John Squires, Rocky Mountain Research Station, Missoula, MT
Current Distribution, Status, and Threats of Canada Lynx in Montana and Wyoming
3
Basis of Assessment - Montana:
a) Lynx Research Program at RMRS initiated in 1998 b) Captured and collared 175 individuals c) Recorded 169,782 GPS and 3043 VHF locations that document lynx movements and resource- use
d) Investigated the following topics regarding lynx in Montana:
4
Basis of Assessment:
• Resource selection Squires, J. R, N. J. DeCesare, J. A. Kolbe, and L. F. Ruggiero. 2008.
Hierarchical den selection of Canada lynx in western Montana. Journal of Wildlife Management 72:1497-1506.
Squires, J. R., N. J. DeCesare, J. A. Kolbe, and L. F. Ruggiero. 2010. Seasonal Resource Selection of Canada Lynx in Managed Forests of the Northern Rocky Mountains. Journal of Wildlife Management 74:1648-1660.
• Prey Selection Squires, J. R. and L. F. Ruggiero. 2007. Winter prey selection of Canada lynx
in northwestern Montana. Journal of Wildlife Management 71:310-315.
• Competition Kolbe, J. A., J. R. Squires, D. H. Pletscher, and R. F. Ruggiero. 2007. The
effect of snowmobile trails on coyote movements within lynx home ranges. Journal of Wildlife Management 71:1409-1418.
5
Basis of Assessment:
• Activity Patterns Kolbe, J. A. and J. R. Squires. 2007. Circadian activity patterns of Canada
lynx in western Montana. Journal of Wildlife Management 71:1607-1611.
Olson, L. E., J. R. Squires, N. J. DeCesare, J. A. Kolbe. 2011. Den use and activity patterns in female Canada lynx (Lynx canadensis) in the Northern Rocky Mountains. Northwest Science 85(3):455-462.
• Detection/Monitoring Squires, J. R., K. S. McKelvey, L. F. Ruggiero. 2004. A snow-tracking protocol
used to delineate local lynx, Lynx canadensis, distributions. Canadian Field-Naturalist 118:583-589.
McKelvey, K. S., J. Von Kienast, K. B. Aubry, G. M. Koehler, B. T. Maletzke, J. R. Squires, E. L. Lindquist, S. Loch, M. K. Schwartz. 2006. DNA analysis of hair and scat collected along snow tracks to document the presence of Canada lynx (Lynx canadensis). Wildlife Society Bulletin 34:451-455.
Squires, J. R., L. E. Olson, D. L. Turner, N. J. DeCesare, and J. A. Kolbe. 2012. Estimating detection probability for Canada lynx using snow-track surveys in the Northern Rocky Mountains. Wildlife Biology 18:215-224.
6
Basis of Assessment:
• Connectivity
Squires, J. R., Nicholas J. DeCesare, Lucretia E. Olson, Jay A. Kolbe, Mark Hebblewhite, and Sean A. Parks. 2013. Combining resource selection and movement behavior to predict corridors of Canada lynx at their southern range periphery. Biological Conservation 157:187–195.
7
Basis of Assessment - Wyoming:
a) Wyoming Game and Fish Department (WGF), in cooperation with the Shoshone National Forest (SNF), initiated lynx surveys during winter 1995/96 - no lynx detected on SNF - Impetus for WGF to fund additional surveys, trapping, and telemetry in the Wyoming Range b) During 1997-98, WGF searched approximately 2055 km of maintained snowmobile routes and 2400 km of non-maintained trails for lynx tracks in 12 areas (Laurion and Oakleaf 1998) – 6 lynx tracks detected
8
Basis of Assessment - Wyoming:
c) During winter 1998-99, three general areas were searched (Laurion and Oakleaf 1999) - 6 tracks located in Wyoming Range.
d) RMRS, in cooperation with WGD, conducted lynx surveys in 2000, 2001, and 2002 - 2000 – Wyoming Range: 5 lynx tracks - kitten and female - 2001 – Wyoming Range: 5 lynx tracks - 2002 – no detections
9
Basis of Assessment - Wyoming:
e) Yellowstone Park Lynx survey from 2001 – 2004 (1,143 km ski-based snow tracks, 749 km snowmobile-based survey, 693 km aircraft survey, and 35 hair snare transects - 105-175 stations; Murphy et al. 2006. Distribution of Canada Lynx in Yellowstone National Park. Northwest
Science 80:199-206)
f) DNA confirmed detections of 3 lynx – 1 female, 1 female with male kitten, and 1 male. All detections on east side of Yellowstone Park – east shore Yellowstone Lake.
10
Basis of Assessment - Wyoming:
g) Endeavor Wildlife Research Foundation conducted track/DNA surveys between 2004-2005 in southern GYA (Bridger-Teton Nat. Forest including Gros Ventre and Teton Wilderness areas) – searched 4,320 miles and detected (DNA confirmed) 18 lynx tracks. Confirmed presence in Wyoming Range.
h) In 2008-2009, Endeavor Wildlife Research searched 2,854 miles for winter tracks throughout the GYA – documented 6 detections on Togwotee Pass, 2 possible detections in Yellowstone National Park, and 1 possible detection on the Beartooth Plateau.
11
Basis of Assessment – Wyoming (telemetry):
e) From 2006-2007, WGF collared 2 lynx (one male, 1 female) in Wyoming Range – monitored throughout the year with conventional ground and aerial telemetry (1996 – 2001) – (N=219 locations – male, N = 212 - female)
f) In 2000, female died. WGF asked RMRS to replace the collar on male with ARGOS (N = 258). Documented summer exploratory movements of male (1999 –2001) based on conventional and Argos telemetry (Squires and Oakleaf. 2005. Movements of a male Canada lynx crossing the Greater Yellowstone Area, including highways. Northwest Science 79:196-201).
12
Status - Montana
13
Status - Montana
Litter size of lynx in western Montana, 1999 – 2007
Seeley Lake
Purcell Mountains
Combined
Year Litters Kittens Kittens / litter
Litters Kittens Kittens / litter
Litters Kittens Kittens / litter
1999 2 4 2.00 2 4 2.00 2000 4 6 1.50 4 6 1.50 2001 3 8 2.67 3 8 2.67 2002 3 4 1.33 3 4 1.33 2003 5 14 2.80 5 14 2.80 2004 5 15 3.00 5 16 3.20 10 31 3.10 2005 5 12 2.40 6 19 3.17 11 31 2.82 2006 3 5 1.67 3 8 2.67 6 13 2.17 2007 3 7 2.33 8 22 2.75 11 29 2.64
MLE Mean 1 33 75 2.24 22 65 2.95 55 140 2.53 MLE Var 1 0.002 0.084 0.008 95% CI 2.21-2.27 2.67-3.23 2.51-2.55
Litter size of lynx in western Montana, 1999 – 2007
Seeley Lake
Purcell Mountains
Combined
Year
Litters
Kittens
Kittens / litter
Litters
Kittens
Kittens / litter
Litters
Kittens
Kittens / litter
1999
2
4
2.00
2
4
2.00
2000
4
6
1.50
4
6
1.50
2001
3
8
2.67
3
8
2.67
2002
3
4
1.33
3
4
1.33
2003
5
14
2.80
5
14
2.80
2004
5
15
3.00
5
16
3.20
10
31
3.10
2005
5
12
2.40
6
19
3.17
11
31
2.82
2006
3
5
1.67
3
8
2.67
6
13
2.17
2007
3
7
2.33
8
22
2.75
11
29
2.64
MLE Mean 1
33
75
2.24
22
65
2.95
55
140
2.53
MLE Var 1
0.002
0.084
0.008
95% CI
2.21-2.27
2.67-3.23
2.51-2.55
14
Status - Montana Proportion of successful adult Canada lynx
Seeley Lake (N=52 breeding-age females)
Purcell Mountains (N=28)
Combined (N = 80)
Year Females Females w kittens
Prop Females Females w kittens
Prop Females Females w kittens
Prop
1999 4 2 0.50 4 2 0.50 2000 6 4 0.67 6 4 0.67 2001 9 3 0.33 9 3 0.33 2002 6 2 0.33 6 2 0.33 2003 5 3 0.60 5 3 0.60 2004 6 4 0.68 5 5 1.00 11 9 0.82 2005 6 5 0.83 7 7 1.00 13 12 0.92 2006 3 3 1.00 6 4 0.67 9 7 0.78 2007 3 3 1.00 10 7 0.70 13 10 0.77
MLE 1 Mean 0.61 0.83 0.67
MLE Var 1 0.01 0.01 0.02
95% CI 0.42-0.81 0.43-0.98 0.45-0.82
Proportion of successful adult Canada lynx
Seeley Lake
(N=52 breeding-age females)
Purcell Mountains
(N=28)
Combined
(N = 80)
Year
Females
Females w kittens
Prop
Females
Females w kittens
Prop
Females
Females w kittens
Prop
1999
4
2
0.50
4
2
0.50
2000
6
4
0.67
6
4
0.67
2001
9
3
0.33
9
3
0.33
2002
6
2
0.33
6
2
0.33
2003
5
3
0.60
5
3
0.60
2004
6
4
0.68
5
5
1.00
11
9
0.82
2005
6
5
0.83
7
7
1.00
13
12
0.92
2006
3
3
1.00
6
4
0.67
9
7
0.78
2007
3
3
1.00
10
7
0.70
13
10
0.77
MLE 1
Mean
0.61
0.83
0.67
MLE Var 1
0.01
0.01
0.02
95% CI
0.42-0.81
0.43-0.98
0.45-0.82
15
We estimated survival based on 125 lynx monitored monthly from 1999-2007 using a staggered entry design; we documented 2376 lynx-use months during this period.
16
Annual Survival Rate of female lynx on the Seeley Lake(1999-2007) and Purcell (2003-2007) Study Areas including a combined estimate.
Female
Subadult Survival 1 Female
Adult Survival 1
Seeley Mean 0.515 0.747
Variance 0.014 0.003
95% CI 0.283 - 0.746 0.648 - 0.846
Purcells Mean 0.683 0.846
Variance 0.017 0.004
95% CI 0.428 - 0.937 0.721 - 0.970
Combined Mean 0.520 0.753
Variance 0.010 0.002
95% CI 0.322 - 0.718 0.659 - 0.847
Annual Survival Rate of female lynx on the Seeley Lake(1999-2007) and Purcell (2003-2007) Study Areas including a combined estimate.
Female
Subadult Survival 1
Female
Adult Survival 1
Seeley
Mean
0.515
0.747
Variance
0.014
0.003
95% CI
0.283 - 0.746
0.648 - 0.846
Purcells
Mean
0.683
0.846
Variance
0.017
0.004
95% CI
0.428 - 0.937
0.721 - 0.970
Combined
Mean
0.520
0.753
Variance
0.010
0.002
95% CI
0.322 - 0.718
0.659 - 0.847
17
Status - Montana
0200400600800
1000120014001600
0.70
0.80
0.90
1.00
1.10
1.20
1.30
1.40
1.50
1.60
Lambda
# of
repl
icat
es
SeeleyPurcellsCombined
Frequency distribution of λ values from 5000 replicates in which lynx vital rates were chosen from a uniform distribution bounded by their 95% confidence intervals
Chart2
0.70.70.7
0.750.750.75
0.80.80.8
0.850.850.85
0.90.90.9
0.950.950.95
111
1.051.051.05
1.11.11.1
1.151.151.15
1.21.21.2
1.251.251.25
1.31.31.3
1.351.351.35
1.41.41.4
1.451.451.45
1.51.51.5
1.551.551.55
1.61.61.6
1.651.651.65
Seeley
Purcells
Combined
Lambda
# of replicates
0
0
0
11
0
0
213
0
7
608
0
178
1080
16
611
1285
86
1079
995
240
1383
549
456
1002
199
680
560
52
817
154
8
798
25
0
708
1
0
499
0
0
356
0
0
204
0
0
92
0
0
39
0
0
7
0
0
2
0
0
0
0
determ_lambdas
Note: This worksheet shows the λ values from our determinisitc models.
λD= the deterministic lambda from the mean matrix for each population, or the matrix in which vital rates are equal to their MLE means
mean λD_rep= the mean lambda from the 5000 replicate matrices in which vital rates were chosen from uniform distributions bound by the 95% CL's for each vital rate
λ from Replicates comes from the the 5000 replicate matrices; the data are stored in each study area's Deterministic folder in a file called "(studyarea)_randomlambdas"
Study Area:SeeleyPurcellsCombinedFrequency Dist. of λ from 5000 replicatesCumulative Dist. of λ from 5000 replicates
λD:0.9231.2230.975
mean λD_rep:0.9251.1680.973λSeeleyPurcellsCombinedλSeeleyPurcellsCombined
std_dev of mean:0.0750.1170.0700.700000.70000
95% UCL:0.9271.1710.0020.7511000.750.2200
95% LCL:0.9231.1650.0000.80213070.804.4800.14
min_λ:0.7200.8530.7800.8560801780.8516.6403.7
max_λ:1.1921.5941.2070.901080166110.9038.240.3215.92
median_λ:0.9241.1630.9720.9512858610790.9563.942.0437.5
λ values from 5000 Replicates:1.21911.01161.0099524013831.0083.846.8465.16
11.07221.14581.08531.0554945610021.0594.8215.9685.2
20.948141.09270.985161.101996805601.1098.829.5696.4
30.905821.23831.01091.15528171541.1599.8445.999.48
40.910631.24121.02461.208798251.2010061.8699.98
50.899351.13891.05571.25070811.2510076.02100
61.11041.16860.982771.30049901.3010086100
70.987181.01840.910251.35035601.3510093.12100
80.972351.12750.862671.40020401.4010097.2100
90.88011.26290.874111.4509201.4510099.04100
100.926211.2271.08111.5003901.5010099.82100
110.882541.20321.03721.550701.5510099.96100
120.927591.19741.02931.600201.60100100100
130.887121.14370.970851.650001.65100100100
140.942971.37970.98392
150.849441.12110.87596
160.909111.17831.12990.70000
170.882181.09430.845970.751100
180.925981.31380.970330.8022407
191.02261.08471.01770.858320185
201.01991.27650.987390.90191216796
210.93561.11161.0720.9531971021875
220.933651.27181.03061.0041923423258
230.863850.994861.00731.0547417984260
240.787711.06730.91121.10494014784820
250.982741.13820.995951.15499222954974
260.908821.33251.07291.20500030934999
270.841681.06990.991031.25500038015000
281.0261.09311.02331.30500043005000
290.862261.31690.995821.35500046565000
300.885011.02450.954191.40500048605000
310.903151.15230.974961.45500049525000
320.989661.1140.971521.50500049915000
330.877481.34450.947221.55500049985000
340.904521.13190.959681.60500050005000
351.00691.09591.0041.65500050005000
360.804261.04741.1463
370.867181.30720.94618
380.810271.14570.83689
390.803571.02921.0378
400.872421.30.93367
410.963851.06130.82504
420.830171.34610.9045
430.971771.41190.9782
440.877161.37641.0758
450.965171.27131.0635
460.883231.09860.96167
470.913971.12141.0812
480.904381.14930.79761
490.839711.12080.98884
500.831651.0540.95876
510.844681.02381.0095
520.934531.08590.99387
530.952591.07440.87838
541.00611.18910.92305
550.950520.917140.91573
560.831331.09790.85254
570.795131.03811.0164
580.894721.02260.97779
590.938521.35230.98565
600.981631.06160.93798
610.925151.30940.9799
620.935151.08860.99776
630.860921.08671.0075
640.930511.0740.82092
650.922721.29070.91295
660.884761.23450.92155
670.939391.14190.97168
680.920251.26250.93703
691.00191.12290.9161
700.841161.37020.94362
710.956841.1511.0146
720.811221.23840.85968
730.921951.30611.055
740.84551.21140.91431
750.825811.18680.98178
761.02931.29410.93738
770.943741.15340.95626
780.956411.10990.94583
791.09661.18560.97578
801.00681.05541.078
810.945341.13370.83511
820.848941.06261.117
830.98031.19281.0667
840.939831.21890.88514
850.924791.13790.83901
860.853741.02630.98255
870.815381.26540.87745
880.850891.43690.91552
890.869221.14780.99491
900.831551.20691.0772
910.935741.27521.0268
920.865471.25451.0193
930.929371.08850.91067
940.986741.17020.87507
950.982321.14031.0074
960.907461.21820.97069
970.852831.11821.0307
980.772651.41780.96541
990.94031.19841.0281
1000.977751.0760.99449
1010.789141.20841.0468
1020.977091.11270.97941
1030.949141.11751.0431
1041.02891.08321.0935
1051.14471.33180.90652
1060.965961.00281.0087
1070.919241.2881.1146
1081.00091.5140.97512
1091.03391.29581.0498
1100.974341.08930.89624
1110.93861.00310.90952
1120.874981.01111.0053
1130.933921.11760.91446
1140.905091.28370.88739
1150.785410.985211.0133
1160.800111.10331.0845
1170.989711.4010.91996
1181.0031.12330.92016
1190.818851.11990.95115
1200.946341.36870.91711
1210.908131.221.0407
1220.86831.28890.9972
1231.05391.15030.9386
1240.866571.3040.86497
1250.977621.26151.0324
1260.923761.21290.92383
1270.790171.26181.0696
1280.855611.19540.83798
1291.09081.22750.96169
1300.869581.36540.9311
1310.911961.11980.93033
1320.905471.25280.90556
1330.839441.38430.9171
1340.865011.15711.0485
1350.884661.2610.97832
1360.849951.30110.99421
1371.00331.36221.0202
1380.895251.25560.94605
1390.969811.29030.97906
1400.927011.18031.055
1410.919651.11131.1016
1421.00631.1120.97294
1430.918211.20171.1256
1440.963881.30441.0468
1450.905151.30430.84398
1460.990871.13911.015
1470.787371.1151.0006
1480.812541.27671.0354
1490.967141.11021.0683
1500.945451.38620.93908
1510.874441.1980.98496
1520.852341.17370.95106
1530.920061.17390.90819
1540.944921.20820.91947
1550.937751.23470.94108
1560.887751.09871.0448
1570.831691.26181.0755
1580.954820.932490.92493
1590.918121.17360.86793
1600.993571.05370.93094
1610.858151.110.96746
1620.975361.33541.0113
1630.962151.33750.9161
1640.99061.31891.0566
1651.01051.40321.0287
1661.00891.10991.0511
1670.882011.08710.90267
1680.921431.13870.89239
1690.924421.19170.92883
1700.801710.980360.983
1710.937291.25850.86821
1720.973951.29940.93049
1730.992181.33351.0063
1740.865351.36480.93471
1750.884521.12410.99329
1760.856381.10840.94299
1770.92190.959260.98652
1780.896581.29890.92373
1790.9831.41171.1038
1800.958231.22060.98936
1810.961981.14421.0913
1821.00871.07780.81181
1831.02271.11440.87117
1840.819941.04581.0254
1850.998431.23530.97976
1860.933881.03341.1629
1871.07981.14790.96356
1880.775831.32380.94831
1890.911051.24351.0218
1900.947581.15561.0281
1910.811821.15620.94609
1920.957791.33061.0961
1930.796321.22981.1324
1940.868081.24770.8981
1951.0131.21750.84481
1960.903651.10621.0038
1970.974691.09591.0128
1980.792781.2160.99659
1990.887191.14380.97214
2000.779111.09981.0515
2011.00441.26840.89833
2020.846531.1380.94539
2031.03321.26031.1056
2040.821441.2331.0299
2050.941221.11051.0716
2060.878061.27220.88033
2070.891291.19740.97583
2080.804511.22780.98685
2090.94941.07930.91425
2100.956571.09790.97221
2110.988771.07251.0172
2120.971691.1271.0137
2130.83271.17250.96876
2140.991911.15341.0116
2150.995410.981950.89298
2161.00851.24460.91787
2170.943791.0260.96212
2180.877941.0580.99448
2190.916491.09531.1326
2200.875091.00331.1523
2210.898821.2431.0602
2220.905791.17420.95258
2230.91681.2750.95241
2240.958871.11131.0728
2250.944110.984580.95289
2260.87031.24260.90668
2270.905841.07761.0439
2280.938341.02720.95157
2290.926681.24531.0161
2301.00441.11941.0463
2310.916761.19361.0072
2321.05751.06170.86868
2330.851.36310.92659
2341.09021.34281.0296
2350.822321.02141.1178
2360.954171.2980.99558
2370.774011.37411.0038
2380.893471.37910.94602
2390.834321.13610.9285
2400.828991.21230.91297
2410.835211.2690.90743
2421.09551.03160.88018
2430.974481.26440.99
2440.848471.05170.93035
2450.956261.03090.99334
2461.03961.03271.0441
2470.990670.98131.0205
2480.857731.38670.99236
2491.0241.13891.0305
2500.95051.22731.0116
2510.918031.0850.95817
2520.869381.03320.98098
2530.880471.32651.0329
2540.83481.26050.90459
2550.992811.17391.071
2560.905491.10240.98019
2570.902211.28690.87163
2580.97311.05181.0137
2591.01591.19931.0403
2600.959231.06810.92144
2610.959331.1520.98653
2620.8681.05930.91701
2630.898231.12791.0162
2640.945221.11150.92401
2650.998071.1420.99881
2660.908611.08450.84298
2671.02871.01590.96118
2681.01931.20150.86678
2690.936311.29410.87872
2700.98931.24120.8669
2711.08631.34390.97072
2720.874421.05330.90332
2730.9341.19590.88756
2740.941581.1290.86591
2751.02931.10391.0375
2761.02830.987540.91478
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