
CHIPPE A DUNN
0 o0 poundAU CLAIRE
~ TREMshyPEAshyLEAU
MICE
ONEIDA
Q LINCOLN
TAYLOR
Q t-~----- MOIRA THON
CLARK
o
10
LA FAVETTE
Hornwor ts amp Liverworts
- None - 1 - 25 taxa
- 26 - 50 tRXshy
- 51 - 75 tax)
- 7c - 100 taxa
FONDOU j LAC ~
DODGE
i O JEfFERSON
WALWORTH
30
Anastrophylltlfl helleranum (Nees) Schust Anastrophyllum michauxil (We~ Buch ~
46 88 136 141 Evans 38 46 141
Anastrophyllum mlnut~ (Schreb) Schuster Aneura plnguls (L) Durn
89 21 384687 88 89 90 (138) (145)
Anthoceros macounil MA Howe ~oceros punctatus L
21 (45) (144) 34
31
Barbilophozia attenuata (Mart) Loeske Barbilophozia barbata (Schmid ~ Schre~
24 141 Loeske 28 39 46 89 90 141
Bazzania trilobata (1) S Gray
1 26 39 46 (57) (59) 79 88 89
90 119 141
BlePbarostoma trichoffiyllum (1) Dumort
1 8 26 39 46 89 90 137 141
Blasia pusilla 1
1 39 46 90
Calypogeja fissa (1) Raddi
90
32
$129geja integristipula Steph C~Ypound2geja muelleriana (Schifm) K Mull
88 89 89
CallpoundO~eja neesiana (Mass amp Carest) K Mull
39 46 89 140 (141)
~pogeja sJlapound2icola (H Arnell tl J bull
Perss) Warnst et Loeske (119) 141
CeIalozia biscuspidata (L) Dum
1 37 I 46 90 t 137 141
CalYpogej~ trichomanis (L) Corda
26 39 (119) 137
33
CeEhalozia catenulata (Hub) Lindb Cephalozia connivens (Dicks) Lindb
39 46 79 141 39 4679 88 (89) 90 119 141
Cephalozia lacinulata Jack ~ Spruce
141 Cephalozia lunulifolia (Dum~ Dum I 28 39 46 79 88 89 90 141
Cepoundhalozia macouni (Aust) Aust Cephalozia ~leniceps (Aust) Lindb 27~ 3746 (66)7987 (139)141 39 46 79 89 141
34
Ceffialoziella 9ivaricata (Sm) Schiffn Cephaloziella ~hista (Jack) Schiffn
46 90 46 (119) 141
Cephaloziella hampeana (Nees) Schiffn
46
Cephaloziella rubella (Nees) Warnst
46 39
Cephaloziella rubella var sullivantii
(Aust) K Mull 46 39
Cephaloziella rubella var bifida (Lindb) Douin
35
46
Chiloscyphus pallescens (Ehrh ~ Hof~ Dum Chiloscyphus pallescens var fragilis
39J 46 88 89J 90 (Roth) K Milll 46
Chiloscyphus E9lyanthos (L) Corda
1 24 39 46
Chiloscyphus polyanthos vax rivularis
(Schrad) Nees 39 46
poundpoundnocepha1um conicum (1) Lindb
12 28 39 46 (49) 87 88 8990 120 121 148
Cololejeunea biddlecomiae (Aust) Evans
39 46 62 87 88 89 90
36
Dipl0]hyllum apicu1atum (Evans) Steph Fossombronia foveo1ata Lindb
(35)41 46 90 140 1 t 38 (39) t 46
Frullania bolanderi Aust
13 46 62 88 90 139 140
Frullanla ari ttoniae Evans
39 46 69 (71) 87 88 89 90
Frullania eboracensis Gott
1 9 28 4 39 46 (57) (59) 69 70 (71) 87 88 89 90 92 93 97 148
Frullania inflata Gott
46 87 88 89 (139)
Frullania oakesiana Aust Frullania plana Sullo
(35) 42 46 87 (139) (140) 92
Frullania ri paria Hampe ~ Lehm
46 69 71 87 89
Frullania ~elwynianna Pears
46 89
Geocalyx SEaveolens (Schrad) Nees
394688 8990 (119) 140
Frullania tamarisci (L) Dum subsp sPagrayana
(Mont) Hatt 28 34 l~6 t (57) 88 89 90
38
Mar pan thus seutatUB ( Web amp Mohr) Spruce Jamesoniella autumnalis (DC) Steph
(35) 39 46 90 1 7 28 39 46 56 (57) 87 8889 90 (119) 137 141
Jungermannia confertissima Nees
137 141
Jungermannia crenuliform1s Aust
79 137 141
Jungermannia ~il1ima Sm
46 90 141 Jungermannia exsert1folia subsp corifo~ (Dum~
Vana (35) 46 79 141
39
Jungermannia hyalina Lyell Jungerrr6nnia 1eiantha Grolle
40 46 79 136 141 39 46 79 88 90 1)7 141
Jungermannia IXlaxis Lindb Jungermannia pumila With
46 67 79 (139) (140) 141 )9 46 90 1)6 141
Jungermannia sphaerocarpa Hook Kurzia setacea (Web) Grolle
4679136140IIH 136
Lejeunea cavifo1ia (Ehrh) Lindb Lepidozia reptans (1) Dum
1 7 27 39 46 88 89 90 137 14010 I ( 27) 46 I 69 (71) 88
lophocolea heterophylla (Schrad) Dum Lophocolea minor Nees
1 )4 39 46 (57) 87 88 89 27 39 46 88 89 90 (119) 90 (93)94 119 136
Lophozia al~stris (Schleich ~ Web) Evans Lophozia ascendens (Warnst) Schust
46 90 137 141 136 140 141
41
loIilozia badensls (Gatt ~ Rabenh)
46 141
lophozia blcrenata (Schmid sectA Hoffm)
Dum 46 141
lophozia collaris (Nees) Dum
39 46 141
lophozla excisa (Dicks) Dum
27 4679 141
lophozia gillmani (Aust) Schust lophozia gIlttulata (Undo tl H Arnell)
36 39 40 46 136 (140) 141 Evans 39 46 136 141
42
141
Lophozia heterocolEDs (Thed) MA Howe IoPhozia ipcisa (Schard) Dum
27 39 46 89 141 8 27 31 46 79 90 140 141
LoQhozia longidens Macoun Lophozia ventricosa
1 46 79 90 141
Lophozia ventricosa var silvicola
(Buch) Jones 141 LoEhozla ventricosa vax longif1ora (Nees)
Macoun 39 46 79 141
43
~~ fragrans (Balbis) Frye et Clark Mannia pi 10sa (Hornem) Frye et Clark
46 (139) (140) 9 46 79 116 139
Mannia triandra (Scop) Gro 11e
46 (140)
Marchantia pound91ymorpha L
1 13 28 3946 (49) 78 87 88 89 90 (Ill) 121 122 147
Mo~rckia hibernica (HookJ Gott
46 89
Metzgeria conjugata Lindb
44
46
Mylia anomala (Hook) S Gray Notothylas orbicularis (Schwein) Sullo
39 46 141 (35)45 46
Nowellia curvifolia (Dicks) Mitt
17 27 3846 88 89 90 141
Odontoschisma denudatuJI1 (Nees ~ Mart)
Dum 46 81 137 141
Pallavicinia lyellii (Hook~ Carruth
90
Odontoschisma macounii (Aust) Under
45 46 (66) (139) 140 141
45
=
Pellia epiphllla (L) CordaPellia endiviifolia (Dicks~ Dum
27 39 46 90 13724 39 46
Pellia neesiana Limp bull Phaeoceros
39 46 89 27 39 45 46 97
Porella pinnata L
13 27 I 46 90 Plagiochila asplenioides (L) Dum
27 I 39 1+6 88 89 I 90 137
46
~e11a platYEhylla (L) Pfeiff Porella platyphylloi~ (Schwein) Lindb
17 27 39 46 69 (71)79 87 88 89 9 34 46 (57) 59 87 88 89 92 90 92 93 77
Preissia quadrata (Seop) Nees
1 36 3946 (71) 88 89 90 121 123
137
Ptllidium ciliare (L) Hampe
1 27 39 46 132 141
Dum
(22) 28 3946 (57) 87 88 89 90 92
Ptilidium ~lcherrium (G Web) Hampe
(9) 13 394656 (57)70 87 88 89 90 93 (119) 136 141
47
Radula obconica Sullo Reboulia hemlsphaerlca (L) Raddi
39 46 88 89 (43) 46 (59) 70 90 t Cf 124 125 (139)
Riccardia ~atifrons Lindb
1 27 39 46 89
Riccardia multifi~ (L) S Gray
39 46 88 145
Riccia beyrichiana Hampe ~ Lehm
46 Riccardia palmata (Hedw) Carruth
39 46 90
48
Riccia bifurca Hoffm Riccia cavernosa Hoffm
48 13739 46 79 137
Riccia fluitans L
242731454679
Riccla frostli Aust
48 79
Riccia sullicantii Aust
79 (137) Riccia sorocarpa Bisch 46 (137)
49
Ricciocarpus natans (1) Corda Sca~ia apiculata Spruce
13 27 46 79 89 110 39 46 (139) 141
ScaJltll1ia carinthica Jack ~ Lindb
137 t 141
Scapania ~ (Mart) Durn
39467990137141
Scapania digenil Schiffn
90
ScaEsectflia cuspiduligera (Nees) K Mull
46 (139) (140) (141)
50
Scapania glaucoeerha1a (Tayl) Aust Seapania gyrrmostomophilia Kaal
39 46 (139) 141 41 44 46 64 (139) (140) 141
ScaP3poundlia irrigua Gott ~ almiddot
7 39 46 79 137 (140L 141
Seapania nemorosa (L) Dum
1 39 46 89 90 137
Scapmi~ ~~ta Smiddotlch
137 141 51
~caPnia poundludicola LDeske amp K Mull Scapania saxicola Schust
46 (66) 141 136 141
Sc~~ scandica (H Arnell ~ Buch) Macv
139 140
Scapania subalpina (Nees) Durn
39 40 46 79 137 141
Scapania undulata (L) Dum
1 42 46 79 141
Scapania umbrosa (Schrad) Dum
137 140 141
52
Trichocolea tomentella (Ehrh) Dum Triomarla exsecta (Schmid) Schiffn
27 (35) 39 46 47 88 137 141 8 28 39 46 90 140 141
Tritomaria exsectiformis (Breidl) Loe~ke Tritomaria Sluinguedentata (Huds) Buch
39 -t6 89 90 136 140 141 7 39 46 141
Trltomaria scitula (Tayl) Joerg
46 136 (139) 141
53
Homalotheciella (Card) Broth subcapillata (Hedw) Broth
Brachythecium BSG acuminatum (Hedw) Awt acutum (Mitt) Sullo calcareum IGndb campestTe (C MuelL) BSG cunum (Lindb) Umpr digastrum C Muell et Kindb ex Macoun et Kindb oxycladon (Brid) Jaeg et Sauerb var oxycladon plumosum (Hedw) BSG populeum (Hedw) BSG var populeum reflexum (Starke ex Web et Mohr) DsG var relexum reiexum (Starke ex Web et Mohr) DSG var pacificum Renet Card ex Roell rlvulare BSG var rivulare tabulum (Hedw) BSC var tabuum salebrosum (Web et Mohr) BSG var salebrosum velutinum (Hedw) BSG Vat velutinum
Bryhniil Kaur graminicolor (Brid) Grout var graminicolor novae-angliae (Sull et Lesq ex sun) Grout
Rhynchostegium BSG senulatum (Hedw) Ja4lg et Sauerb
Rhynchostegiella (BSG) Limpr compacta (C MueU) Loeske
EU1hynchium 8SG hians (Hedw) Sande Lac pulchellum (Hedw) Jenn var pulchelium riparloides (Hedw) Rich
Stokeriella (Kindb) Robins pmelonga (Hedw) Robins var stokesii (Tum) Crum
63 Entodontaceae Entodon C Muell
brevisetus (Hook et Wils ex Wils) Lindb cladorrhizans (Hedw) C Muell seductrlx (Hedw) C MueU var wluctrix
Pleurozium Mitt schreberl (8rid) Mitt var schreberi
64 Plagiotheciaceae Pltlglothecium BSG
CflVifolium (Brid) Iwats var CtWifolium denticulatum (Hedw) 8SG laetum BSG ltztebricola BSG
26
6S SernatophyUaceae Brotherella Loeske ex Fleisch
recumins (Michx) Fleisch tenuirostris (Bruch et Schimp ex Sull) Fleisch
Sematophyllum Mitt demissum (Wils) Mitt
66 Hypnaceae Pl(itygyrium BSG
repens (Brid) BSG
PylaisieUa Kindb intricata (Hedw) Grout polyantha (Hedw) Grout selwynii (Kindb) Crum Steere et Anderson
Homomallium (Schimp) Loeske adnatum (Hedw) Broth
Hypnum Hedw cupressiforme Hedw var cupressiforme cupressifonne Hedw varfiliforme Brid curviolium Hedw fertile Sendtn imponens Hedw lindbergii Mitt palescens (Hedw) P-Beauv pratense Koch ex Brid
Calicladium Crum haldanianum (Grev) Crum
sopterygiopsis Iwats muelleriana (Schimp) Iwats
Isopterygium Mitt e1egans (Brid) lindb pulchellum (Hedw) Jaeg et Sauerb tenerum (Sw) Mitt
Taxiphyllum Fleisch depllQtum (Bruch et Schimp ex sun) Fleisch taxirameum (MitL) Fleisch
Herzogiella Broth seligeri (Brid) Iwats striatella (Brid) Iwats ruracea (Lindh) Iwats
hilium De Not crista-castrenna (Hedw) De Not
27
67 Rhytidiaceae Rhytidium (Sull) Kindb
rugosum (Hedw) Kindb
Rhytidfodelphus (Lindb ex Iimpr) Warnst triquetrus (Hedw) Warnst var triquetrus
68 Hylocomiaceae Hylocomium BSG
pyrenaicum (Spruce) lindb splendens (Hedw) BSG varsplendens
69 Buxbaumiaceae Buxbaumia Hedw
aphyllll Hedw
Diphyscium Mohr follosum (Hedw) Mohr
70 Tetraphidaceae Tetraphi3
pellucida Hedw
71 Polytrichaceae Atrichum PmiddotBeauv
angustatum (Brid) BSG cri8pum (James) Sullo oerstedianum (C Muell) Mitt teneUum (Roehl) BSG undulatum (Hedw) P middotBeauv var unduiatum
Pogonatum P-Beauv alpinum (Hedw) Roehl var alpinum umigerum (Hedw) P-Beauv
Poiytrichum Hedw commune Hedw var commune ormosum Hedw juniperinum Hedw longisetum Brid ohioense Ren et Card piliferum Hedw var pilierum strictum Brid
28
MAPS OF BRYOPHYTE TAXA
REPORTED FOR WISCONSIN
HORNWORTS AND LIVERWORTS
29
bull bull WSHeV~N S vE II
0 ~
e~RRON ~USK
(J~ (j -
CHIPPE A DUNN
0 o0 poundAU CLAIRE
~ TREMshyPEAshyLEAU
MICE
ONEIDA
Q LINCOLN
TAYLOR
Q t-~----- MOIRA THON
CLARK
o
10
LA FAVETTE
Hornwor ts amp Liverworts
- None - 1 - 25 taxa
- 26 - 50 tRXshy
- 51 - 75 tax)
- 7c - 100 taxa
FONDOU j LAC ~
DODGE
i O JEfFERSON
WALWORTH
30
Anastrophylltlfl helleranum (Nees) Schust Anastrophyllum michauxil (We~ Buch ~
46 88 136 141 Evans 38 46 141
Anastrophyllum mlnut~ (Schreb) Schuster Aneura plnguls (L) Durn
89 21 384687 88 89 90 (138) (145)
Anthoceros macounil MA Howe ~oceros punctatus L
21 (45) (144) 34
31
Barbilophozia attenuata (Mart) Loeske Barbilophozia barbata (Schmid ~ Schre~
24 141 Loeske 28 39 46 89 90 141
Bazzania trilobata (1) S Gray
1 26 39 46 (57) (59) 79 88 89
90 119 141
BlePbarostoma trichoffiyllum (1) Dumort
1 8 26 39 46 89 90 137 141
Blasia pusilla 1
1 39 46 90
Calypogeja fissa (1) Raddi
90
32
$129geja integristipula Steph C~Ypound2geja muelleriana (Schifm) K Mull
88 89 89
CallpoundO~eja neesiana (Mass amp Carest) K Mull
39 46 89 140 (141)
~pogeja sJlapound2icola (H Arnell tl J bull
Perss) Warnst et Loeske (119) 141
CeIalozia biscuspidata (L) Dum
1 37 I 46 90 t 137 141
CalYpogej~ trichomanis (L) Corda
26 39 (119) 137
33
CeEhalozia catenulata (Hub) Lindb Cephalozia connivens (Dicks) Lindb
39 46 79 141 39 4679 88 (89) 90 119 141
Cephalozia lacinulata Jack ~ Spruce
141 Cephalozia lunulifolia (Dum~ Dum I 28 39 46 79 88 89 90 141
Cepoundhalozia macouni (Aust) Aust Cephalozia ~leniceps (Aust) Lindb 27~ 3746 (66)7987 (139)141 39 46 79 89 141
34
Ceffialoziella 9ivaricata (Sm) Schiffn Cephaloziella ~hista (Jack) Schiffn
46 90 46 (119) 141
Cephaloziella hampeana (Nees) Schiffn
46
Cephaloziella rubella (Nees) Warnst
46 39
Cephaloziella rubella var sullivantii
(Aust) K Mull 46 39
Cephaloziella rubella var bifida (Lindb) Douin
35
46
Chiloscyphus pallescens (Ehrh ~ Hof~ Dum Chiloscyphus pallescens var fragilis
39J 46 88 89J 90 (Roth) K Milll 46
Chiloscyphus E9lyanthos (L) Corda
1 24 39 46
Chiloscyphus polyanthos vax rivularis
(Schrad) Nees 39 46
poundpoundnocepha1um conicum (1) Lindb
12 28 39 46 (49) 87 88 8990 120 121 148
Cololejeunea biddlecomiae (Aust) Evans
39 46 62 87 88 89 90
36
Dipl0]hyllum apicu1atum (Evans) Steph Fossombronia foveo1ata Lindb
(35)41 46 90 140 1 t 38 (39) t 46
Frullania bolanderi Aust
13 46 62 88 90 139 140
Frullanla ari ttoniae Evans
39 46 69 (71) 87 88 89 90
Frullania eboracensis Gott
1 9 28 4 39 46 (57) (59) 69 70 (71) 87 88 89 90 92 93 97 148
Frullania inflata Gott
46 87 88 89 (139)
Frullania oakesiana Aust Frullania plana Sullo
(35) 42 46 87 (139) (140) 92
Frullania ri paria Hampe ~ Lehm
46 69 71 87 89
Frullania ~elwynianna Pears
46 89
Geocalyx SEaveolens (Schrad) Nees
394688 8990 (119) 140
Frullania tamarisci (L) Dum subsp sPagrayana
(Mont) Hatt 28 34 l~6 t (57) 88 89 90
38
Mar pan thus seutatUB ( Web amp Mohr) Spruce Jamesoniella autumnalis (DC) Steph
(35) 39 46 90 1 7 28 39 46 56 (57) 87 8889 90 (119) 137 141
Jungermannia confertissima Nees
137 141
Jungermannia crenuliform1s Aust
79 137 141
Jungermannia ~il1ima Sm
46 90 141 Jungermannia exsert1folia subsp corifo~ (Dum~
Vana (35) 46 79 141
39
Jungermannia hyalina Lyell Jungerrr6nnia 1eiantha Grolle
40 46 79 136 141 39 46 79 88 90 1)7 141
Jungermannia IXlaxis Lindb Jungermannia pumila With
46 67 79 (139) (140) 141 )9 46 90 1)6 141
Jungermannia sphaerocarpa Hook Kurzia setacea (Web) Grolle
4679136140IIH 136
Lejeunea cavifo1ia (Ehrh) Lindb Lepidozia reptans (1) Dum
1 7 27 39 46 88 89 90 137 14010 I ( 27) 46 I 69 (71) 88
lophocolea heterophylla (Schrad) Dum Lophocolea minor Nees
1 )4 39 46 (57) 87 88 89 27 39 46 88 89 90 (119) 90 (93)94 119 136
Lophozia al~stris (Schleich ~ Web) Evans Lophozia ascendens (Warnst) Schust
46 90 137 141 136 140 141
41
loIilozia badensls (Gatt ~ Rabenh)
46 141
lophozia blcrenata (Schmid sectA Hoffm)
Dum 46 141
lophozia collaris (Nees) Dum
39 46 141
lophozla excisa (Dicks) Dum
27 4679 141
lophozia gillmani (Aust) Schust lophozia gIlttulata (Undo tl H Arnell)
36 39 40 46 136 (140) 141 Evans 39 46 136 141
42
141
Lophozia heterocolEDs (Thed) MA Howe IoPhozia ipcisa (Schard) Dum
27 39 46 89 141 8 27 31 46 79 90 140 141
LoQhozia longidens Macoun Lophozia ventricosa
1 46 79 90 141
Lophozia ventricosa var silvicola
(Buch) Jones 141 LoEhozla ventricosa vax longif1ora (Nees)
Macoun 39 46 79 141
43
~~ fragrans (Balbis) Frye et Clark Mannia pi 10sa (Hornem) Frye et Clark
46 (139) (140) 9 46 79 116 139
Mannia triandra (Scop) Gro 11e
46 (140)
Marchantia pound91ymorpha L
1 13 28 3946 (49) 78 87 88 89 90 (Ill) 121 122 147
Mo~rckia hibernica (HookJ Gott
46 89
Metzgeria conjugata Lindb
44
46
Mylia anomala (Hook) S Gray Notothylas orbicularis (Schwein) Sullo
39 46 141 (35)45 46
Nowellia curvifolia (Dicks) Mitt
17 27 3846 88 89 90 141
Odontoschisma denudatuJI1 (Nees ~ Mart)
Dum 46 81 137 141
Pallavicinia lyellii (Hook~ Carruth
90
Odontoschisma macounii (Aust) Under
45 46 (66) (139) 140 141
45
=
Pellia epiphllla (L) CordaPellia endiviifolia (Dicks~ Dum
27 39 46 90 13724 39 46
Pellia neesiana Limp bull Phaeoceros
39 46 89 27 39 45 46 97
Porella pinnata L
13 27 I 46 90 Plagiochila asplenioides (L) Dum
27 I 39 1+6 88 89 I 90 137
46
~e11a platYEhylla (L) Pfeiff Porella platyphylloi~ (Schwein) Lindb
17 27 39 46 69 (71)79 87 88 89 9 34 46 (57) 59 87 88 89 92 90 92 93 77
Preissia quadrata (Seop) Nees
1 36 3946 (71) 88 89 90 121 123
137
Ptllidium ciliare (L) Hampe
1 27 39 46 132 141
Dum
(22) 28 3946 (57) 87 88 89 90 92
Ptilidium ~lcherrium (G Web) Hampe
(9) 13 394656 (57)70 87 88 89 90 93 (119) 136 141
47
Radula obconica Sullo Reboulia hemlsphaerlca (L) Raddi
39 46 88 89 (43) 46 (59) 70 90 t Cf 124 125 (139)
Riccardia ~atifrons Lindb
1 27 39 46 89
Riccardia multifi~ (L) S Gray
39 46 88 145
Riccia beyrichiana Hampe ~ Lehm
46 Riccardia palmata (Hedw) Carruth
39 46 90
48
Riccia bifurca Hoffm Riccia cavernosa Hoffm
48 13739 46 79 137
Riccia fluitans L
242731454679
Riccla frostli Aust
48 79
Riccia sullicantii Aust
79 (137) Riccia sorocarpa Bisch 46 (137)
49
Ricciocarpus natans (1) Corda Sca~ia apiculata Spruce
13 27 46 79 89 110 39 46 (139) 141
ScaJltll1ia carinthica Jack ~ Lindb
137 t 141
Scapania ~ (Mart) Durn
39467990137141
Scapania digenil Schiffn
90
ScaEsectflia cuspiduligera (Nees) K Mull
46 (139) (140) (141)
50
Scapania glaucoeerha1a (Tayl) Aust Seapania gyrrmostomophilia Kaal
39 46 (139) 141 41 44 46 64 (139) (140) 141
ScaP3poundlia irrigua Gott ~ almiddot
7 39 46 79 137 (140L 141
Seapania nemorosa (L) Dum
1 39 46 89 90 137
Scapmi~ ~~ta Smiddotlch
137 141 51
~caPnia poundludicola LDeske amp K Mull Scapania saxicola Schust
46 (66) 141 136 141
Sc~~ scandica (H Arnell ~ Buch) Macv
139 140
Scapania subalpina (Nees) Durn
39 40 46 79 137 141
Scapania undulata (L) Dum
1 42 46 79 141
Scapania umbrosa (Schrad) Dum
137 140 141
52
Trichocolea tomentella (Ehrh) Dum Triomarla exsecta (Schmid) Schiffn
27 (35) 39 46 47 88 137 141 8 28 39 46 90 140 141
Tritomaria exsectiformis (Breidl) Loe~ke Tritomaria Sluinguedentata (Huds) Buch
39 -t6 89 90 136 140 141 7 39 46 141
Trltomaria scitula (Tayl) Joerg
46 136 (139) 141
53
6S SernatophyUaceae Brotherella Loeske ex Fleisch
recumins (Michx) Fleisch tenuirostris (Bruch et Schimp ex Sull) Fleisch
Sematophyllum Mitt demissum (Wils) Mitt
66 Hypnaceae Pl(itygyrium BSG
repens (Brid) BSG
PylaisieUa Kindb intricata (Hedw) Grout polyantha (Hedw) Grout selwynii (Kindb) Crum Steere et Anderson
Homomallium (Schimp) Loeske adnatum (Hedw) Broth
Hypnum Hedw cupressiforme Hedw var cupressiforme cupressifonne Hedw varfiliforme Brid curviolium Hedw fertile Sendtn imponens Hedw lindbergii Mitt palescens (Hedw) P-Beauv pratense Koch ex Brid
Calicladium Crum haldanianum (Grev) Crum
sopterygiopsis Iwats muelleriana (Schimp) Iwats
Isopterygium Mitt e1egans (Brid) lindb pulchellum (Hedw) Jaeg et Sauerb tenerum (Sw) Mitt
Taxiphyllum Fleisch depllQtum (Bruch et Schimp ex sun) Fleisch taxirameum (MitL) Fleisch
Herzogiella Broth seligeri (Brid) Iwats striatella (Brid) Iwats ruracea (Lindh) Iwats
hilium De Not crista-castrenna (Hedw) De Not
27
67 Rhytidiaceae Rhytidium (Sull) Kindb
rugosum (Hedw) Kindb
Rhytidfodelphus (Lindb ex Iimpr) Warnst triquetrus (Hedw) Warnst var triquetrus
68 Hylocomiaceae Hylocomium BSG
pyrenaicum (Spruce) lindb splendens (Hedw) BSG varsplendens
69 Buxbaumiaceae Buxbaumia Hedw
aphyllll Hedw
Diphyscium Mohr follosum (Hedw) Mohr
70 Tetraphidaceae Tetraphi3
pellucida Hedw
71 Polytrichaceae Atrichum PmiddotBeauv
angustatum (Brid) BSG cri8pum (James) Sullo oerstedianum (C Muell) Mitt teneUum (Roehl) BSG undulatum (Hedw) P middotBeauv var unduiatum
Pogonatum P-Beauv alpinum (Hedw) Roehl var alpinum umigerum (Hedw) P-Beauv
Poiytrichum Hedw commune Hedw var commune ormosum Hedw juniperinum Hedw longisetum Brid ohioense Ren et Card piliferum Hedw var pilierum strictum Brid
28
MAPS OF BRYOPHYTE TAXA
REPORTED FOR WISCONSIN
HORNWORTS AND LIVERWORTS
29
bull bull WSHeV~N S vE II
0 ~
e~RRON ~USK
(J~ (j -
CHIPPE A DUNN
0 o0 poundAU CLAIRE
~ TREMshyPEAshyLEAU
MICE
ONEIDA
Q LINCOLN
TAYLOR
Q t-~----- MOIRA THON
CLARK
o
10
LA FAVETTE
Hornwor ts amp Liverworts
- None - 1 - 25 taxa
- 26 - 50 tRXshy
- 51 - 75 tax)
- 7c - 100 taxa
FONDOU j LAC ~
DODGE
i O JEfFERSON
WALWORTH
30
Anastrophylltlfl helleranum (Nees) Schust Anastrophyllum michauxil (We~ Buch ~
46 88 136 141 Evans 38 46 141
Anastrophyllum mlnut~ (Schreb) Schuster Aneura plnguls (L) Durn
89 21 384687 88 89 90 (138) (145)
Anthoceros macounil MA Howe ~oceros punctatus L
21 (45) (144) 34
31
Barbilophozia attenuata (Mart) Loeske Barbilophozia barbata (Schmid ~ Schre~
24 141 Loeske 28 39 46 89 90 141
Bazzania trilobata (1) S Gray
1 26 39 46 (57) (59) 79 88 89
90 119 141
BlePbarostoma trichoffiyllum (1) Dumort
1 8 26 39 46 89 90 137 141
Blasia pusilla 1
1 39 46 90
Calypogeja fissa (1) Raddi
90
32
$129geja integristipula Steph C~Ypound2geja muelleriana (Schifm) K Mull
88 89 89
CallpoundO~eja neesiana (Mass amp Carest) K Mull
39 46 89 140 (141)
~pogeja sJlapound2icola (H Arnell tl J bull
Perss) Warnst et Loeske (119) 141
CeIalozia biscuspidata (L) Dum
1 37 I 46 90 t 137 141
CalYpogej~ trichomanis (L) Corda
26 39 (119) 137
33
CeEhalozia catenulata (Hub) Lindb Cephalozia connivens (Dicks) Lindb
39 46 79 141 39 4679 88 (89) 90 119 141
Cephalozia lacinulata Jack ~ Spruce
141 Cephalozia lunulifolia (Dum~ Dum I 28 39 46 79 88 89 90 141
Cepoundhalozia macouni (Aust) Aust Cephalozia ~leniceps (Aust) Lindb 27~ 3746 (66)7987 (139)141 39 46 79 89 141
34
Ceffialoziella 9ivaricata (Sm) Schiffn Cephaloziella ~hista (Jack) Schiffn
46 90 46 (119) 141
Cephaloziella hampeana (Nees) Schiffn
46
Cephaloziella rubella (Nees) Warnst
46 39
Cephaloziella rubella var sullivantii
(Aust) K Mull 46 39
Cephaloziella rubella var bifida (Lindb) Douin
35
46
Chiloscyphus pallescens (Ehrh ~ Hof~ Dum Chiloscyphus pallescens var fragilis
39J 46 88 89J 90 (Roth) K Milll 46
Chiloscyphus E9lyanthos (L) Corda
1 24 39 46
Chiloscyphus polyanthos vax rivularis
(Schrad) Nees 39 46
poundpoundnocepha1um conicum (1) Lindb
12 28 39 46 (49) 87 88 8990 120 121 148
Cololejeunea biddlecomiae (Aust) Evans
39 46 62 87 88 89 90
36
Dipl0]hyllum apicu1atum (Evans) Steph Fossombronia foveo1ata Lindb
(35)41 46 90 140 1 t 38 (39) t 46
Frullania bolanderi Aust
13 46 62 88 90 139 140
Frullanla ari ttoniae Evans
39 46 69 (71) 87 88 89 90
Frullania eboracensis Gott
1 9 28 4 39 46 (57) (59) 69 70 (71) 87 88 89 90 92 93 97 148
Frullania inflata Gott
46 87 88 89 (139)
Frullania oakesiana Aust Frullania plana Sullo
(35) 42 46 87 (139) (140) 92
Frullania ri paria Hampe ~ Lehm
46 69 71 87 89
Frullania ~elwynianna Pears
46 89
Geocalyx SEaveolens (Schrad) Nees
394688 8990 (119) 140
Frullania tamarisci (L) Dum subsp sPagrayana
(Mont) Hatt 28 34 l~6 t (57) 88 89 90
38
Mar pan thus seutatUB ( Web amp Mohr) Spruce Jamesoniella autumnalis (DC) Steph
(35) 39 46 90 1 7 28 39 46 56 (57) 87 8889 90 (119) 137 141
Jungermannia confertissima Nees
137 141
Jungermannia crenuliform1s Aust
79 137 141
Jungermannia ~il1ima Sm
46 90 141 Jungermannia exsert1folia subsp corifo~ (Dum~
Vana (35) 46 79 141
39
Jungermannia hyalina Lyell Jungerrr6nnia 1eiantha Grolle
40 46 79 136 141 39 46 79 88 90 1)7 141
Jungermannia IXlaxis Lindb Jungermannia pumila With
46 67 79 (139) (140) 141 )9 46 90 1)6 141
Jungermannia sphaerocarpa Hook Kurzia setacea (Web) Grolle
4679136140IIH 136
Lejeunea cavifo1ia (Ehrh) Lindb Lepidozia reptans (1) Dum
1 7 27 39 46 88 89 90 137 14010 I ( 27) 46 I 69 (71) 88
lophocolea heterophylla (Schrad) Dum Lophocolea minor Nees
1 )4 39 46 (57) 87 88 89 27 39 46 88 89 90 (119) 90 (93)94 119 136
Lophozia al~stris (Schleich ~ Web) Evans Lophozia ascendens (Warnst) Schust
46 90 137 141 136 140 141
41
loIilozia badensls (Gatt ~ Rabenh)
46 141
lophozia blcrenata (Schmid sectA Hoffm)
Dum 46 141
lophozia collaris (Nees) Dum
39 46 141
lophozla excisa (Dicks) Dum
27 4679 141
lophozia gillmani (Aust) Schust lophozia gIlttulata (Undo tl H Arnell)
36 39 40 46 136 (140) 141 Evans 39 46 136 141
42
141
Lophozia heterocolEDs (Thed) MA Howe IoPhozia ipcisa (Schard) Dum
27 39 46 89 141 8 27 31 46 79 90 140 141
LoQhozia longidens Macoun Lophozia ventricosa
1 46 79 90 141
Lophozia ventricosa var silvicola
(Buch) Jones 141 LoEhozla ventricosa vax longif1ora (Nees)
Macoun 39 46 79 141
43
~~ fragrans (Balbis) Frye et Clark Mannia pi 10sa (Hornem) Frye et Clark
46 (139) (140) 9 46 79 116 139
Mannia triandra (Scop) Gro 11e
46 (140)
Marchantia pound91ymorpha L
1 13 28 3946 (49) 78 87 88 89 90 (Ill) 121 122 147
Mo~rckia hibernica (HookJ Gott
46 89
Metzgeria conjugata Lindb
44
46
Mylia anomala (Hook) S Gray Notothylas orbicularis (Schwein) Sullo
39 46 141 (35)45 46
Nowellia curvifolia (Dicks) Mitt
17 27 3846 88 89 90 141
Odontoschisma denudatuJI1 (Nees ~ Mart)
Dum 46 81 137 141
Pallavicinia lyellii (Hook~ Carruth
90
Odontoschisma macounii (Aust) Under
45 46 (66) (139) 140 141
45
=
Pellia epiphllla (L) CordaPellia endiviifolia (Dicks~ Dum
27 39 46 90 13724 39 46
Pellia neesiana Limp bull Phaeoceros
39 46 89 27 39 45 46 97
Porella pinnata L
13 27 I 46 90 Plagiochila asplenioides (L) Dum
27 I 39 1+6 88 89 I 90 137
46
~e11a platYEhylla (L) Pfeiff Porella platyphylloi~ (Schwein) Lindb
17 27 39 46 69 (71)79 87 88 89 9 34 46 (57) 59 87 88 89 92 90 92 93 77
Preissia quadrata (Seop) Nees
1 36 3946 (71) 88 89 90 121 123
137
Ptllidium ciliare (L) Hampe
1 27 39 46 132 141
Dum
(22) 28 3946 (57) 87 88 89 90 92
Ptilidium ~lcherrium (G Web) Hampe
(9) 13 394656 (57)70 87 88 89 90 93 (119) 136 141
47
Radula obconica Sullo Reboulia hemlsphaerlca (L) Raddi
39 46 88 89 (43) 46 (59) 70 90 t Cf 124 125 (139)
Riccardia ~atifrons Lindb
1 27 39 46 89
Riccardia multifi~ (L) S Gray
39 46 88 145
Riccia beyrichiana Hampe ~ Lehm
46 Riccardia palmata (Hedw) Carruth
39 46 90
48
Riccia bifurca Hoffm Riccia cavernosa Hoffm
48 13739 46 79 137
Riccia fluitans L
242731454679
Riccla frostli Aust
48 79
Riccia sullicantii Aust
79 (137) Riccia sorocarpa Bisch 46 (137)
49
Ricciocarpus natans (1) Corda Sca~ia apiculata Spruce
13 27 46 79 89 110 39 46 (139) 141
ScaJltll1ia carinthica Jack ~ Lindb
137 t 141
Scapania ~ (Mart) Durn
39467990137141
Scapania digenil Schiffn
90
ScaEsectflia cuspiduligera (Nees) K Mull
46 (139) (140) (141)
50
Scapania glaucoeerha1a (Tayl) Aust Seapania gyrrmostomophilia Kaal
39 46 (139) 141 41 44 46 64 (139) (140) 141
ScaP3poundlia irrigua Gott ~ almiddot
7 39 46 79 137 (140L 141
Seapania nemorosa (L) Dum
1 39 46 89 90 137
Scapmi~ ~~ta Smiddotlch
137 141 51
~caPnia poundludicola LDeske amp K Mull Scapania saxicola Schust
46 (66) 141 136 141
Sc~~ scandica (H Arnell ~ Buch) Macv
139 140
Scapania subalpina (Nees) Durn
39 40 46 79 137 141
Scapania undulata (L) Dum
1 42 46 79 141
Scapania umbrosa (Schrad) Dum
137 140 141
52
Trichocolea tomentella (Ehrh) Dum Triomarla exsecta (Schmid) Schiffn
27 (35) 39 46 47 88 137 141 8 28 39 46 90 140 141
Tritomaria exsectiformis (Breidl) Loe~ke Tritomaria Sluinguedentata (Huds) Buch
39 -t6 89 90 136 140 141 7 39 46 141
Trltomaria scitula (Tayl) Joerg
46 136 (139) 141
53
67 Rhytidiaceae Rhytidium (Sull) Kindb
rugosum (Hedw) Kindb
Rhytidfodelphus (Lindb ex Iimpr) Warnst triquetrus (Hedw) Warnst var triquetrus
68 Hylocomiaceae Hylocomium BSG
pyrenaicum (Spruce) lindb splendens (Hedw) BSG varsplendens
69 Buxbaumiaceae Buxbaumia Hedw
aphyllll Hedw
Diphyscium Mohr follosum (Hedw) Mohr
70 Tetraphidaceae Tetraphi3
pellucida Hedw
71 Polytrichaceae Atrichum PmiddotBeauv
angustatum (Brid) BSG cri8pum (James) Sullo oerstedianum (C Muell) Mitt teneUum (Roehl) BSG undulatum (Hedw) P middotBeauv var unduiatum
Pogonatum P-Beauv alpinum (Hedw) Roehl var alpinum umigerum (Hedw) P-Beauv
Poiytrichum Hedw commune Hedw var commune ormosum Hedw juniperinum Hedw longisetum Brid ohioense Ren et Card piliferum Hedw var pilierum strictum Brid
28
MAPS OF BRYOPHYTE TAXA
REPORTED FOR WISCONSIN
HORNWORTS AND LIVERWORTS
29
bull bull WSHeV~N S vE II
0 ~
e~RRON ~USK
(J~ (j -
CHIPPE A DUNN
0 o0 poundAU CLAIRE
~ TREMshyPEAshyLEAU
MICE
ONEIDA
Q LINCOLN
TAYLOR
Q t-~----- MOIRA THON
CLARK
o
10
LA FAVETTE
Hornwor ts amp Liverworts
- None - 1 - 25 taxa
- 26 - 50 tRXshy
- 51 - 75 tax)
- 7c - 100 taxa
FONDOU j LAC ~
DODGE
i O JEfFERSON
WALWORTH
30
Anastrophylltlfl helleranum (Nees) Schust Anastrophyllum michauxil (We~ Buch ~
46 88 136 141 Evans 38 46 141
Anastrophyllum mlnut~ (Schreb) Schuster Aneura plnguls (L) Durn
89 21 384687 88 89 90 (138) (145)
Anthoceros macounil MA Howe ~oceros punctatus L
21 (45) (144) 34
31
Barbilophozia attenuata (Mart) Loeske Barbilophozia barbata (Schmid ~ Schre~
24 141 Loeske 28 39 46 89 90 141
Bazzania trilobata (1) S Gray
1 26 39 46 (57) (59) 79 88 89
90 119 141
BlePbarostoma trichoffiyllum (1) Dumort
1 8 26 39 46 89 90 137 141
Blasia pusilla 1
1 39 46 90
Calypogeja fissa (1) Raddi
90
32
$129geja integristipula Steph C~Ypound2geja muelleriana (Schifm) K Mull
88 89 89
CallpoundO~eja neesiana (Mass amp Carest) K Mull
39 46 89 140 (141)
~pogeja sJlapound2icola (H Arnell tl J bull
Perss) Warnst et Loeske (119) 141
CeIalozia biscuspidata (L) Dum
1 37 I 46 90 t 137 141
CalYpogej~ trichomanis (L) Corda
26 39 (119) 137
33
CeEhalozia catenulata (Hub) Lindb Cephalozia connivens (Dicks) Lindb
39 46 79 141 39 4679 88 (89) 90 119 141
Cephalozia lacinulata Jack ~ Spruce
141 Cephalozia lunulifolia (Dum~ Dum I 28 39 46 79 88 89 90 141
Cepoundhalozia macouni (Aust) Aust Cephalozia ~leniceps (Aust) Lindb 27~ 3746 (66)7987 (139)141 39 46 79 89 141
34
Ceffialoziella 9ivaricata (Sm) Schiffn Cephaloziella ~hista (Jack) Schiffn
46 90 46 (119) 141
Cephaloziella hampeana (Nees) Schiffn
46
Cephaloziella rubella (Nees) Warnst
46 39
Cephaloziella rubella var sullivantii
(Aust) K Mull 46 39
Cephaloziella rubella var bifida (Lindb) Douin
35
46
Chiloscyphus pallescens (Ehrh ~ Hof~ Dum Chiloscyphus pallescens var fragilis
39J 46 88 89J 90 (Roth) K Milll 46
Chiloscyphus E9lyanthos (L) Corda
1 24 39 46
Chiloscyphus polyanthos vax rivularis
(Schrad) Nees 39 46
poundpoundnocepha1um conicum (1) Lindb
12 28 39 46 (49) 87 88 8990 120 121 148
Cololejeunea biddlecomiae (Aust) Evans
39 46 62 87 88 89 90
36
Dipl0]hyllum apicu1atum (Evans) Steph Fossombronia foveo1ata Lindb
(35)41 46 90 140 1 t 38 (39) t 46
Frullania bolanderi Aust
13 46 62 88 90 139 140
Frullanla ari ttoniae Evans
39 46 69 (71) 87 88 89 90
Frullania eboracensis Gott
1 9 28 4 39 46 (57) (59) 69 70 (71) 87 88 89 90 92 93 97 148
Frullania inflata Gott
46 87 88 89 (139)
Frullania oakesiana Aust Frullania plana Sullo
(35) 42 46 87 (139) (140) 92
Frullania ri paria Hampe ~ Lehm
46 69 71 87 89
Frullania ~elwynianna Pears
46 89
Geocalyx SEaveolens (Schrad) Nees
394688 8990 (119) 140
Frullania tamarisci (L) Dum subsp sPagrayana
(Mont) Hatt 28 34 l~6 t (57) 88 89 90
38
Mar pan thus seutatUB ( Web amp Mohr) Spruce Jamesoniella autumnalis (DC) Steph
(35) 39 46 90 1 7 28 39 46 56 (57) 87 8889 90 (119) 137 141
Jungermannia confertissima Nees
137 141
Jungermannia crenuliform1s Aust
79 137 141
Jungermannia ~il1ima Sm
46 90 141 Jungermannia exsert1folia subsp corifo~ (Dum~
Vana (35) 46 79 141
39
Jungermannia hyalina Lyell Jungerrr6nnia 1eiantha Grolle
40 46 79 136 141 39 46 79 88 90 1)7 141
Jungermannia IXlaxis Lindb Jungermannia pumila With
46 67 79 (139) (140) 141 )9 46 90 1)6 141
Jungermannia sphaerocarpa Hook Kurzia setacea (Web) Grolle
4679136140IIH 136
Lejeunea cavifo1ia (Ehrh) Lindb Lepidozia reptans (1) Dum
1 7 27 39 46 88 89 90 137 14010 I ( 27) 46 I 69 (71) 88
lophocolea heterophylla (Schrad) Dum Lophocolea minor Nees
1 )4 39 46 (57) 87 88 89 27 39 46 88 89 90 (119) 90 (93)94 119 136
Lophozia al~stris (Schleich ~ Web) Evans Lophozia ascendens (Warnst) Schust
46 90 137 141 136 140 141
41
loIilozia badensls (Gatt ~ Rabenh)
46 141
lophozia blcrenata (Schmid sectA Hoffm)
Dum 46 141
lophozia collaris (Nees) Dum
39 46 141
lophozla excisa (Dicks) Dum
27 4679 141
lophozia gillmani (Aust) Schust lophozia gIlttulata (Undo tl H Arnell)
36 39 40 46 136 (140) 141 Evans 39 46 136 141
42
141
Lophozia heterocolEDs (Thed) MA Howe IoPhozia ipcisa (Schard) Dum
27 39 46 89 141 8 27 31 46 79 90 140 141
LoQhozia longidens Macoun Lophozia ventricosa
1 46 79 90 141
Lophozia ventricosa var silvicola
(Buch) Jones 141 LoEhozla ventricosa vax longif1ora (Nees)
Macoun 39 46 79 141
43
~~ fragrans (Balbis) Frye et Clark Mannia pi 10sa (Hornem) Frye et Clark
46 (139) (140) 9 46 79 116 139
Mannia triandra (Scop) Gro 11e
46 (140)
Marchantia pound91ymorpha L
1 13 28 3946 (49) 78 87 88 89 90 (Ill) 121 122 147
Mo~rckia hibernica (HookJ Gott
46 89
Metzgeria conjugata Lindb
44
46
Mylia anomala (Hook) S Gray Notothylas orbicularis (Schwein) Sullo
39 46 141 (35)45 46
Nowellia curvifolia (Dicks) Mitt
17 27 3846 88 89 90 141
Odontoschisma denudatuJI1 (Nees ~ Mart)
Dum 46 81 137 141
Pallavicinia lyellii (Hook~ Carruth
90
Odontoschisma macounii (Aust) Under
45 46 (66) (139) 140 141
45
=
Pellia epiphllla (L) CordaPellia endiviifolia (Dicks~ Dum
27 39 46 90 13724 39 46
Pellia neesiana Limp bull Phaeoceros
39 46 89 27 39 45 46 97
Porella pinnata L
13 27 I 46 90 Plagiochila asplenioides (L) Dum
27 I 39 1+6 88 89 I 90 137
46
~e11a platYEhylla (L) Pfeiff Porella platyphylloi~ (Schwein) Lindb
17 27 39 46 69 (71)79 87 88 89 9 34 46 (57) 59 87 88 89 92 90 92 93 77
Preissia quadrata (Seop) Nees
1 36 3946 (71) 88 89 90 121 123
137
Ptllidium ciliare (L) Hampe
1 27 39 46 132 141
Dum
(22) 28 3946 (57) 87 88 89 90 92
Ptilidium ~lcherrium (G Web) Hampe
(9) 13 394656 (57)70 87 88 89 90 93 (119) 136 141
47
Radula obconica Sullo Reboulia hemlsphaerlca (L) Raddi
39 46 88 89 (43) 46 (59) 70 90 t Cf 124 125 (139)
Riccardia ~atifrons Lindb
1 27 39 46 89
Riccardia multifi~ (L) S Gray
39 46 88 145
Riccia beyrichiana Hampe ~ Lehm
46 Riccardia palmata (Hedw) Carruth
39 46 90
48
Riccia bifurca Hoffm Riccia cavernosa Hoffm
48 13739 46 79 137
Riccia fluitans L
242731454679
Riccla frostli Aust
48 79
Riccia sullicantii Aust
79 (137) Riccia sorocarpa Bisch 46 (137)
49
Ricciocarpus natans (1) Corda Sca~ia apiculata Spruce
13 27 46 79 89 110 39 46 (139) 141
ScaJltll1ia carinthica Jack ~ Lindb
137 t 141
Scapania ~ (Mart) Durn
39467990137141
Scapania digenil Schiffn
90
ScaEsectflia cuspiduligera (Nees) K Mull
46 (139) (140) (141)
50
Scapania glaucoeerha1a (Tayl) Aust Seapania gyrrmostomophilia Kaal
39 46 (139) 141 41 44 46 64 (139) (140) 141
ScaP3poundlia irrigua Gott ~ almiddot
7 39 46 79 137 (140L 141
Seapania nemorosa (L) Dum
1 39 46 89 90 137
Scapmi~ ~~ta Smiddotlch
137 141 51
~caPnia poundludicola LDeske amp K Mull Scapania saxicola Schust
46 (66) 141 136 141
Sc~~ scandica (H Arnell ~ Buch) Macv
139 140
Scapania subalpina (Nees) Durn
39 40 46 79 137 141
Scapania undulata (L) Dum
1 42 46 79 141
Scapania umbrosa (Schrad) Dum
137 140 141
52
Trichocolea tomentella (Ehrh) Dum Triomarla exsecta (Schmid) Schiffn
27 (35) 39 46 47 88 137 141 8 28 39 46 90 140 141
Tritomaria exsectiformis (Breidl) Loe~ke Tritomaria Sluinguedentata (Huds) Buch
39 -t6 89 90 136 140 141 7 39 46 141
Trltomaria scitula (Tayl) Joerg
46 136 (139) 141
53
MAPS OF BRYOPHYTE TAXA
REPORTED FOR WISCONSIN
HORNWORTS AND LIVERWORTS
29
bull bull WSHeV~N S vE II
0 ~
e~RRON ~USK
(J~ (j -
CHIPPE A DUNN
0 o0 poundAU CLAIRE
~ TREMshyPEAshyLEAU
MICE
ONEIDA
Q LINCOLN
TAYLOR
Q t-~----- MOIRA THON
CLARK
o
10
LA FAVETTE
Hornwor ts amp Liverworts
- None - 1 - 25 taxa
- 26 - 50 tRXshy
- 51 - 75 tax)
- 7c - 100 taxa
FONDOU j LAC ~
DODGE
i O JEfFERSON
WALWORTH
30
Anastrophylltlfl helleranum (Nees) Schust Anastrophyllum michauxil (We~ Buch ~
46 88 136 141 Evans 38 46 141
Anastrophyllum mlnut~ (Schreb) Schuster Aneura plnguls (L) Durn
89 21 384687 88 89 90 (138) (145)
Anthoceros macounil MA Howe ~oceros punctatus L
21 (45) (144) 34
31
Barbilophozia attenuata (Mart) Loeske Barbilophozia barbata (Schmid ~ Schre~
24 141 Loeske 28 39 46 89 90 141
Bazzania trilobata (1) S Gray
1 26 39 46 (57) (59) 79 88 89
90 119 141
BlePbarostoma trichoffiyllum (1) Dumort
1 8 26 39 46 89 90 137 141
Blasia pusilla 1
1 39 46 90
Calypogeja fissa (1) Raddi
90
32
$129geja integristipula Steph C~Ypound2geja muelleriana (Schifm) K Mull
88 89 89
CallpoundO~eja neesiana (Mass amp Carest) K Mull
39 46 89 140 (141)
~pogeja sJlapound2icola (H Arnell tl J bull
Perss) Warnst et Loeske (119) 141
CeIalozia biscuspidata (L) Dum
1 37 I 46 90 t 137 141
CalYpogej~ trichomanis (L) Corda
26 39 (119) 137
33
CeEhalozia catenulata (Hub) Lindb Cephalozia connivens (Dicks) Lindb
39 46 79 141 39 4679 88 (89) 90 119 141
Cephalozia lacinulata Jack ~ Spruce
141 Cephalozia lunulifolia (Dum~ Dum I 28 39 46 79 88 89 90 141
Cepoundhalozia macouni (Aust) Aust Cephalozia ~leniceps (Aust) Lindb 27~ 3746 (66)7987 (139)141 39 46 79 89 141
34
Ceffialoziella 9ivaricata (Sm) Schiffn Cephaloziella ~hista (Jack) Schiffn
46 90 46 (119) 141
Cephaloziella hampeana (Nees) Schiffn
46
Cephaloziella rubella (Nees) Warnst
46 39
Cephaloziella rubella var sullivantii
(Aust) K Mull 46 39
Cephaloziella rubella var bifida (Lindb) Douin
35
46
Chiloscyphus pallescens (Ehrh ~ Hof~ Dum Chiloscyphus pallescens var fragilis
39J 46 88 89J 90 (Roth) K Milll 46
Chiloscyphus E9lyanthos (L) Corda
1 24 39 46
Chiloscyphus polyanthos vax rivularis
(Schrad) Nees 39 46
poundpoundnocepha1um conicum (1) Lindb
12 28 39 46 (49) 87 88 8990 120 121 148
Cololejeunea biddlecomiae (Aust) Evans
39 46 62 87 88 89 90
36
Dipl0]hyllum apicu1atum (Evans) Steph Fossombronia foveo1ata Lindb
(35)41 46 90 140 1 t 38 (39) t 46
Frullania bolanderi Aust
13 46 62 88 90 139 140
Frullanla ari ttoniae Evans
39 46 69 (71) 87 88 89 90
Frullania eboracensis Gott
1 9 28 4 39 46 (57) (59) 69 70 (71) 87 88 89 90 92 93 97 148
Frullania inflata Gott
46 87 88 89 (139)
Frullania oakesiana Aust Frullania plana Sullo
(35) 42 46 87 (139) (140) 92
Frullania ri paria Hampe ~ Lehm
46 69 71 87 89
Frullania ~elwynianna Pears
46 89
Geocalyx SEaveolens (Schrad) Nees
394688 8990 (119) 140
Frullania tamarisci (L) Dum subsp sPagrayana
(Mont) Hatt 28 34 l~6 t (57) 88 89 90
38
Mar pan thus seutatUB ( Web amp Mohr) Spruce Jamesoniella autumnalis (DC) Steph
(35) 39 46 90 1 7 28 39 46 56 (57) 87 8889 90 (119) 137 141
Jungermannia confertissima Nees
137 141
Jungermannia crenuliform1s Aust
79 137 141
Jungermannia ~il1ima Sm
46 90 141 Jungermannia exsert1folia subsp corifo~ (Dum~
Vana (35) 46 79 141
39
Jungermannia hyalina Lyell Jungerrr6nnia 1eiantha Grolle
40 46 79 136 141 39 46 79 88 90 1)7 141
Jungermannia IXlaxis Lindb Jungermannia pumila With
46 67 79 (139) (140) 141 )9 46 90 1)6 141
Jungermannia sphaerocarpa Hook Kurzia setacea (Web) Grolle
4679136140IIH 136
Lejeunea cavifo1ia (Ehrh) Lindb Lepidozia reptans (1) Dum
1 7 27 39 46 88 89 90 137 14010 I ( 27) 46 I 69 (71) 88
lophocolea heterophylla (Schrad) Dum Lophocolea minor Nees
1 )4 39 46 (57) 87 88 89 27 39 46 88 89 90 (119) 90 (93)94 119 136
Lophozia al~stris (Schleich ~ Web) Evans Lophozia ascendens (Warnst) Schust
46 90 137 141 136 140 141
41
loIilozia badensls (Gatt ~ Rabenh)
46 141
lophozia blcrenata (Schmid sectA Hoffm)
Dum 46 141
lophozia collaris (Nees) Dum
39 46 141
lophozla excisa (Dicks) Dum
27 4679 141
lophozia gillmani (Aust) Schust lophozia gIlttulata (Undo tl H Arnell)
36 39 40 46 136 (140) 141 Evans 39 46 136 141
42
141
Lophozia heterocolEDs (Thed) MA Howe IoPhozia ipcisa (Schard) Dum
27 39 46 89 141 8 27 31 46 79 90 140 141
LoQhozia longidens Macoun Lophozia ventricosa
1 46 79 90 141
Lophozia ventricosa var silvicola
(Buch) Jones 141 LoEhozla ventricosa vax longif1ora (Nees)
Macoun 39 46 79 141
43
~~ fragrans (Balbis) Frye et Clark Mannia pi 10sa (Hornem) Frye et Clark
46 (139) (140) 9 46 79 116 139
Mannia triandra (Scop) Gro 11e
46 (140)
Marchantia pound91ymorpha L
1 13 28 3946 (49) 78 87 88 89 90 (Ill) 121 122 147
Mo~rckia hibernica (HookJ Gott
46 89
Metzgeria conjugata Lindb
44
46
Mylia anomala (Hook) S Gray Notothylas orbicularis (Schwein) Sullo
39 46 141 (35)45 46
Nowellia curvifolia (Dicks) Mitt
17 27 3846 88 89 90 141
Odontoschisma denudatuJI1 (Nees ~ Mart)
Dum 46 81 137 141
Pallavicinia lyellii (Hook~ Carruth
90
Odontoschisma macounii (Aust) Under
45 46 (66) (139) 140 141
45
=
Pellia epiphllla (L) CordaPellia endiviifolia (Dicks~ Dum
27 39 46 90 13724 39 46
Pellia neesiana Limp bull Phaeoceros
39 46 89 27 39 45 46 97
Porella pinnata L
13 27 I 46 90 Plagiochila asplenioides (L) Dum
27 I 39 1+6 88 89 I 90 137
46
~e11a platYEhylla (L) Pfeiff Porella platyphylloi~ (Schwein) Lindb
17 27 39 46 69 (71)79 87 88 89 9 34 46 (57) 59 87 88 89 92 90 92 93 77
Preissia quadrata (Seop) Nees
1 36 3946 (71) 88 89 90 121 123
137
Ptllidium ciliare (L) Hampe
1 27 39 46 132 141
Dum
(22) 28 3946 (57) 87 88 89 90 92
Ptilidium ~lcherrium (G Web) Hampe
(9) 13 394656 (57)70 87 88 89 90 93 (119) 136 141
47
Radula obconica Sullo Reboulia hemlsphaerlca (L) Raddi
39 46 88 89 (43) 46 (59) 70 90 t Cf 124 125 (139)
Riccardia ~atifrons Lindb
1 27 39 46 89
Riccardia multifi~ (L) S Gray
39 46 88 145
Riccia beyrichiana Hampe ~ Lehm
46 Riccardia palmata (Hedw) Carruth
39 46 90
48
Riccia bifurca Hoffm Riccia cavernosa Hoffm
48 13739 46 79 137
Riccia fluitans L
242731454679
Riccla frostli Aust
48 79
Riccia sullicantii Aust
79 (137) Riccia sorocarpa Bisch 46 (137)
49
Ricciocarpus natans (1) Corda Sca~ia apiculata Spruce
13 27 46 79 89 110 39 46 (139) 141
ScaJltll1ia carinthica Jack ~ Lindb
137 t 141
Scapania ~ (Mart) Durn
39467990137141
Scapania digenil Schiffn
90
ScaEsectflia cuspiduligera (Nees) K Mull
46 (139) (140) (141)
50
Scapania glaucoeerha1a (Tayl) Aust Seapania gyrrmostomophilia Kaal
39 46 (139) 141 41 44 46 64 (139) (140) 141
ScaP3poundlia irrigua Gott ~ almiddot
7 39 46 79 137 (140L 141
Seapania nemorosa (L) Dum
1 39 46 89 90 137
Scapmi~ ~~ta Smiddotlch
137 141 51
~caPnia poundludicola LDeske amp K Mull Scapania saxicola Schust
46 (66) 141 136 141
Sc~~ scandica (H Arnell ~ Buch) Macv
139 140
Scapania subalpina (Nees) Durn
39 40 46 79 137 141
Scapania undulata (L) Dum
1 42 46 79 141
Scapania umbrosa (Schrad) Dum
137 140 141
52
Trichocolea tomentella (Ehrh) Dum Triomarla exsecta (Schmid) Schiffn
27 (35) 39 46 47 88 137 141 8 28 39 46 90 140 141
Tritomaria exsectiformis (Breidl) Loe~ke Tritomaria Sluinguedentata (Huds) Buch
39 -t6 89 90 136 140 141 7 39 46 141
Trltomaria scitula (Tayl) Joerg
46 136 (139) 141
53
bull bull WSHeV~N S vE II
0 ~
e~RRON ~USK
(J~ (j -
CHIPPE A DUNN
0 o0 poundAU CLAIRE
~ TREMshyPEAshyLEAU
MICE
ONEIDA
Q LINCOLN
TAYLOR
Q t-~----- MOIRA THON
CLARK
o
10
LA FAVETTE
Hornwor ts amp Liverworts
- None - 1 - 25 taxa
- 26 - 50 tRXshy
- 51 - 75 tax)
- 7c - 100 taxa
FONDOU j LAC ~
DODGE
i O JEfFERSON
WALWORTH
30
Anastrophylltlfl helleranum (Nees) Schust Anastrophyllum michauxil (We~ Buch ~
46 88 136 141 Evans 38 46 141
Anastrophyllum mlnut~ (Schreb) Schuster Aneura plnguls (L) Durn
89 21 384687 88 89 90 (138) (145)
Anthoceros macounil MA Howe ~oceros punctatus L
21 (45) (144) 34
31
Barbilophozia attenuata (Mart) Loeske Barbilophozia barbata (Schmid ~ Schre~
24 141 Loeske 28 39 46 89 90 141
Bazzania trilobata (1) S Gray
1 26 39 46 (57) (59) 79 88 89
90 119 141
BlePbarostoma trichoffiyllum (1) Dumort
1 8 26 39 46 89 90 137 141
Blasia pusilla 1
1 39 46 90
Calypogeja fissa (1) Raddi
90
32
$129geja integristipula Steph C~Ypound2geja muelleriana (Schifm) K Mull
88 89 89
CallpoundO~eja neesiana (Mass amp Carest) K Mull
39 46 89 140 (141)
~pogeja sJlapound2icola (H Arnell tl J bull
Perss) Warnst et Loeske (119) 141
CeIalozia biscuspidata (L) Dum
1 37 I 46 90 t 137 141
CalYpogej~ trichomanis (L) Corda
26 39 (119) 137
33
CeEhalozia catenulata (Hub) Lindb Cephalozia connivens (Dicks) Lindb
39 46 79 141 39 4679 88 (89) 90 119 141
Cephalozia lacinulata Jack ~ Spruce
141 Cephalozia lunulifolia (Dum~ Dum I 28 39 46 79 88 89 90 141
Cepoundhalozia macouni (Aust) Aust Cephalozia ~leniceps (Aust) Lindb 27~ 3746 (66)7987 (139)141 39 46 79 89 141
34
Ceffialoziella 9ivaricata (Sm) Schiffn Cephaloziella ~hista (Jack) Schiffn
46 90 46 (119) 141
Cephaloziella hampeana (Nees) Schiffn
46
Cephaloziella rubella (Nees) Warnst
46 39
Cephaloziella rubella var sullivantii
(Aust) K Mull 46 39
Cephaloziella rubella var bifida (Lindb) Douin
35
46
Chiloscyphus pallescens (Ehrh ~ Hof~ Dum Chiloscyphus pallescens var fragilis
39J 46 88 89J 90 (Roth) K Milll 46
Chiloscyphus E9lyanthos (L) Corda
1 24 39 46
Chiloscyphus polyanthos vax rivularis
(Schrad) Nees 39 46
poundpoundnocepha1um conicum (1) Lindb
12 28 39 46 (49) 87 88 8990 120 121 148
Cololejeunea biddlecomiae (Aust) Evans
39 46 62 87 88 89 90
36
Dipl0]hyllum apicu1atum (Evans) Steph Fossombronia foveo1ata Lindb
(35)41 46 90 140 1 t 38 (39) t 46
Frullania bolanderi Aust
13 46 62 88 90 139 140
Frullanla ari ttoniae Evans
39 46 69 (71) 87 88 89 90
Frullania eboracensis Gott
1 9 28 4 39 46 (57) (59) 69 70 (71) 87 88 89 90 92 93 97 148
Frullania inflata Gott
46 87 88 89 (139)
Frullania oakesiana Aust Frullania plana Sullo
(35) 42 46 87 (139) (140) 92
Frullania ri paria Hampe ~ Lehm
46 69 71 87 89
Frullania ~elwynianna Pears
46 89
Geocalyx SEaveolens (Schrad) Nees
394688 8990 (119) 140
Frullania tamarisci (L) Dum subsp sPagrayana
(Mont) Hatt 28 34 l~6 t (57) 88 89 90
38
Mar pan thus seutatUB ( Web amp Mohr) Spruce Jamesoniella autumnalis (DC) Steph
(35) 39 46 90 1 7 28 39 46 56 (57) 87 8889 90 (119) 137 141
Jungermannia confertissima Nees
137 141
Jungermannia crenuliform1s Aust
79 137 141
Jungermannia ~il1ima Sm
46 90 141 Jungermannia exsert1folia subsp corifo~ (Dum~
Vana (35) 46 79 141
39
Jungermannia hyalina Lyell Jungerrr6nnia 1eiantha Grolle
40 46 79 136 141 39 46 79 88 90 1)7 141
Jungermannia IXlaxis Lindb Jungermannia pumila With
46 67 79 (139) (140) 141 )9 46 90 1)6 141
Jungermannia sphaerocarpa Hook Kurzia setacea (Web) Grolle
4679136140IIH 136
Lejeunea cavifo1ia (Ehrh) Lindb Lepidozia reptans (1) Dum
1 7 27 39 46 88 89 90 137 14010 I ( 27) 46 I 69 (71) 88
lophocolea heterophylla (Schrad) Dum Lophocolea minor Nees
1 )4 39 46 (57) 87 88 89 27 39 46 88 89 90 (119) 90 (93)94 119 136
Lophozia al~stris (Schleich ~ Web) Evans Lophozia ascendens (Warnst) Schust
46 90 137 141 136 140 141
41
loIilozia badensls (Gatt ~ Rabenh)
46 141
lophozia blcrenata (Schmid sectA Hoffm)
Dum 46 141
lophozia collaris (Nees) Dum
39 46 141
lophozla excisa (Dicks) Dum
27 4679 141
lophozia gillmani (Aust) Schust lophozia gIlttulata (Undo tl H Arnell)
36 39 40 46 136 (140) 141 Evans 39 46 136 141
42
141
Lophozia heterocolEDs (Thed) MA Howe IoPhozia ipcisa (Schard) Dum
27 39 46 89 141 8 27 31 46 79 90 140 141
LoQhozia longidens Macoun Lophozia ventricosa
1 46 79 90 141
Lophozia ventricosa var silvicola
(Buch) Jones 141 LoEhozla ventricosa vax longif1ora (Nees)
Macoun 39 46 79 141
43
~~ fragrans (Balbis) Frye et Clark Mannia pi 10sa (Hornem) Frye et Clark
46 (139) (140) 9 46 79 116 139
Mannia triandra (Scop) Gro 11e
46 (140)
Marchantia pound91ymorpha L
1 13 28 3946 (49) 78 87 88 89 90 (Ill) 121 122 147
Mo~rckia hibernica (HookJ Gott
46 89
Metzgeria conjugata Lindb
44
46
Mylia anomala (Hook) S Gray Notothylas orbicularis (Schwein) Sullo
39 46 141 (35)45 46
Nowellia curvifolia (Dicks) Mitt
17 27 3846 88 89 90 141
Odontoschisma denudatuJI1 (Nees ~ Mart)
Dum 46 81 137 141
Pallavicinia lyellii (Hook~ Carruth
90
Odontoschisma macounii (Aust) Under
45 46 (66) (139) 140 141
45
=
Pellia epiphllla (L) CordaPellia endiviifolia (Dicks~ Dum
27 39 46 90 13724 39 46
Pellia neesiana Limp bull Phaeoceros
39 46 89 27 39 45 46 97
Porella pinnata L
13 27 I 46 90 Plagiochila asplenioides (L) Dum
27 I 39 1+6 88 89 I 90 137
46
~e11a platYEhylla (L) Pfeiff Porella platyphylloi~ (Schwein) Lindb
17 27 39 46 69 (71)79 87 88 89 9 34 46 (57) 59 87 88 89 92 90 92 93 77
Preissia quadrata (Seop) Nees
1 36 3946 (71) 88 89 90 121 123
137
Ptllidium ciliare (L) Hampe
1 27 39 46 132 141
Dum
(22) 28 3946 (57) 87 88 89 90 92
Ptilidium ~lcherrium (G Web) Hampe
(9) 13 394656 (57)70 87 88 89 90 93 (119) 136 141
47
Radula obconica Sullo Reboulia hemlsphaerlca (L) Raddi
39 46 88 89 (43) 46 (59) 70 90 t Cf 124 125 (139)
Riccardia ~atifrons Lindb
1 27 39 46 89
Riccardia multifi~ (L) S Gray
39 46 88 145
Riccia beyrichiana Hampe ~ Lehm
46 Riccardia palmata (Hedw) Carruth
39 46 90
48
Riccia bifurca Hoffm Riccia cavernosa Hoffm
48 13739 46 79 137
Riccia fluitans L
242731454679
Riccla frostli Aust
48 79
Riccia sullicantii Aust
79 (137) Riccia sorocarpa Bisch 46 (137)
49
Ricciocarpus natans (1) Corda Sca~ia apiculata Spruce
13 27 46 79 89 110 39 46 (139) 141
ScaJltll1ia carinthica Jack ~ Lindb
137 t 141
Scapania ~ (Mart) Durn
39467990137141
Scapania digenil Schiffn
90
ScaEsectflia cuspiduligera (Nees) K Mull
46 (139) (140) (141)
50
Scapania glaucoeerha1a (Tayl) Aust Seapania gyrrmostomophilia Kaal
39 46 (139) 141 41 44 46 64 (139) (140) 141
ScaP3poundlia irrigua Gott ~ almiddot
7 39 46 79 137 (140L 141
Seapania nemorosa (L) Dum
1 39 46 89 90 137
Scapmi~ ~~ta Smiddotlch
137 141 51
~caPnia poundludicola LDeske amp K Mull Scapania saxicola Schust
46 (66) 141 136 141
Sc~~ scandica (H Arnell ~ Buch) Macv
139 140
Scapania subalpina (Nees) Durn
39 40 46 79 137 141
Scapania undulata (L) Dum
1 42 46 79 141
Scapania umbrosa (Schrad) Dum
137 140 141
52
Trichocolea tomentella (Ehrh) Dum Triomarla exsecta (Schmid) Schiffn
27 (35) 39 46 47 88 137 141 8 28 39 46 90 140 141
Tritomaria exsectiformis (Breidl) Loe~ke Tritomaria Sluinguedentata (Huds) Buch
39 -t6 89 90 136 140 141 7 39 46 141
Trltomaria scitula (Tayl) Joerg
46 136 (139) 141
53
Anastrophylltlfl helleranum (Nees) Schust Anastrophyllum michauxil (We~ Buch ~
46 88 136 141 Evans 38 46 141
Anastrophyllum mlnut~ (Schreb) Schuster Aneura plnguls (L) Durn
89 21 384687 88 89 90 (138) (145)
Anthoceros macounil MA Howe ~oceros punctatus L
21 (45) (144) 34
31
Barbilophozia attenuata (Mart) Loeske Barbilophozia barbata (Schmid ~ Schre~
24 141 Loeske 28 39 46 89 90 141
Bazzania trilobata (1) S Gray
1 26 39 46 (57) (59) 79 88 89
90 119 141
BlePbarostoma trichoffiyllum (1) Dumort
1 8 26 39 46 89 90 137 141
Blasia pusilla 1
1 39 46 90
Calypogeja fissa (1) Raddi
90
32
$129geja integristipula Steph C~Ypound2geja muelleriana (Schifm) K Mull
88 89 89
CallpoundO~eja neesiana (Mass amp Carest) K Mull
39 46 89 140 (141)
~pogeja sJlapound2icola (H Arnell tl J bull
Perss) Warnst et Loeske (119) 141
CeIalozia biscuspidata (L) Dum
1 37 I 46 90 t 137 141
CalYpogej~ trichomanis (L) Corda
26 39 (119) 137
33
CeEhalozia catenulata (Hub) Lindb Cephalozia connivens (Dicks) Lindb
39 46 79 141 39 4679 88 (89) 90 119 141
Cephalozia lacinulata Jack ~ Spruce
141 Cephalozia lunulifolia (Dum~ Dum I 28 39 46 79 88 89 90 141
Cepoundhalozia macouni (Aust) Aust Cephalozia ~leniceps (Aust) Lindb 27~ 3746 (66)7987 (139)141 39 46 79 89 141
34
Ceffialoziella 9ivaricata (Sm) Schiffn Cephaloziella ~hista (Jack) Schiffn
46 90 46 (119) 141
Cephaloziella hampeana (Nees) Schiffn
46
Cephaloziella rubella (Nees) Warnst
46 39
Cephaloziella rubella var sullivantii
(Aust) K Mull 46 39
Cephaloziella rubella var bifida (Lindb) Douin
35
46
Chiloscyphus pallescens (Ehrh ~ Hof~ Dum Chiloscyphus pallescens var fragilis
39J 46 88 89J 90 (Roth) K Milll 46
Chiloscyphus E9lyanthos (L) Corda
1 24 39 46
Chiloscyphus polyanthos vax rivularis
(Schrad) Nees 39 46
poundpoundnocepha1um conicum (1) Lindb
12 28 39 46 (49) 87 88 8990 120 121 148
Cololejeunea biddlecomiae (Aust) Evans
39 46 62 87 88 89 90
36
Dipl0]hyllum apicu1atum (Evans) Steph Fossombronia foveo1ata Lindb
(35)41 46 90 140 1 t 38 (39) t 46
Frullania bolanderi Aust
13 46 62 88 90 139 140
Frullanla ari ttoniae Evans
39 46 69 (71) 87 88 89 90
Frullania eboracensis Gott
1 9 28 4 39 46 (57) (59) 69 70 (71) 87 88 89 90 92 93 97 148
Frullania inflata Gott
46 87 88 89 (139)
Frullania oakesiana Aust Frullania plana Sullo
(35) 42 46 87 (139) (140) 92
Frullania ri paria Hampe ~ Lehm
46 69 71 87 89
Frullania ~elwynianna Pears
46 89
Geocalyx SEaveolens (Schrad) Nees
394688 8990 (119) 140
Frullania tamarisci (L) Dum subsp sPagrayana
(Mont) Hatt 28 34 l~6 t (57) 88 89 90
38
Mar pan thus seutatUB ( Web amp Mohr) Spruce Jamesoniella autumnalis (DC) Steph
(35) 39 46 90 1 7 28 39 46 56 (57) 87 8889 90 (119) 137 141
Jungermannia confertissima Nees
137 141
Jungermannia crenuliform1s Aust
79 137 141
Jungermannia ~il1ima Sm
46 90 141 Jungermannia exsert1folia subsp corifo~ (Dum~
Vana (35) 46 79 141
39
Jungermannia hyalina Lyell Jungerrr6nnia 1eiantha Grolle
40 46 79 136 141 39 46 79 88 90 1)7 141
Jungermannia IXlaxis Lindb Jungermannia pumila With
46 67 79 (139) (140) 141 )9 46 90 1)6 141
Jungermannia sphaerocarpa Hook Kurzia setacea (Web) Grolle
4679136140IIH 136
Lejeunea cavifo1ia (Ehrh) Lindb Lepidozia reptans (1) Dum
1 7 27 39 46 88 89 90 137 14010 I ( 27) 46 I 69 (71) 88
lophocolea heterophylla (Schrad) Dum Lophocolea minor Nees
1 )4 39 46 (57) 87 88 89 27 39 46 88 89 90 (119) 90 (93)94 119 136
Lophozia al~stris (Schleich ~ Web) Evans Lophozia ascendens (Warnst) Schust
46 90 137 141 136 140 141
41
loIilozia badensls (Gatt ~ Rabenh)
46 141
lophozia blcrenata (Schmid sectA Hoffm)
Dum 46 141
lophozia collaris (Nees) Dum
39 46 141
lophozla excisa (Dicks) Dum
27 4679 141
lophozia gillmani (Aust) Schust lophozia gIlttulata (Undo tl H Arnell)
36 39 40 46 136 (140) 141 Evans 39 46 136 141
42
141
Lophozia heterocolEDs (Thed) MA Howe IoPhozia ipcisa (Schard) Dum
27 39 46 89 141 8 27 31 46 79 90 140 141
LoQhozia longidens Macoun Lophozia ventricosa
1 46 79 90 141
Lophozia ventricosa var silvicola
(Buch) Jones 141 LoEhozla ventricosa vax longif1ora (Nees)
Macoun 39 46 79 141
43
~~ fragrans (Balbis) Frye et Clark Mannia pi 10sa (Hornem) Frye et Clark
46 (139) (140) 9 46 79 116 139
Mannia triandra (Scop) Gro 11e
46 (140)
Marchantia pound91ymorpha L
1 13 28 3946 (49) 78 87 88 89 90 (Ill) 121 122 147
Mo~rckia hibernica (HookJ Gott
46 89
Metzgeria conjugata Lindb
44
46
Mylia anomala (Hook) S Gray Notothylas orbicularis (Schwein) Sullo
39 46 141 (35)45 46
Nowellia curvifolia (Dicks) Mitt
17 27 3846 88 89 90 141
Odontoschisma denudatuJI1 (Nees ~ Mart)
Dum 46 81 137 141
Pallavicinia lyellii (Hook~ Carruth
90
Odontoschisma macounii (Aust) Under
45 46 (66) (139) 140 141
45
=
Pellia epiphllla (L) CordaPellia endiviifolia (Dicks~ Dum
27 39 46 90 13724 39 46
Pellia neesiana Limp bull Phaeoceros
39 46 89 27 39 45 46 97
Porella pinnata L
13 27 I 46 90 Plagiochila asplenioides (L) Dum
27 I 39 1+6 88 89 I 90 137
46
~e11a platYEhylla (L) Pfeiff Porella platyphylloi~ (Schwein) Lindb
17 27 39 46 69 (71)79 87 88 89 9 34 46 (57) 59 87 88 89 92 90 92 93 77
Preissia quadrata (Seop) Nees
1 36 3946 (71) 88 89 90 121 123
137
Ptllidium ciliare (L) Hampe
1 27 39 46 132 141
Dum
(22) 28 3946 (57) 87 88 89 90 92
Ptilidium ~lcherrium (G Web) Hampe
(9) 13 394656 (57)70 87 88 89 90 93 (119) 136 141
47
Radula obconica Sullo Reboulia hemlsphaerlca (L) Raddi
39 46 88 89 (43) 46 (59) 70 90 t Cf 124 125 (139)
Riccardia ~atifrons Lindb
1 27 39 46 89
Riccardia multifi~ (L) S Gray
39 46 88 145
Riccia beyrichiana Hampe ~ Lehm
46 Riccardia palmata (Hedw) Carruth
39 46 90
48
Riccia bifurca Hoffm Riccia cavernosa Hoffm
48 13739 46 79 137
Riccia fluitans L
242731454679
Riccla frostli Aust
48 79
Riccia sullicantii Aust
79 (137) Riccia sorocarpa Bisch 46 (137)
49
Ricciocarpus natans (1) Corda Sca~ia apiculata Spruce
13 27 46 79 89 110 39 46 (139) 141
ScaJltll1ia carinthica Jack ~ Lindb
137 t 141
Scapania ~ (Mart) Durn
39467990137141
Scapania digenil Schiffn
90
ScaEsectflia cuspiduligera (Nees) K Mull
46 (139) (140) (141)
50
Scapania glaucoeerha1a (Tayl) Aust Seapania gyrrmostomophilia Kaal
39 46 (139) 141 41 44 46 64 (139) (140) 141
ScaP3poundlia irrigua Gott ~ almiddot
7 39 46 79 137 (140L 141
Seapania nemorosa (L) Dum
1 39 46 89 90 137
Scapmi~ ~~ta Smiddotlch
137 141 51
~caPnia poundludicola LDeske amp K Mull Scapania saxicola Schust
46 (66) 141 136 141
Sc~~ scandica (H Arnell ~ Buch) Macv
139 140
Scapania subalpina (Nees) Durn
39 40 46 79 137 141
Scapania undulata (L) Dum
1 42 46 79 141
Scapania umbrosa (Schrad) Dum
137 140 141
52
Trichocolea tomentella (Ehrh) Dum Triomarla exsecta (Schmid) Schiffn
27 (35) 39 46 47 88 137 141 8 28 39 46 90 140 141
Tritomaria exsectiformis (Breidl) Loe~ke Tritomaria Sluinguedentata (Huds) Buch
39 -t6 89 90 136 140 141 7 39 46 141
Trltomaria scitula (Tayl) Joerg
46 136 (139) 141
53
Barbilophozia attenuata (Mart) Loeske Barbilophozia barbata (Schmid ~ Schre~
24 141 Loeske 28 39 46 89 90 141
Bazzania trilobata (1) S Gray
1 26 39 46 (57) (59) 79 88 89
90 119 141
BlePbarostoma trichoffiyllum (1) Dumort
1 8 26 39 46 89 90 137 141
Blasia pusilla 1
1 39 46 90
Calypogeja fissa (1) Raddi
90
32
$129geja integristipula Steph C~Ypound2geja muelleriana (Schifm) K Mull
88 89 89
CallpoundO~eja neesiana (Mass amp Carest) K Mull
39 46 89 140 (141)
~pogeja sJlapound2icola (H Arnell tl J bull
Perss) Warnst et Loeske (119) 141
CeIalozia biscuspidata (L) Dum
1 37 I 46 90 t 137 141
CalYpogej~ trichomanis (L) Corda
26 39 (119) 137
33
CeEhalozia catenulata (Hub) Lindb Cephalozia connivens (Dicks) Lindb
39 46 79 141 39 4679 88 (89) 90 119 141
Cephalozia lacinulata Jack ~ Spruce
141 Cephalozia lunulifolia (Dum~ Dum I 28 39 46 79 88 89 90 141
Cepoundhalozia macouni (Aust) Aust Cephalozia ~leniceps (Aust) Lindb 27~ 3746 (66)7987 (139)141 39 46 79 89 141
34
Ceffialoziella 9ivaricata (Sm) Schiffn Cephaloziella ~hista (Jack) Schiffn
46 90 46 (119) 141
Cephaloziella hampeana (Nees) Schiffn
46
Cephaloziella rubella (Nees) Warnst
46 39
Cephaloziella rubella var sullivantii
(Aust) K Mull 46 39
Cephaloziella rubella var bifida (Lindb) Douin
35
46
Chiloscyphus pallescens (Ehrh ~ Hof~ Dum Chiloscyphus pallescens var fragilis
39J 46 88 89J 90 (Roth) K Milll 46
Chiloscyphus E9lyanthos (L) Corda
1 24 39 46
Chiloscyphus polyanthos vax rivularis
(Schrad) Nees 39 46
poundpoundnocepha1um conicum (1) Lindb
12 28 39 46 (49) 87 88 8990 120 121 148
Cololejeunea biddlecomiae (Aust) Evans
39 46 62 87 88 89 90
36
Dipl0]hyllum apicu1atum (Evans) Steph Fossombronia foveo1ata Lindb
(35)41 46 90 140 1 t 38 (39) t 46
Frullania bolanderi Aust
13 46 62 88 90 139 140
Frullanla ari ttoniae Evans
39 46 69 (71) 87 88 89 90
Frullania eboracensis Gott
1 9 28 4 39 46 (57) (59) 69 70 (71) 87 88 89 90 92 93 97 148
Frullania inflata Gott
46 87 88 89 (139)
Frullania oakesiana Aust Frullania plana Sullo
(35) 42 46 87 (139) (140) 92
Frullania ri paria Hampe ~ Lehm
46 69 71 87 89
Frullania ~elwynianna Pears
46 89
Geocalyx SEaveolens (Schrad) Nees
394688 8990 (119) 140
Frullania tamarisci (L) Dum subsp sPagrayana
(Mont) Hatt 28 34 l~6 t (57) 88 89 90
38
Mar pan thus seutatUB ( Web amp Mohr) Spruce Jamesoniella autumnalis (DC) Steph
(35) 39 46 90 1 7 28 39 46 56 (57) 87 8889 90 (119) 137 141
Jungermannia confertissima Nees
137 141
Jungermannia crenuliform1s Aust
79 137 141
Jungermannia ~il1ima Sm
46 90 141 Jungermannia exsert1folia subsp corifo~ (Dum~
Vana (35) 46 79 141
39
Jungermannia hyalina Lyell Jungerrr6nnia 1eiantha Grolle
40 46 79 136 141 39 46 79 88 90 1)7 141
Jungermannia IXlaxis Lindb Jungermannia pumila With
46 67 79 (139) (140) 141 )9 46 90 1)6 141
Jungermannia sphaerocarpa Hook Kurzia setacea (Web) Grolle
4679136140IIH 136
Lejeunea cavifo1ia (Ehrh) Lindb Lepidozia reptans (1) Dum
1 7 27 39 46 88 89 90 137 14010 I ( 27) 46 I 69 (71) 88
lophocolea heterophylla (Schrad) Dum Lophocolea minor Nees
1 )4 39 46 (57) 87 88 89 27 39 46 88 89 90 (119) 90 (93)94 119 136
Lophozia al~stris (Schleich ~ Web) Evans Lophozia ascendens (Warnst) Schust
46 90 137 141 136 140 141
41
loIilozia badensls (Gatt ~ Rabenh)
46 141
lophozia blcrenata (Schmid sectA Hoffm)
Dum 46 141
lophozia collaris (Nees) Dum
39 46 141
lophozla excisa (Dicks) Dum
27 4679 141
lophozia gillmani (Aust) Schust lophozia gIlttulata (Undo tl H Arnell)
36 39 40 46 136 (140) 141 Evans 39 46 136 141
42
141
Lophozia heterocolEDs (Thed) MA Howe IoPhozia ipcisa (Schard) Dum
27 39 46 89 141 8 27 31 46 79 90 140 141
LoQhozia longidens Macoun Lophozia ventricosa
1 46 79 90 141
Lophozia ventricosa var silvicola
(Buch) Jones 141 LoEhozla ventricosa vax longif1ora (Nees)
Macoun 39 46 79 141
43
~~ fragrans (Balbis) Frye et Clark Mannia pi 10sa (Hornem) Frye et Clark
46 (139) (140) 9 46 79 116 139
Mannia triandra (Scop) Gro 11e
46 (140)
Marchantia pound91ymorpha L
1 13 28 3946 (49) 78 87 88 89 90 (Ill) 121 122 147
Mo~rckia hibernica (HookJ Gott
46 89
Metzgeria conjugata Lindb
44
46
Mylia anomala (Hook) S Gray Notothylas orbicularis (Schwein) Sullo
39 46 141 (35)45 46
Nowellia curvifolia (Dicks) Mitt
17 27 3846 88 89 90 141
Odontoschisma denudatuJI1 (Nees ~ Mart)
Dum 46 81 137 141
Pallavicinia lyellii (Hook~ Carruth
90
Odontoschisma macounii (Aust) Under
45 46 (66) (139) 140 141
45
=
Pellia epiphllla (L) CordaPellia endiviifolia (Dicks~ Dum
27 39 46 90 13724 39 46
Pellia neesiana Limp bull Phaeoceros
39 46 89 27 39 45 46 97
Porella pinnata L
13 27 I 46 90 Plagiochila asplenioides (L) Dum
27 I 39 1+6 88 89 I 90 137
46
~e11a platYEhylla (L) Pfeiff Porella platyphylloi~ (Schwein) Lindb
17 27 39 46 69 (71)79 87 88 89 9 34 46 (57) 59 87 88 89 92 90 92 93 77
Preissia quadrata (Seop) Nees
1 36 3946 (71) 88 89 90 121 123
137
Ptllidium ciliare (L) Hampe
1 27 39 46 132 141
Dum
(22) 28 3946 (57) 87 88 89 90 92
Ptilidium ~lcherrium (G Web) Hampe
(9) 13 394656 (57)70 87 88 89 90 93 (119) 136 141
47
Radula obconica Sullo Reboulia hemlsphaerlca (L) Raddi
39 46 88 89 (43) 46 (59) 70 90 t Cf 124 125 (139)
Riccardia ~atifrons Lindb
1 27 39 46 89
Riccardia multifi~ (L) S Gray
39 46 88 145
Riccia beyrichiana Hampe ~ Lehm
46 Riccardia palmata (Hedw) Carruth
39 46 90
48
Riccia bifurca Hoffm Riccia cavernosa Hoffm
48 13739 46 79 137
Riccia fluitans L
242731454679
Riccla frostli Aust
48 79
Riccia sullicantii Aust
79 (137) Riccia sorocarpa Bisch 46 (137)
49
Ricciocarpus natans (1) Corda Sca~ia apiculata Spruce
13 27 46 79 89 110 39 46 (139) 141
ScaJltll1ia carinthica Jack ~ Lindb
137 t 141
Scapania ~ (Mart) Durn
39467990137141
Scapania digenil Schiffn
90
ScaEsectflia cuspiduligera (Nees) K Mull
46 (139) (140) (141)
50
Scapania glaucoeerha1a (Tayl) Aust Seapania gyrrmostomophilia Kaal
39 46 (139) 141 41 44 46 64 (139) (140) 141
ScaP3poundlia irrigua Gott ~ almiddot
7 39 46 79 137 (140L 141
Seapania nemorosa (L) Dum
1 39 46 89 90 137
Scapmi~ ~~ta Smiddotlch
137 141 51
~caPnia poundludicola LDeske amp K Mull Scapania saxicola Schust
46 (66) 141 136 141
Sc~~ scandica (H Arnell ~ Buch) Macv
139 140
Scapania subalpina (Nees) Durn
39 40 46 79 137 141
Scapania undulata (L) Dum
1 42 46 79 141
Scapania umbrosa (Schrad) Dum
137 140 141
52
Trichocolea tomentella (Ehrh) Dum Triomarla exsecta (Schmid) Schiffn
27 (35) 39 46 47 88 137 141 8 28 39 46 90 140 141
Tritomaria exsectiformis (Breidl) Loe~ke Tritomaria Sluinguedentata (Huds) Buch
39 -t6 89 90 136 140 141 7 39 46 141
Trltomaria scitula (Tayl) Joerg
46 136 (139) 141
53
$129geja integristipula Steph C~Ypound2geja muelleriana (Schifm) K Mull
88 89 89
CallpoundO~eja neesiana (Mass amp Carest) K Mull
39 46 89 140 (141)
~pogeja sJlapound2icola (H Arnell tl J bull
Perss) Warnst et Loeske (119) 141
CeIalozia biscuspidata (L) Dum
1 37 I 46 90 t 137 141
CalYpogej~ trichomanis (L) Corda
26 39 (119) 137
33
CeEhalozia catenulata (Hub) Lindb Cephalozia connivens (Dicks) Lindb
39 46 79 141 39 4679 88 (89) 90 119 141
Cephalozia lacinulata Jack ~ Spruce
141 Cephalozia lunulifolia (Dum~ Dum I 28 39 46 79 88 89 90 141
Cepoundhalozia macouni (Aust) Aust Cephalozia ~leniceps (Aust) Lindb 27~ 3746 (66)7987 (139)141 39 46 79 89 141
34
Ceffialoziella 9ivaricata (Sm) Schiffn Cephaloziella ~hista (Jack) Schiffn
46 90 46 (119) 141
Cephaloziella hampeana (Nees) Schiffn
46
Cephaloziella rubella (Nees) Warnst
46 39
Cephaloziella rubella var sullivantii
(Aust) K Mull 46 39
Cephaloziella rubella var bifida (Lindb) Douin
35
46
Chiloscyphus pallescens (Ehrh ~ Hof~ Dum Chiloscyphus pallescens var fragilis
39J 46 88 89J 90 (Roth) K Milll 46
Chiloscyphus E9lyanthos (L) Corda
1 24 39 46
Chiloscyphus polyanthos vax rivularis
(Schrad) Nees 39 46
poundpoundnocepha1um conicum (1) Lindb
12 28 39 46 (49) 87 88 8990 120 121 148
Cololejeunea biddlecomiae (Aust) Evans
39 46 62 87 88 89 90
36
Dipl0]hyllum apicu1atum (Evans) Steph Fossombronia foveo1ata Lindb
(35)41 46 90 140 1 t 38 (39) t 46
Frullania bolanderi Aust
13 46 62 88 90 139 140
Frullanla ari ttoniae Evans
39 46 69 (71) 87 88 89 90
Frullania eboracensis Gott
1 9 28 4 39 46 (57) (59) 69 70 (71) 87 88 89 90 92 93 97 148
Frullania inflata Gott
46 87 88 89 (139)
Frullania oakesiana Aust Frullania plana Sullo
(35) 42 46 87 (139) (140) 92
Frullania ri paria Hampe ~ Lehm
46 69 71 87 89
Frullania ~elwynianna Pears
46 89
Geocalyx SEaveolens (Schrad) Nees
394688 8990 (119) 140
Frullania tamarisci (L) Dum subsp sPagrayana
(Mont) Hatt 28 34 l~6 t (57) 88 89 90
38
Mar pan thus seutatUB ( Web amp Mohr) Spruce Jamesoniella autumnalis (DC) Steph
(35) 39 46 90 1 7 28 39 46 56 (57) 87 8889 90 (119) 137 141
Jungermannia confertissima Nees
137 141
Jungermannia crenuliform1s Aust
79 137 141
Jungermannia ~il1ima Sm
46 90 141 Jungermannia exsert1folia subsp corifo~ (Dum~
Vana (35) 46 79 141
39
Jungermannia hyalina Lyell Jungerrr6nnia 1eiantha Grolle
40 46 79 136 141 39 46 79 88 90 1)7 141
Jungermannia IXlaxis Lindb Jungermannia pumila With
46 67 79 (139) (140) 141 )9 46 90 1)6 141
Jungermannia sphaerocarpa Hook Kurzia setacea (Web) Grolle
4679136140IIH 136
Lejeunea cavifo1ia (Ehrh) Lindb Lepidozia reptans (1) Dum
1 7 27 39 46 88 89 90 137 14010 I ( 27) 46 I 69 (71) 88
lophocolea heterophylla (Schrad) Dum Lophocolea minor Nees
1 )4 39 46 (57) 87 88 89 27 39 46 88 89 90 (119) 90 (93)94 119 136
Lophozia al~stris (Schleich ~ Web) Evans Lophozia ascendens (Warnst) Schust
46 90 137 141 136 140 141
41
loIilozia badensls (Gatt ~ Rabenh)
46 141
lophozia blcrenata (Schmid sectA Hoffm)
Dum 46 141
lophozia collaris (Nees) Dum
39 46 141
lophozla excisa (Dicks) Dum
27 4679 141
lophozia gillmani (Aust) Schust lophozia gIlttulata (Undo tl H Arnell)
36 39 40 46 136 (140) 141 Evans 39 46 136 141
42
141
Lophozia heterocolEDs (Thed) MA Howe IoPhozia ipcisa (Schard) Dum
27 39 46 89 141 8 27 31 46 79 90 140 141
LoQhozia longidens Macoun Lophozia ventricosa
1 46 79 90 141
Lophozia ventricosa var silvicola
(Buch) Jones 141 LoEhozla ventricosa vax longif1ora (Nees)
Macoun 39 46 79 141
43
~~ fragrans (Balbis) Frye et Clark Mannia pi 10sa (Hornem) Frye et Clark
46 (139) (140) 9 46 79 116 139
Mannia triandra (Scop) Gro 11e
46 (140)
Marchantia pound91ymorpha L
1 13 28 3946 (49) 78 87 88 89 90 (Ill) 121 122 147
Mo~rckia hibernica (HookJ Gott
46 89
Metzgeria conjugata Lindb
44
46
Mylia anomala (Hook) S Gray Notothylas orbicularis (Schwein) Sullo
39 46 141 (35)45 46
Nowellia curvifolia (Dicks) Mitt
17 27 3846 88 89 90 141
Odontoschisma denudatuJI1 (Nees ~ Mart)
Dum 46 81 137 141
Pallavicinia lyellii (Hook~ Carruth
90
Odontoschisma macounii (Aust) Under
45 46 (66) (139) 140 141
45
=
Pellia epiphllla (L) CordaPellia endiviifolia (Dicks~ Dum
27 39 46 90 13724 39 46
Pellia neesiana Limp bull Phaeoceros
39 46 89 27 39 45 46 97
Porella pinnata L
13 27 I 46 90 Plagiochila asplenioides (L) Dum
27 I 39 1+6 88 89 I 90 137
46
~e11a platYEhylla (L) Pfeiff Porella platyphylloi~ (Schwein) Lindb
17 27 39 46 69 (71)79 87 88 89 9 34 46 (57) 59 87 88 89 92 90 92 93 77
Preissia quadrata (Seop) Nees
1 36 3946 (71) 88 89 90 121 123
137
Ptllidium ciliare (L) Hampe
1 27 39 46 132 141
Dum
(22) 28 3946 (57) 87 88 89 90 92
Ptilidium ~lcherrium (G Web) Hampe
(9) 13 394656 (57)70 87 88 89 90 93 (119) 136 141
47
Radula obconica Sullo Reboulia hemlsphaerlca (L) Raddi
39 46 88 89 (43) 46 (59) 70 90 t Cf 124 125 (139)
Riccardia ~atifrons Lindb
1 27 39 46 89
Riccardia multifi~ (L) S Gray
39 46 88 145
Riccia beyrichiana Hampe ~ Lehm
46 Riccardia palmata (Hedw) Carruth
39 46 90
48
Riccia bifurca Hoffm Riccia cavernosa Hoffm
48 13739 46 79 137
Riccia fluitans L
242731454679
Riccla frostli Aust
48 79
Riccia sullicantii Aust
79 (137) Riccia sorocarpa Bisch 46 (137)
49
Ricciocarpus natans (1) Corda Sca~ia apiculata Spruce
13 27 46 79 89 110 39 46 (139) 141
ScaJltll1ia carinthica Jack ~ Lindb
137 t 141
Scapania ~ (Mart) Durn
39467990137141
Scapania digenil Schiffn
90
ScaEsectflia cuspiduligera (Nees) K Mull
46 (139) (140) (141)
50
Scapania glaucoeerha1a (Tayl) Aust Seapania gyrrmostomophilia Kaal
39 46 (139) 141 41 44 46 64 (139) (140) 141
ScaP3poundlia irrigua Gott ~ almiddot
7 39 46 79 137 (140L 141
Seapania nemorosa (L) Dum
1 39 46 89 90 137
Scapmi~ ~~ta Smiddotlch
137 141 51
~caPnia poundludicola LDeske amp K Mull Scapania saxicola Schust
46 (66) 141 136 141
Sc~~ scandica (H Arnell ~ Buch) Macv
139 140
Scapania subalpina (Nees) Durn
39 40 46 79 137 141
Scapania undulata (L) Dum
1 42 46 79 141
Scapania umbrosa (Schrad) Dum
137 140 141
52
Trichocolea tomentella (Ehrh) Dum Triomarla exsecta (Schmid) Schiffn
27 (35) 39 46 47 88 137 141 8 28 39 46 90 140 141
Tritomaria exsectiformis (Breidl) Loe~ke Tritomaria Sluinguedentata (Huds) Buch
39 -t6 89 90 136 140 141 7 39 46 141
Trltomaria scitula (Tayl) Joerg
46 136 (139) 141
53
CeEhalozia catenulata (Hub) Lindb Cephalozia connivens (Dicks) Lindb
39 46 79 141 39 4679 88 (89) 90 119 141
Cephalozia lacinulata Jack ~ Spruce
141 Cephalozia lunulifolia (Dum~ Dum I 28 39 46 79 88 89 90 141
Cepoundhalozia macouni (Aust) Aust Cephalozia ~leniceps (Aust) Lindb 27~ 3746 (66)7987 (139)141 39 46 79 89 141
34
Ceffialoziella 9ivaricata (Sm) Schiffn Cephaloziella ~hista (Jack) Schiffn
46 90 46 (119) 141
Cephaloziella hampeana (Nees) Schiffn
46
Cephaloziella rubella (Nees) Warnst
46 39
Cephaloziella rubella var sullivantii
(Aust) K Mull 46 39
Cephaloziella rubella var bifida (Lindb) Douin
35
46
Chiloscyphus pallescens (Ehrh ~ Hof~ Dum Chiloscyphus pallescens var fragilis
39J 46 88 89J 90 (Roth) K Milll 46
Chiloscyphus E9lyanthos (L) Corda
1 24 39 46
Chiloscyphus polyanthos vax rivularis
(Schrad) Nees 39 46
poundpoundnocepha1um conicum (1) Lindb
12 28 39 46 (49) 87 88 8990 120 121 148
Cololejeunea biddlecomiae (Aust) Evans
39 46 62 87 88 89 90
36
Dipl0]hyllum apicu1atum (Evans) Steph Fossombronia foveo1ata Lindb
(35)41 46 90 140 1 t 38 (39) t 46
Frullania bolanderi Aust
13 46 62 88 90 139 140
Frullanla ari ttoniae Evans
39 46 69 (71) 87 88 89 90
Frullania eboracensis Gott
1 9 28 4 39 46 (57) (59) 69 70 (71) 87 88 89 90 92 93 97 148
Frullania inflata Gott
46 87 88 89 (139)
Frullania oakesiana Aust Frullania plana Sullo
(35) 42 46 87 (139) (140) 92
Frullania ri paria Hampe ~ Lehm
46 69 71 87 89
Frullania ~elwynianna Pears
46 89
Geocalyx SEaveolens (Schrad) Nees
394688 8990 (119) 140
Frullania tamarisci (L) Dum subsp sPagrayana
(Mont) Hatt 28 34 l~6 t (57) 88 89 90
38
Mar pan thus seutatUB ( Web amp Mohr) Spruce Jamesoniella autumnalis (DC) Steph
(35) 39 46 90 1 7 28 39 46 56 (57) 87 8889 90 (119) 137 141
Jungermannia confertissima Nees
137 141
Jungermannia crenuliform1s Aust
79 137 141
Jungermannia ~il1ima Sm
46 90 141 Jungermannia exsert1folia subsp corifo~ (Dum~
Vana (35) 46 79 141
39
Jungermannia hyalina Lyell Jungerrr6nnia 1eiantha Grolle
40 46 79 136 141 39 46 79 88 90 1)7 141
Jungermannia IXlaxis Lindb Jungermannia pumila With
46 67 79 (139) (140) 141 )9 46 90 1)6 141
Jungermannia sphaerocarpa Hook Kurzia setacea (Web) Grolle
4679136140IIH 136
Lejeunea cavifo1ia (Ehrh) Lindb Lepidozia reptans (1) Dum
1 7 27 39 46 88 89 90 137 14010 I ( 27) 46 I 69 (71) 88
lophocolea heterophylla (Schrad) Dum Lophocolea minor Nees
1 )4 39 46 (57) 87 88 89 27 39 46 88 89 90 (119) 90 (93)94 119 136
Lophozia al~stris (Schleich ~ Web) Evans Lophozia ascendens (Warnst) Schust
46 90 137 141 136 140 141
41
loIilozia badensls (Gatt ~ Rabenh)
46 141
lophozia blcrenata (Schmid sectA Hoffm)
Dum 46 141
lophozia collaris (Nees) Dum
39 46 141
lophozla excisa (Dicks) Dum
27 4679 141
lophozia gillmani (Aust) Schust lophozia gIlttulata (Undo tl H Arnell)
36 39 40 46 136 (140) 141 Evans 39 46 136 141
42
141
Lophozia heterocolEDs (Thed) MA Howe IoPhozia ipcisa (Schard) Dum
27 39 46 89 141 8 27 31 46 79 90 140 141
LoQhozia longidens Macoun Lophozia ventricosa
1 46 79 90 141
Lophozia ventricosa var silvicola
(Buch) Jones 141 LoEhozla ventricosa vax longif1ora (Nees)
Macoun 39 46 79 141
43
~~ fragrans (Balbis) Frye et Clark Mannia pi 10sa (Hornem) Frye et Clark
46 (139) (140) 9 46 79 116 139
Mannia triandra (Scop) Gro 11e
46 (140)
Marchantia pound91ymorpha L
1 13 28 3946 (49) 78 87 88 89 90 (Ill) 121 122 147
Mo~rckia hibernica (HookJ Gott
46 89
Metzgeria conjugata Lindb
44
46
Mylia anomala (Hook) S Gray Notothylas orbicularis (Schwein) Sullo
39 46 141 (35)45 46
Nowellia curvifolia (Dicks) Mitt
17 27 3846 88 89 90 141
Odontoschisma denudatuJI1 (Nees ~ Mart)
Dum 46 81 137 141
Pallavicinia lyellii (Hook~ Carruth
90
Odontoschisma macounii (Aust) Under
45 46 (66) (139) 140 141
45
=
Pellia epiphllla (L) CordaPellia endiviifolia (Dicks~ Dum
27 39 46 90 13724 39 46
Pellia neesiana Limp bull Phaeoceros
39 46 89 27 39 45 46 97
Porella pinnata L
13 27 I 46 90 Plagiochila asplenioides (L) Dum
27 I 39 1+6 88 89 I 90 137
46
~e11a platYEhylla (L) Pfeiff Porella platyphylloi~ (Schwein) Lindb
17 27 39 46 69 (71)79 87 88 89 9 34 46 (57) 59 87 88 89 92 90 92 93 77
Preissia quadrata (Seop) Nees
1 36 3946 (71) 88 89 90 121 123
137
Ptllidium ciliare (L) Hampe
1 27 39 46 132 141
Dum
(22) 28 3946 (57) 87 88 89 90 92
Ptilidium ~lcherrium (G Web) Hampe
(9) 13 394656 (57)70 87 88 89 90 93 (119) 136 141
47
Radula obconica Sullo Reboulia hemlsphaerlca (L) Raddi
39 46 88 89 (43) 46 (59) 70 90 t Cf 124 125 (139)
Riccardia ~atifrons Lindb
1 27 39 46 89
Riccardia multifi~ (L) S Gray
39 46 88 145
Riccia beyrichiana Hampe ~ Lehm
46 Riccardia palmata (Hedw) Carruth
39 46 90
48
Riccia bifurca Hoffm Riccia cavernosa Hoffm
48 13739 46 79 137
Riccia fluitans L
242731454679
Riccla frostli Aust
48 79
Riccia sullicantii Aust
79 (137) Riccia sorocarpa Bisch 46 (137)
49
Ricciocarpus natans (1) Corda Sca~ia apiculata Spruce
13 27 46 79 89 110 39 46 (139) 141
ScaJltll1ia carinthica Jack ~ Lindb
137 t 141
Scapania ~ (Mart) Durn
39467990137141
Scapania digenil Schiffn
90
ScaEsectflia cuspiduligera (Nees) K Mull
46 (139) (140) (141)
50
Scapania glaucoeerha1a (Tayl) Aust Seapania gyrrmostomophilia Kaal
39 46 (139) 141 41 44 46 64 (139) (140) 141
ScaP3poundlia irrigua Gott ~ almiddot
7 39 46 79 137 (140L 141
Seapania nemorosa (L) Dum
1 39 46 89 90 137
Scapmi~ ~~ta Smiddotlch
137 141 51
~caPnia poundludicola LDeske amp K Mull Scapania saxicola Schust
46 (66) 141 136 141
Sc~~ scandica (H Arnell ~ Buch) Macv
139 140
Scapania subalpina (Nees) Durn
39 40 46 79 137 141
Scapania undulata (L) Dum
1 42 46 79 141
Scapania umbrosa (Schrad) Dum
137 140 141
52
Trichocolea tomentella (Ehrh) Dum Triomarla exsecta (Schmid) Schiffn
27 (35) 39 46 47 88 137 141 8 28 39 46 90 140 141
Tritomaria exsectiformis (Breidl) Loe~ke Tritomaria Sluinguedentata (Huds) Buch
39 -t6 89 90 136 140 141 7 39 46 141
Trltomaria scitula (Tayl) Joerg
46 136 (139) 141
53
Ceffialoziella 9ivaricata (Sm) Schiffn Cephaloziella ~hista (Jack) Schiffn
46 90 46 (119) 141
Cephaloziella hampeana (Nees) Schiffn
46
Cephaloziella rubella (Nees) Warnst
46 39
Cephaloziella rubella var sullivantii
(Aust) K Mull 46 39
Cephaloziella rubella var bifida (Lindb) Douin
35
46
Chiloscyphus pallescens (Ehrh ~ Hof~ Dum Chiloscyphus pallescens var fragilis
39J 46 88 89J 90 (Roth) K Milll 46
Chiloscyphus E9lyanthos (L) Corda
1 24 39 46
Chiloscyphus polyanthos vax rivularis
(Schrad) Nees 39 46
poundpoundnocepha1um conicum (1) Lindb
12 28 39 46 (49) 87 88 8990 120 121 148
Cololejeunea biddlecomiae (Aust) Evans
39 46 62 87 88 89 90
36
Dipl0]hyllum apicu1atum (Evans) Steph Fossombronia foveo1ata Lindb
(35)41 46 90 140 1 t 38 (39) t 46
Frullania bolanderi Aust
13 46 62 88 90 139 140
Frullanla ari ttoniae Evans
39 46 69 (71) 87 88 89 90
Frullania eboracensis Gott
1 9 28 4 39 46 (57) (59) 69 70 (71) 87 88 89 90 92 93 97 148
Frullania inflata Gott
46 87 88 89 (139)
Frullania oakesiana Aust Frullania plana Sullo
(35) 42 46 87 (139) (140) 92
Frullania ri paria Hampe ~ Lehm
46 69 71 87 89
Frullania ~elwynianna Pears
46 89
Geocalyx SEaveolens (Schrad) Nees
394688 8990 (119) 140
Frullania tamarisci (L) Dum subsp sPagrayana
(Mont) Hatt 28 34 l~6 t (57) 88 89 90
38
Mar pan thus seutatUB ( Web amp Mohr) Spruce Jamesoniella autumnalis (DC) Steph
(35) 39 46 90 1 7 28 39 46 56 (57) 87 8889 90 (119) 137 141
Jungermannia confertissima Nees
137 141
Jungermannia crenuliform1s Aust
79 137 141
Jungermannia ~il1ima Sm
46 90 141 Jungermannia exsert1folia subsp corifo~ (Dum~
Vana (35) 46 79 141
39
Jungermannia hyalina Lyell Jungerrr6nnia 1eiantha Grolle
40 46 79 136 141 39 46 79 88 90 1)7 141
Jungermannia IXlaxis Lindb Jungermannia pumila With
46 67 79 (139) (140) 141 )9 46 90 1)6 141
Jungermannia sphaerocarpa Hook Kurzia setacea (Web) Grolle
4679136140IIH 136
Lejeunea cavifo1ia (Ehrh) Lindb Lepidozia reptans (1) Dum
1 7 27 39 46 88 89 90 137 14010 I ( 27) 46 I 69 (71) 88
lophocolea heterophylla (Schrad) Dum Lophocolea minor Nees
1 )4 39 46 (57) 87 88 89 27 39 46 88 89 90 (119) 90 (93)94 119 136
Lophozia al~stris (Schleich ~ Web) Evans Lophozia ascendens (Warnst) Schust
46 90 137 141 136 140 141
41
loIilozia badensls (Gatt ~ Rabenh)
46 141
lophozia blcrenata (Schmid sectA Hoffm)
Dum 46 141
lophozia collaris (Nees) Dum
39 46 141
lophozla excisa (Dicks) Dum
27 4679 141
lophozia gillmani (Aust) Schust lophozia gIlttulata (Undo tl H Arnell)
36 39 40 46 136 (140) 141 Evans 39 46 136 141
42
141
Lophozia heterocolEDs (Thed) MA Howe IoPhozia ipcisa (Schard) Dum
27 39 46 89 141 8 27 31 46 79 90 140 141
LoQhozia longidens Macoun Lophozia ventricosa
1 46 79 90 141
Lophozia ventricosa var silvicola
(Buch) Jones 141 LoEhozla ventricosa vax longif1ora (Nees)
Macoun 39 46 79 141
43
~~ fragrans (Balbis) Frye et Clark Mannia pi 10sa (Hornem) Frye et Clark
46 (139) (140) 9 46 79 116 139
Mannia triandra (Scop) Gro 11e
46 (140)
Marchantia pound91ymorpha L
1 13 28 3946 (49) 78 87 88 89 90 (Ill) 121 122 147
Mo~rckia hibernica (HookJ Gott
46 89
Metzgeria conjugata Lindb
44
46
Mylia anomala (Hook) S Gray Notothylas orbicularis (Schwein) Sullo
39 46 141 (35)45 46
Nowellia curvifolia (Dicks) Mitt
17 27 3846 88 89 90 141
Odontoschisma denudatuJI1 (Nees ~ Mart)
Dum 46 81 137 141
Pallavicinia lyellii (Hook~ Carruth
90
Odontoschisma macounii (Aust) Under
45 46 (66) (139) 140 141
45
=
Pellia epiphllla (L) CordaPellia endiviifolia (Dicks~ Dum
27 39 46 90 13724 39 46
Pellia neesiana Limp bull Phaeoceros
39 46 89 27 39 45 46 97
Porella pinnata L
13 27 I 46 90 Plagiochila asplenioides (L) Dum
27 I 39 1+6 88 89 I 90 137
46
~e11a platYEhylla (L) Pfeiff Porella platyphylloi~ (Schwein) Lindb
17 27 39 46 69 (71)79 87 88 89 9 34 46 (57) 59 87 88 89 92 90 92 93 77
Preissia quadrata (Seop) Nees
1 36 3946 (71) 88 89 90 121 123
137
Ptllidium ciliare (L) Hampe
1 27 39 46 132 141
Dum
(22) 28 3946 (57) 87 88 89 90 92
Ptilidium ~lcherrium (G Web) Hampe
(9) 13 394656 (57)70 87 88 89 90 93 (119) 136 141
47
Radula obconica Sullo Reboulia hemlsphaerlca (L) Raddi
39 46 88 89 (43) 46 (59) 70 90 t Cf 124 125 (139)
Riccardia ~atifrons Lindb
1 27 39 46 89
Riccardia multifi~ (L) S Gray
39 46 88 145
Riccia beyrichiana Hampe ~ Lehm
46 Riccardia palmata (Hedw) Carruth
39 46 90
48
Riccia bifurca Hoffm Riccia cavernosa Hoffm
48 13739 46 79 137
Riccia fluitans L
242731454679
Riccla frostli Aust
48 79
Riccia sullicantii Aust
79 (137) Riccia sorocarpa Bisch 46 (137)
49
Ricciocarpus natans (1) Corda Sca~ia apiculata Spruce
13 27 46 79 89 110 39 46 (139) 141
ScaJltll1ia carinthica Jack ~ Lindb
137 t 141
Scapania ~ (Mart) Durn
39467990137141
Scapania digenil Schiffn
90
ScaEsectflia cuspiduligera (Nees) K Mull
46 (139) (140) (141)
50
Scapania glaucoeerha1a (Tayl) Aust Seapania gyrrmostomophilia Kaal
39 46 (139) 141 41 44 46 64 (139) (140) 141
ScaP3poundlia irrigua Gott ~ almiddot
7 39 46 79 137 (140L 141
Seapania nemorosa (L) Dum
1 39 46 89 90 137
Scapmi~ ~~ta Smiddotlch
137 141 51
~caPnia poundludicola LDeske amp K Mull Scapania saxicola Schust
46 (66) 141 136 141
Sc~~ scandica (H Arnell ~ Buch) Macv
139 140
Scapania subalpina (Nees) Durn
39 40 46 79 137 141
Scapania undulata (L) Dum
1 42 46 79 141
Scapania umbrosa (Schrad) Dum
137 140 141
52
Trichocolea tomentella (Ehrh) Dum Triomarla exsecta (Schmid) Schiffn
27 (35) 39 46 47 88 137 141 8 28 39 46 90 140 141
Tritomaria exsectiformis (Breidl) Loe~ke Tritomaria Sluinguedentata (Huds) Buch
39 -t6 89 90 136 140 141 7 39 46 141
Trltomaria scitula (Tayl) Joerg
46 136 (139) 141
53
Chiloscyphus pallescens (Ehrh ~ Hof~ Dum Chiloscyphus pallescens var fragilis
39J 46 88 89J 90 (Roth) K Milll 46
Chiloscyphus E9lyanthos (L) Corda
1 24 39 46
Chiloscyphus polyanthos vax rivularis
(Schrad) Nees 39 46
poundpoundnocepha1um conicum (1) Lindb
12 28 39 46 (49) 87 88 8990 120 121 148
Cololejeunea biddlecomiae (Aust) Evans
39 46 62 87 88 89 90
36
Dipl0]hyllum apicu1atum (Evans) Steph Fossombronia foveo1ata Lindb
(35)41 46 90 140 1 t 38 (39) t 46
Frullania bolanderi Aust
13 46 62 88 90 139 140
Frullanla ari ttoniae Evans
39 46 69 (71) 87 88 89 90
Frullania eboracensis Gott
1 9 28 4 39 46 (57) (59) 69 70 (71) 87 88 89 90 92 93 97 148
Frullania inflata Gott
46 87 88 89 (139)
Frullania oakesiana Aust Frullania plana Sullo
(35) 42 46 87 (139) (140) 92
Frullania ri paria Hampe ~ Lehm
46 69 71 87 89
Frullania ~elwynianna Pears
46 89
Geocalyx SEaveolens (Schrad) Nees
394688 8990 (119) 140
Frullania tamarisci (L) Dum subsp sPagrayana
(Mont) Hatt 28 34 l~6 t (57) 88 89 90
38
Mar pan thus seutatUB ( Web amp Mohr) Spruce Jamesoniella autumnalis (DC) Steph
(35) 39 46 90 1 7 28 39 46 56 (57) 87 8889 90 (119) 137 141
Jungermannia confertissima Nees
137 141
Jungermannia crenuliform1s Aust
79 137 141
Jungermannia ~il1ima Sm
46 90 141 Jungermannia exsert1folia subsp corifo~ (Dum~
Vana (35) 46 79 141
39
Jungermannia hyalina Lyell Jungerrr6nnia 1eiantha Grolle
40 46 79 136 141 39 46 79 88 90 1)7 141
Jungermannia IXlaxis Lindb Jungermannia pumila With
46 67 79 (139) (140) 141 )9 46 90 1)6 141
Jungermannia sphaerocarpa Hook Kurzia setacea (Web) Grolle
4679136140IIH 136
Lejeunea cavifo1ia (Ehrh) Lindb Lepidozia reptans (1) Dum
1 7 27 39 46 88 89 90 137 14010 I ( 27) 46 I 69 (71) 88
lophocolea heterophylla (Schrad) Dum Lophocolea minor Nees
1 )4 39 46 (57) 87 88 89 27 39 46 88 89 90 (119) 90 (93)94 119 136
Lophozia al~stris (Schleich ~ Web) Evans Lophozia ascendens (Warnst) Schust
46 90 137 141 136 140 141
41
loIilozia badensls (Gatt ~ Rabenh)
46 141
lophozia blcrenata (Schmid sectA Hoffm)
Dum 46 141
lophozia collaris (Nees) Dum
39 46 141
lophozla excisa (Dicks) Dum
27 4679 141
lophozia gillmani (Aust) Schust lophozia gIlttulata (Undo tl H Arnell)
36 39 40 46 136 (140) 141 Evans 39 46 136 141
42
141
Lophozia heterocolEDs (Thed) MA Howe IoPhozia ipcisa (Schard) Dum
27 39 46 89 141 8 27 31 46 79 90 140 141
LoQhozia longidens Macoun Lophozia ventricosa
1 46 79 90 141
Lophozia ventricosa var silvicola
(Buch) Jones 141 LoEhozla ventricosa vax longif1ora (Nees)
Macoun 39 46 79 141
43
~~ fragrans (Balbis) Frye et Clark Mannia pi 10sa (Hornem) Frye et Clark
46 (139) (140) 9 46 79 116 139
Mannia triandra (Scop) Gro 11e
46 (140)
Marchantia pound91ymorpha L
1 13 28 3946 (49) 78 87 88 89 90 (Ill) 121 122 147
Mo~rckia hibernica (HookJ Gott
46 89
Metzgeria conjugata Lindb
44
46
Mylia anomala (Hook) S Gray Notothylas orbicularis (Schwein) Sullo
39 46 141 (35)45 46
Nowellia curvifolia (Dicks) Mitt
17 27 3846 88 89 90 141
Odontoschisma denudatuJI1 (Nees ~ Mart)
Dum 46 81 137 141
Pallavicinia lyellii (Hook~ Carruth
90
Odontoschisma macounii (Aust) Under
45 46 (66) (139) 140 141
45
=
Pellia epiphllla (L) CordaPellia endiviifolia (Dicks~ Dum
27 39 46 90 13724 39 46
Pellia neesiana Limp bull Phaeoceros
39 46 89 27 39 45 46 97
Porella pinnata L
13 27 I 46 90 Plagiochila asplenioides (L) Dum
27 I 39 1+6 88 89 I 90 137
46
~e11a platYEhylla (L) Pfeiff Porella platyphylloi~ (Schwein) Lindb
17 27 39 46 69 (71)79 87 88 89 9 34 46 (57) 59 87 88 89 92 90 92 93 77
Preissia quadrata (Seop) Nees
1 36 3946 (71) 88 89 90 121 123
137
Ptllidium ciliare (L) Hampe
1 27 39 46 132 141
Dum
(22) 28 3946 (57) 87 88 89 90 92
Ptilidium ~lcherrium (G Web) Hampe
(9) 13 394656 (57)70 87 88 89 90 93 (119) 136 141
47
Radula obconica Sullo Reboulia hemlsphaerlca (L) Raddi
39 46 88 89 (43) 46 (59) 70 90 t Cf 124 125 (139)
Riccardia ~atifrons Lindb
1 27 39 46 89
Riccardia multifi~ (L) S Gray
39 46 88 145
Riccia beyrichiana Hampe ~ Lehm
46 Riccardia palmata (Hedw) Carruth
39 46 90
48
Riccia bifurca Hoffm Riccia cavernosa Hoffm
48 13739 46 79 137
Riccia fluitans L
242731454679
Riccla frostli Aust
48 79
Riccia sullicantii Aust
79 (137) Riccia sorocarpa Bisch 46 (137)
49
Ricciocarpus natans (1) Corda Sca~ia apiculata Spruce
13 27 46 79 89 110 39 46 (139) 141
ScaJltll1ia carinthica Jack ~ Lindb
137 t 141
Scapania ~ (Mart) Durn
39467990137141
Scapania digenil Schiffn
90
ScaEsectflia cuspiduligera (Nees) K Mull
46 (139) (140) (141)
50
Scapania glaucoeerha1a (Tayl) Aust Seapania gyrrmostomophilia Kaal
39 46 (139) 141 41 44 46 64 (139) (140) 141
ScaP3poundlia irrigua Gott ~ almiddot
7 39 46 79 137 (140L 141
Seapania nemorosa (L) Dum
1 39 46 89 90 137
Scapmi~ ~~ta Smiddotlch
137 141 51
~caPnia poundludicola LDeske amp K Mull Scapania saxicola Schust
46 (66) 141 136 141
Sc~~ scandica (H Arnell ~ Buch) Macv
139 140
Scapania subalpina (Nees) Durn
39 40 46 79 137 141
Scapania undulata (L) Dum
1 42 46 79 141
Scapania umbrosa (Schrad) Dum
137 140 141
52
Trichocolea tomentella (Ehrh) Dum Triomarla exsecta (Schmid) Schiffn
27 (35) 39 46 47 88 137 141 8 28 39 46 90 140 141
Tritomaria exsectiformis (Breidl) Loe~ke Tritomaria Sluinguedentata (Huds) Buch
39 -t6 89 90 136 140 141 7 39 46 141
Trltomaria scitula (Tayl) Joerg
46 136 (139) 141
53
Dipl0]hyllum apicu1atum (Evans) Steph Fossombronia foveo1ata Lindb
(35)41 46 90 140 1 t 38 (39) t 46
Frullania bolanderi Aust
13 46 62 88 90 139 140
Frullanla ari ttoniae Evans
39 46 69 (71) 87 88 89 90
Frullania eboracensis Gott
1 9 28 4 39 46 (57) (59) 69 70 (71) 87 88 89 90 92 93 97 148
Frullania inflata Gott
46 87 88 89 (139)
Frullania oakesiana Aust Frullania plana Sullo
(35) 42 46 87 (139) (140) 92
Frullania ri paria Hampe ~ Lehm
46 69 71 87 89
Frullania ~elwynianna Pears
46 89
Geocalyx SEaveolens (Schrad) Nees
394688 8990 (119) 140
Frullania tamarisci (L) Dum subsp sPagrayana
(Mont) Hatt 28 34 l~6 t (57) 88 89 90
38
Mar pan thus seutatUB ( Web amp Mohr) Spruce Jamesoniella autumnalis (DC) Steph
(35) 39 46 90 1 7 28 39 46 56 (57) 87 8889 90 (119) 137 141
Jungermannia confertissima Nees
137 141
Jungermannia crenuliform1s Aust
79 137 141
Jungermannia ~il1ima Sm
46 90 141 Jungermannia exsert1folia subsp corifo~ (Dum~
Vana (35) 46 79 141
39
Jungermannia hyalina Lyell Jungerrr6nnia 1eiantha Grolle
40 46 79 136 141 39 46 79 88 90 1)7 141
Jungermannia IXlaxis Lindb Jungermannia pumila With
46 67 79 (139) (140) 141 )9 46 90 1)6 141
Jungermannia sphaerocarpa Hook Kurzia setacea (Web) Grolle
4679136140IIH 136
Lejeunea cavifo1ia (Ehrh) Lindb Lepidozia reptans (1) Dum
1 7 27 39 46 88 89 90 137 14010 I ( 27) 46 I 69 (71) 88
lophocolea heterophylla (Schrad) Dum Lophocolea minor Nees
1 )4 39 46 (57) 87 88 89 27 39 46 88 89 90 (119) 90 (93)94 119 136
Lophozia al~stris (Schleich ~ Web) Evans Lophozia ascendens (Warnst) Schust
46 90 137 141 136 140 141
41
loIilozia badensls (Gatt ~ Rabenh)
46 141
lophozia blcrenata (Schmid sectA Hoffm)
Dum 46 141
lophozia collaris (Nees) Dum
39 46 141
lophozla excisa (Dicks) Dum
27 4679 141
lophozia gillmani (Aust) Schust lophozia gIlttulata (Undo tl H Arnell)
36 39 40 46 136 (140) 141 Evans 39 46 136 141
42
141
Lophozia heterocolEDs (Thed) MA Howe IoPhozia ipcisa (Schard) Dum
27 39 46 89 141 8 27 31 46 79 90 140 141
LoQhozia longidens Macoun Lophozia ventricosa
1 46 79 90 141
Lophozia ventricosa var silvicola
(Buch) Jones 141 LoEhozla ventricosa vax longif1ora (Nees)
Macoun 39 46 79 141
43
~~ fragrans (Balbis) Frye et Clark Mannia pi 10sa (Hornem) Frye et Clark
46 (139) (140) 9 46 79 116 139
Mannia triandra (Scop) Gro 11e
46 (140)
Marchantia pound91ymorpha L
1 13 28 3946 (49) 78 87 88 89 90 (Ill) 121 122 147
Mo~rckia hibernica (HookJ Gott
46 89
Metzgeria conjugata Lindb
44
46
Mylia anomala (Hook) S Gray Notothylas orbicularis (Schwein) Sullo
39 46 141 (35)45 46
Nowellia curvifolia (Dicks) Mitt
17 27 3846 88 89 90 141
Odontoschisma denudatuJI1 (Nees ~ Mart)
Dum 46 81 137 141
Pallavicinia lyellii (Hook~ Carruth
90
Odontoschisma macounii (Aust) Under
45 46 (66) (139) 140 141
45
=
Pellia epiphllla (L) CordaPellia endiviifolia (Dicks~ Dum
27 39 46 90 13724 39 46
Pellia neesiana Limp bull Phaeoceros
39 46 89 27 39 45 46 97
Porella pinnata L
13 27 I 46 90 Plagiochila asplenioides (L) Dum
27 I 39 1+6 88 89 I 90 137
46
~e11a platYEhylla (L) Pfeiff Porella platyphylloi~ (Schwein) Lindb
17 27 39 46 69 (71)79 87 88 89 9 34 46 (57) 59 87 88 89 92 90 92 93 77
Preissia quadrata (Seop) Nees
1 36 3946 (71) 88 89 90 121 123
137
Ptllidium ciliare (L) Hampe
1 27 39 46 132 141
Dum
(22) 28 3946 (57) 87 88 89 90 92
Ptilidium ~lcherrium (G Web) Hampe
(9) 13 394656 (57)70 87 88 89 90 93 (119) 136 141
47
Radula obconica Sullo Reboulia hemlsphaerlca (L) Raddi
39 46 88 89 (43) 46 (59) 70 90 t Cf 124 125 (139)
Riccardia ~atifrons Lindb
1 27 39 46 89
Riccardia multifi~ (L) S Gray
39 46 88 145
Riccia beyrichiana Hampe ~ Lehm
46 Riccardia palmata (Hedw) Carruth
39 46 90
48
Riccia bifurca Hoffm Riccia cavernosa Hoffm
48 13739 46 79 137
Riccia fluitans L
242731454679
Riccla frostli Aust
48 79
Riccia sullicantii Aust
79 (137) Riccia sorocarpa Bisch 46 (137)
49
Ricciocarpus natans (1) Corda Sca~ia apiculata Spruce
13 27 46 79 89 110 39 46 (139) 141
ScaJltll1ia carinthica Jack ~ Lindb
137 t 141
Scapania ~ (Mart) Durn
39467990137141
Scapania digenil Schiffn
90
ScaEsectflia cuspiduligera (Nees) K Mull
46 (139) (140) (141)
50
Scapania glaucoeerha1a (Tayl) Aust Seapania gyrrmostomophilia Kaal
39 46 (139) 141 41 44 46 64 (139) (140) 141
ScaP3poundlia irrigua Gott ~ almiddot
7 39 46 79 137 (140L 141
Seapania nemorosa (L) Dum
1 39 46 89 90 137
Scapmi~ ~~ta Smiddotlch
137 141 51
~caPnia poundludicola LDeske amp K Mull Scapania saxicola Schust
46 (66) 141 136 141
Sc~~ scandica (H Arnell ~ Buch) Macv
139 140
Scapania subalpina (Nees) Durn
39 40 46 79 137 141
Scapania undulata (L) Dum
1 42 46 79 141
Scapania umbrosa (Schrad) Dum
137 140 141
52
Trichocolea tomentella (Ehrh) Dum Triomarla exsecta (Schmid) Schiffn
27 (35) 39 46 47 88 137 141 8 28 39 46 90 140 141
Tritomaria exsectiformis (Breidl) Loe~ke Tritomaria Sluinguedentata (Huds) Buch
39 -t6 89 90 136 140 141 7 39 46 141
Trltomaria scitula (Tayl) Joerg
46 136 (139) 141
53
Frullania oakesiana Aust Frullania plana Sullo
(35) 42 46 87 (139) (140) 92
Frullania ri paria Hampe ~ Lehm
46 69 71 87 89
Frullania ~elwynianna Pears
46 89
Geocalyx SEaveolens (Schrad) Nees
394688 8990 (119) 140
Frullania tamarisci (L) Dum subsp sPagrayana
(Mont) Hatt 28 34 l~6 t (57) 88 89 90
38
Mar pan thus seutatUB ( Web amp Mohr) Spruce Jamesoniella autumnalis (DC) Steph
(35) 39 46 90 1 7 28 39 46 56 (57) 87 8889 90 (119) 137 141
Jungermannia confertissima Nees
137 141
Jungermannia crenuliform1s Aust
79 137 141
Jungermannia ~il1ima Sm
46 90 141 Jungermannia exsert1folia subsp corifo~ (Dum~
Vana (35) 46 79 141
39
Jungermannia hyalina Lyell Jungerrr6nnia 1eiantha Grolle
40 46 79 136 141 39 46 79 88 90 1)7 141
Jungermannia IXlaxis Lindb Jungermannia pumila With
46 67 79 (139) (140) 141 )9 46 90 1)6 141
Jungermannia sphaerocarpa Hook Kurzia setacea (Web) Grolle
4679136140IIH 136
Lejeunea cavifo1ia (Ehrh) Lindb Lepidozia reptans (1) Dum
1 7 27 39 46 88 89 90 137 14010 I ( 27) 46 I 69 (71) 88
lophocolea heterophylla (Schrad) Dum Lophocolea minor Nees
1 )4 39 46 (57) 87 88 89 27 39 46 88 89 90 (119) 90 (93)94 119 136
Lophozia al~stris (Schleich ~ Web) Evans Lophozia ascendens (Warnst) Schust
46 90 137 141 136 140 141
41
loIilozia badensls (Gatt ~ Rabenh)
46 141
lophozia blcrenata (Schmid sectA Hoffm)
Dum 46 141
lophozia collaris (Nees) Dum
39 46 141
lophozla excisa (Dicks) Dum
27 4679 141
lophozia gillmani (Aust) Schust lophozia gIlttulata (Undo tl H Arnell)
36 39 40 46 136 (140) 141 Evans 39 46 136 141
42
141
Lophozia heterocolEDs (Thed) MA Howe IoPhozia ipcisa (Schard) Dum
27 39 46 89 141 8 27 31 46 79 90 140 141
LoQhozia longidens Macoun Lophozia ventricosa
1 46 79 90 141
Lophozia ventricosa var silvicola
(Buch) Jones 141 LoEhozla ventricosa vax longif1ora (Nees)
Macoun 39 46 79 141
43
~~ fragrans (Balbis) Frye et Clark Mannia pi 10sa (Hornem) Frye et Clark
46 (139) (140) 9 46 79 116 139
Mannia triandra (Scop) Gro 11e
46 (140)
Marchantia pound91ymorpha L
1 13 28 3946 (49) 78 87 88 89 90 (Ill) 121 122 147
Mo~rckia hibernica (HookJ Gott
46 89
Metzgeria conjugata Lindb
44
46
Mylia anomala (Hook) S Gray Notothylas orbicularis (Schwein) Sullo
39 46 141 (35)45 46
Nowellia curvifolia (Dicks) Mitt
17 27 3846 88 89 90 141
Odontoschisma denudatuJI1 (Nees ~ Mart)
Dum 46 81 137 141
Pallavicinia lyellii (Hook~ Carruth
90
Odontoschisma macounii (Aust) Under
45 46 (66) (139) 140 141
45
=
Pellia epiphllla (L) CordaPellia endiviifolia (Dicks~ Dum
27 39 46 90 13724 39 46
Pellia neesiana Limp bull Phaeoceros
39 46 89 27 39 45 46 97
Porella pinnata L
13 27 I 46 90 Plagiochila asplenioides (L) Dum
27 I 39 1+6 88 89 I 90 137
46
~e11a platYEhylla (L) Pfeiff Porella platyphylloi~ (Schwein) Lindb
17 27 39 46 69 (71)79 87 88 89 9 34 46 (57) 59 87 88 89 92 90 92 93 77
Preissia quadrata (Seop) Nees
1 36 3946 (71) 88 89 90 121 123
137
Ptllidium ciliare (L) Hampe
1 27 39 46 132 141
Dum
(22) 28 3946 (57) 87 88 89 90 92
Ptilidium ~lcherrium (G Web) Hampe
(9) 13 394656 (57)70 87 88 89 90 93 (119) 136 141
47
Radula obconica Sullo Reboulia hemlsphaerlca (L) Raddi
39 46 88 89 (43) 46 (59) 70 90 t Cf 124 125 (139)
Riccardia ~atifrons Lindb
1 27 39 46 89
Riccardia multifi~ (L) S Gray
39 46 88 145
Riccia beyrichiana Hampe ~ Lehm
46 Riccardia palmata (Hedw) Carruth
39 46 90
48
Riccia bifurca Hoffm Riccia cavernosa Hoffm
48 13739 46 79 137
Riccia fluitans L
242731454679
Riccla frostli Aust
48 79
Riccia sullicantii Aust
79 (137) Riccia sorocarpa Bisch 46 (137)
49
Ricciocarpus natans (1) Corda Sca~ia apiculata Spruce
13 27 46 79 89 110 39 46 (139) 141
ScaJltll1ia carinthica Jack ~ Lindb
137 t 141
Scapania ~ (Mart) Durn
39467990137141
Scapania digenil Schiffn
90
ScaEsectflia cuspiduligera (Nees) K Mull
46 (139) (140) (141)
50
Scapania glaucoeerha1a (Tayl) Aust Seapania gyrrmostomophilia Kaal
39 46 (139) 141 41 44 46 64 (139) (140) 141
ScaP3poundlia irrigua Gott ~ almiddot
7 39 46 79 137 (140L 141
Seapania nemorosa (L) Dum
1 39 46 89 90 137
Scapmi~ ~~ta Smiddotlch
137 141 51
~caPnia poundludicola LDeske amp K Mull Scapania saxicola Schust
46 (66) 141 136 141
Sc~~ scandica (H Arnell ~ Buch) Macv
139 140
Scapania subalpina (Nees) Durn
39 40 46 79 137 141
Scapania undulata (L) Dum
1 42 46 79 141
Scapania umbrosa (Schrad) Dum
137 140 141
52
Trichocolea tomentella (Ehrh) Dum Triomarla exsecta (Schmid) Schiffn
27 (35) 39 46 47 88 137 141 8 28 39 46 90 140 141
Tritomaria exsectiformis (Breidl) Loe~ke Tritomaria Sluinguedentata (Huds) Buch
39 -t6 89 90 136 140 141 7 39 46 141
Trltomaria scitula (Tayl) Joerg
46 136 (139) 141
53
Mar pan thus seutatUB ( Web amp Mohr) Spruce Jamesoniella autumnalis (DC) Steph
(35) 39 46 90 1 7 28 39 46 56 (57) 87 8889 90 (119) 137 141
Jungermannia confertissima Nees
137 141
Jungermannia crenuliform1s Aust
79 137 141
Jungermannia ~il1ima Sm
46 90 141 Jungermannia exsert1folia subsp corifo~ (Dum~
Vana (35) 46 79 141
39
Jungermannia hyalina Lyell Jungerrr6nnia 1eiantha Grolle
40 46 79 136 141 39 46 79 88 90 1)7 141
Jungermannia IXlaxis Lindb Jungermannia pumila With
46 67 79 (139) (140) 141 )9 46 90 1)6 141
Jungermannia sphaerocarpa Hook Kurzia setacea (Web) Grolle
4679136140IIH 136
Lejeunea cavifo1ia (Ehrh) Lindb Lepidozia reptans (1) Dum
1 7 27 39 46 88 89 90 137 14010 I ( 27) 46 I 69 (71) 88
lophocolea heterophylla (Schrad) Dum Lophocolea minor Nees
1 )4 39 46 (57) 87 88 89 27 39 46 88 89 90 (119) 90 (93)94 119 136
Lophozia al~stris (Schleich ~ Web) Evans Lophozia ascendens (Warnst) Schust
46 90 137 141 136 140 141
41
loIilozia badensls (Gatt ~ Rabenh)
46 141
lophozia blcrenata (Schmid sectA Hoffm)
Dum 46 141
lophozia collaris (Nees) Dum
39 46 141
lophozla excisa (Dicks) Dum
27 4679 141
lophozia gillmani (Aust) Schust lophozia gIlttulata (Undo tl H Arnell)
36 39 40 46 136 (140) 141 Evans 39 46 136 141
42
141
Lophozia heterocolEDs (Thed) MA Howe IoPhozia ipcisa (Schard) Dum
27 39 46 89 141 8 27 31 46 79 90 140 141
LoQhozia longidens Macoun Lophozia ventricosa
1 46 79 90 141
Lophozia ventricosa var silvicola
(Buch) Jones 141 LoEhozla ventricosa vax longif1ora (Nees)
Macoun 39 46 79 141
43
~~ fragrans (Balbis) Frye et Clark Mannia pi 10sa (Hornem) Frye et Clark
46 (139) (140) 9 46 79 116 139
Mannia triandra (Scop) Gro 11e
46 (140)
Marchantia pound91ymorpha L
1 13 28 3946 (49) 78 87 88 89 90 (Ill) 121 122 147
Mo~rckia hibernica (HookJ Gott
46 89
Metzgeria conjugata Lindb
44
46
Mylia anomala (Hook) S Gray Notothylas orbicularis (Schwein) Sullo
39 46 141 (35)45 46
Nowellia curvifolia (Dicks) Mitt
17 27 3846 88 89 90 141
Odontoschisma denudatuJI1 (Nees ~ Mart)
Dum 46 81 137 141
Pallavicinia lyellii (Hook~ Carruth
90
Odontoschisma macounii (Aust) Under
45 46 (66) (139) 140 141
45
=
Pellia epiphllla (L) CordaPellia endiviifolia (Dicks~ Dum
27 39 46 90 13724 39 46
Pellia neesiana Limp bull Phaeoceros
39 46 89 27 39 45 46 97
Porella pinnata L
13 27 I 46 90 Plagiochila asplenioides (L) Dum
27 I 39 1+6 88 89 I 90 137
46
~e11a platYEhylla (L) Pfeiff Porella platyphylloi~ (Schwein) Lindb
17 27 39 46 69 (71)79 87 88 89 9 34 46 (57) 59 87 88 89 92 90 92 93 77
Preissia quadrata (Seop) Nees
1 36 3946 (71) 88 89 90 121 123
137
Ptllidium ciliare (L) Hampe
1 27 39 46 132 141
Dum
(22) 28 3946 (57) 87 88 89 90 92
Ptilidium ~lcherrium (G Web) Hampe
(9) 13 394656 (57)70 87 88 89 90 93 (119) 136 141
47
Radula obconica Sullo Reboulia hemlsphaerlca (L) Raddi
39 46 88 89 (43) 46 (59) 70 90 t Cf 124 125 (139)
Riccardia ~atifrons Lindb
1 27 39 46 89
Riccardia multifi~ (L) S Gray
39 46 88 145
Riccia beyrichiana Hampe ~ Lehm
46 Riccardia palmata (Hedw) Carruth
39 46 90
48
Riccia bifurca Hoffm Riccia cavernosa Hoffm
48 13739 46 79 137
Riccia fluitans L
242731454679
Riccla frostli Aust
48 79
Riccia sullicantii Aust
79 (137) Riccia sorocarpa Bisch 46 (137)
49
Ricciocarpus natans (1) Corda Sca~ia apiculata Spruce
13 27 46 79 89 110 39 46 (139) 141
ScaJltll1ia carinthica Jack ~ Lindb
137 t 141
Scapania ~ (Mart) Durn
39467990137141
Scapania digenil Schiffn
90
ScaEsectflia cuspiduligera (Nees) K Mull
46 (139) (140) (141)
50
Scapania glaucoeerha1a (Tayl) Aust Seapania gyrrmostomophilia Kaal
39 46 (139) 141 41 44 46 64 (139) (140) 141
ScaP3poundlia irrigua Gott ~ almiddot
7 39 46 79 137 (140L 141
Seapania nemorosa (L) Dum
1 39 46 89 90 137
Scapmi~ ~~ta Smiddotlch
137 141 51
~caPnia poundludicola LDeske amp K Mull Scapania saxicola Schust
46 (66) 141 136 141
Sc~~ scandica (H Arnell ~ Buch) Macv
139 140
Scapania subalpina (Nees) Durn
39 40 46 79 137 141
Scapania undulata (L) Dum
1 42 46 79 141
Scapania umbrosa (Schrad) Dum
137 140 141
52
Trichocolea tomentella (Ehrh) Dum Triomarla exsecta (Schmid) Schiffn
27 (35) 39 46 47 88 137 141 8 28 39 46 90 140 141
Tritomaria exsectiformis (Breidl) Loe~ke Tritomaria Sluinguedentata (Huds) Buch
39 -t6 89 90 136 140 141 7 39 46 141
Trltomaria scitula (Tayl) Joerg
46 136 (139) 141
53
Jungermannia hyalina Lyell Jungerrr6nnia 1eiantha Grolle
40 46 79 136 141 39 46 79 88 90 1)7 141
Jungermannia IXlaxis Lindb Jungermannia pumila With
46 67 79 (139) (140) 141 )9 46 90 1)6 141
Jungermannia sphaerocarpa Hook Kurzia setacea (Web) Grolle
4679136140IIH 136
Lejeunea cavifo1ia (Ehrh) Lindb Lepidozia reptans (1) Dum
1 7 27 39 46 88 89 90 137 14010 I ( 27) 46 I 69 (71) 88
lophocolea heterophylla (Schrad) Dum Lophocolea minor Nees
1 )4 39 46 (57) 87 88 89 27 39 46 88 89 90 (119) 90 (93)94 119 136
Lophozia al~stris (Schleich ~ Web) Evans Lophozia ascendens (Warnst) Schust
46 90 137 141 136 140 141
41
loIilozia badensls (Gatt ~ Rabenh)
46 141
lophozia blcrenata (Schmid sectA Hoffm)
Dum 46 141
lophozia collaris (Nees) Dum
39 46 141
lophozla excisa (Dicks) Dum
27 4679 141
lophozia gillmani (Aust) Schust lophozia gIlttulata (Undo tl H Arnell)
36 39 40 46 136 (140) 141 Evans 39 46 136 141
42
141
Lophozia heterocolEDs (Thed) MA Howe IoPhozia ipcisa (Schard) Dum
27 39 46 89 141 8 27 31 46 79 90 140 141
LoQhozia longidens Macoun Lophozia ventricosa
1 46 79 90 141
Lophozia ventricosa var silvicola
(Buch) Jones 141 LoEhozla ventricosa vax longif1ora (Nees)
Macoun 39 46 79 141
43
~~ fragrans (Balbis) Frye et Clark Mannia pi 10sa (Hornem) Frye et Clark
46 (139) (140) 9 46 79 116 139
Mannia triandra (Scop) Gro 11e
46 (140)
Marchantia pound91ymorpha L
1 13 28 3946 (49) 78 87 88 89 90 (Ill) 121 122 147
Mo~rckia hibernica (HookJ Gott
46 89
Metzgeria conjugata Lindb
44
46
Mylia anomala (Hook) S Gray Notothylas orbicularis (Schwein) Sullo
39 46 141 (35)45 46
Nowellia curvifolia (Dicks) Mitt
17 27 3846 88 89 90 141
Odontoschisma denudatuJI1 (Nees ~ Mart)
Dum 46 81 137 141
Pallavicinia lyellii (Hook~ Carruth
90
Odontoschisma macounii (Aust) Under
45 46 (66) (139) 140 141
45
=
Pellia epiphllla (L) CordaPellia endiviifolia (Dicks~ Dum
27 39 46 90 13724 39 46
Pellia neesiana Limp bull Phaeoceros
39 46 89 27 39 45 46 97
Porella pinnata L
13 27 I 46 90 Plagiochila asplenioides (L) Dum
27 I 39 1+6 88 89 I 90 137
46
~e11a platYEhylla (L) Pfeiff Porella platyphylloi~ (Schwein) Lindb
17 27 39 46 69 (71)79 87 88 89 9 34 46 (57) 59 87 88 89 92 90 92 93 77
Preissia quadrata (Seop) Nees
1 36 3946 (71) 88 89 90 121 123
137
Ptllidium ciliare (L) Hampe
1 27 39 46 132 141
Dum
(22) 28 3946 (57) 87 88 89 90 92
Ptilidium ~lcherrium (G Web) Hampe
(9) 13 394656 (57)70 87 88 89 90 93 (119) 136 141
47
Radula obconica Sullo Reboulia hemlsphaerlca (L) Raddi
39 46 88 89 (43) 46 (59) 70 90 t Cf 124 125 (139)
Riccardia ~atifrons Lindb
1 27 39 46 89
Riccardia multifi~ (L) S Gray
39 46 88 145
Riccia beyrichiana Hampe ~ Lehm
46 Riccardia palmata (Hedw) Carruth
39 46 90
48
Riccia bifurca Hoffm Riccia cavernosa Hoffm
48 13739 46 79 137
Riccia fluitans L
242731454679
Riccla frostli Aust
48 79
Riccia sullicantii Aust
79 (137) Riccia sorocarpa Bisch 46 (137)
49
Ricciocarpus natans (1) Corda Sca~ia apiculata Spruce
13 27 46 79 89 110 39 46 (139) 141
ScaJltll1ia carinthica Jack ~ Lindb
137 t 141
Scapania ~ (Mart) Durn
39467990137141
Scapania digenil Schiffn
90
ScaEsectflia cuspiduligera (Nees) K Mull
46 (139) (140) (141)
50
Scapania glaucoeerha1a (Tayl) Aust Seapania gyrrmostomophilia Kaal
39 46 (139) 141 41 44 46 64 (139) (140) 141
ScaP3poundlia irrigua Gott ~ almiddot
7 39 46 79 137 (140L 141
Seapania nemorosa (L) Dum
1 39 46 89 90 137
Scapmi~ ~~ta Smiddotlch
137 141 51
~caPnia poundludicola LDeske amp K Mull Scapania saxicola Schust
46 (66) 141 136 141
Sc~~ scandica (H Arnell ~ Buch) Macv
139 140
Scapania subalpina (Nees) Durn
39 40 46 79 137 141
Scapania undulata (L) Dum
1 42 46 79 141
Scapania umbrosa (Schrad) Dum
137 140 141
52
Trichocolea tomentella (Ehrh) Dum Triomarla exsecta (Schmid) Schiffn
27 (35) 39 46 47 88 137 141 8 28 39 46 90 140 141
Tritomaria exsectiformis (Breidl) Loe~ke Tritomaria Sluinguedentata (Huds) Buch
39 -t6 89 90 136 140 141 7 39 46 141
Trltomaria scitula (Tayl) Joerg
46 136 (139) 141
53
Lejeunea cavifo1ia (Ehrh) Lindb Lepidozia reptans (1) Dum
1 7 27 39 46 88 89 90 137 14010 I ( 27) 46 I 69 (71) 88
lophocolea heterophylla (Schrad) Dum Lophocolea minor Nees
1 )4 39 46 (57) 87 88 89 27 39 46 88 89 90 (119) 90 (93)94 119 136
Lophozia al~stris (Schleich ~ Web) Evans Lophozia ascendens (Warnst) Schust
46 90 137 141 136 140 141
41
loIilozia badensls (Gatt ~ Rabenh)
46 141
lophozia blcrenata (Schmid sectA Hoffm)
Dum 46 141
lophozia collaris (Nees) Dum
39 46 141
lophozla excisa (Dicks) Dum
27 4679 141
lophozia gillmani (Aust) Schust lophozia gIlttulata (Undo tl H Arnell)
36 39 40 46 136 (140) 141 Evans 39 46 136 141
42
141
Lophozia heterocolEDs (Thed) MA Howe IoPhozia ipcisa (Schard) Dum
27 39 46 89 141 8 27 31 46 79 90 140 141
LoQhozia longidens Macoun Lophozia ventricosa
1 46 79 90 141
Lophozia ventricosa var silvicola
(Buch) Jones 141 LoEhozla ventricosa vax longif1ora (Nees)
Macoun 39 46 79 141
43
~~ fragrans (Balbis) Frye et Clark Mannia pi 10sa (Hornem) Frye et Clark
46 (139) (140) 9 46 79 116 139
Mannia triandra (Scop) Gro 11e
46 (140)
Marchantia pound91ymorpha L
1 13 28 3946 (49) 78 87 88 89 90 (Ill) 121 122 147
Mo~rckia hibernica (HookJ Gott
46 89
Metzgeria conjugata Lindb
44
46
Mylia anomala (Hook) S Gray Notothylas orbicularis (Schwein) Sullo
39 46 141 (35)45 46
Nowellia curvifolia (Dicks) Mitt
17 27 3846 88 89 90 141
Odontoschisma denudatuJI1 (Nees ~ Mart)
Dum 46 81 137 141
Pallavicinia lyellii (Hook~ Carruth
90
Odontoschisma macounii (Aust) Under
45 46 (66) (139) 140 141
45
=
Pellia epiphllla (L) CordaPellia endiviifolia (Dicks~ Dum
27 39 46 90 13724 39 46
Pellia neesiana Limp bull Phaeoceros
39 46 89 27 39 45 46 97
Porella pinnata L
13 27 I 46 90 Plagiochila asplenioides (L) Dum
27 I 39 1+6 88 89 I 90 137
46
~e11a platYEhylla (L) Pfeiff Porella platyphylloi~ (Schwein) Lindb
17 27 39 46 69 (71)79 87 88 89 9 34 46 (57) 59 87 88 89 92 90 92 93 77
Preissia quadrata (Seop) Nees
1 36 3946 (71) 88 89 90 121 123
137
Ptllidium ciliare (L) Hampe
1 27 39 46 132 141
Dum
(22) 28 3946 (57) 87 88 89 90 92
Ptilidium ~lcherrium (G Web) Hampe
(9) 13 394656 (57)70 87 88 89 90 93 (119) 136 141
47
Radula obconica Sullo Reboulia hemlsphaerlca (L) Raddi
39 46 88 89 (43) 46 (59) 70 90 t Cf 124 125 (139)
Riccardia ~atifrons Lindb
1 27 39 46 89
Riccardia multifi~ (L) S Gray
39 46 88 145
Riccia beyrichiana Hampe ~ Lehm
46 Riccardia palmata (Hedw) Carruth
39 46 90
48
Riccia bifurca Hoffm Riccia cavernosa Hoffm
48 13739 46 79 137
Riccia fluitans L
242731454679
Riccla frostli Aust
48 79
Riccia sullicantii Aust
79 (137) Riccia sorocarpa Bisch 46 (137)
49
Ricciocarpus natans (1) Corda Sca~ia apiculata Spruce
13 27 46 79 89 110 39 46 (139) 141
ScaJltll1ia carinthica Jack ~ Lindb
137 t 141
Scapania ~ (Mart) Durn
39467990137141
Scapania digenil Schiffn
90
ScaEsectflia cuspiduligera (Nees) K Mull
46 (139) (140) (141)
50
Scapania glaucoeerha1a (Tayl) Aust Seapania gyrrmostomophilia Kaal
39 46 (139) 141 41 44 46 64 (139) (140) 141
ScaP3poundlia irrigua Gott ~ almiddot
7 39 46 79 137 (140L 141
Seapania nemorosa (L) Dum
1 39 46 89 90 137
Scapmi~ ~~ta Smiddotlch
137 141 51
~caPnia poundludicola LDeske amp K Mull Scapania saxicola Schust
46 (66) 141 136 141
Sc~~ scandica (H Arnell ~ Buch) Macv
139 140
Scapania subalpina (Nees) Durn
39 40 46 79 137 141
Scapania undulata (L) Dum
1 42 46 79 141
Scapania umbrosa (Schrad) Dum
137 140 141
52
Trichocolea tomentella (Ehrh) Dum Triomarla exsecta (Schmid) Schiffn
27 (35) 39 46 47 88 137 141 8 28 39 46 90 140 141
Tritomaria exsectiformis (Breidl) Loe~ke Tritomaria Sluinguedentata (Huds) Buch
39 -t6 89 90 136 140 141 7 39 46 141
Trltomaria scitula (Tayl) Joerg
46 136 (139) 141
53
loIilozia badensls (Gatt ~ Rabenh)
46 141
lophozia blcrenata (Schmid sectA Hoffm)
Dum 46 141
lophozia collaris (Nees) Dum
39 46 141
lophozla excisa (Dicks) Dum
27 4679 141
lophozia gillmani (Aust) Schust lophozia gIlttulata (Undo tl H Arnell)
36 39 40 46 136 (140) 141 Evans 39 46 136 141
42
141
Lophozia heterocolEDs (Thed) MA Howe IoPhozia ipcisa (Schard) Dum
27 39 46 89 141 8 27 31 46 79 90 140 141
LoQhozia longidens Macoun Lophozia ventricosa
1 46 79 90 141
Lophozia ventricosa var silvicola
(Buch) Jones 141 LoEhozla ventricosa vax longif1ora (Nees)
Macoun 39 46 79 141
43
~~ fragrans (Balbis) Frye et Clark Mannia pi 10sa (Hornem) Frye et Clark
46 (139) (140) 9 46 79 116 139
Mannia triandra (Scop) Gro 11e
46 (140)
Marchantia pound91ymorpha L
1 13 28 3946 (49) 78 87 88 89 90 (Ill) 121 122 147
Mo~rckia hibernica (HookJ Gott
46 89
Metzgeria conjugata Lindb
44
46
Mylia anomala (Hook) S Gray Notothylas orbicularis (Schwein) Sullo
39 46 141 (35)45 46
Nowellia curvifolia (Dicks) Mitt
17 27 3846 88 89 90 141
Odontoschisma denudatuJI1 (Nees ~ Mart)
Dum 46 81 137 141
Pallavicinia lyellii (Hook~ Carruth
90
Odontoschisma macounii (Aust) Under
45 46 (66) (139) 140 141
45
=
Pellia epiphllla (L) CordaPellia endiviifolia (Dicks~ Dum
27 39 46 90 13724 39 46
Pellia neesiana Limp bull Phaeoceros
39 46 89 27 39 45 46 97
Porella pinnata L
13 27 I 46 90 Plagiochila asplenioides (L) Dum
27 I 39 1+6 88 89 I 90 137
46
~e11a platYEhylla (L) Pfeiff Porella platyphylloi~ (Schwein) Lindb
17 27 39 46 69 (71)79 87 88 89 9 34 46 (57) 59 87 88 89 92 90 92 93 77
Preissia quadrata (Seop) Nees
1 36 3946 (71) 88 89 90 121 123
137
Ptllidium ciliare (L) Hampe
1 27 39 46 132 141
Dum
(22) 28 3946 (57) 87 88 89 90 92
Ptilidium ~lcherrium (G Web) Hampe
(9) 13 394656 (57)70 87 88 89 90 93 (119) 136 141
47
Radula obconica Sullo Reboulia hemlsphaerlca (L) Raddi
39 46 88 89 (43) 46 (59) 70 90 t Cf 124 125 (139)
Riccardia ~atifrons Lindb
1 27 39 46 89
Riccardia multifi~ (L) S Gray
39 46 88 145
Riccia beyrichiana Hampe ~ Lehm
46 Riccardia palmata (Hedw) Carruth
39 46 90
48
Riccia bifurca Hoffm Riccia cavernosa Hoffm
48 13739 46 79 137
Riccia fluitans L
242731454679
Riccla frostli Aust
48 79
Riccia sullicantii Aust
79 (137) Riccia sorocarpa Bisch 46 (137)
49
Ricciocarpus natans (1) Corda Sca~ia apiculata Spruce
13 27 46 79 89 110 39 46 (139) 141
ScaJltll1ia carinthica Jack ~ Lindb
137 t 141
Scapania ~ (Mart) Durn
39467990137141
Scapania digenil Schiffn
90
ScaEsectflia cuspiduligera (Nees) K Mull
46 (139) (140) (141)
50
Scapania glaucoeerha1a (Tayl) Aust Seapania gyrrmostomophilia Kaal
39 46 (139) 141 41 44 46 64 (139) (140) 141
ScaP3poundlia irrigua Gott ~ almiddot
7 39 46 79 137 (140L 141
Seapania nemorosa (L) Dum
1 39 46 89 90 137
Scapmi~ ~~ta Smiddotlch
137 141 51
~caPnia poundludicola LDeske amp K Mull Scapania saxicola Schust
46 (66) 141 136 141
Sc~~ scandica (H Arnell ~ Buch) Macv
139 140
Scapania subalpina (Nees) Durn
39 40 46 79 137 141
Scapania undulata (L) Dum
1 42 46 79 141
Scapania umbrosa (Schrad) Dum
137 140 141
52
Trichocolea tomentella (Ehrh) Dum Triomarla exsecta (Schmid) Schiffn
27 (35) 39 46 47 88 137 141 8 28 39 46 90 140 141
Tritomaria exsectiformis (Breidl) Loe~ke Tritomaria Sluinguedentata (Huds) Buch
39 -t6 89 90 136 140 141 7 39 46 141
Trltomaria scitula (Tayl) Joerg
46 136 (139) 141
53
141
Lophozia heterocolEDs (Thed) MA Howe IoPhozia ipcisa (Schard) Dum
27 39 46 89 141 8 27 31 46 79 90 140 141
LoQhozia longidens Macoun Lophozia ventricosa
1 46 79 90 141
Lophozia ventricosa var silvicola
(Buch) Jones 141 LoEhozla ventricosa vax longif1ora (Nees)
Macoun 39 46 79 141
43
~~ fragrans (Balbis) Frye et Clark Mannia pi 10sa (Hornem) Frye et Clark
46 (139) (140) 9 46 79 116 139
Mannia triandra (Scop) Gro 11e
46 (140)
Marchantia pound91ymorpha L
1 13 28 3946 (49) 78 87 88 89 90 (Ill) 121 122 147
Mo~rckia hibernica (HookJ Gott
46 89
Metzgeria conjugata Lindb
44
46
Mylia anomala (Hook) S Gray Notothylas orbicularis (Schwein) Sullo
39 46 141 (35)45 46
Nowellia curvifolia (Dicks) Mitt
17 27 3846 88 89 90 141
Odontoschisma denudatuJI1 (Nees ~ Mart)
Dum 46 81 137 141
Pallavicinia lyellii (Hook~ Carruth
90
Odontoschisma macounii (Aust) Under
45 46 (66) (139) 140 141
45
=
Pellia epiphllla (L) CordaPellia endiviifolia (Dicks~ Dum
27 39 46 90 13724 39 46
Pellia neesiana Limp bull Phaeoceros
39 46 89 27 39 45 46 97
Porella pinnata L
13 27 I 46 90 Plagiochila asplenioides (L) Dum
27 I 39 1+6 88 89 I 90 137
46
~e11a platYEhylla (L) Pfeiff Porella platyphylloi~ (Schwein) Lindb
17 27 39 46 69 (71)79 87 88 89 9 34 46 (57) 59 87 88 89 92 90 92 93 77
Preissia quadrata (Seop) Nees
1 36 3946 (71) 88 89 90 121 123
137
Ptllidium ciliare (L) Hampe
1 27 39 46 132 141
Dum
(22) 28 3946 (57) 87 88 89 90 92
Ptilidium ~lcherrium (G Web) Hampe
(9) 13 394656 (57)70 87 88 89 90 93 (119) 136 141
47
Radula obconica Sullo Reboulia hemlsphaerlca (L) Raddi
39 46 88 89 (43) 46 (59) 70 90 t Cf 124 125 (139)
Riccardia ~atifrons Lindb
1 27 39 46 89
Riccardia multifi~ (L) S Gray
39 46 88 145
Riccia beyrichiana Hampe ~ Lehm
46 Riccardia palmata (Hedw) Carruth
39 46 90
48
Riccia bifurca Hoffm Riccia cavernosa Hoffm
48 13739 46 79 137
Riccia fluitans L
242731454679
Riccla frostli Aust
48 79
Riccia sullicantii Aust
79 (137) Riccia sorocarpa Bisch 46 (137)
49
Ricciocarpus natans (1) Corda Sca~ia apiculata Spruce
13 27 46 79 89 110 39 46 (139) 141
ScaJltll1ia carinthica Jack ~ Lindb
137 t 141
Scapania ~ (Mart) Durn
39467990137141
Scapania digenil Schiffn
90
ScaEsectflia cuspiduligera (Nees) K Mull
46 (139) (140) (141)
50
Scapania glaucoeerha1a (Tayl) Aust Seapania gyrrmostomophilia Kaal
39 46 (139) 141 41 44 46 64 (139) (140) 141
ScaP3poundlia irrigua Gott ~ almiddot
7 39 46 79 137 (140L 141
Seapania nemorosa (L) Dum
1 39 46 89 90 137
Scapmi~ ~~ta Smiddotlch
137 141 51
~caPnia poundludicola LDeske amp K Mull Scapania saxicola Schust
46 (66) 141 136 141
Sc~~ scandica (H Arnell ~ Buch) Macv
139 140
Scapania subalpina (Nees) Durn
39 40 46 79 137 141
Scapania undulata (L) Dum
1 42 46 79 141
Scapania umbrosa (Schrad) Dum
137 140 141
52
Trichocolea tomentella (Ehrh) Dum Triomarla exsecta (Schmid) Schiffn
27 (35) 39 46 47 88 137 141 8 28 39 46 90 140 141
Tritomaria exsectiformis (Breidl) Loe~ke Tritomaria Sluinguedentata (Huds) Buch
39 -t6 89 90 136 140 141 7 39 46 141
Trltomaria scitula (Tayl) Joerg
46 136 (139) 141
53
~~ fragrans (Balbis) Frye et Clark Mannia pi 10sa (Hornem) Frye et Clark
46 (139) (140) 9 46 79 116 139
Mannia triandra (Scop) Gro 11e
46 (140)
Marchantia pound91ymorpha L
1 13 28 3946 (49) 78 87 88 89 90 (Ill) 121 122 147
Mo~rckia hibernica (HookJ Gott
46 89
Metzgeria conjugata Lindb
44
46
Mylia anomala (Hook) S Gray Notothylas orbicularis (Schwein) Sullo
39 46 141 (35)45 46
Nowellia curvifolia (Dicks) Mitt
17 27 3846 88 89 90 141
Odontoschisma denudatuJI1 (Nees ~ Mart)
Dum 46 81 137 141
Pallavicinia lyellii (Hook~ Carruth
90
Odontoschisma macounii (Aust) Under
45 46 (66) (139) 140 141
45
=
Pellia epiphllla (L) CordaPellia endiviifolia (Dicks~ Dum
27 39 46 90 13724 39 46
Pellia neesiana Limp bull Phaeoceros
39 46 89 27 39 45 46 97
Porella pinnata L
13 27 I 46 90 Plagiochila asplenioides (L) Dum
27 I 39 1+6 88 89 I 90 137
46
~e11a platYEhylla (L) Pfeiff Porella platyphylloi~ (Schwein) Lindb
17 27 39 46 69 (71)79 87 88 89 9 34 46 (57) 59 87 88 89 92 90 92 93 77
Preissia quadrata (Seop) Nees
1 36 3946 (71) 88 89 90 121 123
137
Ptllidium ciliare (L) Hampe
1 27 39 46 132 141
Dum
(22) 28 3946 (57) 87 88 89 90 92
Ptilidium ~lcherrium (G Web) Hampe
(9) 13 394656 (57)70 87 88 89 90 93 (119) 136 141
47
Radula obconica Sullo Reboulia hemlsphaerlca (L) Raddi
39 46 88 89 (43) 46 (59) 70 90 t Cf 124 125 (139)
Riccardia ~atifrons Lindb
1 27 39 46 89
Riccardia multifi~ (L) S Gray
39 46 88 145
Riccia beyrichiana Hampe ~ Lehm
46 Riccardia palmata (Hedw) Carruth
39 46 90
48
Riccia bifurca Hoffm Riccia cavernosa Hoffm
48 13739 46 79 137
Riccia fluitans L
242731454679
Riccla frostli Aust
48 79
Riccia sullicantii Aust
79 (137) Riccia sorocarpa Bisch 46 (137)
49
Ricciocarpus natans (1) Corda Sca~ia apiculata Spruce
13 27 46 79 89 110 39 46 (139) 141
ScaJltll1ia carinthica Jack ~ Lindb
137 t 141
Scapania ~ (Mart) Durn
39467990137141
Scapania digenil Schiffn
90
ScaEsectflia cuspiduligera (Nees) K Mull
46 (139) (140) (141)
50
Scapania glaucoeerha1a (Tayl) Aust Seapania gyrrmostomophilia Kaal
39 46 (139) 141 41 44 46 64 (139) (140) 141
ScaP3poundlia irrigua Gott ~ almiddot
7 39 46 79 137 (140L 141
Seapania nemorosa (L) Dum
1 39 46 89 90 137
Scapmi~ ~~ta Smiddotlch
137 141 51
~caPnia poundludicola LDeske amp K Mull Scapania saxicola Schust
46 (66) 141 136 141
Sc~~ scandica (H Arnell ~ Buch) Macv
139 140
Scapania subalpina (Nees) Durn
39 40 46 79 137 141
Scapania undulata (L) Dum
1 42 46 79 141
Scapania umbrosa (Schrad) Dum
137 140 141
52
Trichocolea tomentella (Ehrh) Dum Triomarla exsecta (Schmid) Schiffn
27 (35) 39 46 47 88 137 141 8 28 39 46 90 140 141
Tritomaria exsectiformis (Breidl) Loe~ke Tritomaria Sluinguedentata (Huds) Buch
39 -t6 89 90 136 140 141 7 39 46 141
Trltomaria scitula (Tayl) Joerg
46 136 (139) 141
53
Mylia anomala (Hook) S Gray Notothylas orbicularis (Schwein) Sullo
39 46 141 (35)45 46
Nowellia curvifolia (Dicks) Mitt
17 27 3846 88 89 90 141
Odontoschisma denudatuJI1 (Nees ~ Mart)
Dum 46 81 137 141
Pallavicinia lyellii (Hook~ Carruth
90
Odontoschisma macounii (Aust) Under
45 46 (66) (139) 140 141
45
=
Pellia epiphllla (L) CordaPellia endiviifolia (Dicks~ Dum
27 39 46 90 13724 39 46
Pellia neesiana Limp bull Phaeoceros
39 46 89 27 39 45 46 97
Porella pinnata L
13 27 I 46 90 Plagiochila asplenioides (L) Dum
27 I 39 1+6 88 89 I 90 137
46
~e11a platYEhylla (L) Pfeiff Porella platyphylloi~ (Schwein) Lindb
17 27 39 46 69 (71)79 87 88 89 9 34 46 (57) 59 87 88 89 92 90 92 93 77
Preissia quadrata (Seop) Nees
1 36 3946 (71) 88 89 90 121 123
137
Ptllidium ciliare (L) Hampe
1 27 39 46 132 141
Dum
(22) 28 3946 (57) 87 88 89 90 92
Ptilidium ~lcherrium (G Web) Hampe
(9) 13 394656 (57)70 87 88 89 90 93 (119) 136 141
47
Radula obconica Sullo Reboulia hemlsphaerlca (L) Raddi
39 46 88 89 (43) 46 (59) 70 90 t Cf 124 125 (139)
Riccardia ~atifrons Lindb
1 27 39 46 89
Riccardia multifi~ (L) S Gray
39 46 88 145
Riccia beyrichiana Hampe ~ Lehm
46 Riccardia palmata (Hedw) Carruth
39 46 90
48
Riccia bifurca Hoffm Riccia cavernosa Hoffm
48 13739 46 79 137
Riccia fluitans L
242731454679
Riccla frostli Aust
48 79
Riccia sullicantii Aust
79 (137) Riccia sorocarpa Bisch 46 (137)
49
Ricciocarpus natans (1) Corda Sca~ia apiculata Spruce
13 27 46 79 89 110 39 46 (139) 141
ScaJltll1ia carinthica Jack ~ Lindb
137 t 141
Scapania ~ (Mart) Durn
39467990137141
Scapania digenil Schiffn
90
ScaEsectflia cuspiduligera (Nees) K Mull
46 (139) (140) (141)
50
Scapania glaucoeerha1a (Tayl) Aust Seapania gyrrmostomophilia Kaal
39 46 (139) 141 41 44 46 64 (139) (140) 141
ScaP3poundlia irrigua Gott ~ almiddot
7 39 46 79 137 (140L 141
Seapania nemorosa (L) Dum
1 39 46 89 90 137
Scapmi~ ~~ta Smiddotlch
137 141 51
~caPnia poundludicola LDeske amp K Mull Scapania saxicola Schust
46 (66) 141 136 141
Sc~~ scandica (H Arnell ~ Buch) Macv
139 140
Scapania subalpina (Nees) Durn
39 40 46 79 137 141
Scapania undulata (L) Dum
1 42 46 79 141
Scapania umbrosa (Schrad) Dum
137 140 141
52
Trichocolea tomentella (Ehrh) Dum Triomarla exsecta (Schmid) Schiffn
27 (35) 39 46 47 88 137 141 8 28 39 46 90 140 141
Tritomaria exsectiformis (Breidl) Loe~ke Tritomaria Sluinguedentata (Huds) Buch
39 -t6 89 90 136 140 141 7 39 46 141
Trltomaria scitula (Tayl) Joerg
46 136 (139) 141
53
=
Pellia epiphllla (L) CordaPellia endiviifolia (Dicks~ Dum
27 39 46 90 13724 39 46
Pellia neesiana Limp bull Phaeoceros
39 46 89 27 39 45 46 97
Porella pinnata L
13 27 I 46 90 Plagiochila asplenioides (L) Dum
27 I 39 1+6 88 89 I 90 137
46
~e11a platYEhylla (L) Pfeiff Porella platyphylloi~ (Schwein) Lindb
17 27 39 46 69 (71)79 87 88 89 9 34 46 (57) 59 87 88 89 92 90 92 93 77
Preissia quadrata (Seop) Nees
1 36 3946 (71) 88 89 90 121 123
137
Ptllidium ciliare (L) Hampe
1 27 39 46 132 141
Dum
(22) 28 3946 (57) 87 88 89 90 92
Ptilidium ~lcherrium (G Web) Hampe
(9) 13 394656 (57)70 87 88 89 90 93 (119) 136 141
47
Radula obconica Sullo Reboulia hemlsphaerlca (L) Raddi
39 46 88 89 (43) 46 (59) 70 90 t Cf 124 125 (139)
Riccardia ~atifrons Lindb
1 27 39 46 89
Riccardia multifi~ (L) S Gray
39 46 88 145
Riccia beyrichiana Hampe ~ Lehm
46 Riccardia palmata (Hedw) Carruth
39 46 90
48
Riccia bifurca Hoffm Riccia cavernosa Hoffm
48 13739 46 79 137
Riccia fluitans L
242731454679
Riccla frostli Aust
48 79
Riccia sullicantii Aust
79 (137) Riccia sorocarpa Bisch 46 (137)
49
Ricciocarpus natans (1) Corda Sca~ia apiculata Spruce
13 27 46 79 89 110 39 46 (139) 141
ScaJltll1ia carinthica Jack ~ Lindb
137 t 141
Scapania ~ (Mart) Durn
39467990137141
Scapania digenil Schiffn
90
ScaEsectflia cuspiduligera (Nees) K Mull
46 (139) (140) (141)
50
Scapania glaucoeerha1a (Tayl) Aust Seapania gyrrmostomophilia Kaal
39 46 (139) 141 41 44 46 64 (139) (140) 141
ScaP3poundlia irrigua Gott ~ almiddot
7 39 46 79 137 (140L 141
Seapania nemorosa (L) Dum
1 39 46 89 90 137
Scapmi~ ~~ta Smiddotlch
137 141 51
~caPnia poundludicola LDeske amp K Mull Scapania saxicola Schust
46 (66) 141 136 141
Sc~~ scandica (H Arnell ~ Buch) Macv
139 140
Scapania subalpina (Nees) Durn
39 40 46 79 137 141
Scapania undulata (L) Dum
1 42 46 79 141
Scapania umbrosa (Schrad) Dum
137 140 141
52
Trichocolea tomentella (Ehrh) Dum Triomarla exsecta (Schmid) Schiffn
27 (35) 39 46 47 88 137 141 8 28 39 46 90 140 141
Tritomaria exsectiformis (Breidl) Loe~ke Tritomaria Sluinguedentata (Huds) Buch
39 -t6 89 90 136 140 141 7 39 46 141
Trltomaria scitula (Tayl) Joerg
46 136 (139) 141
53
~e11a platYEhylla (L) Pfeiff Porella platyphylloi~ (Schwein) Lindb
17 27 39 46 69 (71)79 87 88 89 9 34 46 (57) 59 87 88 89 92 90 92 93 77
Preissia quadrata (Seop) Nees
1 36 3946 (71) 88 89 90 121 123
137
Ptllidium ciliare (L) Hampe
1 27 39 46 132 141
Dum
(22) 28 3946 (57) 87 88 89 90 92
Ptilidium ~lcherrium (G Web) Hampe
(9) 13 394656 (57)70 87 88 89 90 93 (119) 136 141
47
Radula obconica Sullo Reboulia hemlsphaerlca (L) Raddi
39 46 88 89 (43) 46 (59) 70 90 t Cf 124 125 (139)
Riccardia ~atifrons Lindb
1 27 39 46 89
Riccardia multifi~ (L) S Gray
39 46 88 145
Riccia beyrichiana Hampe ~ Lehm
46 Riccardia palmata (Hedw) Carruth
39 46 90
48
Riccia bifurca Hoffm Riccia cavernosa Hoffm
48 13739 46 79 137
Riccia fluitans L
242731454679
Riccla frostli Aust
48 79
Riccia sullicantii Aust
79 (137) Riccia sorocarpa Bisch 46 (137)
49
Ricciocarpus natans (1) Corda Sca~ia apiculata Spruce
13 27 46 79 89 110 39 46 (139) 141
ScaJltll1ia carinthica Jack ~ Lindb
137 t 141
Scapania ~ (Mart) Durn
39467990137141
Scapania digenil Schiffn
90
ScaEsectflia cuspiduligera (Nees) K Mull
46 (139) (140) (141)
50
Scapania glaucoeerha1a (Tayl) Aust Seapania gyrrmostomophilia Kaal
39 46 (139) 141 41 44 46 64 (139) (140) 141
ScaP3poundlia irrigua Gott ~ almiddot
7 39 46 79 137 (140L 141
Seapania nemorosa (L) Dum
1 39 46 89 90 137
Scapmi~ ~~ta Smiddotlch
137 141 51
~caPnia poundludicola LDeske amp K Mull Scapania saxicola Schust
46 (66) 141 136 141
Sc~~ scandica (H Arnell ~ Buch) Macv
139 140
Scapania subalpina (Nees) Durn
39 40 46 79 137 141
Scapania undulata (L) Dum
1 42 46 79 141
Scapania umbrosa (Schrad) Dum
137 140 141
52
Trichocolea tomentella (Ehrh) Dum Triomarla exsecta (Schmid) Schiffn
27 (35) 39 46 47 88 137 141 8 28 39 46 90 140 141
Tritomaria exsectiformis (Breidl) Loe~ke Tritomaria Sluinguedentata (Huds) Buch
39 -t6 89 90 136 140 141 7 39 46 141
Trltomaria scitula (Tayl) Joerg
46 136 (139) 141
53
Radula obconica Sullo Reboulia hemlsphaerlca (L) Raddi
39 46 88 89 (43) 46 (59) 70 90 t Cf 124 125 (139)
Riccardia ~atifrons Lindb
1 27 39 46 89
Riccardia multifi~ (L) S Gray
39 46 88 145
Riccia beyrichiana Hampe ~ Lehm
46 Riccardia palmata (Hedw) Carruth
39 46 90
48
Riccia bifurca Hoffm Riccia cavernosa Hoffm
48 13739 46 79 137
Riccia fluitans L
242731454679
Riccla frostli Aust
48 79
Riccia sullicantii Aust
79 (137) Riccia sorocarpa Bisch 46 (137)
49
Ricciocarpus natans (1) Corda Sca~ia apiculata Spruce
13 27 46 79 89 110 39 46 (139) 141
ScaJltll1ia carinthica Jack ~ Lindb
137 t 141
Scapania ~ (Mart) Durn
39467990137141
Scapania digenil Schiffn
90
ScaEsectflia cuspiduligera (Nees) K Mull
46 (139) (140) (141)
50
Scapania glaucoeerha1a (Tayl) Aust Seapania gyrrmostomophilia Kaal
39 46 (139) 141 41 44 46 64 (139) (140) 141
ScaP3poundlia irrigua Gott ~ almiddot
7 39 46 79 137 (140L 141
Seapania nemorosa (L) Dum
1 39 46 89 90 137
Scapmi~ ~~ta Smiddotlch
137 141 51
~caPnia poundludicola LDeske amp K Mull Scapania saxicola Schust
46 (66) 141 136 141
Sc~~ scandica (H Arnell ~ Buch) Macv
139 140
Scapania subalpina (Nees) Durn
39 40 46 79 137 141
Scapania undulata (L) Dum
1 42 46 79 141
Scapania umbrosa (Schrad) Dum
137 140 141
52
Trichocolea tomentella (Ehrh) Dum Triomarla exsecta (Schmid) Schiffn
27 (35) 39 46 47 88 137 141 8 28 39 46 90 140 141
Tritomaria exsectiformis (Breidl) Loe~ke Tritomaria Sluinguedentata (Huds) Buch
39 -t6 89 90 136 140 141 7 39 46 141
Trltomaria scitula (Tayl) Joerg
46 136 (139) 141
53
Riccia bifurca Hoffm Riccia cavernosa Hoffm
48 13739 46 79 137
Riccia fluitans L
242731454679
Riccla frostli Aust
48 79
Riccia sullicantii Aust
79 (137) Riccia sorocarpa Bisch 46 (137)
49
Ricciocarpus natans (1) Corda Sca~ia apiculata Spruce
13 27 46 79 89 110 39 46 (139) 141
ScaJltll1ia carinthica Jack ~ Lindb
137 t 141
Scapania ~ (Mart) Durn
39467990137141
Scapania digenil Schiffn
90
ScaEsectflia cuspiduligera (Nees) K Mull
46 (139) (140) (141)
50
Scapania glaucoeerha1a (Tayl) Aust Seapania gyrrmostomophilia Kaal
39 46 (139) 141 41 44 46 64 (139) (140) 141
ScaP3poundlia irrigua Gott ~ almiddot
7 39 46 79 137 (140L 141
Seapania nemorosa (L) Dum
1 39 46 89 90 137
Scapmi~ ~~ta Smiddotlch
137 141 51
~caPnia poundludicola LDeske amp K Mull Scapania saxicola Schust
46 (66) 141 136 141
Sc~~ scandica (H Arnell ~ Buch) Macv
139 140
Scapania subalpina (Nees) Durn
39 40 46 79 137 141
Scapania undulata (L) Dum
1 42 46 79 141
Scapania umbrosa (Schrad) Dum
137 140 141
52
Trichocolea tomentella (Ehrh) Dum Triomarla exsecta (Schmid) Schiffn
27 (35) 39 46 47 88 137 141 8 28 39 46 90 140 141
Tritomaria exsectiformis (Breidl) Loe~ke Tritomaria Sluinguedentata (Huds) Buch
39 -t6 89 90 136 140 141 7 39 46 141
Trltomaria scitula (Tayl) Joerg
46 136 (139) 141
53
Ricciocarpus natans (1) Corda Sca~ia apiculata Spruce
13 27 46 79 89 110 39 46 (139) 141
ScaJltll1ia carinthica Jack ~ Lindb
137 t 141
Scapania ~ (Mart) Durn
39467990137141
Scapania digenil Schiffn
90
ScaEsectflia cuspiduligera (Nees) K Mull
46 (139) (140) (141)
50
Scapania glaucoeerha1a (Tayl) Aust Seapania gyrrmostomophilia Kaal
39 46 (139) 141 41 44 46 64 (139) (140) 141
ScaP3poundlia irrigua Gott ~ almiddot
7 39 46 79 137 (140L 141
Seapania nemorosa (L) Dum
1 39 46 89 90 137
Scapmi~ ~~ta Smiddotlch
137 141 51
~caPnia poundludicola LDeske amp K Mull Scapania saxicola Schust
46 (66) 141 136 141
Sc~~ scandica (H Arnell ~ Buch) Macv
139 140
Scapania subalpina (Nees) Durn
39 40 46 79 137 141
Scapania undulata (L) Dum
1 42 46 79 141
Scapania umbrosa (Schrad) Dum
137 140 141
52
Trichocolea tomentella (Ehrh) Dum Triomarla exsecta (Schmid) Schiffn
27 (35) 39 46 47 88 137 141 8 28 39 46 90 140 141
Tritomaria exsectiformis (Breidl) Loe~ke Tritomaria Sluinguedentata (Huds) Buch
39 -t6 89 90 136 140 141 7 39 46 141
Trltomaria scitula (Tayl) Joerg
46 136 (139) 141
53
Scapania glaucoeerha1a (Tayl) Aust Seapania gyrrmostomophilia Kaal
39 46 (139) 141 41 44 46 64 (139) (140) 141
ScaP3poundlia irrigua Gott ~ almiddot
7 39 46 79 137 (140L 141
Seapania nemorosa (L) Dum
1 39 46 89 90 137
Scapmi~ ~~ta Smiddotlch
137 141 51
~caPnia poundludicola LDeske amp K Mull Scapania saxicola Schust
46 (66) 141 136 141
Sc~~ scandica (H Arnell ~ Buch) Macv
139 140
Scapania subalpina (Nees) Durn
39 40 46 79 137 141
Scapania undulata (L) Dum
1 42 46 79 141
Scapania umbrosa (Schrad) Dum
137 140 141
52
Trichocolea tomentella (Ehrh) Dum Triomarla exsecta (Schmid) Schiffn
27 (35) 39 46 47 88 137 141 8 28 39 46 90 140 141
Tritomaria exsectiformis (Breidl) Loe~ke Tritomaria Sluinguedentata (Huds) Buch
39 -t6 89 90 136 140 141 7 39 46 141
Trltomaria scitula (Tayl) Joerg
46 136 (139) 141
53
~caPnia poundludicola LDeske amp K Mull Scapania saxicola Schust
46 (66) 141 136 141
Sc~~ scandica (H Arnell ~ Buch) Macv
139 140
Scapania subalpina (Nees) Durn
39 40 46 79 137 141
Scapania undulata (L) Dum
1 42 46 79 141
Scapania umbrosa (Schrad) Dum
137 140 141
52
Trichocolea tomentella (Ehrh) Dum Triomarla exsecta (Schmid) Schiffn
27 (35) 39 46 47 88 137 141 8 28 39 46 90 140 141
Tritomaria exsectiformis (Breidl) Loe~ke Tritomaria Sluinguedentata (Huds) Buch
39 -t6 89 90 136 140 141 7 39 46 141
Trltomaria scitula (Tayl) Joerg
46 136 (139) 141
53
Trichocolea tomentella (Ehrh) Dum Triomarla exsecta (Schmid) Schiffn
27 (35) 39 46 47 88 137 141 8 28 39 46 90 140 141
Tritomaria exsectiformis (Breidl) Loe~ke Tritomaria Sluinguedentata (Huds) Buch
39 -t6 89 90 136 140 141 7 39 46 141
Trltomaria scitula (Tayl) Joerg
46 136 (139) 141
53