New Species Discoveries and Records in Bhutan...

© Hartmann, Barclay & Weipert: Biodiversität und Naturausstattung im Himalaya VI. - Erfurt 2018

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Introduction

Bhutan is located in the eastern Himalayas, with an area of 38,394 km², bordering China to the north and India to the south, east and west (NBSAP 2014). The country has 70.46 % forest cover (LCMP 2010). The lowest elevation is at around 100 metres above sea level (masl) in southern Bhutan, and the highest peak, Gangkhar Phuensum, rises to 7,570 masl. Most researches has been focused on the three taxonomic groups, mammals, birds, and plants. By far the largest biodiversity is among invertebrate groups such as molluscs, dragonflies and damselflies, beetles, bees and wasps, true flies, moths and butterflies. However, these groups remain largely unstudied. Some research on certain taxonomic groups has been initiated, for example on fungi, fishes, amphibians and reptiles (Gyelt-shen et al. 2017a, Gurung et al. 2013, Wangyel 2013, Mata et al. 2010). Some taxonomic groups are even less studied and thus lack baseline data. We expect many new records and new species to be discovered from these groups.

Currently, peer-reviewed scientific documentation of new spe-cies records and discoveries are poor. New records are more commonly published through local media and press releases. It would be more efficient to document new records and species discoveries through scientific publications and documenta-tion through online platforms such as the Bhutan Biodiversity Portal (www.biodiversity.bt). This data portal is a consortium based online platform to document the country’s biodiversity using Citizen Science with taxonomic expert participation.

The insect fauna of Bhutan is poorly studied. Thus, an inverte-brate project in Bhutan was initiated in 2012, which aimed to enhance the research facilities for the study of invertebrates. This led to the discoveries of numerous new records and spe-cies new to science almost every year. The National Biodi-

versity Centre currently houses specimens of some groups of invertebrates, which will serve as permanent collections and the basis for further research and public education.

A checklist of new discoveries and new records of species from 2009-2017 is provided; however not all new records and discoveries may have been provided due to the lack of access to scientific peer-reviewed publications, and some may have been overlooked. This article only documents the new discoveries and new records from peer-reviewed publications accessible to the authors.

Materials and Methods

To generate this article, many institutions and researchers were consulted for information on specific taxonomic groups. Species data were obtained from peer-reviewed publications by the National Biodiversity Centre (NBC), Sherubtse College, College of Natural Resources (CNR), and Naturalis Biodiver-sity Center (The Netherlands), Ugyen Wangchuck Institute for Conservation and Environmental Research (UWICER), De-partment of Forests and Park Services (DoFPs), National Plant Protection Centre (NPPC), and other individual researchers and institutions (Gittenberger et al. 2017a, 2017b, Gyelt-shen et al. 2017a, Dorji et al. 2017, Yoshida Yangzom & Long. 2017, Dalstrom et al. 2017). Additional information on species data were obtained from the Bhutan Biodiversity Portal and the National Insect Repository Centre at the Na-tional Biodiversity Centre in Thimphu. New species validation and data curation has been conducted to ensure the quality of the information presented here. Species rediscoveries are not taken into account.

New Species Discoveries and Records in Bhutan Himalaya

Choki Gyeltshen, Kezang Tobgay, Nima Gyeltshen, Tshering Dorji & Sangay Dema

Abstract

Bhutan is well known for its rich biodiversity and efficient environmental conservation; however certain taxonomic groups are data deficient. The number of new species records and discoveries is expected to increase, since many of the less accessible areas of high biodiversity are still unexplored. This assumption is further substantiated by the fact that a number of new species have been described and new records have been found in the country over the years despite a very low level of taxonomic work. This paper therefore attempts to take stock of the recently discovered and described species as well as new species records for Bhutan from 2009 - 2017. The objectives of this compilation is to provide updated and comprehensive information on biodiversity statistics on new species records and discoveries in Bhutan. At least 31 species new to science were discovered in Bhutan during the last nine years. The new discoveries include 16 species of plants, four moths, four molluscs, one dragonfly, three fishes, one stonefly, and two beetle species. The new species discoveries have occurred at a rate of three species per year. A total of 566 new species of flora and fauna for Bhutan have been recorded.

Zusammenfassung

Bhutan ist bekannt für seinen Artenreichtum und seinen effizienten Umweltschutz, jedoch ist das Wissen um viele Artengruppen noch defizitär. Die Zahl der nachgewiesenen Arten steigt stetig, viele Gebiete mit großem Artenreichtum sind noch unerforscht. Dies hängt auch mit dem bisher noch unzureichenden Stand der taxonomischen und faunistischen Arbeiten im Land zusammen. Die vorliegende Arbeit fasst alle Arten zusammen, die von 2009 bis 2017 entweder neu aus Bhutan beschrieben oder neu nachgewiesen wurden. In den letzten neun Jahren konnten 31 Arten als neu für die Wissenschaft beschrieben werden, darunter 16 Pflanzenarten, vier Schmetterlingsarten, vier Schnecken-arten, drei Fischarten und je eine Art von Libellen, Steinfliegen und zwei Käferarten. Dies ist eine Rate von drei Arten pro Jahr. Bisher sind insgesamt im betrachteten Zeitraum 566 Arten neu für Bhutan nachgewiesen.

Key words: Biodiversity, statistics, checklists, Himalaya, Bhutan

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Hartmann, M.; M.V.L. Barclay & J. Weipert: Biodiversität und Naturausstattung im Himalaya VI

Results

Species new to science

According to the World Wide Fund for Nature, 2009 report, at least 353 new species have been discovered in the eastern Himalayas from 1998-2008, which include 242 plants, 16

amphibians, 16 reptiles, 14 fishes, two birds and two mammals and at least 61 new invertebrates. Of these, 21 species were discovered in Bhutan from 1998-2008.

This article discusses, at least 31 species new to science, dis-covered in Bhutan from 2009-2017, comprising 16 species of plants, four moths, four molluscs, one dragonfly, three fishes,

Fig. 3. Roscoea megalantha discovered from eastern Bhutan in 2017. (© Choki Gyeltshen).

Fig. 4. Meconopsis merakensis, new to science. (© Rinchen Yangzom).

Fig. 1. Spathoglottis jetsuniae discovered from central Bhutan in 2017. (© Kezang Tobgay, NBC).

Fig. 2. Meconopsis gakyidiana, national flower of Bhutan. (© T. Yoshida).

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Gyeltshen, Ch. et al.: New Species Discoveries and Records in Bhutan Himalaya ...

one stonefly, and two beetle species (Table. 1). The new species discoveries have occurred at a rate of three species per year. The following is a brief summary of new species described from Bhutan from 2009 to 2017:

Plants (Angiosperms)

In 2017, a new species of orchid was discovered, which was de-scribed by a team from NBC, and was named in honour of Her Majesty Gyaltsuen Jetsuen Pema Wangchuck as Spathoglottis jetsuniae (Gyeltshen et al. 2017b; Fig. 1). Spathoglottis jetsuniae was first discovered in Zhemgang and is currently also known from two other localities in Samdrup Jongkhar and Samtse. Fur-thermore, in the same year, two more species of Meconopsis were described, namely, Meconopsis merakensis (Fig. 4) (Yoshida, Yangzom & Long 2017) from eastern Bhutan and M. elongata (Yoshida et al. 2016) from Haa district in western Bhutan. In-terestingly, the national flower of Bhutan, which was previously considered to be a form of M. grandis, was found to be a different species. Thus, the national flower, from eastern Bhutan, was rec-ognised as a separate species and named M. gakyidiana (Yoshida, Yangzom & Long 2017; Fig. 2).

In addition, in 2017 a new species Roscoea megalantha (Yoshi-da, Yangzom & Newman 2017; Fig. 3) was discovered and described from eastern Bhutan. This species was previously recognized as Roscoea purpurea by other regional herbaria. Cur-rently, Bhutan has three species of Roscoea species: R. alpina, R. bhutanica, and R. megalantha. A blue poppy, Meconopsis bhutanica (Yoshida & Grey-Wilson 2012) was discovered as a new species from western Bhutan in 2012. Also a new species, Aconitum bhutanobulbilliferum (Kadota 2010) was described from Gasa, northern Bhutan. The holotype of this species is deposited at the National Herbarium THIM, NBC.

A new species of cherry tree, Prunus harae (Ohba & Akiya-ma 2010) was discovered from central Bhutan, and is known only from Bhutan at 2500-3200 masl. The epithet honours the late Professor Hiroshi Hara of the University of Tokyo, who collected this specimen and also participated in the study of the flora of Bhutan (Ohba & Akiyama 2010). A common white-beam tree, Sorbus karchungii (Rushforth 2009) was described in 2009 and named after Karchung Wangchuk for his assistance during field trips. S. karchungii is cultivated in Britain, with the seeds flown in from Trashigang. Two endemic plants, Cotoneaster bumthangensis (Fryer & Hylmo 2009), and C. thimphuensis (Fryer & Hylmo 2009) were described in 2009. Little is known of these species especially in the wild. C. bumthangensis is found only in Tang valley, Bumthang district, thus meriting this species as endemic to Tang valley. C. thimphuensis is known from a type collection made in 1994, and it ranges from Paro to Chelela Pass.

Molluscs (Mollusca)

The year 2013 was marked by the discovery and description of one of the world’s smallest snail species, from Paro, western Bhutan. Truncatellina bhutanensis (Gittenberger et al. 2013; Fig. 5) is commonly found in calcareous areas at an elevation around 2500 masl. Three more snail species were discovered and described in 2017 namely: Rahula kleini (Gittenberger,

ary Consul of Bhutan in the Netherlands. Rahula trongsaensis was discovered from Trongsa, central Bhutan at an altitude of 2210 masl. The epithet refers to Trongsa district.

Dragonflies/Damselflies (Odonata)

The major highlight in invertebrate discoveries was the discovery of a new species of damselfly (family Synlestidae) from Trongsa in 2017 at an elevation of 2450 masl. This species was named in honour of the Crown Prince of Bhutan, His Royal Highness Gyalsey Jigme Namgyel Wangchuck of Bhutan, and called the Gyalsey Emerald Spreadwing (Fig. 9), scientifically known as Megalestes gyalsey (Gyeltshen et al. 2017a; cDNA: RMNH; Holotype: Bhutan). Currently, three species of Mega lestes are known from Bhutan: M. major, M. irma and M. gyalsey.

Fishes (Pisces)

A new species of torrent catfish Parachiloglanis bhutanensis (family Sisoridae) (Fig. 6) was discovered in 2013 at an eleva-tion of 2211 masl in Trashigang, and is the first endemic fish species scientifically described from Bhutan (Thoni & Gurung 2014). In 2013 two new species, Garra bimaculacauda from central Bhutan and Garra parastenorhynchus (family Cyprini-dae) were described from southern Bhutan (Thoni et al. 2016).

Stoneflies (Plecoptera)A species of stoneflies (Plecoptera), Claassenia drukpa (family Perlidae) was described in 2010 from Punakha, Bhutan at an elevation of 1370 masl; the specimen was collected in 2006 (Stark & Sivec 2010).

Beetles (Coleoptera)

A new species of beetle, Trilophidius gemmatus (family Carabi-dae) was described from Thrumshingla National Park at an altitude of around 2500 masl. (Balkenohl 2017). The holotype is depo-sited at the NMBE (Natural History Museum Bern, Switzerland).

Fig. 5. One of the smallest snails in the world, Truncatellina bhutanen-sis. (© E. Gittenberger).

Leda & Sherub; Holotype NBC), R. trongsaensis (Fig. 8) (Gittenberger, Leda & Sherub 2017a; Holotype NBC), and Erhaia wangchu-ki (Gittenberger, Sherub & Stelbrink 2017c; Holotype NBC). E. wangchuki was dis-covered in Genekha, Thim-phu at an altitude of 2825 masl. The species epithet is after Jigme Wangchuk, who collected the specimens.

All the specimens of molluscs are deposited in the National Insect Repository at NBC, Thimphu. Rahula kleini was discovered from Mongar, Eastern Bhutan at an altitude of 2300 masl. The epithet re-fers to Cornelis Klein, Honor-

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Moths (Lepidoptera)

The new moth species Notodonta dedmazai (Schintlmeister 2013) (family Notodontidae) was described in 2013 from Nob-ding and Punakha region at an altitudinal range from 1500 to 1970 masl. Thitarodes namnai and T. caligophilus (family Hepialidae) were described in 2010 (Maczey et al. 2010). Both Thitarodes moths are hosts of Ophiocordyceps sinensis (Chinese Caterpillar Fungus). In 2009, a new species of the family Saturniidae, Salassa bhutanensis, was described (Brechlin 2009).

New Records to Bhutan

Several species have been recorded as new to the country, which may be due to most of the forests being intact and pristine with low accessibility. There have been numerous new records an-

nounced or released through local media and press releases. However, we provide a checklist only of new species records that are published in peer-reviewed publications (Table 2).

A staggering 119 moths (Lepidoptera), 108 fishes (Pisces), 101 vascular plants (Angiosperms), 65 species of bees and wasps (Hymenoptera), 59 species of dragonflies and damsel-flies (Odonata), 44 species of amphibians and reptiles, 35 spe-cies of snails and slugs (Molluscs), 14 crabs (Decapods), five cicadas, five birds, four beetles (Coleoptera), four butterflies, two mosses (Bryophytes), and one species of true fly (Diptera) were recorded new in Bhutan during the period are discussed (Table 2). A total of 566 new records for Bhutan are reported.For example, a new record of bee Megacompsomeris cochi-nensis was recorded in Bhutan in 2017 and similarly, an orchid species Bulbophyllum obrienianum was recorded as new to Bhutan in 2017 (Fig. 7 and Fig. 10).

Table 1: Summary of the species new to science described from Bhutan from 2009 to 2017.S. N. Species Name Year Reference/JournalPLANTS

1 Meconopsis gakyidiana Yoshida, Yangzom & Long 2017 Sibbaldia: The journal of Botanic Garden Horticulture, No. 14. 69-96

2 Meconopsis merakensis Yoshida, Yangzom & Long 2017 Sibbaldia: The journal of Botanic Garden Horticulture, No. 14. 69-96

3 Spathoglottis jetsuniae Gyeltshen, Tobgyel & Dalstrom 2017 Lankesteriana 17 (3) 4 Roscoea megalantha Yoshida & Yangzom 2017 Edinburgh Journal of Botany: 1-95 Meconopsis elongata Yoshida, Yangzom & Long 2016 The Plantsman, September: 177-1816 Meconopsis bhutanica Yoshida & Grey-Wilson 2012 The Plantsman, July: 96-1017 Dysphania bhutanica Sukhor 2012 Willdenowia 42 (2):169-180. https://doi.org/10.3372/

wi.42.422028 Prunus gongshanensis Wen 2012 PhytoKeys 11: 39-59 doi: 10.3897/phytokeys.11.28259 Aconitum bhutanobulbilliferum Kadota 2010 Journal of Japanese Botany 85: 8-24 10 Prunus harae Ohba & Akiyama 2010 Bulletin of the National Museum of Natural Science, Ser. B 36

(4): 133-14011 Astragalus paroensis Podlech 2010 Pakistan Journal of Botany, Special Issue 42: 27-3412 Dactylicapnos platycarpa Liden 2010 Nordic Journal of Botany 28: 656-660, doi: 10.1111/j.1756-

1051.2010.00872.x13 Cotoneaster bumthangensis Fryer & Hylmo 2009 Cotoneasters Comprehensive Guide 308 (159; pl. 89)14 Cotoneaster thimphuensis Fryer & Hylmo 2009 Cotoneasters Comprehensive Guide 308 (159; pl. 89)15 Cotoneaster hicksii Fryer & Hylmo 2009 Cotoneasters; 313, pl. 9216 Sorbus karchungii Rushforth 2009 Trees: International Dendrology Society, Year Book: 74-102

MOLLUSCS17 Rahula kleini Gittenberger, Leda & Sherub 2017 Journal of Conchology, Vol. 42, No.6 413-41818 Rahula trongsaensis Gittenberger, Leda & Sherub 2017 Journal of Conchology, Vol. 42, No.6 413-41819 Erhaia wangchuki Gittenberger, Leda & Stelbrink 2017 ZooKeys 679: 21-28 20 Truncatellina bhutanensis Gittenberger, Leda & Sherub 2013 Basteria 77 (1-3): 29-32

DRAGONFLIES21 Megalestes gyalsey Gyeltshen, Kalkman & Orr 2017 Zootaxa 4244 (4): 588-594

PLECOPTERA22 Claassenia drukpa Stark & Sivec 2010 Illiesia, 6 (24): 303-314

BEETLES23 Trilophidius gemmatus Balkenohl 2017 Alpine Entomology 1: 51-5624 Thanasimus bhutanensis Gerstmeier 2009 Biodiversität und Naturausstattung im Himalaya III: 351-358

MOTHS25 Notodonta dedmazai Schintlmeister 2013 Zootaxa 3608 (4): 283-28426 Thitarodes namnai Maczey, Dhendup, Cannon, Hywel-Jones &

Rai2010 Zootaxa 2412: 42-52

27 Thitarodes caligophilus Maczey, Dhendup, Cannon, Hywel-Jones & Rai

2010 Zootaxa 2412: 42-52

28 Salassa bhutanensis Brechlin 2009 Entomo-Satsphingia. 2 (1): 45-55FISHES

29 Garra bimaculacauda Thoni, Gurung & Mayden 2016 Zootaxa. 4169 (1): 115-13230 Garra parastenorhynchus Thoni, Gurung & Mayden 2016 Zootaxa. 4169 (1): 115-13231 Parachiloglanis bhutanensis Thoni & Gurung 2014 Zootaxa 3869 (3): 306-312

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Conclusion

From 2009-2017, 31 species were described as new to science from Bhutan, of which 16 were plants, four molluscs, four moths, three fishes, one dragonfly, two beetle, and one stonefly. Around three species are described each year from Bhutan. A total of 566 species have been recorded as new to Bhutan, despite much less effort having been applied to the exploration of the lower taxonomic groups.

Acknowledgements

We would like to thank Dr. Tashi Yangzome Dorji, Program Director, NBC, for her constant motivation and encourage-ment. We are also grateful to Matthias Hartmann (Erfurt, Ger-many) for support and encouragement to come up with this article, and Maxwell Barclay (London, UK) for the language review. Our heartfelt gratitude goes to all the data contribu-tors: Dr. Vincent J. Kalkman (Naturalis, The Netherlands), Prof. Edmund (Edi) Gittenberger (Naturalis), Mrs. Rinchen Yangzom (NBC), Dr. D. B. Gurung (CNR), Pema Leda (NBC), Stig Dalstrom (NBC), Jigme Tsheltrum Wangyel (DoFPs), Dr. Michael Thomas (University of Hawaii), Dr. Sherub Sherub (UWICER), Mani P. Nirola (NBC), Karma Wangdi (UWI-CER), Thinley Gyeltshen (Sherubste), Phurba Dorji (RSPN), Cheten Dorji (CNR), Dawa Penjor (Mushroom Centre), Dr. Karma Wangchuk (Sherubtse), Dr. David Long (UK), Dr. Cees Gielis (Naturalis), Dr. Oscar Vorst (Naturalis), Jigme Wangchuck (UWICER), Tshering Nidup (Sherubtse), Dani-elle Northey, Tshering Dorji (both NBC) and Lhab Tshering (NBC).

Fig. 6. Parachiloglanis bhutanensis, a new and endemic fish species to Bhutan. (© D. B. Gurung).

Fig. 7: Megacompsomeris cochinensis, new record for Bhutan 2017 (© Phurba Dorji).

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References

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Brechlin, R. (2009): An annoted checklist of the Saturniidae and Brahmaeidae (Lepidoptera) of the kingdom of Bhutan. Entomo-satsphingia 2 (1): 47-55.

Dalstrom, S.; C. Gyeltshen & N. Gyeltshen (2017): A century of new orchid records of Bhutan. - National Biodiversity Centre, Thimphu, Bhutan.

Dorji, P.; W. Klein & T. Nidup (2017): Taxonomic study of social ves-pid wasps (Hymenoptera: Vespidae: Vespinae & Polistinae) in Bhu-tan. - Journal of Insect Biodiversity and Systematics 3 (2): 91-104.

Fryer, J. & B. Hylmo (2009): Cotoneasters: Comprehensive Guide 308 (159; pl. 89).

Gittenberger, E.; P. Leda & S. Sherub (2013). Bhutanese snails, the smallest one: Truncatellina bhutanensis spec. nov. (Gastropoda, Pulmonata, Vertiginidae). - Basteria 77 (1-3): 29-32.

Gittenberger, E.; P. Leda & S. Sherub (2017a): Gastropods in Bhu-tan, the genus Rahula (Pulmonata: Helicarionidae). - Journal of Conchology 42, 6: 413-418.

Gittenberger, E.; P. Leda, C. Gyeltshen, S. Sherub & S. Dema (2017b): A field guide to the common molluscs of Bhutan. - Natio-nal Biodiversity Centre, Thimphu, Bhutan.

Gittenberger, E.; S. Sherub & B. Stelbrink (2017c). Erhaia Davis & Kuo (Gastropoda, Rissooidea, Amnicolidae) also in Bhutan. - ZooKeys 679: 21-28. https://doi.org/10.3897/zookeys.679.13326.

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Gyeltshen, T.; V. J. Kalkman & A. G. Orr (2017a): Honouring His Royal Highness the Crown Prince of Bhutan: Megalestes gyalsey (Odonata: Synlestidae). - Zootaxa 4244 (4): 58-594. DOI: https://doi.org/10.11646/zootaxa.4244.4.9.

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Gyeltshen, N.; K. Tobgyel & S. Dalstrom (2017): A new and striking Spathoglottis (Orchidaceae: Collabiinae), Honoring Her Majesty the Queen of Bhutan. - Lankesteriana 17 (3): 395-401. DOI: http://dx.doi.org/10.15517/lank.v17i3.31575.

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Fig. 8: Rahula trongsaensis, new to science, discovered from Trongsa (© Edi Gittenberger).

Fig. 9: Megalestes gyalsey (Gyalsey Emerald Spreadwing), new to science. (© Albert Orr).

Fig. 10: Bulbophyllum obrienianum, new record for Bhutan, discovered from Zhemgang (© Stig Dalstrom).

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Schintlmeister, A. (2013). Notodonta dedmazai sp. nov., a new noto-dontid moth from Bhutan (Lepidoptera: Notodontidae). - Zootaxa 3608 (4): 283-284. http://dx.doi.org/10.11646/zootaxa.3608.4.8

Stark, B. P. & I. Sivec (2010): Systematic notes on the genus Claas-senia Wu (Plecoptera: Perilidae), with description of a new spe-cies. - Illiesia 6 (24): 303-314. http://www2.pms-lj.si/illiesia/Illie-sia06-24.pdf.

Thoni, R. J. & D. B. Gurung (2014): Parachiloglanis bhutanensis, a new species of torrent catfish (Siluriformes: Sisoridae) from Bhu-tan. - Zootaxa 3869 (3): 306-312.

Thoni, R. J.; D. B. Gurung & R. L. Mayden (2016): A review of the genus Garra Hamilton 1822 of Bhutan, including the descriptions of two new species and three additional records (Cypriniformes: Cyprinidae). - Zootaxa 4169 (1): 115-132.

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Yoshida, T.; R. Yangzom & D. Long (2016). The opening of an area to tourist has resulted in the discovery of a new blue poppy. - The Plantsman, September 2016: 177-181.

Yoshida, T.; R. Yangzom & D. Long (2017). Dancing butterflies of the east Himalayas – New Meconopsis species from east Bhutan, Aru-nachal Pradesh and south Tibet. - Sibbaldia: The Journal of Botanic Garden Horticulture 14: 69-96.

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Authors addresses:

Choki Gyeltshen*Kezang TobgayNima GyeltshenTshering DorjiSangay Dema*National Biodiversity CentreMinistry of Agriculture and ForestsSerbithangThimphu 11001, Bhutan

corresponding addresses: [email protected] [email protected]

66


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m, S

., G

yelts

hen,

C. &

Gye

ltshe

n, N

.20

17A

cen

tury

of n

ew o

rchi

d re

cord

s of B

huta

n

4Ap

hyllo

rchi

s alp

ina

D

alst

rom

, S.,

Gye

ltshe

n, C

. & G

yelts

hen,

N.

2017

A c

entu

ry o

f new

orc

hid

reco

rds o

f Bhu

tan

5Bu

lbop

hyllu

m c

f. st

riat

um

Dal

stro

m, S

., G

yelts

hen,

C. &

Gye

ltshe

n, N

.20

17A

cen

tury

of n

ew o

rchi

d re

cord

s of B

huta

n

6Bu

lbop

hyllu

m b

ifurc

atofl

oren

s D

alst

rom

, S.,

Gye

ltshe

n, C

. & G

yelts

hen,

N.

2017

A c

entu

ry o

f new

orc

hid

reco

rds o

f Bhu

tan

7Bu

lbop

hyllu

m b

iset

um

Dal

stro

m, S

., G

yelts

hen,

C. &

Gye

ltshe

n, N

.20

17A

cen

tury

of n

ew o

rchi

d re

cord

s of B

huta

n

8Bu

lbop

hyllu

m c

ornu

-cer

vi

Dal

stro

m, S

., G

yelts

hen,

C. &

Gye

ltshe

n, N

.20

17A

cen

tury

of n

ew o

rchi

d re

cord

s of B

huta

n

9Bu

lbop

hyllu

m c

ylin

drac

eum

D

alst

rom

, S.,

Gye

ltshe

n, C

. & G

yelts

hen,

N.

2017

A c

entu

ry o

f new

orc

hid

reco

rds o

f Bhu

tan

10Bu

lbop

hyllu

m d

epre

ssum

D

alst

rom

, S.,

Gye

ltshe

n, C

. & G

yelts

hen,

N.

2017

A c

entu

ry o

f new

orc

hid

reco

rds o

f Bhu

tan

11Bu

lbop

hyllu

m h

irtu

m

Dal

stro

m, S

., G

yelts

hen,

C. &

Gye

ltshe

n, N

.20

17A

cen

tury

of n

ew o

rchi

d re

cord

s of B

huta

n

12Bu

lbop

hyllu

m in

terp

ositu

m

Dal

stro

m, S

., G

yelts

hen,

C. &

Gye

ltshe

n, N

.20

17A

cen

tury

of n

ew o

rchi

d re

cord

s of B

huta

n

13Bu

lbop

hyllu

m le

ptan

thum

D

alst

rom

, S.,

Gye

ltshe

n, C

. & G

yelts

hen,

N.

2017

A c

entu

ry o

f new

orc

hid

reco

rds o

f Bhu

tan

14Bu

lbop

hyllu

m lo

ngifl

orum

D

alst

rom

, S.,

Gye

ltshe

n, C

. & G

yelts

hen,

N.

2017

A c

entu

ry o

f new

orc

hid

reco

rds o

f Bhu

tan

15Bu

lbop

hyllu

m m

ucro

natu

m

Dal

stro

m, S

., G

yelts

hen,

C. &

Gye

ltshe

n, N

.20

17A

cen

tury

of n

ew o

rchi

d re

cord

s of B

huta

n

16Bu

lbop

hyllu

m n

ipon

dhii

Dal

stro

m, S

., G

yelts

hen,

C. &

Gye

ltshe

n, N

.20

17A

cen

tury

of n

ew o

rchi

d re

cord

s of B

huta

n

17Bu

lbop

hyllu

m o

brie

nian

um

Dal

stro

m, S

., G

yelts

hen,

C. &

Gye

ltshe

n, N

.20

17A

cen

tury

of n

ew o

rchi

d re

cord

s of B

huta

n

18Bu

lbop

hyllu

m ra

skot

ii D

alst

rom

, S.,

Gye

ltshe

n, C

. & G

yelts

hen,

N.

2017

A c

entu

ry o

f new

orc

hid

reco

rds o

f Bhu

tan

19Bu

lbop

hyllu

m ri

gidu

m

Dal

stro

m, S

., G

yelts

hen,

C. &

Gye

ltshe

n, N

.20

17A

cen

tury

of n

ew o

rchi

d re

cord

s of B

huta

n

20Bu

lbop

hyllu

m sa

rcop

hyllo

ides

D

alst

rom

, S.,

Gye

ltshe

n, C

. & G

yelts

hen,

N.

2017

A c

entu

ry o

f new

orc

hid

reco

rds o

f Bhu

tan

21Bu

lbop

hyllu

m sa

rcop

hyllu

m

Dal

stro

m, S

., G

yelts

hen,

C. &

Gye

ltshe

n, N

.20

17A

cen

tury

of n

ew o

rchi

d re

cord

s of B

huta

n

22Bu

lbop

hyllu

m st

erile

D

alst

rom

, S.,

Gye

ltshe

n, C

. & G

yelts

hen,

N.

2017

A c

entu

ry o

f new

orc

hid

reco

rds o

f Bhu

tan

23Bu

lbop

hyllu

m su

nipi

a D

alst

rom

, S.,

Gye

ltshe

n, C

. & G

yelts

hen,

N.

2017

A c

entu

ry o

f new

orc

hid

reco

rds o

f Bhu

tan

24Bu

lbop

hyllu

m v

irid

iflor

um

Dal

stro

m, S

., G

yelts

hen,

C. &

Gye

ltshe

n, N

.20

17A

cen

tury

of n

ew o

rchi

d re

cord

s of B

huta

n

25C

alan

the

davi

dii

Dal

stro

m, S

., G

yelts

hen,

C. &

Gye

ltshe

n, N

.20

17A

cen

tury

of n

ew o

rchi

d re

cord

s of B

huta

n

26C

alan

the

dens

iflor

a D

alst

rom

, S.,

Gye

ltshe

n, C

. & G

yelts

hen,

N.

2017

A c

entu

ry o

f new

orc

hid

reco

rds o

f Bhu

tan

27C

alan

the

yuks

omne

nsis

D

alst

rom

, S.,

Gye

ltshe

n, C

. & G

yelts

hen,

N.

2017

A c

entu

ry o

f new

orc

hid

reco

rds o

f Bhu

tan

28C

allo

styl

is b

ambu

sifo

liaD

alst

rom

, S.,

Gye

ltshe

n, C

. & G

yelts

hen,

N.

2017

A c

entu

ry o

f new

orc

hid

reco

rds o

f Bhu

tan

29C

allo

styl

is ri

gida

Blu

me

Dal

stro

m, S

., G

yelts

hen,

C. &

Gye

ltshe

n, N

.20

17A

cen

tury

of n

ew o

rchi

d re

cord

s of B

huta

n

30C

ham

aega

stro

dia

shik

okia

na

Dal

stro

m, S

., G

yelts

hen,

C. &

Gye

ltshe

n, N

.20

17A

cen

tury

of n

ew o

rchi

d re

cord

s of B

huta

n

67


31C

heiro

styl

is g

riffi

thii

Dal

stro

m, S

., G

yelts

hen,

C. &

Gye

ltshe

n, N

.20

17A

cen

tury

of n

ew o

rchi

d re

cord

s of B

huta

n

32C

heiro

styl

is y

unna

nens

is

Dal

stro

m, S

., G

yelts

hen,

C. &

Gye

ltshe

n, N

.20

17A

cen

tury

of n

ew o

rchi

d re

cord

s of B

huta

n

33C

hrys

oglo

ssum

orn

atum

D

alst

rom

, S.,

Gye

ltshe

n, C

. & G

yelts

hen,

N.

2017

A c

entu

ry o

f new

orc

hid

reco

rds o

f Bhu

tan

34C

leis

osto

ma

disc

olor

Dal

stro

m, S

., G

yelts

hen,

C. &

Gye

ltshe

n, N

.20

17A

cen

tury

of n

ew o

rchi

d re

cord

s of B

huta

n

35C

leis

osto

ma

filifo

rme

Dal

stro

m, S

., G

yelts

hen,

C. &

Gye

ltshe

n, N

.20

17A

cen

tury

of n

ew o

rchi

d re

cord

s of B

huta

n

36C

leis

osto

ma

sim

ondi

i D

alst

rom

, S.,

Gye

ltshe

n, C

. & G

yelts

hen,

N.

2017

A c

entu

ry o

f new

orc

hid

reco

rds o

f Bhu

tan

37C

onch

idiu

m p

usill

um

Dal

stro

m, S

., G

yelts

hen,

C. &

Gye

ltshe

n, N

.20

17A

cen

tury

of n

ew o

rchi

d re

cord

s of B

huta

n

38C

repi

dium

jose

phia

num

D

alst

rom

, S.,

Gye

ltshe

n, C

. & G

yelts

hen,

N.

2017

A c

entu

ry o

f new

orc

hid

reco

rds o

f Bhu

tan

39C

repi

dium

kha

sian

um

Dal

stro

m, S

., G

yelts

hen,

C. &

Gye

ltshe

n, N

.20

17A

cen

tury

of n

ew o

rchi

d re

cord

s of B

huta

n

40C

repi

dium

pur

pure

umD

alst

rom

, S.,

Gye

ltshe

n, C

. & G

yelts

hen,

N.

2017

A c

entu

ry o

f new

orc

hid

reco

rds o

f Bhu

tan

41C

ymbi

dium

alo

ifoliu

m

Dal

stro

m, S

., G

yelts

hen,

C. &

Gye

ltshe

n, N

.20

17A

cen

tury

of n

ew o

rchi

d re

cord

s of B

huta

n

42C

ymbi

dium

coc

hlea

re

Dal

stro

m, S

., G

yelts

hen,

C. &

Gye

ltshe

n, N

.20

17A

cen

tury

of n

ew o

rchi

d re

cord

s of B

huta

n

43C

ymbi

dium

ebu

rneu

m

Dal

stro

m, S

., G

yelts

hen,

C. &

Gye

ltshe

n, N

.20

17A

cen

tury

of n

ew o

rchi

d re

cord

s of B

huta

n

44C

ymbi

dium

mac

rorh

izon

D

alst

rom

, S.,

Gye

ltshe

n, C

. & G

yelts

hen,

N.

2017

A c

entu

ry o

f new

orc

hid

reco

rds o

f Bhu

tan

45D

endr

obiu

m je

nkin

sii

Dal

stro

m, S

., G

yelts

hen,

C. &

Gye

ltshe

n, N

.20

17A

cen

tury

of n

ew o

rchi

d re

cord

s of B

huta

n

46D

endr

obiu

m m

ontic

ola

Dal

stro

m, S

., G

yelts

hen,

C. &

Gye

ltshe

n, N

.20

17A

cen

tury

of n

ew o

rchi

d re

cord

s of B

huta

n

47D

endr

obiu

m p

orph

yroc

hilu

m

Dal

stro

m, S

., G

yelts

hen,

C. &

Gye

ltshe

n, N

.20

17A

cen

tury

of n

ew o

rchi

d re

cord

s of B

huta

n

48D

endr

obiu

m ra

bani

i D

alst

rom

, S.,

Gye

ltshe

n, C

. & G

yelts

hen,

N.

2017

A c

entu

ry o

f new

orc

hid

reco

rds o

f Bhu

tan

49D

endr

obiu

m ru

cker

i D

alst

rom

, S.,

Gye

ltshe

n, C

. & G

yelts

hen,

N.

2017

A c

entu

ry o

f new

orc

hid

reco

rds o

f Bhu

tan

50D

idym

ople

xis p

alle

ns

Dal

stro

m, S

., G

yelts

hen,

C. &

Gye

ltshe

n, N

.20

17A

cen

tury

of n

ew o

rchi

d re

cord

s of B

huta

n

51Ep

ipog

ium

japo

nicu

m

Dal

stro

m, S

., G

yelts

hen,

C. &

Gye

ltshe

n, N

.20

17A

cen

tury

of n

ew o

rchi

d re

cord

s of B

huta

n

52Ep

ipog

ium

rose

umD

alst

rom

, S.,

Gye

ltshe

n, C

. & G

yelts

hen,

N.

2017

A c

entu

ry o

f new

orc

hid

reco

rds o

f Bhu

tan

53Er

ia c

f. cl

avic

aulis

D

alst

rom

, S.,

Gye

ltshe

n, C

. & G

yelts

hen,

N.

2017

A c

entu

ry o

f new

orc

hid

reco

rds o

f Bhu

tan

54Er

ia b

huta

nica

D

alst

rom

, S.,

Gye

ltshe

n, C

. & G

yelts

hen,

N.

2017

A c

entu

ry o

f new

orc

hid

reco

rds o

f Bhu

tan

55Er

ia b

iflor

a D

alst

rom

, S.,

Gye

ltshe

n, C

. & G

yelts

hen,

N.

2017

A c

entu

ry o

f new

orc

hid

reco

rds o

f Bhu

tan

56Er

ia v

ittat

a D

alst

rom

, S.,

Gye

ltshe

n, C

. & G

yelts

hen,

N.

2017

A c

entu

ry o

f new

orc

hid

reco

rds o

f Bhu

tan

57Eu

loph

ia zo

lling

eri

Dal

stro

m, S

., G

yelts

hen,

C. &

Gye

ltshe

n, N

.20

17A

cen

tury

of n

ew o

rchi

d re

cord

s of B

huta

n

58G

astro

chilu

s cf.

linea

rifo

lius

Dal

stro

m, S

., G

yelts

hen,

C. &

Gye

ltshe

n, N

.20

17A

cen

tury

of n

ew o

rchi

d re

cord

s of B

huta

n

59G

eodo

rum

den

siflo

rum

D

alst

rom

, S.,

Gye

ltshe

n, C

. & G

yelts

hen,

N.

2017

A c

entu

ry o

f new

orc

hid

reco

rds o

f Bhu

tan

60G

oody

era

biflo

ra

Dal

stro

m, S

., G

yelts

hen,

C. &

Gye

ltshe

n, N

.20

17A

cen

tury

of n

ew o

rchi

d re

cord

s of B

huta

n

61H

aben

aria

furc

ifera

D

alst

rom

, S.,

Gye

ltshe

n, C

. & G

yelts

hen,

N.

2017

A c

entu

ry o

f new

orc

hid

reco

rds o

f Bhu

tan

62H

aben

aria

pec

tinat

a D

alst

rom

, S.,

Gye

ltshe

n, C

. & G

yelts

hen,

N.

2017

A c

entu

ry o

f new

orc

hid

reco

rds o

f Bhu

tan

63H

erm

iniu

m m

onor

chis

D

alst

rom

, S.,

Gye

ltshe

n, C

. & G

yelts

hen,

N.

2017

A c

entu

ry o

f new

orc

hid

reco

rds o

f Bhu

tan

64Li

pari

s bis

tria

ta

Dal

stro

m, S

., G

yelts

hen,

C. &

Gye

ltshe

n, N

.20

17A

cen

tury

of n

ew o

rchi

d re

cord

s of B

huta

n

65Li

pari

s defl

exa

Dal

stro

m, S

., G

yelts

hen,

C. &

Gye

ltshe

n, N

.20

17A

cen

tury

of n

ew o

rchi

d re

cord

s of B

huta

n

66Li

pari

s elli

ptic

a D

alst

rom

, S.,

Gye

ltshe

n, C

. & G

yelts

hen,

N.

2017

A c

entu

ry o

f new

orc

hid

reco

rds o

f Bhu

tan

68


67Li

pari

s fiss

ipet

ala

Dal

stro

m, S

., G

yelts

hen,

C. &

Gye

ltshe

n, N

.20

17A

cen

tury

of n

ew o

rchi

d re

cord

s of B

huta

n

68Li

pari

s pyg

mae

a D

alst

rom

, S.,

Gye

ltshe

n, C

. & G

yelts

hen,

N.

2017

A c

entu

ry o

f new

orc

hid

reco

rds o

f Bhu

tan

69Li

pari

s ros

trat

a D

alst

rom

, S.,

Gye

ltshe

n, C

. & G

yelts

hen,

N.

2017

A c

entu

ry o

f new

orc

hid

reco

rds o

f Bhu

tan

70N

eotti

a al

tern

ifolia

D

alst

rom

, S.,

Gye

ltshe

n, C

. & G

yelts

hen,

N.

2017

A c

entu

ry o

f new

orc

hid

reco

rds o

f Bhu

tan

71N

eotti

a m

icro

glot

tisD

alst

rom

, S.,

Gye

ltshe

n, C

. & G

yelts

hen,

N.

2017

A c

entu

ry o

f new

orc

hid

reco

rds o

f Bhu

tan

72N

ephe

laph

yllu

m p

ulch

rum

Dal

stro

m, S

., G

yelts

hen,

C. &

Gye

ltshe

n, N

.20

17A

cen

tury

of n

ew o

rchi

d re

cord

s of B

huta

n

73O

bero

nia

max

ima

Dal

stro

m, S

., G

yelts

hen,

C. &

Gye

ltshe

n, N

.20

17A

cen

tury

of n

ew o

rchi

d re

cord

s of B

huta

n

74O

dont

ochi

lus g

rand

iflor

usD

alst

rom

, S.,

Gye

ltshe

n, C

. & G

yelts

hen,

N.

2017

A c

entu

ry o

f new

orc

hid

reco

rds o

f Bhu

tan

75O

dont

ochi

lus p

oila

nei

Dal

stro

m, S

., G

yelts

hen,

C. &

Gye

ltshe

n, N

.20

17A

cen

tury

of n

ew o

rchi

d re

cord

s of B

huta

n

76O

toch

ilus a

lbus

D

alst

rom

, S.,

Gye

ltshe

n, C

. & G

yelts

hen,

N.

2017

A c

entu

ry o

f new

orc

hid

reco

rds o

f Bhu

tan

77Pa

nise

a yu

nnan

ensi

s D

alst

rom

, S.,

Gye

ltshe

n, C

. & G

yelts

hen,

N.

2017

A c

entu

ry o

f new

orc

hid

reco

rds o

f Bhu

tan

78Pa

phio

pedi

lum

ven

ustu

m

Dal

stro

m, S

., G

yelts

hen,

C. &

Gye

ltshe

n, N

.20

17A

cen

tury

of n

ew o

rchi

d re

cord

s of B

huta

n

79Pe

rist

ylus

bie

rman

nian

usD

alst

rom

, S.,

Gye

ltshe

n, C

. & G

yelts

hen,

N.

2017

A c

entu

ry o

f new

orc

hid

reco

rds o

f Bhu

tan

80Ph

aius

wal

lichi

i D

alst

rom

, S.,

Gye

ltshe

n, C

. & G

yelts

hen,

N.

2017

A c

entu

ry o

f new

orc

hid

reco

rds o

f Bhu

tan

81Ph

alae

nops

is lo

bbii

Dal

stro

m, S

., G

yelts

hen,

C. &

Gye

ltshe

n, N

.20

17A

cen

tury

of n

ew o

rchi

d re

cord

s of B

huta

n

82Ph

reat

ia e

lega

ns

Dal

stro

m, S

., G

yelts

hen,

C. &

Gye

ltshe

n, N

.20

17A

cen

tury

of n

ew o

rchi

d re

cord

s of B

huta

n

83Pl

atan

ther

a al

bom

argi

nata

D

alst

rom

, S.,

Gye

ltshe

n, C

. & G

yelts

hen,

N.

2017

A c

entu

ry o

f new

orc

hid

reco

rds o

f Bhu

tan

84Pl

atan

ther

a dy

eria

na

Dal

stro

m, S

., G

yelts

hen,

C. &

Gye

ltshe

n, N

.20

17A

cen

tury

of n

ew o

rchi

d re

cord

s of B

huta

n

85Pl

atan

ther

a si

kkim

ensi

sD

alst

rom

, S.,

Gye

ltshe

n, C

. & G

yelts

hen,

N.

2017

A c

entu

ry o

f new

orc

hid

reco

rds o

f Bhu

tan

86Pt

eroc

eras

tere

s D

alst

rom

, S.,

Gye

ltshe

n, C

. & G

yelts

hen,

N.

2017

A c

entu

ry o

f new

orc

hid

reco

rds o

f Bhu

tan

87Rh

ombo

da a

runa

chal

ensi

s D

alst

rom

, S.,

Gye

ltshe

n, C

. & G

yelts

hen,

N.

2017

A c

entu

ry o

f new

orc

hid

reco

rds o

f Bhu

tan

88Rh

ombo

da la

nceo

lata

D

alst

rom

, S.,

Gye

ltshe

n, C

. & G

yelts

hen,

N.

2017

A c

entu

ry o

f new

orc

hid

reco

rds o

f Bhu

tan

89So

rbus

hed

lund

iiSc

hnei

der,

C. K

.20

09Tr

ees:

Inte

rnat

iona

l Den

drol

ogy

Soci

ety

90Sp

iran

thes

cf.

hong

kong

ensi

sD

alst

rom

, S.,

Gye

ltshe

n, C

. & G

yelts

hen,

N.

2017

A c

entu

ry o

f new

orc

hid

reco

rds o

f Bhu

tan

91St

ereo

chilu

s hir

tus

Dal

stro

m, S

., G

yelts

hen,

C. &

Gye

ltshe

n, N

.20

17A

cen

tury

of n

ew o

rchi

d re

cord

s of B

huta

n

92Ta

inia

min

or

Dal

stro

m, S

., G

yelts

hen,

C. &

Gye

ltshe

n, N

.20

17A

cen

tury

of n

ew o

rchi

d re

cord

s of B

huta

n

93Ta

inia

pen

angi

ana

Dal

stro

m, S

., G

yelts

hen,

C. &

Gye

ltshe

n, N

.20

17A

cen

tury

of n

ew o

rchi

d re

cord

s of B

huta

n

94Th

elas

is p

ygm

aea

Dal

stro

m, S

., G

yelts

hen,

C. &

Gye

ltshe

n, N

.20

17A

cen

tury

of n

ew o

rchi

d re

cord

s of B

huta

n

95Th

rixs

perm

um ja

poni

cum

D

alst

rom

, S.,

Gye

ltshe

n, C

. & G

yelts

hen,

N.

2017

A c

entu

ry o

f new

orc

hid

reco

rds o

f Bhu

tan

96Th

rixs

perm

um m

usci

floru

m

Dal

stro

m, S

., G

yelts

hen,

C. &

Gye

ltshe

n, N

.20

17A

cen

tury

of n

ew o

rchi

d re

cord

s of B

huta

n

97Tr

icho

tosi

a da

syph

ylla

Dal

stro

m, S

., G

yelts

hen,

C. &

Gye

ltshe

n, N

.20

17A

cen

tury

of n

ew o

rchi

d re

cord

s of B

huta

n

98Tr

icho

tosi

a pu

lvin

ata

Dal

stro

m, S

., G

yelts

hen,

C. &

Gye

ltshe

n, N

.20

17A

cen

tury

of n

ew o

rchi

d re

cord

s of B

huta

n

99Tr

opid

ia n

amas

iae

Dal

stro

m, S

., G

yelts

hen,

C. &

Gye

ltshe

n, N

.20

17A

cen

tury

of n

ew o

rchi

d re

cord

s of B

huta

n

100

Unc

ifera

acu

min

ata

Dal

stro

m, S

., G

yelts

hen,

C. &

Gye

ltshe

n, N

.20

17A

cen

tury

of n

ew o

rchi

d re

cord

s of B

huta

n

101

Zeux

ine

affini

s D

alst

rom

, S.,

Gye

ltshe

n, C

. & G

yelts

hen,

N.

2017

A c

entu

ry o

f new

orc

hid

reco

rds o

f Bhu

tan

Mol

lusc

a (S

nails

and

Slu

gs)

69


S.N

o.Sp

ecie

sSc

ient

ist (

s)Ye

arR

efer

ence

1Al

lope

as c

lavu

linum

Gitt

enbe

rger

, E.,

Leda

, P.,

Gye

ltshe

n, C

., Sh

erub

, S. &

Dem

a, S

.20

17A

fiel

d gu

ide

to th

e co

mm

on m

ollu

scs o

f Bhu

tan

2Ar

ioph

anta

inte

rrup

taG

itten

berg

er, E

., Le

da, P

., G

yelts

hen,

C.,

Sher

ub, S

. & D

ema,

S.

2017

A fi

eld

guid

e to

the

com

mon

mol

lusc

s of B

huta

n

3Be

llam

ya (F

ilopa

ludi

na) b

enga

lens

isG

itten

berg

er, E

., Le

da, P

., W

angd

i, K

. & S

heru

b, S

.20

17B

iodi

vers

ity Jo

urna

l 8 (4

)

4Be

nson

ella

plic

iden

sG

itten

berg

er, E

., Le

da, P

., G

yelts

hen,

C.,

Sher

ub, S

. & D

ema,

S.

2017

A fi

eld

guid

e to

the

com

mon

mol

lusc

s of B

huta

n

5Br

adyb

aena

radi

cico

laG

itten

berg

er, E

., Le

da, P

., G

yelts

hen,

C.,

Sher

ub, S

. & D

ema,

S.

2017

A fi

eld

guid

e to

the

com

mon

mol

lusc

s of B

huta

n

6Br

otia

cos

tula

Gitt

enbe

rger

, E.,

Leda

, P.,

Wan

gdi,

K. &

She

rub,

S.

2017

Bio

dive

rsity

Jour

nal 8

(4)

7C

aryc

hium

indi

cum

Gitt

enbe

rger

, E.,

Leda

, P.,

Win

ter,

A.J.

& H

ochu

m, A

.20

17B

aste

ria 8

1 (4

-6):

107-

110

8C

urve

lla k

hasi

ana

Gitt

enbe

rger

, E.,

Leda

, P.,

Gye

ltshe

n, C

., Sh

erub

, S. &

Dem

a, S

.20

17A

fiel

d gu

ide

to th

e co

mm

on m

ollu

scs o

f Bhu

tan

9C

yclo

phor

us a

uror

aG

itten

berg

er, E

., Le

da, P

., G

yelts

hen,

C.,

Sher

ub, S

. & D

ema,

S.

2017

A fi

eld

guid

e to

the

com

mon

mol

lusc

s of B

huta

n

10C

yclo

phor

us p

ears

oni

Gitt

enbe

rger

, E.,

Leda

, P.,

Gye

ltshe

n, C

., Sh

erub

, S. &

Dem

a, S

.20

17A

fiel

d gu

ide

to th

e co

mm

on m

ollu

scs o

f Bhu

tan

11G

alba

trun

catu

laG

itten

berg

er, E

., Le

da, P

., W

angd

i, K

. & S

heru

b, S

.20

17B

iodi

vers

ity Jo

urna

l 8 (4

)

12G

astro

copt

a hu

ttoni

ana

Gitt

enbe

rger

, E.,

Leda

, P.,

Gye

ltshe

n, C

., Sh

erub

, S. &

Dem

a, S

.20

17A

fiel

d gu

ide

to th

e co

mm

on m

ollu

scs o

f Bhu

tan

13G

ulel

la b

icol

orG

itten

berg

er, E

., Le

da, P

., G

yelts

hen,

C.,

Sher

ub, S

. & D

ema,

S.

2017

A fi

eld

guid

e to

the

com

mon

mol

lusc

s of B

huta

n

14G

yrau

lus r

otul

aG

itten

berg

er, E

., Le

da, P

., W

angd

i, K

. & S

heru

b, S

.20

17B

iodi

vers

ity Jo

urna

l 8 (4

)

15G

yrau

lus s

ival

ensi

sG

itten

berg

er, E

., Le

da, P

., W

angd

i, K

. & S

heru

b, S

.20

17B

iodi

vers

ity Jo

urna

l 8 (4

)

16K

alie

lla b

arra

kpor

ensi

sG

itten

berg

er, E

., Le

da, P

., G

yelts

hen,

C.,

Sher

ub, S

. & D

ema,

S.

2017

A fi

eld

guid

e to

the

com

mon

mol

lusc

s of B

huta

n

17La

mel

liden

s jen

kins

ianu

sG

itten

berg

er, E

., Le

da, P

., G

yelts

hen,

C.,

Sher

ub, S

. & D

ema,

S.

2017

A fi

eld

guid

e to

the

com

mon

mol

lusc

s of B

huta

n

18La

ndou

ria

hutto

niG

itten

berg

er, E

., Le

da, P

., G

yelts

hen,

C.,

Sher

ub, S

. & D

ema,

S.

2017

A fi

eld

guid

e to

the

com

mon

mol

lusc

s of B

huta

n

19Li

ssac

hatin

a fu

lica

Gitt

enbe

rger

, E.,

Leda

, P.,

Gye

ltshe

n, C

., Sh

erub

, S. &

Dem

a, S

.20

17A

fiel

d gu

ide

to th

e co

mm

on m

ollu

scs o

f Bhu

tan

20M

acro

chla

mys

indi

caG

itten

berg

er, E

., Le

da, P

., G

yelts

hen,

C.,

Sher

ub, S

. & D

ema,

S.

2017

A fi

eld

guid

e to

the

com

mon

mol

lusc

s of B

huta

n

21M

elan

oide

s tub

ercu

lata

Gitt

enbe

rger

, E.,

Leda

, P.,

Wan

gdi,

K. &

She

rub,

S.

2017

Bio

dive

rsity

Jour

nal 8

(4)

22M

irus

nila

giri

cus

Gitt

enbe

rger

, E.,

Leda

, P.,

Gye

ltshe

n, C

., Sh

erub

, S. &

Dem

a, S

.20

17A

fiel

d gu

ide

to th

e co

mm

on m

ollu

scs o

f Bhu

tan

23Pa

ludo

mus

con

ica

Gitt

enbe

rger

, E.,

Leda

, P.,

Wan

gdi,

K. &

She

rub,

S.

2017

Bio

dive

rsity

Jour

nal 8

(4)

24Pe

arso

nia

naga

ensi

sG

itten

berg

er, E

., Le

da, P

., G

yelts

hen,

C.,

Sher

ub, S

. & D

ema,

S.

2017

A fi

eld

guid

e to

the

com

mon

mol

lusc

s of B

huta

n

25Pu

pilla

turc

men

ica

Gitt

enbe

rger

, E.,

Leda

, P.,

Gye

ltshe

n, C

., Sh

erub

, S. &

Dem

a, S

.20

17A

fiel

d gu

ide

to th

e co

mm

on m

ollu

scs o

f Bhu

tan

26Ra

dix

acum

inat

aG

itten

berg

er, E

., Le

da, P

., W

angd

i, K

. & S

heru

b, S

.20

17B

iodi

vers

ity Jo

urna

l 8 (4

)

27Ra

dix

ande

rson

iana

Gitt

enbe

rger

, E.,

Leda

, P.,

Wan

gdi,

K. &

She

rub,

S.

2017

Bio

dive

rsity

Jour

nal 8

(4)

28Ra

hula

bas

caud

aG

itten

berg

er, E

., Te

nzin

, U. &

She

rub,

S.

2017

Bas

teria

81

(4-6

): 11

9-12

2

29Sc

hist

olom

a fu

nicu

lalu

mG

itten

berg

er, E

., Le

da, P

., G

yelts

hen,

C.,

Sher

ub, S

. & D

ema,

S.

2017

A fi

eld

guid

e to

the

com

mon

mol

lusc

s of B

huta

n

30St

rept

aulu

s bla

nfor

diG

itten

berg

er, E

., Le

da, P

., G

yelts

hen,

C.,

Sher

ub, S

. & D

ema,

S.

2017

A fi

eld

guid

e to

the

com

mon

mol

lusc

s of B

huta

n

31Th

iara

(Tar

ebia

) gra

nife

raG

itten

berg

er, E

., Le

da, P

., W

angd

i, K

. & S

heru

b, S

.20

17B

iodi

vers

ity Jo

urna

l 8 (4

)

32Th

iara

(Tar

ebia

) lin

eata

Gitt

enbe

rger

, E.,

Leda

, P.,

Wan

gdi,

K. &

She

rub,

S.

2017

Bio

dive

rsity

Jour

nal 8

(4)

33Th

iara

(Thi

ara)

scab

raG

itten

berg

er, E

., Le

da, P

., W

angd

i, K

. & S

heru

b, S

.20

17B

iodi

vers

ity Jo

urna

l 8 (4

)

34Tr

achi

a de

libra

taG

itten

berg

er, E

., Le

da, P

., G

yelts

hen,

C.,

Sher

ub, S

. & D

ema,

S.

2017

A fi

eld

guid

e to

the

com

mon

mol

lusc

s of B

huta

n

35Va

lloni

a co

stoh

imal

aG

itten

berg

er, E

., Le

da, P

., G

yelts

hen,

C.,

Sher

ub, S

. & D

ema,

S.

2017

A fi

eld

guid

e to

the

com

mon

mol

lusc

s of B

huta

n

70


Dec

apod

a (C

rabs

)

S.N

o.Sp

ecie

sSc

ient

ist (

s)Ye

arR

efer

ence

1Ac

anth

opot

amon

fung

osum

Dor

ji, K

.20

14C

olle

ge o

f Nat

ural

Res

ourc

es, L

obes

a

2Ac

anth

opot

amon

mar

tens

iD

orji,

K.

2014

Col

lege

of N

atur

al R

esou

rces

, Lob

esa

3Ac

anth

opot

amon

pan

ning

iD

orji,

K.

2014

Col

lege

of N

atur

al R

esou

rces

, Lob

esa

4Al

com

on lo

phoc

arpu

sD

orji,

K.

2014

Col

lege

of N

atur

al R

esou

rces

, Lob

esa

5Al

com

on su

perc

ilios

umD

orji,

K.

2014

Col

lege

of N

atur

al R

esou

rces

, Lob

esa

6Ba

ryth

elph

usa

cuni

cula

ris

Dor

ji, K

.20

14C

olle

ge o

f Nat

ural

Res

ourc

es, L

obes

a

7Ba

ryth

elph

usa

lugu

bris

Dor

ji, K

.20

14C

olle

ge o

f Nat

ural

Res

ourc

es, L

obes

a

8H

imal

ayap

otam

on a

tkin

soni

anum

Dor

ji, K

.20

14C

olle

ge o

f Nat

ural

Res

ourc

es, L

obes

a

9H

imal

ayap

otam

on e

mph

yset

umSa

mdr

up, T

.20

17C

olle

ge o

f Nat

ural

Res

ourc

es, L

obes

a

10Li

otel

phus

a qu

adra

taSa

mdr

up, T

.20

18C

olle

ge o

f Nat

ural

Res

ourc

es, L

obes

a

11Li

othe

lphu

sa la

evis

Dor

ji, K

.20

14C

olle

ge o

f Nat

ural

Res

ourc

es, L

obes

a

12M

acro

brac

hium

ass

amen

seD

orji,

K.

2014

Col

lege

of N

atur

al R

esou

rces

, Lob

esa

13M

ayde

lliat

help

husa

lugu

bris

Dor

ji, K

.20

14C

olle

ge o

f Nat

ural

Res

ourc

es, L

obes

a

14Po

tam

iscu

s sik

kim

ense

Dor

ji, K

.20

14C

olle

ge o

f Nat

ural

Res

ourc

es, L

obes

a

Odo

nata

(Dra

gonfl

ies a

nd D

amse

lflie

s)

S.N

o.Sp

ecie

sSc

ient

ist (

s)Ye

arR

efer

ence

1Ac

iagr

ion

palli

dum

Kal

kman

, V. J

. & G

yelts

hen,

T.

2016

Jour

nal o

f the

Inte

rnat

iona

l Dra

gonfl

y Fu

nd, 9

4

2Ae

shna

shen

nong

Gye

ltshe

n, T

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alkm

an, V

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2017

Not

ulae

odo

nato

logi

cae

8 (9

): 31

9-37

4

3Ag

rioc

nem

is c

laus

eni

Mitr

a, A

., D

em.,

Gye

ltshe

n, K

., D

orji,

L.,

Puri,

N.K

., Ts

herin

g, P

., W

angd

i, P.

, Ach

arya

, P.,

Nam

gyel

, R.,

Dor

ji, S

., Ph

unts

ho, S

. & L

hade

n20

14Jo

urna

l of E

ntom

olog

y an

d Zo

olog

y St

udie

s 2 (2

): 11

-15

4Ag

rioc

nem

is fe

min

aM

itra,

A.,

Cho

den,

K.,

Dor

ji, Y

., Pe

njor

, T.,

Dor

ji, R

., Su

bedi

, K. &

Dor

ji, P

.20

12B

huta

n Jo

urna

l of R

esea

rch

& D

evel

opm

ent,

Aut

umn

5An

acia

esch

na ja

spid

eaM

itra,

A.,

Cho

den,

K.,

Dor

ji, Y

., Pe

njor

, T.,

Dor

ji, R

., Su

bedi

, K. &

Dor

ji, P

.20

12B

huta

n Jo

urna

l of R

esea

rch

& D

evel

opm

ent,

Aut

umn

6An

ax g

utta

tus

Gye

ltshe

n, T

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alkm

an, V

. J.

2017

Not

ulae

odo

nato

logi

cae

8 (9

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4

7An

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dicu

sM

itra,

A.,

Dem

., G

yelts

hen,

K.,

Dor

ji, L

., Pu

ri, N

.K.,

Tshe

ring,

P.,

Wan

gdi,

P., A

char

ya, P

., N

amgy

el, R

., D

orji,

S.,

Phun

tsho

. S. &

Lha

den

2014

Jour

nal o

f Ent

omol

ogy

and

Zool

ogy

Stud

ies 2

(2):

11-1

5

8An

isog

omph

us b

ivitt

atus

Mitr

a, A

.20

13A

dvan

ces i

n En

viro

nmen

tal S

cien

ce. O

xfor

d B

ook

Com

pany

, Ja

ipur

, vi+

272p

p

9An

isog

omph

us o

ccip

italis

Gye

ltshe

n, T

.20

17N

otul

ae o

dona

tolo

gica

e 8

(9):

319-

374

10An

isop

leur

a le

stoi

des

Kal

kman

, V. J

. & G

yelts

hen,

T.

2016

Jour

nal o

f the

Inte

rnat

iona

l Dra

gonfl

y Fu

nd 9

4

11Ar

gioc

nem

is ru

besc

ens r

ubeo

laM

itra,

A.,

Cho

den,

K.,

Dor

ji, Y

., Pe

njor

, T.,

Dor

ji, R

., Su

bedi

, K. &

Dor

ji, P

.20

12B

huta

n Jo

urna

l of R

esea

rch

& D

evel

opm

ent,

Aut

umn

12Ar

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cyph

a (R

hino

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a) c

unea

taD

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2014

Jour

nal o

f Ent

omol

ogy

and

Zool

ogy

Stud

ies 2

(4):

51-5

7

13As

iago

mph

us o

done

liG

yelts

hen,

T. &

Kal

kman

, V. J

.20

17N

otul

ae o

dona

tolo

gica

e 8

(9):

319-

374

14Br

achy

dipl

ax so

brin

aM

itra,

A.,

Dem

., G

yelts

hen,

K.,

Dor

ji, L

., Pu

ri, N

.K.,

Tshe

ring,

P.,

Wan

gdi,

P., A

char

ya, P

., N

amgy

el, R

., D

orji,

S.,

Phun

tsho

. S. &

Lha

den

2014

Jour

nal o

f Ent

omol

ogy

and

Zool

ogy

Stud

ies 2

(2):

11-1

5

15C

alic

nem

ia m

inia

taM

itra,

A.,

Cho

den,

K.,

Dor

ji, Y

., Pe

njor

, T.,

Dor

ji, R

., Su

bedi

, K. &

Dor

ji, P

.20

12B

huta

n Jo

urna

l of R

esea

rch

& D

evel

opm

ent,

Aut

umn

71


16C

amac

inia

gig

ante

aG

yelts

hen,

T.,

Nid

up, T

., D

orji,

P.,

Dor

ji, T

. & K

alkm

an, V

. J.

2017

Jour

nal o

f Thr

eate

ned

Taxa

9 (1

): 97

43-9

747

17C

eria

grio

n co

rom

ande

lianu

mM

itra,

A.,

Cho

den,

K.,

Dor

ji, Y

., Pe

njor

, T.,

Dor

ji, R

., Su

bedi

, K. &

Dor

ji, P

.20

12B

huta

n Jo

urna

l of R

esea

rch

& D

evel

opm

ent,

Aut

umn

18C

eria

grio

n ru

biae

Mitr

a, A

., C

hode

n, K

., D

orji,

Y.,

Penj

or, T

., D

orji,

R.,

Sube

di, K

. & D

orji,

P.

2012

Bhu

tan

Jour

nal o

f Res

earc

h &

Dev

elop

men

t, A

utum

n

19C

hlor

ogom

phus

mor

toni

Mitr

a, A

., C

hode

n, K

., D

orji,

Y.,

Penj

or, T

., D

orji,

R.,

Sube

di, K

. & D

orji,

P.

2012

Bhu

tan

Jour

nal o

f Res

earc

h &

Dev

elop

men

t, A

utum

n

20C

hlor

ogom

phus

pre

cios

usG

yelts

hen,

T.

2017

Not

ulae

odo

nato

logi

cae

8 (9

): 31

9-37

4

21C

oelic

cia

svih

leri

Mitr

a, A

., C

hode

n, K

., D

orji,

Y.,

Penj

or, T

., D

orji,

R.,

Sube

di, K

. & D

orji,

P.

2012

Bhu

tan

Jour

nal o

f Res

earc

h &

Dev

elop

men

t, A

utum

n

22C

oper

a vi

ttata

ass

amen

sis

Mitr

a, A

., C

hode

n, K

., D

orji,

Y.,

Penj

or, T

., D

orji,

R.,

Sube

di, K

. & D

orji,

P.

2012

Bhu

tan

Jour

nal o

f Res

earc

h &

Dev

elop

men

t, A

utum

n

23C

ratil

la li

neat

aM

itra,

A.,

Cho

den,

K.,

Dor

ji, Y

., Pe

njor

, T.,

Dor

ji, R

., Su

bedi

, K. &

Dor

ji, P

.20

12B

huta

n Jo

urna

l of R

esea

rch

& D

evel

opm

ent,

Aut

umn

24C

roco

them

is e

ryth

raea

Kal

kman

, V. J

. & G

yelts

hen,

T.

2016

Jour

nal o

f the

Inte

rnat

iona

l Dra

gonfl

y Fu

nd 9

4

25D

avid

ius z

allo

rens

isG

yelts

hen,

T.

2017

Not

ulae

odo

nato

logi

cae

8 (9

): 31

9-37

4

26D

ysph

aea

glor

iosa

Mitr

a, A

., C

hode

n, K

., D

orji,

Y.,

Penj

or, T

., D

orji,

R.,

Sube

di, K

. & D

orji,

P.

2012

Bhu

tan

Jour

nal o

f Res

earc

h &

Dev

elop

men

t, A

utum

n

27En

alla

gma

parv

umM

itra,

A.,

Cho

den,

K.,

Dor

ji, Y

., Pe

njor

, T.,

Dor

ji, R

., Su

bedi

, K. &

Dor

ji, P

.20

12B

huta

n Jo

urna

l of R

esea

rch

& D

evel

opm

ent,

Aut

umn

28Ep

ioph

lebi

a la

idla

wi

Bro

ckha

us, T

. & H

artm

ann,

A.

2009

Odo

nato

logi

ca 3

8 (3

): 20

3-21

5

29Eu

phae

a oc

hrac

ea b

runn

eaM

itra,

A.,

Cho

den,

K.,

Dor

ji, Y

., Pe

njor

, T.,

Dor

ji, R

., Su

bedi

, K. &

Dor

ji, P

.20

12B

huta

n Jo

urna

l of R

esea

rch

& D

evel

opm

ent,

Aut

umn

30G

ynac

anth

a in

cisu

raK

alkm

an, V

. J. &

Gye

ltshe

n, T

.20

16Jo

urna

l of t

he In

tern

atio

nal D

rago

nfly

Fund

,94

31G

ynac

anth

a kh

asia

caK

alkm

an, V

. J. &

Gye

ltshe

n, T

.20

16Jo

urna

l of t

he In

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atio

nal D

rago

nfly

Fund

94

32G

ynac

anth

a su

bint

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ltshe

n, T

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2017

Not

ulae

odo

nato

logi

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8 (9

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9-37

4

33G

ynac

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aesc

hna

sikk

ima

Kal

kman

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yelts

hen,

T.

2016

Jour

nal o

f the

Inte

rnat

iona

l Dra

gonfl

y Fu

nd, 9

4

34H

imal

agri

on e

xcla

mat

ione

Mitr

a, A

., D

em.,

Gye

ltshe

n, K

., D

orji,

L.,

Puri,

N.K

., Ts

herin

g, P

., W

angd

i, P.

, Ach

arya

, P.,

Nam

gyel

, R.,

Dor

ji, S

., Ph

unts

ho. S

. & L

hade

n20

14Jo

urna

l of E

ntom

olog

y an

d Zo

olog

y St

udie

s 2 (2

): 11

-15

35H

uoso

ma

tinct

ipen

neM

itra,

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2013

Adv

ance

s in

Envi

ronm

enta

l Sci

ence

. Oxf

ord

Boo

k C

ompa

ny,

Jaip

ur, v

i+27

2pp

36La

mel

ligom

phus

risi

Kal

kman

, V. J

. & G

yelts

hen,

T.

2016

Jour

nal o

f the

Inte

rnat

iona

l Dra

gonfl

y Fu

nd 9

4

37Le

stes

pra

emor

sus

Mitr

a, A

., D

em.,

Gye

ltshe

n, K

., D

orji,

L.,

Puri,

N.K

., Ts

herin

g, P

., W

angd

i, P.

, Ach

arya

, P.,

Nam

gyel

, R.,

Dor

ji, S

., Ph

unts

ho. S

. & L

hade

n20

14Jo

urna

l of E

ntom

olog

y an

d Zo

olog

y St

udie

s 2 (2

): 11

-15

38Le

stes

thor

acic

usM

itra,

A.,

Cho

den,

K.,

Dor

ji, Y

., Pe

njor

, T.,

Dor

ji, R

., Su

bedi

, K. &

Dor

ji, P

.20

12B

huta

n Jo

urna

l of R

esea

rch

& D

evel

opm

ent,

Aut

umn

39Li

bella

go li

neat

aG

yelts

hen,

T.,

Nid

up, T

., D

orji,

P.,

Dor

ji, T

. & K

alkm

an, V

. J.

2017

Jour

nal o

f Thr

eate

ned

Taxa

9 (1

): 97

43-9

747

40Ly

riot

hem

is b

ivitt

ata

Gye

ltshe

n, T

.20

17N

otul

ae o

dona

tolo

gica

e 8

(9):

319-

374

41M

egal

este

s irm

aK

alkm

an, V

. J. &

Gye

ltshe

n, T

.20

16Jo

urna

l of t

he In

tern

atio

nal D

rago

nfly

Fund

94

42N

eallo

gast

er h

erm

iona

eG

yelts

hen,

T. &

Kal

kman

, V. J

.20

17N

otul

ae o

dona

tolo

gica

e 8

(9):

319-

374

43N

euro

them

is in

term

edia

ata

lant

aM

itra,

A.,

Dem

., G

yelts

hen,

K.,

Dor

ji, L

., Pu

ri, N

.K.,

Tshe

ring,

P.,

Wan

gdi,

P., A

char

ya, P

., N

amgy

el, R

., D

orji,

S.,

Phun

tsho

. S. &

Lha

den

2014

Jour

nal o

f Ent

omol

ogy

and

Zool

ogy

Stud

ies 2

(2):

11-1

5

44Pa

rago

mph

us li

neat

usM

itra,

A.,

Cho

den,

K.,

Dor

ji, Y

., Pe

njor

, T.,

Dor

ji, R

., Su

bedi

, K. &

Dor

ji, P

.20

12B

huta

n Jo

urna

l of R

esea

rch

& D

evel

opm

ent,

Aut

umn

45Pe

riae

schn

a m

agda

lena

Mitr

a, A

.20

13A

dvan

ces i

n En

viro

nmen

tal S

cien

ce. O

xfor

d B

ook

Com

pany

, Ja

ipur

, vi+

272p

p

46Pe

riss

ogom

phus

stev

ensi

Mitr

a, A

., D

em.,

Gye

ltshe

n, K

., D

orji,

L.,

Puri,

N.K

., Ts

herin

g, P

., W

angd

i, P.

, Ach

arya

, P.,

Nam

gyel

, R.,

Dor

ji, S

., Ph

unts

ho. S

. & L

hade

n20

14Jo

urna

l of E

ntom

olog

y an

d Zo

olog

y St

udie

s 2 (2

): 11

-15

47Ph

iloga

nga

mon

tana

Gye

ltshe

n, T

.20

17N

otul

ae o

dona

tolo

gica

e 8

(9):

319-

374

72


48Po

lyca

ntha

gyna

ery

thro

mel

asM

itra,

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2013

Adv

ance

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Envi

ronm

enta

l Sci

ence

. Oxf

ord

Boo

k C

ompa

ny,

Jaip

ur, v

i+27

2pp

49Po

tam

arch

a co

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erG

yelts

hen,

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2017

Not

ulae

odo

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logi

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8 (9

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4

50Pr

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alai

caM

itra,

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den,

K.,

Dor

ji, Y

., Pe

njor

, T.,

Dor

ji, R

., Su

bedi

, K. &

Dor

ji, P

.20

12B

huta

n Jo

urna

l of R

esea

rch

& D

evel

opm

ent,

Aut

umn

51Rh

inoc

ypha

qua

drim

acul

ata

Mitr

a, A

., C

hode

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., D

orji,

Y.,

Penj

or, T

., D

orji,

R.,

Sube

di, K

. & D

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Bhu

tan

Jour

nal o

f Res

earc

h &

Dev

elop

men

t, A

utum

n

52Rh

yoth

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Gye

ltshe

n, T

. & K

alkm

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2017

Not

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odo

nato

logi

cae

8 (9

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4

53So

mat

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ora

davi

esi

Kal

kman

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. & G

yelts

hen,

T.

2016

Jour

nal o

f the

Inte

rnat

iona

l Dra

gonfl

y Fu

nd 9

4

54St

ylog

omph

us in

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iG

yelts

hen,

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2017

Not

ulae

odo

nato

logi

cae

8 (9

): 31

9-37

4

55Sy

mpe

trum

fons

colo

mbi

iK

alkm

an, V

. J. &

Gye

ltshe

n, T

.20

16Jo

urna

l of t

he In

tern

atio

nal D

rago

nfly

Fund

94

56Th

olym

is ti

llarg

aK

alkm

an, V

. J. &

Gye

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n, T

.20

16Jo

urna

l of t

he In

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atio

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nfly

Fund

94

57Tr

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ilari

sM

itra,

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., G

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hen,

K.,

Dor

ji, L

., Pu

ri, N

.K.,

Tshe

ring,

P.,

Wan

gdi,

P., A

char

ya, P

., N

amgy

el, R

., D

orji,

S.,

Phun

tsho

. S. &

Lha

den

2014

Jour

nal o

f Ent

omol

ogy

and

Zool

ogy

Stud

ies 2

(2):

11-1

5

58Ve

stal

is g

raci

lis g

raci

lisM

itra,

A.,

Cho

den,

K.,

Dor

ji, Y

., Pe

njor

, T.,

Dor

ji, R

., Su

bedi

, K. &

Dor

ji, P

.20

12B

huta

n Jo

urna

l of R

esea

rch

& D

evel

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ent,

Aut

umn

59Zy

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yelts

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2017

Not

ulae

odo

nato

logi

cae

8 (9

): 31

9-37

4

Cic

ada

S.N

o.Sp

ecie

sSc

ient

ist (

s)Ye

arR

efer

ence

1H

aphs

a su

laiy

aiD

orji,

P.

2016

Jour

nal o

f Ent

omol

ogy

and

Zool

ogy

Stud

ies 4

(4):

381-

383

2H

uesc

hys s

angu

inea

Dor

ji, P

.20

16Jo

urna

l of E

ntom

olog

y an

d Zo

olog

y St

udie

s 4 (4

): 38

1-38

3

3Po

lyne

ura

duca

lisD

orji,

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2016

Jour

nal o

f Ent

omol

ogy

and

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ogy

Stud

ies 4

(4):

381-

383

4Ta

lain

ga b

ingh

ami

Dor

ji, P

.20

16Jo

urna

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d Zo

olog

y St

udie

s 4 (4

): 38

1-38

3

5Ta

nna

thal

iaD

orji,

P.

2016

Jour

nal o

f Ent

omol

ogy

and

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ogy

Stud

ies 4

(4):

381-

383

Hym

enop

tera

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ps)

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ar

Ref

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amps

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iella

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ulim

eris

) ann

ula-

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ata

Nid

up, T

., K

lein

, W.,

Kum

ar, P

. G. &

Dor

ji, P

.20

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alte

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: 9-1

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psom

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up, T

., K

lein

, W.,

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ar, P

. G. &

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ji, P

.20

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con

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umN

idup

, T.,

Gye

ltshe

n, T

., K

umar

, P. G

., K

lein

, W. &

Dor

ji, P

.20

16Jo

urna

l of T

hrea

tene

d Ta

xa 8

(12)

: 946

7-94

72

4D

elta

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rien

sN

idup

, T.,

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ltshe

n, T

., K

umar

, P. G

., K

lein

, W. &

Dor

ji, P

.20

16Jo

urna

l of T

hrea

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xa 8

(12)

: 946

7-94

72

5D

elta

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iform

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rme

Nid

up, T

., G

yelts

hen,

T.,

Kum

ar, P

. G.,

Kle

in, W

. & D

orji,

P.

2016

Jour

nal o

f Thr

eate

ned

Taxa

8 (1

2): 9

467-

9472

6D

olic

hove

spul

a la

ma

Dor

ji, P

., K

lein

, W. &

Nid

up, T

.20

17Jo

urna

l of I

nsec

t Bio

dive

rsity

and

Sys

tem

atic

s 3 (2

): 91

-104

7Eu

men

es g

ibbo

sus

Nid

up, T

., G

yelts

hen,

T.,

Kum

ar, P

. G.,

Kle

in, W

. & D

orji,

P.

2016

Jour

nal o

f Thr

eate

ned

Taxa

8 (1

2): 9

467-

9472

8Eu

sten

ogas

ter s

citu

laN

idup

, T.,

Kle

in, W

. & D

orji,

P.

2017

Jour

nal o

f Thr

eate

ned

Taxa

9 (7

): 10

487-

1048

9

9La

bus p

usill

usN

idup

, T.,

Gye

ltshe

n, T

., K

umar

, P. G

., K

lein

, W. &

Dor

ji, P

.20

16Jo

urna

l of T

hrea

tene

d Ta

xa 8

(12)

: 946

7-94

72

10Li

acos

ery

thro

som

a er

ythr

osom

aN

idup

, T.,

Kle

in, W

., K

umar

, P. G

. & D

orji,

P.

2017

Hal

tere

s 8: 9

-19

11M

egac

amps

omer

is c

ochi

nens

isN

idup

, T.,

Kle

in, W

., K

umar

, P. G

. & D

orji,

P.

2017

Hal

tere

s 8: 9

-19

12M

egac

amps

omer

is sh

illon

gens

isN

idup

, T.,

Kle

in, W

., K

umar

, P. G

. & D

orji,

P.

2017

Hal

tere

s 8: 9

-19

73


13M

egas

colia

(Reg

isco

lia) a

zure

a ch

ris-

tiana

Nid

up, T

., K

lein

, W.,

Kum

ar, P

. G. &

Dor

ji, P

.20

17H

alte

res 8

: 9-1

9

14M

egas

colia

(Reg

isco

lia) a

zure

a hi

ndo-

stan

aN

idup

, T.,

Kle

in, W

., K

umar

, P. G

. & D

orji,

P.

2017

Hal

tere

s 8: 9

-19

15Pa

rapo

lybi

a no

dosa

Dor

ji, P

., K

lein

, W. &

Nid

up, T

.20

17Jo

urna

l of I

nsec

t Bio

dive

rsity

and

Sys

tem

atic

s 3 (2

): 91

-104

16Pa

rapo

lybi

a va

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Dor

ji, P

., K

lein

, W. &

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up, T

.20

17Jo

urna

l of I

nsec

t Bio

dive

rsity

and

Sys

tem

atic

s 3 (2

): 91

-104

17Pa

reum

enes

qua

dris

pino

sus a

cutu

sN

idup

, T.,

Gye

ltshe

n, T

., K

umar

, P. G

., K

lein

, W. &

Dor

ji, P

.20

16Jo

urna

l of T

hrea

tene

d Ta

xa 8

(12)

: 946

7-94

72

18Pa

risc

hnog

aste

r mel

lyi

Nid

up, T

., K

lein

, W. &

Dor

ji, P

.20

17Jo

urna

l of T

hrea

tene

d Ta

xa 9

(7):

1048

7-10

489

19Ph

aler

imer

is p

hale

rata

pha

lera

taN

idup

, T.,

Kle

in, W

. & D

orji,

P.

2017

Hal

tere

s 8: 9

-19

20Po

liste

s (G

yros

tom

a) o

livac

eus

Dor

ji, P

., K

lein

, W. &

Nid

up, T

.20

16B

io B

ulle

tin 2

(1):

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136

21Po

liste

s (G

yros

tom

a) ro

thne

yi si

kki-

men

sis

Dor

ji, P

., K

lein

, W. &

Nid

up, T

.20

16B

io B

ulle

tin 2

(1):

128-

136

22Po

liste

s (G

yros

tom

a) te

nebr

icos

us

sulc

atus

Dor

ji, P

., K

lein

, W. &

Nid

up, T

.20

16B

io B

ulle

tin 2

(1):

128-

136

23Po

liste

s (Po

liste

lla) n

igri

tars

usD

orji,

P.,

Kle

in, W

. & N

idup

, T.

2017

Jour

nal o

f Ins

ect B

iodi

vers

ity a

nd S

yste

mat

ics 3

(2):

91-1

04

24Po

liste

s (Po

liste

lla) s

anto

shae

Dor

ji, P

., K

lein

, W. &

Nid

up, T

.20

16B

io B

ulle

tin 2

(1):

128-

136

25Pr

oves

pa b

arth

elem

yiD

orji,

P.,

Gye

ltshe

n, T

., K

lein

, W. &

Nid

up, T

.20

17Jo

urna

l of T

hrea

tene

d Ta

xa 9

(4):

1010

2-10

108

26Ro

palid

ia a

rtife

xD

orji,

P.,

Kle

in, W

. & N

idup

, T.

2017

Jour

nal o

f Ins

ect B

iodi

vers

ity a

nd S

yste

mat

ics 3

(2):

91-1

04

27Ro

palid

ia fa

scia

taD

orji,

P.,

Kle

in, W

. & N

idup

, T.

2016

Bio

Bul

letin

2 (1

): 12

8-14

0

28Ro

palid

ia jo

cobs

oni

Dor

ji, P

., K

lein

, W. &

Nid

up, T

.20

16B

io B

ulle

tin 2

(1):

128-

140

29Ro

palid

ia o

rnat

icep

sD

orji,

P.,

Kle

in, W

. & N

idup

, T.

2017

Jour

nal o

f Ins

ect B

iodi

vers

ity a

nd S

yste

mat

ics 3

(2):

91-1

04

30Ro

palid

ia ru

foco

llari

sD

orji,

P.,

Kle

in, W

. & N

idup

, T.

2016

Bio

Bul

letin

2 (1

): 12

8-13

6

31Ro

palid

ia ru

fopl

agia

ta g

rave

lyi

Dor

ji, P

., K

lein

, W. &

Nid

up, T

.20

17Jo

urna

l of I

nsec

t Bio

dive

rsity

and

Sys

tem

atic

s 3 (2

): 91

-104

32Ro

palid

ia sa

ntos

hae

Dor

ji, P

., K

lein

, W. &

Nid

up, T

.20

16B

io B

ulle

tin 2

(1):

128-

140

33Ro

palid

ia st

igm

aD

orji,

P.,

Kle

in, W

. & N

idup

, T.

2017

Jour

nal o

f Ins

ect B

iodi

vers

ity a

nd S

yste

mat

ics 3

(2):

91-1

04

34Sc

olia

(Dis

colia

) bin

otat

a bi

nota

taN

idup

, T.,

Kle

in, W

., K

umar

, P. G

. & D

orji,

P.

2017

Hal

tere

s 8: 9

-19

35Sc

olia

(Dis

colia

) cly

peat

a ru

fuhi

rta

Nid

up, T

., K

lein

, W.,

Kum

ar, P

. G. &

Dor

ji, P

.20

17H

alte

res 8

: 9-1

9

36Sc

olia

(Dis

colia

) deh

raen

sis

Nid

up, T

., K

lein

, W.,

Kum

ar, P

. G. &

Dor

ji, P

.20

17H

alte

res 8

: 9-1

9

37Sc

olia

(Dis

colia

) des

idio

saN

idup

, T.,

Kle

in, W

., K

umar

, P. G

. & D

orji,

P.

2017

Hal

tere

s 8: 9

-19

38Sc

olia

(Dis

colia

) eliz

abet

hae

Nid

up, T

., K

lein

, W.,

Kum

ar, P

. G. &

Dor

ji, P

.20

17H

alte

res 8

: 9-1

9

39Sc

olia

(Dis

colia

) fas

ciat

opun

ctat

a du

nens

isN

idup

, T.,

Kle

in, W

., K

umar

, P. G

. & D

orji,

P.

2017

Hal

tere

s 8: 9

-19

40Sc

olia

(Dis

colia

) kam

enge

nsis

Nid

up, T

., K

lein

, W.,

Kum

ar, P

. G. &

Dor

ji, P

.20

17H

alte

res 8

: 9-1

9

41Sc

olia

(Dis

colia

) rug

ifron

sN

idup

, T.,

Kle

in, W

., K

umar

, P. G

. & D

orji,

P.

2017

Hal

tere

s 8: 9

-19

42Sc

olia

(Dis

colia

) ven

usta

Nid

up, T

., K

lein

, W.,

Kum

ar, P

. G. &

Dor

ji, P

.20

17H

alte

res 8

: 9-1

9

43Ve

spa

affini

s affi

nis

Dor

ji, P

., G

yelts

hen,

T.,

Kle

in, W

. & N

idup

, T.

2017

Jour

nal o

f Thr

eate

ned

Taxa

9 (4

): 10

102-

1010

8

44Ve

spa

affini

s ind

osin

ensi

sD

orji,

P.,

Gye

ltshe

n, T

., K

lein

, W. &

Nid

up, T

.20

17Jo

urna

l of T

hrea

tene

d Ta

xa 9

(4):

1010

2-10

108

45Ve

spa

anal

is n

igra

nsD

orji,

P.,

Gye

ltshe

n, T

., K

lein

, W. &

Nid

up, T

.20

17Jo

urna

l of T

hrea

tene

d Ta

xa 9

(4):

1010

2-10

108

74


46Ve

spa

basa

lisD

orji,

P.,

Gye

ltshe

n, T

., K

lein

, W. &

Nid

up, T

.20

17Jo

urna

l of T

hrea

tene

d Ta

xa 9

(4):

1010

2-10

108

47Ve

spa

bico

lor

Dor

ji, P

., G

yelts

hen,

T.,

Kle

in, W

. & N

idup

, T.

2017

Jour

nal o

f Thr

eate

ned

Taxa

9 (4

): 10

102-

1010

8

48Ve

spa

fum

ida

Dor

ji, P

., K

lein

, W. &

Nid

up, T

.20

17Jo

urna

l of I

nsec

t Bio

dive

rsity

and

Sys

tem

atic

s 3 (2

): 91

-104

49Ve

spa

man

dari

nia

mag

nific

aD

orji,

P.,

Gye

ltshe

n, T

., K

lein

, W. &

Nid

up, T

.20

17Jo

urna

l of T

hrea

tene

d Ta

xa 9

(4):

1010

2-10

108

50Ve

spa

tropi

ca le

efm

ansi

Dor

ji, P

., G

yelts

hen,

T.,

Kle

in, W

. & N

idup

, T.

2017

Jour

nal o

f Thr

eate

ned

Taxa

9 (4

): 10

102-

1010

8

51Ve

spa

velu

tina

nigr

ithor

axD

orji,

P.,

Gye

ltshe

n, T

., K

lein

, W. &

Nid

up, T

.20

17Jo

urna

l of T

hrea

tene

d Ta

xa 9

(4):

1010

2-10

108

52Ve

spa

velu

tina

vari

ana

Dor

ji, P

., K

lein

, W. &

Nid

up, T

.20

17Jo

urna

l of I

nsec

t Bio

dive

rsity

and

Sys

tem

atic

s 3 (2

): 91

-104

53Ve

spa

viva

xD

orji,

P.,

Kle

in, W

. & N

idup

, T.

2017

Jour

nal o

f Ins

ect B

iodi

vers

ity a

nd S

yste

mat

ics 3

(2):

91-1

04

54Ve

spul

a fla

vice

ps fl

avic

eps

Dor

ji, P

., K

lein

, W. &

Nid

up, T

.20

17Jo

urna

l of I

nsec

t Bio

dive

rsity

and

Sys

tem

atic

s 3 (2

): 91

-104

55Ve

spul

a nu

rsei

Dor

ji, P

., K

lein

, W. &

Nid

up, T

.20

17Jo

urna

l of I

nsec

t Bio

dive

rsity

and

Sys

tem

atic

s 3 (2

): 91

-104

56Ve

spul

a st

ruct

orD

orji,

P.,

Kle

in, W

. & N

idup

, T.

2017

Jour

nal o

f Ins

ect B

iodi

vers

ity a

nd S

yste

mat

ics 3

(2):

91-1

04

57Ve

spul

a vu

lgar

isD

orji,

P.,

Kle

in, W

. & N

idup

, T.

2017

Jour

nal o

f Ins

ect B

iodi

vers

ity a

nd S

yste

mat

ics 3

(2):

91-1

04

58Xy

loco

pa (C

teno

poda

) fen

estr

ata

Dor

ji, P

., K

lein

, W. &

Nid

up, T

.20

16Jo

urna

l of N

ew B

iolo

gica

l Rep

orts

5 (a

) 106

-113

59Xy

loco

pa (C

yane

oder

es) a

cutip

enni

sD

orji,

P.,

Kle

in, W

. & N

idup

, T.

2016

Jour

nal o

f New

Bio

logi

cal R

epor

ts 5

(a) 1

06-1

13

60Xy

loco

pa (N

ycto

mel

itta)

tran

queb

aric

aD

orji,

P.,

Kle

in, W

. & N

idup

, T.

2016

Jour

nal o

f New

Bio

logi

cal R

epor

ts 5

(a) 1

06-1

13

61Xy

loco

pa (O

rbite

lla) a

esta

uns

Dor

ji, P

., K

lein

, W. &

Nid

up, T

.20

16Jo

urna

l of N

ew B

iolo

gica

l Rep

orts

5 (a

) 106

-113

62Xy

loco

pa (P

laty

nopo

da) l

atip

esD

orji,

P.,

Kle

in, W

. & N

idup

, T.

2016

Jour

nal o

f New

Bio

logi

cal R

epor

ts 5

(a) 1

06-1

13

63Xy

loco

pa (P

laty

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da) m

agni

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Dor

ji, P

., K

lein

, W. &

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up, T

.20

16Jo

urna

l of N

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gica

l Rep

orts

5 (a

) 106

-113

64Xy

loco

pa (Z

onoh

irsu

ta) c

olla

ris b

ing-

ham

iD

orji,

P.,

Kle

in, W

. & N

idup

, T.

2016

Jour

nal o

f New

Bio

logi

cal R

epor

ts 5

(a) 1

06-1

13

65Ze

thus

dol

osus

Nid

up, T

., G

yelts

hen,

T.,

Kum

ar, P

. G.,

Kle

in, W

. & D

orji,

P.

2016

Jour

nal o

f Thr

eate

ned

Taxa

8 (1

2): 9

467-

9472

Col

eopt

era

(Bee

tles)

S.N

o.Sp

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ient

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ncta

tus l

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Rec

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2H

ydat

icus

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atoi

sato

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hosh

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anda

l, G

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13R

ecor

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oolo

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vey

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a 11

3 (P

art-3

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Rec

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terfl

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efer

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ostic

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angd

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e, Y

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terfl

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opal

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2Eu

thal

ia a

mpl

ifasc

iaW

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i, S.

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gdi,

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aguc

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12B

utte

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s (Te

inop

alpu

s) N

o. 6

2

3N

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phyr

us su

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Wan

gdi,

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i, K

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12B

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alpu

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2

4U

na u

sta

Poel

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Lep

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S.N

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arR

efer

ence

1Ac

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A F

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Com

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Mot

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f Bhu

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am, J

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Wan

gdi,

K.

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Jour

nal o

f Asi

a-Pa

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Bio

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atho

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Irun

gbam

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hib,

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i, K

.20

16Jo

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Paci

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iodi

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a ze

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Jour

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dive

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ra b

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aIr

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Jour

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Bio

dive

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.20

17A

Fie

ld G

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e C

omm

on M

oths

of B

huta

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7Ar

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blep

sis r

ubid

aIr

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Jour

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f Asi

a-Pa

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Irun

gbam

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angd

i, K

.20

16Jo

urna

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sia-

Paci

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iodi

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ity, 1

e32

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cari

cae

Gie

lis, C

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angd

i, K

.20

17A

Fie

ld G

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to th

e C

omm

on M

oths

of B

huta

n

10Be

lippa

hor

rida

Irun

gbam

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hib,

M. S

. & S

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yev,

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.20

17Jo

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hrea

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d Ta

xa 9

(2):

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-981

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11Bo

tyod

es a

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isIr

ungb

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Jour

nal o

f Asi

a-Pa

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Bio

dive

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es c

aldu

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Irun

gbam

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angd

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.20

16Jo

urna

l of A

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Paci

fic B

iodi

vers

ity, 1

e32

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tyod

es c

roco

pter

alis

Irun

gbam

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angd

i, K

.20

16Jo

urna

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sia-

Paci

fic B

iodi

vers

ity, 1

e32

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adin

a di

agon

alis

Irun

gbam

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hib,

M. S

. & W

angd

i, K

.20

16Jo

urna

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Paci

fic B

iodi

vers

ity, 1

e32

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alam

otro

pha

late

llus

Irun

gbam

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., C

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M. S

. & W

angd

i, K

.20

16Jo

urna

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Paci

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iodi

vers

ity, 1

e32

16C

hero

met

tia fe

rrug

inea

Irun

gbam

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hib,

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yev,

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.20

17Jo

urna

l of T

hrea

tene

d Ta

xa 9

(2):

9795

-981

3

17C

hrys

orab

dia

bivi

ttaG

ielis

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K.

2017

A F

ield

Gui

de to

the

Com

mon

Mot

hs o

f Bhu

tan

18C

hrys

oteu

chia

div

isel

laG

ielis

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Wan

gdi,

K.

2017

A F

ield

Gui

de to

the

Com

mon

Mot

hs o

f Bhu

tan

19C

irrh

ochr

ista

bri

zoal

isIr

ungb

am, J

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Chi

b, M

. S. &

Wan

gdi,

K20

16Jo

urna

l of A

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Paci

fic B

iodi

vers

ity, 1

e32

20C

irrh

ochr

ista

fum

ipal

pis

Irun

gbam

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., C

hib,

M. S

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angd

i, K

2016

Jour

nal o

f Asi

a-Pa

cific

Bio

dive

rsity

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2

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irrh

ochr

ista

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usa

Irun

gbam

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M. S

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angd

i, K

2016

Jour

nal o

f Asi

a-Pa

cific

Bio

dive

rsity

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2

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lani

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erio

nIr

ungb

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.20

14Jo

urna

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hrea

tene

d Ta

xa 6

(1):

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23C

naph

aloc

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ungb

am, J

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b, M

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Wan

gdi,

K20

16Jo

urna

l of A

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Paci

fic B

iodi

vers

ity, 1

e32

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onog

ethe

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Irun

gbam

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hib,

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angd

i, K

2016

Jour

nal o

f Asi

a-Pa

cific

Bio

dive

rsity

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2

25C

otac

hena

nep

alen

sis

Irun

gbam

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M. S

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angd

i, K

2016

Jour

nal o

f Asi

a-Pa

cific

Bio

dive

rsity

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2

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otac

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pub

esce

nsIr

ungb

am, J

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Chi

b, M

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Wan

gdi,

K20

16Jo

urna

l of A

sia-

Paci

fic B

iodi

vers

ity, 1

e32

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roci

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olal

isIr

ungb

am, J

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b, M

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gdi,

K20

16Jo

urna

l of A

sia-

Paci

fic B

iodi

vers

ity, 1

e32

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roci

doph

ora

fasc

iata

Irun

gbam

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angd

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2016

Jour

nal o

f Asi

a-Pa

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Bio

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aG

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A F

ield

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the

Com

mon

Mot

hs o

f Bhu

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ydal

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gdi,

K20

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urna

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Paci

fic B

iodi

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ity, 1

e32

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iaph

nia

indi

caIr

ungb

am, J

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Chi

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Wan

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K20

16Jo

urna

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Paci

fic B

iodi

vers

ity, 1

e32

32D

iath

raus

ta p

rofu

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ungb

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Chi

b, M

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gdi,

K20

16Jo

urna

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Paci

fic B

iodi

vers

ity, 1

e32

33D

icho

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is d

efini

taIr

ungb

am, J

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gdi,

K20

16Jo

urna

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Paci

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iodi

vers

ity, 1

e32

34D

icho

croc

is ze

bral

isIr

ungb

am, J

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Chi

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Wan

gdi,

K20

16Jo

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Paci

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vers

ity, 1

e32

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35D

icho

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lisIr

ungb

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gdi,

K20

16Jo

urna

l of A

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Paci

fic B

iodi

vers

ity, 1

e32

36D

repa

na g

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arip

enni

sG

ielis

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2017

A F

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Gui

de to

the

Com

mon

Mot

hs o

f Bhu

tan

37En

docr

osis

flav

ibas

alis

Irun

gbam

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hib,

M. S

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angd

i, K

2016

Jour

nal o

f Asi

a-Pa

cific

Bio

dive

rsity

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2

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dotr

icha

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inal

isIr

ungb

am, J

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Chi

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K20

16Jo

urna

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Paci

fic B

iodi

vers

ity, 1

e32

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la p

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ocal

isIr

ungb

am, J

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Chi

b, M

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Wan

gdi,

K20

16Jo

urna

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Paci

fic B

iodi

vers

ity, 1

e32

40Eo

ophy

la se

junc

talis

Irun

gbam

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hib,

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angd

i, K

2016

Jour

nal o

f Asi

a-Pa

cific

Bio

dive

rsity

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2

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ipar

batti

a gl

orio

salis

wha

lleyi

Irun

gbam

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hib,

M. S

. & W

angd

i, K

2016

Jour

nal o

f Asi

a-Pa

cific

Bio

dive

rsity

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2

42Er

iste

na b

ifurc

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Irun

gbam

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hib,

M. S

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angd

i, K

2016

Jour

nal o

f Asi

a-Pa

cific

Bio

dive

rsity

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2

43Er

pis m

acul

aris

Irun

gbam

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hib,

M. S

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angd

i, K

2016

Jour

nal o

f Asi

a-Pa

cific

Bio

dive

rsity

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2

44Eu

doci

ma

sala

min

iaG

ielis

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Wan

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2017

A F

ield

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de to

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Com

mon

Mot

hs o

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tan

45Fi

lode

s ful

vido

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isIr

ungb

am, J

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Chi

b, M

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Wan

gdi,

K20

16Jo

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Paci

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ity, 1

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46G

lyph

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biv

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ungb

am, J

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Chi

b, M

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Wan

gdi,

K20

16Jo

urna

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Paci

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iodi

vers

ity, 1

e32

47G

lyph

odes

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Irun

gbam

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M. S

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angd

i, K

2016

Jour

nal o

f Asi

a-Pa

cific

Bio

dive

rsity

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2

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lyph

odes

cri

thea

lisIr

ungb

am, J

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Chi

b, M

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Wan

gdi,

K20

16Jo

urna

l of A

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Paci

fic B

iodi

vers

ity, 1

e32

49G

lyph

odes

har

utai

Irun

gbam

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hib,

M. S

. & W

angd

i, K

2016

Jour

nal o

f Asi

a-Pa

cific

Bio

dive

rsity

, 1e3

2

50G

lyph

odes

ony

chin

alis

Irun

gbam

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hib,

M. S

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angd

i, K

2016

Jour

nal o

f Asi

a-Pa

cific

Bio

dive

rsity

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2

51G

lyph

odes

stol

alis

Irun

gbam

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., C

hib,

M. S

. & W

angd

i, K

2016

Jour

nal o

f Asi

a-Pa

cific

Bio

dive

rsity

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2

52H

arita

lode

s der

ogat

aIr

ungb

am, J

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Chi

b, M

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Wan

gdi,

K20

16Jo

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Paci

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iodi

vers

ity, 1

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53H

eort

ia v

itess

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sIr

ungb

am, J

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Chi

b, M

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Wan

gdi,

K20

16Jo

urna

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Paci

fic B

iodi

vers

ity, 1

e32

54H

yalo

bath

ra m

inio

salis

Irun

gbam

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hib,

M. S

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angd

i, K

2016

Jour

nal o

f Asi

a-Pa

cific

Bio

dive

rsity

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2

55H

yalo

plag

a pu

lchr

alis

Irun

gbam

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hib,

M. S

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angd

i, K

2016

Jour

nal o

f Asi

a-Pa

cific

Bio

dive

rsity

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2

56H

ypso

pygi

a m

auri

tialis

Irun

gbam

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hib,

M. S

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angd

i, K

2016

Jour

nal o

f Asi

a-Pa

cific

Bio

dive

rsity

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2

57H

ypso

pygi

a ni

tidic

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ungb

am, J

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Chi

b, M

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Wan

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K20

16Jo

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vers

ity, 1

e32

58Io

taph

ora

irid

icol

orG

ielis

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A F

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de to

the

Com

mon

Mot

hs o

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tan

59Ir

aga

rugo

saIr

ungb

am, J

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b, M

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Sol

ovye

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2017

Jour

nal o

f Thr

eate

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95-9

813

60Is

chyj

a he

mip

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Gie

lis, C

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e C

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oths

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61La

mor

ia v

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Irun

gbam

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hib,

M. S

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angd

i, K

2016

Jour

nal o

f Asi

a-Pa

cific

Bio

dive

rsity

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2

62Li

sta

ficki

Irun

gbam

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hib,

M. S

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angd

i, K

2016

Jour

nal o

f Asi

a-Pa

cific

Bio

dive

rsity

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2

63Lu

mar

ia p

robo

lias

Gie

lis, C

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.20

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64M

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ungb

am, J

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Chi

b, M

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16Jo

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Paci

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ity, 1

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65M

iresa

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ida

Irun

gbam

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17Jo

urna

l of T

hrea

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d Ta

xa 9

(2):

9795

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3

66N

acol

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chry

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cta

Irun

gbam

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hib,

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angd

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2016

Jour

nal o

f Asi

a-Pa

cific

Bio

dive

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2

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taIr

ungb

am, J

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Chi

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gdi,

K20

16Jo

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Paci

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ity, 1

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68N

ausi

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lisIr

ungb

am, J

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K20

16Jo

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Paci

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iodi

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ity, 1

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ausi

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2016

Jour

nal o

f Asi

a-Pa

cific

Bio

dive

rsity

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2

70N

evri

na a

boe

Gie

lis, C

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angd

i, K

.20

17A

Fie

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to th

e C

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on M

oths

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77


71N

evri

na p

roco

pia

Irun

gbam

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hib,

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angd

i, K

2016

Jour

nal o

f Asi

a-Pa

cific

Bio

dive

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2

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A F

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Com

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Mot

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16Jo

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Paci

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A F

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75O

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16Jo

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Paci

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76O

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ungb

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Chi

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16Jo

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16Jo

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78O

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Jour

nal o

f Asi

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Bio

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Irun

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Jour

nal o

f Asi

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Bio

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16Jo

urna

l of A

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Paci

fic B

iodi

vers

ity, 1

e32

81Pa

chyn

oa sa

belia

lisIr

ungb

am, J

. S.,

Chi

b, M

. S. &

Wan

gdi,

K20

16Jo

urna

l of A

sia-

Paci

fic B

iodi

vers

ity, 1

e32

82Pa

chyn

oa x

anth

ochy

taIr

ungb

am, J

. S.,

Chi

b, M

. S. &

Wan

gdi,

K20

16Jo

urna

l of A

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Paci

fic B

iodi

vers

ity, 1

e32

83Pa

gyda

salv

alis

Irun

gbam

, J. S

., C

hib,

M. S

. & W

angd

i, K

2016

Jour

nal o

f Asi

a-Pa

cific

Bio

dive

rsity

, 1e3

2

84Pa

lpita

ann

ulife

rIr

ungb

am, J

. S.,

Chi

b, M

. S. &

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gdi,

K20

16Jo

urna

l of A

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Paci

fic B

iodi

vers

ity, 1

e32

85Pa

lpita

asi

atic

alis

Irun

gbam

, J. S

., C

hib,

M. S

. & W

angd

i, K

2016

Jour

nal o

f Asi

a-Pa

cific

Bio

dive

rsity

, 1e3

2

86Pa

lpita

war

rena

lisIr

ungb

am, J

. S.,

Chi

b, M

. S. &

Wan

gdi,

K20

16Jo

urna

l of A

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Paci

fic B

iodi

vers

ity, 1

e32

87Pa

rapo

nyx

fluct

uosa

lisIr

ungb

am, J

. S.,

Chi

b, M

. S. &

Wan

gdi,

K20

16Jo

urna

l of A

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Paci

fic B

iodi

vers

ity, 1

e32

88Pa

rapo

nyx

stag

nalis

Irun

gbam

, J. S

., C

hib,

M. S

. & W

angd

i, K

2016

Jour

nal o

f Asi

a-Pa

cific

Bio

dive

rsity

, 1e3

2

89Pa

rotis

mar

gina

taIr

ungb

am, J

. S.,

Chi

b, M

. S. &

Wan

gdi,

K20

16Jo

urna

l of A

sia-

Paci

fic B

iodi

vers

ity, 1

e34

90Pa

tani

a ca

leto

ralis

Irun

gbam

, J. S

., C

hib,

M. S

. & W

angd

i, K

2016

Jour

nal o

f Asi

a-Pa

cific

Bio

dive

rsity

, 1e3

4

91Po

lyth

lipta

cer

ealis

Irun

gbam

, J. S

., C

hib,

M. S

. & W

angd

i, K

2016

Jour

nal o

f Asi

a-Pa

cific

Bio

dive

rsity

, 1e3

4

92Po

lyth

lipta

div

aric

ata

Irun

gbam

, J. S

., C

hib,

M. S

. & W

angd

i, K

2016

Jour

nal o

f Asi

a-Pa

cific

Bio

dive

rsity

, 1e3

4

93Pr

ooed

ema

insc

isal

isIr

ungb

am, J

. S.,

Chi

b, M

. S. &

Wan

gdi,

K20

16Jo

urna

l of A

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Paci

fic B

iodi

vers

ity, 1

e34

94Pr

opha

ntis

adu

sta

Irun

gbam

, J. S

., C

hib,

M. S

. & W

angd

i, K

2016

Jour

nal o

f Asi

a-Pa

cific

Bio

dive

rsity

1e3

2

95Py

cnar

mon

lact

ifera

lisIr

ungb

am, J

. S.,

Chi

b, M

. S. &

Wan

gdi,

K20

16Jo

urna

l of A

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Paci

fic B

iodi

vers

ity, 1

e32

96Py

gosp

ila ty

res

Irun

gbam

, J. S

., C

hib,

M. S

. & W

angd

i, K

2016

Jour

nal o

f Asi

a-Pa

cific

Bio

dive

rsity

, 1e3

2

97Py

ralis

pic

talis

Irun

gbam

, J. S

., C

hib,

M. S

. & W

angd

i, K

2016

Jour

nal o

f Asi

a-Pa

cific

Bio

dive

rsity

, 1e3

2

98Rh

agob

a oc

tom

acul

alis

Irun

gbam

, J. S

., C

hib,

M. S

. & W

angd

i, K

2016

Jour

nal o

f Asi

a-Pa

cific

Bio

dive

rsity

, 1e3

2

99Rh

imph

alea

trog

usal

isIr

ungb

am, J

. S.,

Chi

b, M

. S. &

Wan

gdi,

K20

16Jo

urna

l of A

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Paci

fic B

iodi

vers

ity, 1

e32

100

Sam

eode

s can

cella

lisIr

ungb

am, J

. S.,

Chi

b, M

. S. &

Wan

gdi,

K20

16Jo

urna

l of A

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Paci

fic B

iodi

vers

ity, 1

e32

101

Sans

area

circ

ulife

raIr

ungb

am, J

. S.,

Chi

b, M

. S. &

Sol

ovye

v, A

. V.

2017

Jour

nal o

f Thr

eate

ned

Taxa

9 (2

): 97

95-9

813

102

Scop

elod

es te

stac

eaIr

ungb

am, J

. S.,

Chi

b, M

. S. &

Sol

ovye

v, A

. V.

2017

Jour

nal o

f Thr

eate

ned

Taxa

9 (2

): 97

95-9

813

103

Seto

ra fl

etch

eri

Irun

gbam

, J. S

., C

hib,

M. S

. & S

olov

yev,

A. V

.20

17Jo

urna

l of T

hrea

tene

d Ta

xa 9

(2):

9795

-981

3

104

Seto

ra p

osto

rnat

aIr

ungb

am, J

. S.,

Chi

b, M

. S. &

Sol

ovye

v, A

. V.

2017

Jour

nal o

f Thr

eate

ned

Taxa

9 (2

): 97

95-9

813

105

Spol

adea

recu

rval

isIr

ungb

am, J

. S.,

Chi

b, M

. S. &

Wan

gdi,

K20

16Jo

urna

l of A

sia-

Paci

fic B

iodi

vers

ity, 1

e32

106

Stre

psin

oma

croe

susa

lisIr

ungb

am, J

. S.,

Chi

b, M

. S. &

Wan

gdi,

K20

16Jo

urna

l of A

sia-

Paci

fic B

iodi

vers

ity, 1

e32

78


107

Susi

ca h

imal

ayan

aIr

ungb

am, J

. S.,

Chi

b, M

. S. &

Sol

ovye

v, A

. V.

2017

Jour

nal o

f Thr

eate

ned

Taxa

9 (2

): 97

95-9

813

108

Sylle

pte

gast

ralis

Irun

gbam

, J. S

., C

hib,

M. S

. & W

angd

i, K

2016

Jour

nal o

f Asi

a-Pa

cific

Bio

dive

rsity

, 1e3

2

109

Tala

nga

sexp

unct

alis

Irun

gbam

, J. S

., C

hib,

M. S

. & W

angd

i, K

2016

Jour

nal o

f Asi

a-Pa

cific

Bio

dive

rsity

, 1e3

2

110

Tenn

ya p

ropo

liaIr

ungb

am, J

. S.,

Chi

b, M

. S. &

Sol

ovye

v, A

. V.

2017

Jour

nal o

f Thr

eate

ned

Taxa

9 (2

): 97

95-9

813

111

Tera

stia

egi

alea

lisIr

ungb

am, J

. S.,

Chi

b, M

. S. &

Wan

gdi,

K20

16Jo

urna

l of A

sia-

Paci

fic B

iodi

vers

ity, 1

e32

112

Thos

ea m

agna

Irun

gbam

, J. S

., C

hib,

M. S

. & S

olov

yev,

A. V

.20

17Jo

urna

l of T

hrea

tene

d Ta

xa 9

(2):

9795

-981

3

113

Tysp

anod

es li

neal

isIr

ungb

am, J

. S.,

Chi

b, M

. S. &

Wan

gdi,

K20

16Jo

urna

l of A

sia-

Paci

fic B

iodi

vers

ity, 1

e32

114

Tysp

anod

es n

igro

linea

lisIr

ungb

am, J

. S.,

Chi

b, M

. S. &

Wan

gdi,

K20

16Jo

urna

l of A

sia-

Paci

fic B

iodi

vers

ity, 1

e32

115

Unc

obot

yode

s pat

ulal

isIr

ungb

am, J

. S.,

Chi

b, M

. S. &

Wan

gdi,

K20

16Jo

urna

l of A

sia-

Paci

fic B

iodi

vers

ity, 1

e32

116

Uth

inia

alb

isig

nalis

Irun

gbam

, J. S

., C

hib,

M. S

. & W

angd

i, K

2016

Jour

nal o

f Asi

a-Pa

cific

Bio

dive

rsity

, 1e3

2

117

Vanl

angi

a un

iform

isIr

ungb

am, J

. S.,

Chi

b, M

. S. &

Sol

ovye

v, A

. V.

2017

Jour

nal o

f Thr

eate

ned

Taxa

9 (2

): 97

95-9

813

118

Vite

ssa

sura

deva

Irun

gbam

, J. S

., C

hib,

M. S

. & W

angd

i, K

2016

Jour

nal o

f Asi

a-Pa

cific

Bio

dive

rsity

, 1e3

2

119

Zith

a to

rrid

alis

Irun

gbam

, J. S

., C

hib,

M. S

. & W

angd

i, K

2016

Jour

nal o

f Asi

a-Pa

cific

Bio

dive

rsity

, 1e3

2

Dip

tera

(Tru

e Fl

ies)

S.N

o.Sp

ecie

sSc

ient

ist (

s)Ye

arR

efer

ence

1C

ulex

(Cul

icio

myi

a) sa

sai

Phan

itcha

kun,

T.,

Wila

i, P.

, Sai

ngam

sook

, J.,

Nam

gay,

R.,

Dru

kpa,

T.,

Tsud

a, Y

., W

alto

n, C

., H

ar-

back

, R.E

. & S

ombo

om, P

.20

17A

cta

Trop

ica

4266

Pisc

es (F

ishe

s)

S.N

o.Sp

ecie

sSc

ient

ist (

s)Ye

arR

efer

ence

1Ab

oric

hthy

s gar

oens

is

Gur

ung,

D. B

., D

orji,

S.,

Tshe

ring,

U. &

Wan

gyel

, J. T

.20

13Jo

urna

l of T

hrea

tene

d Ta

xa 5

(14)

: 488

0-48

86

2Ac

anth

ocob

itis b

otia

G

urun

g, D

. B.,

Dor

ji, S

., Ts

herin

g, U

. & W

angy

el, J

. T.

2013

Jour

nal o

f Thr

eate

ned

Taxa

5 (1

4): 4

880-

4886

3Ai

lia c

oila

G

urun

g, D

. B.,

Dor

ji, S

., Ts

herin

g, U

. & W

angy

el, J

. T.

2013

Jour

nal o

f Thr

eate

ned

Taxa

5 (1

4): 4

880-

4886

4Am

blyc

eps a

pang

iG

urun

g, D

. B.,

Dor

ji, S

., Ts

herin

g, U

. & W

angy

el, J

. T.

2013

Jour

nal o

f Thr

eate

ned

Taxa

5 (1

4): 4

880-

4886

5Am

blyc

eps c

erin

umTs

herin

g, S

., W

angc

huk,

K.,

Dor

ji, S

. & N

orbu

, P.

2017

Fiel

d gu

ide

to F

ishe

s of W

este

rn B

huta

n, N

RC

RLF

, Bhu

tan

6Am

blyc

eps c

f. ar

unac

hale

nsis

Tshe

ring,

S.,

Wan

gchu

k, K

., D

orji,

S. &

Nor

bu, P

.20

17Fi

eld

guid

e to

Fis

hes o

f Wes

tern

Bhu

tan,

NR

CR

LF, B

huta

n

7Am

blyc

eps m

ango

isG

urun

g, D

. B.,

Dor

ji, S

., Ts

herin

g, U

. & W

angy

el, J

. T.

2013

Jour

nal o

f Thr

eate

ned

Taxa

5 (1

4): 4

880-

4886

8An

guill

a be

ngal

ensi

s G

urun

g, D

. B.,

Dor

ji, S

., Ts

herin

g, U

. & W

angy

el, J

. T.

2013

Jour

nal o

f Thr

eate

ned

Taxa

5 (1

4): 4

880-

4886

9As

pido

pari

a ja

yaTs

herin

g, S

., W

angc

huk,

K.,

Dor

ji, S

. & N

orbu

, P.

2017

Fiel

d gu

ide

to F

ishe

s of W

este

rn B

huta

n, N

RC

RLF

, Bhu

tan

10As

pido

pari

a m

orar

G

urun

g, D

. B.,

Dor

ji, S

., Ts

herin

g, U

. & W

angy

el, J

. T.

2013

Jour

nal o

f Thr

eate

ned

Taxa

5 (1

4): 4

880-

4886

11Ba

dis b

adis

G

urun

g, D

. B.,

Dor

ji, S

., Ts

herin

g, U

. & W

angy

el, J

. T.

2013

Jour

nal o

f Thr

eate

ned

Taxa

5 (1

4): 4

880-

4886

12Ba

dis d

ibru

ensi

sTs

herin

g, S

., W

angc

huk,

K.,

Dor

ji, S

. & N

orbu

, P.

2017

Fiel

d gu

ide

to F

ishe

s of W

este

rn B

huta

n, N

RC

RLF

, Bhu

tan

13Ba

gari

us b

agar

ius

Gur

ung,

D. B

., D

orji,

S.,

Tshe

ring,

U. &

Wan

gyel

, J. T

.20

13Jo

urna

l of T

hrea

tene

d Ta

xa 5

(14)

: 488

0-48

86

14Ba

ngan

a de

roG

urun

g, D

. B.,

Dor

ji, S

., Ts

herin

g, U

. & W

angy

el, J

. T.

2013

Jour

nal o

f Thr

eate

ned

Taxa

5 (1

4): 4

880-

4886

15Ba

riliu

s bar

naG

urun

g, D

. B.,

Dor

ji, S

., Ts

herin

g, U

. & W

angy

el, J

. T.

2013

Jour

nal o

f Thr

eate

ned

Taxa

5 (1

4): 4

880-

4886

16Ba

riliu

s ben

delis

is

Gur

ung,

D. B

., D

orji,

S.,

Tshe

ring,

U. &

Wan

gyel

, J. T

.20

13Jo

urna

l of T

hrea

tene

d Ta

xa 5

(14)

: 488

0-48

86

17Ba

riliu

s sha

cra

Tshe

ring,

S.,

Wan

gchu

k, K

., D

orji,

S. &

Nor

bu, P

.20

17Fi

eld

guid

e to

Fis

hes o

f Wes

tern

Bhu

tan,

NR

CR

LF, B

huta

n

79


18Ba

riliu

s vag

ra

Gur

ung,

D. B

., D

orji,

S.,

Tshe

ring,

U. &

Wan

gyel

, J. T

.20

13Jo

urna

l of T

hrea

tene

d Ta

xa 5

(14)

: 488

0-48

86

19Ba

tasi

o ba

tasi

o G

urun

g, D

. B.,

Dor

ji, S

., Ts

herin

g, U

. & W

angy

el, J

. T.

2013

Jour

nal o

f Thr

eate

ned

Taxa

5 (1

4): 4

880-

4886

20Ba

tasi

o fa

scio

latu

sG

urun

g, D

. B.,

Dor

ji, S

., Ts

herin

g, U

. & W

angy

el, J

. T.

2013

Jour

nal o

f Thr

eate

ned

Taxa

5 (1

4): 4

880-

4886

21Ba

tasi

o m

eria

nien

sis

Gur

ung,

D. B

., D

orji,

S.,

Tshe

ring,

U. &

Wan

gyel

, J. T

.20

13Jo

urna

l of T

hrea

tene

d Ta

xa 5

(14)

: 488

0-48

86

22Bo

tia a

lmor

hae

Gur

ung,

D. B

., D

orji,

S.,

Tshe

ring,

U. &

Wan

gyel

, J. T

.20

13Jo

urna

l of T

hrea

tene

d Ta

xa 5

(14)

: 488

0-48

86

23Bo

tia d

ario

G

urun

g, D

. B.,

Dor

ji, S

., Ts

herin

g, U

. & W

angy

el, J

. T.

2013

Jour

nal o

f Thr

eate

ned

Taxa

5 (1

4): 4

880-

4886

24C

atla

cat

laG

urun

g, D

. B.,

Dor

ji, S

., Ts

herin

g, U

. & W

angy

el, J

. T.

2013

Jour

nal o

f Thr

eate

ned

Taxa

5 (1

4): 4

880-

4886

25C

hagu

nius

cha

guni

o G

urun

g, D

. B.,

Dor

ji, S

., Ts

herin

g, U

. & W

angy

el, J

. T.

2013

Jour

nal o

f Thr

eate

ned

Taxa

5 (1

4): 4

880-

4886

26C

hann

a am

phib

eus

Gur

ung,

D. B

., D

orji,

S.,

Tshe

ring,

U. &

Wan

gyel

, J. T

.20

13Jo

urna

l of T

hrea

tene

d Ta

xa 5

(14)

: 488

0-48

86

27C

hann

a ga

chua

G

urun

g, D

. B.,

Dor

ji, S

., Ts

herin

g, U

. & W

angy

el, J

. T.

2013

Jour

nal o

f Thr

eate

ned

Taxa

5 (1

4): 4

880-

4886

28C

hann

a m

elan

ostig

ma

Tshe

ring,

S.,

Wan

gchu

k, K

., D

orji,

S. &

Nor

bu, P

.20

17Fi

eld

guid

e to

Fis

hes o

f Wes

tern

Bhu

tan,

NR

CR

LF, B

huta

n

29C

hann

a pu

ncta

ta

Gur

ung,

D. B

., D

orji,

S.,

Tshe

ring,

U. &

Wan

gyel

, J. T

.20

13Jo

urna

l of T

hrea

tene

d Ta

xa 5

(14)

: 488

0-48

86

30C

hann

a st

ewar

tii

Gur

ung,

D. B

., D

orji,

S.,

Tshe

ring,

U. &

Wan

gyel

, J. T

.20

13Jo

urna

l of T

hrea

tene

d Ta

xa 5

(14)

: 488

0-48

86

31C

hann

a st

riat

aG

urun

g, D

. B.,

Dor

ji, S

., Ts

herin

g, U

. & W

angy

el, J

. T.

2013

Jour

nal o

f Thr

eate

ned

Taxa

5 (1

4): 4

880-

4886

32C

irrh

inus

mri

gala

G

urun

g, D

. B.,

Dor

ji, S

., Ts

herin

g, U

. & W

angy

el, J

. T.

2013

Jour

nal o

f Thr

eate

ned

Taxa

5 (1

4): 4

880-

4886

33C

lari

as m

agur

Gur

ung,

D. B

., D

orji,

S.,

Tshe

ring,

U. &

Wan

gyel

, J. T

.20

13Jo

urna

l of T

hrea

tene

d Ta

xa 5

(14)

: 488

0-48

86

34C

ross

oche

ilus l

atiu

s G

urun

g, D

. B.,

Dor

ji, S

., Ts

herin

g, U

. & W

angy

el, J

. T.

2013

Jour

nal o

f Thr

eate

ned

Taxa

5 (1

4): 4

880-

4886

35C

teno

phar

yngo

don

idel

la

Gur

ung,

D. B

., D

orji,

S.,

Tshe

ring,

U. &

Wan

gyel

, J. T

.20

13Jo

urna

l of T

hrea

tene

d Ta

xa 5

(14)

: 488

0-48

86

36D

aino

dan

gila

Gur

ung,

D. B

., D

orji,

S.,

Tshe

ring,

U. &

Wan

gyel

, J. T

.20

13Jo

urna

l of T

hrea

tene

d Ta

xa 5

(14)

: 488

0-48

86

37D

anio

cf.

assa

mila

Tshe

ring,

S.,

Wan

gchu

k, K

., D

orji,

S. &

Nor

bu, P

.20

17Fi

eld

guid

e to

Fis

hes o

f Wes

tern

Bhu

tan,

NR

CR

LF, B

huta

n

38D

anio

reri

oG

urun

g, D

. B.,

Dor

ji, S

., Ts

herin

g, U

. & W

angy

el, J

. T.

2013

Jour

nal o

f Thr

eate

ned

Taxa

5 (1

4): 4

880-

4886

39D

evar

io a

equi

pinn

atus

Gur

ung,

D. B

., D

orji,

S.,

Tshe

ring,

U. &

Wan

gyel

, J. T

.20

13Jo

urna

l of T

hrea

tene

d Ta

xa 5

(14)

: 488

0-48

86

40D

evar

io a

ssam

ensi

sTs

herin

g, S

., W

angc

huk,

K.,

Dor

ji, S

. & N

orbu

, P.

2017

Fiel

d gu

ide

to F

ishe

s of W

este

rn B

huta

n, N

RC

RLF

, Bhu

tan

41Ex

osto

ma

labi

atum

G

urun

g, D

. B.,

Dor

ji, S

., Ts

herin

g, U

. & W

angy

el, J

. T.

2013

Jour

nal o

f Thr

eate

ned

Taxa

5 (1

4): 4

880-

4886

42G

agat

a ce

nia

Gur

ung,

D. B

., D

orji,

S.,

Tshe

ring,

U. &

Wan

gyel

, J. T

.20

13Jo

urna

l of T

hrea

tene

d Ta

xa 5

(14)

: 488

0-48

86

43G

arra

ann

anda

lei

Gur

ung,

D. B

., D

orji,

S.,

Tshe

ring,

U. &

Wan

gyel

, J. T

.20

13Jo

urna

l of T

hrea

tene

d Ta

xa 5

(14)

: 488

0-48

86

44G

arra

aru

nach

alen

sis

Tshe

ring,

S.,

Wan

gchu

k, K

., D

orji,

S. &

Nor

bu, P

.20

17Fi

eld

guid

e to

Fis

hes o

f Wes

tern

Bhu

tan,

NR

CR

LF, B

huta

n

45G

arra

aru

piR

.J. T

honi

, D.B

. Gur

ung

& R

.L. M

ayde

n20

16Zo

otax

a 41

69 (1

): 11

5-13

2

46G

arra

biro

stri

sR

.J. T

honi

, D.B

. Gur

ung

& R

.L. M

ayde

n20

16Zo

otax

a 41

69 (1

): 11

5-13

2

47G

arra

got

yla

Gur

ung,

D. B

., D

orji,

S.,

Tshe

ring,

U. &

Wan

gyel

, J. T

.20

13Jo

urna

l of T

hrea

tene

d Ta

xa 5

(14)

: 488

0-48

86

48G

arra

lam

taTs

herin

g, S

., W

angc

huk,

K.,

Dor

ji, S

. & N

orbu

, P.

2017

Fiel

d gu

ide

to F

ishe

s of W

este

rn B

huta

n, N

RC

RLF

, Bhu

tan

49G

arra

liss

orhy

nchu

sR

.J. T

honi

, D.B

. Gur

ung

& R

.L. M

ayde

n20

16Zo

otax

a 41

69 (1

): 11

5-13

4

50G

arra

qua

drat

irost

ris

Tshe

ring,

S.,

Wan

gchu

k, K

., D

orji,

S. &

Nor

bu, P

.20

17Fi

eld

guid

e to

Fis

hes o

f Wes

tern

Bhu

tan,

NR

CR

LF, B

huta

n

51G

loss

ogob

ius g

iuri

sTs

herin

g, S

., W

angc

huk,

K.,

Dor

ji, S

. & N

orbu

, P.

2017

Fiel

d gu

ide

to F

ishe

s of W

este

rn B

huta

n, N

RC

RLF

, Bhu

tan

52G

lypt

otho

rax

botiu

sTs

herin

g, S

., W

angc

huk,

K.,

Dor

ji, S

. & N

orbu

, P.

2017

Fiel

d gu

ide

to F

ishe

s of W

este

rn B

huta

n, N

RC

RLF

, Bhu

tan

53G

lypt

otho

rax

cavi

a G

urun

g, D

. B.,

Dor

ji, S

., Ts

herin

g, U

. & W

angy

el, J

. T.

2013

Jour

nal o

f Thr

eate

ned

Taxa

5 (1

4): 4

880-

4886

80


54G

lypt

otho

rax

cf. p

anda

Tshe

ring,

S.,

Wan

gchu

k, K

., D

orji,

S. &

Nor

bu, P

.20

17Fi

eld

guid

e to

Fis

hes o

f Wes

tern

Bhu

tan,

NR

CR

LF, B

huta

n

55G

lypt

otho

rax

cf. t

elch

itta

Gur

ung,

D. B

., D

orji,

S.,

Tshe

ring,

U. &

Wan

gyel

, J. T

.20

13Jo

urna

l of T

hrea

tene

d Ta

xa 5

(14)

: 488

0-48

86

56G

lypt

otho

rax

stri

atus

Gur

ung,

D. B

., D

orji,

S.,

Tshe

ring,

U. &

Wan

gyel

, J. T

.20

13Jo

urna

l of T

hrea

tene

d Ta

xa 5

(14)

: 488

0-48

86

57G

ogan

gra

viri

desc

ens

Gur

ung,

D. B

., D

orji,

S.,

Tshe

ring,

U. &

Wan

gyel

, J. T

.20

13Jo

urna

l of T

hrea

tene

d Ta

xa 5

(14)

: 488

0-48

86

58H

eter

opne

uste

s fos

silis

G

urun

g, D

. B.,

Dor

ji, S

., Ts

herin

g, U

. & W

angy

el, J

. T.

2013

Jour

nal o

f Thr

eate

ned

Taxa

5 (1

4): 4

880-

4886

59H

ypop

htha

lmic

hthy

s mol

itrix

G

urun

g, D

. B.,

Dor

ji, S

., Ts

herin

g, U

. & W

angy

el, J

. T.

2013

Jour

nal o

f Thr

eate

ned

Taxa

5 (1

4): 4

880-

4886

60H

ypop

htha

lmic

hthy

s nob

ilis

Gur

ung,

D. B

., D

orji,

S.,

Tshe

ring,

U. &

Wan

gyel

, J. T

.20

13Jo

urna

l of T

hrea

tene

d Ta

xa 5

(14)

: 488

0-48

86

61La

beo

dyoc

heilu

s G

urun

g, D

. B.,

Dor

ji, S

., Ts

herin

g, U

. & W

angy

el, J

. T.

2013

Jour

nal o

f Thr

eate

ned

Taxa

5 (1

4): 4

880-

4886

62La

beo

pang

usia

G

urun

g, D

. B.,

Dor

ji, S

., Ts

herin

g, U

. & W

angy

el, J

. T.

2013

Jour

nal o

f Thr

eate

ned

Taxa

5 (1

4): 4

880-

4886

63La

beo

rohi

ta

Gur

ung,

D. B

., D

orji,

S.,

Tshe

ring,

U. &

Wan

gyel

, J. T

.20

13Jo

urna

l of T

hrea

tene

d Ta

xa 5

(14)

: 488

0-48

86

64Le

pido

ceph

alic

hthy

s gun

tea

Gur

ung,

D. B

., D

orji,

S.,

Tshe

ring,

U. &

Wan

gyel

, J. T

.20

13Jo

urna

l of T

hrea

tene

d Ta

xa 5

(14)

: 488

0-48

86

65M

acro

gnat

hus m

oreh

ensi

s G

urun

g, D

. B.,

Dor

ji, S

., Ts

herin

g, U

. & W

angy

el, J

. T.

2013

Jour

nal o

f Thr

eate

ned

Taxa

5 (1

4): 4

880-

4886

66M

asta

cem

belu

s arm

atus

Gur

ung,

D. B

., D

orji,

S.,

Tshe

ring,

U. &

Wan

gyel

, J. T

.20

13Jo

urna

l of T

hrea

tene

d Ta

xa 5

(14)

: 488

0-48

86

67M

ystu

s ble

eker

i G

urun

g, D

. B.,

Dor

ji, S

., Ts

herin

g, U

. & W

angy

el, J

. T.

2013

Jour

nal o

f Thr

eate

ned

Taxa

5 (1

4): 4

880-

4886

68M

ystu

s cf.

dibr

ugar

ensi

s G

urun

g, D

. B.,

Dor

ji, S

., Ts

herin

g, U

. & W

angy

el, J

. T.

2013

Jour

nal o

f Thr

eate

ned

Taxa

5 (1

4): 4

880-

4886

69M

ystu

s vitt

atus

G

urun

g, D

. B.,

Dor

ji, S

., Ts

herin

g, U

. & W

angy

el, J

. T.

2013

Jour

nal o

f Thr

eate

ned

Taxa

5 (1

4): 4

880-

4886

70N

andu

s nan

dus

Gur

ung,

D. B

., D

orji,

S.,

Tshe

ring,

U. &

Wan

gyel

, J. T

.20

13Jo

urna

l of T

hrea

tene

d Ta

xa 5

(14)

: 488

0-48

86

71N

eoeu

cirr

hich

thys

may

delli

Tshe

ring,

S.,

Wan

gchu

k, K

., D

orji,

S. &

Nor

bu, P

.20

17Fi

eld

guid

e to

Fis

hes o

f Wes

tern

Bhu

tan,

NR

CR

LF, B

huta

n

72N

eolis

soch

ilus d

ukai

Tshe

ring,

S.,

Wan

gchu

k, K

., D

orji,

S. &

Nor

bu, P

.20

17Fi

eld

guid

e to

Fis

hes o

f Wes

tern

Bhu

tan,

NR

CR

LF, B

huta

n

73N

eolis

soch

ilus h

exag

onol

epis

Gur

ung,

D. B

., D

orji,

S.,

Tshe

ring,

U. &

Wan

gyel

, J. T

.20

13Jo

urna

l of T

hrea

tene

d Ta

xa 5

(14)

: 488

0-48

86

74N

eolis

soch

ilus h

exas

ticus

Tshe

ring,

S.,

Wan

gchu

k, K

., D

orji,

S. &

Nor

bu, P

.20

17Fi

eld

guid

e to

Fis

hes o

f Wes

tern

Bhu

tan,

NR

CR

LF, B

huta

n

75O

lyra

cf.

kem

pi

Gur

ung,

D. B

., D

orji,

S.,

Tshe

ring,

U. &

Wan

gyel

, J. T

.20

13Jo

urna

l of T

hrea

tene

d Ta

xa 5

(14)

: 488

0-48

86

76O

lyra

pra

estig

iosa

Tshe

ring,

S.,

Wan

gchu

k, K

., D

orji,

S. &

Nor

bu, P

.20

17Fi

eld

guid

e to

Fis

hes o

f Wes

tern

Bhu

tan,

NR

CR

LF, B

huta

n

77O

mpo

k pa

bda

Gur

ung,

D. B

., D

orji,

S.,

Tshe

ring,

U. &

Wan

gyel

, J. T

.20

13Jo

urna

l of T

hrea

tene

d Ta

xa 5

(14)

: 488

0-48

86

78O

ncor

hync

hus m

ykis

s G

urun

g, D

. B.,

Dor

ji, S

., Ts

herin

g, U

. & W

angy

el, J

. T.

2013

Jour

nal o

f Thr

eate

ned

Taxa

5 (1

4): 4

880-

4886

79O

reic

hthy

s cre

nuch

oide

sTs

herin

g, S

., W

angc

huk,

K.,

Dor

ji, S

. & N

orbu

, P.

2017

Fiel

d gu

ide

to F

ishe

s of W

este

rn B

huta

n, N

RC

RLF

, Bhu

tan

80Pa

raca

ntho

cobi

tis a

butw

ebi

Tshe

ring,

S.,

Wan

gchu

k, K

., D

orji,

S. &

Nor

bu, P

.20

17Fi

eld

guid

e to

Fis

hes o

f Wes

tern

Bhu

tan,

NR

CR

LF, B

huta

n

81Pe

thia

con

chon

ius

Tshe

ring,

S.,

Wan

gchu

k, K

., D

orji,

S. &

Nor

bu, P

.20

17Fi

eld

guid

e to

Fis

hes o

f Wes

tern

Bhu

tan,

NR

CR

LF, B

huta

n

82Pe

thia

tict

oTs

herin

g, S

., W

angc

huk,

K.,

Dor

ji, S

. & N

orbu

, P.

2017

Fiel

d gu

ide

to F

ishe

s of W

este

rn B

huta

n, N

RC

RLF

, Bhu

tan

83Ps

eude

chen

eis s

ireni

caTs

herin

g, S

., W

angc

huk,

K.,

Dor

ji, S

. & N

orbu

, P.

2017

Fiel

d gu

ide

to F

ishe

s of W

este

rn B

huta

n, N

RC

RLF

, Bhu

tan

84Ps

eude

chen

eis s

ulca

taG

urun

g, D

. B.,

Dor

ji, S

., Ts

herin

g, U

. & W

angy

el, J

. T.

2013

Jour

nal o

f Thr

eate

ned

Taxa

5 (1

4): 4

880-

4886

85Ps

eudo

lagu

via

feru

la

Gur

ung,

D. B

., D

orji,

S.,

Tshe

ring,

U. &

Wan

gyel

, J. T

.20

13Jo

urna

l of T

hrea

tene

d Ta

xa 5

(14)

: 488

0-48

86

86Ps

ilorh

ynch

us b

alito

ra

Gur

ung,

D. B

., D

orji,

S.,

Tshe

ring,

U. &

Wan

gyel

, J. T

.20

13Jo

urna

l of T

hrea

tene

d Ta

xa 5

(14)

: 488

0-48

86

87Ps

ilorh

ynch

us c

f. ar

unac

hale

nsis

Tshe

ring,

S.,

Wan

gchu

k, K

., D

orji,

S. &

Nor

bu, P

.20

17Fi

eld

guid

e to

Fis

hes o

f Wes

tern

Bhu

tan,

NR

CR

LF, B

huta

n

88Pt

eroc

rypt

is b

arak

ensi

sTs

herin

g, S

., W

angc

huk,

K.,

Dor

ji, S

. & N

orbu

, P.

2017

Fiel

d gu

ide

to F

ishe

s of W

este

rn B

huta

n, N

RC

RLF

, Bhu

tan

89Pu

ntiu

s cho

la

Gur

ung,

D. B

., D

orji,

S.,

Tshe

ring,

U. &

Wan

gyel

, J. T

.20

13Jo

urna

l of T

hrea

tene

d Ta

xa 5

(14)

: 488

0-48

86

81


90Pu

ntiu

s sar

ana

Gur

ung,

D. B

., D

orji,

S.,

Tshe

ring,

U. &

Wan

gyel

, J. T

.20

13Jo

urna

l of T

hrea

tene

d Ta

xa 5

(14)

: 488

0-48

86

91Pu

ntiu

s sop

hore

Tshe

ring,

S.,

Wan

gchu

k, K

., D

orji,

S. &

Nor

bu, P

.20

17Fi

eld

guid

e to

Fis

hes o

f Wes

tern

Bhu

tan,

NR

CR

LF, B

huta

n

92Pu

ntiu

s tic

toG

urun

g, D

. B.,

Dor

ji, S

., Ts

herin

g, U

. & W

angy

el, J

. T.

2013

Jour

nal o

f Thr

eate

ned

Taxa

5 (1

4): 4

880-

4886

93Ra

iam

as b

ola

G

urun

g, D

. B.,

Dor

ji, S

., Ts

herin

g, U

. & W

angy

el, J

. T.

2013

Jour

nal o

f Thr

eate

ned

Taxa

5 (1

4): 4

880-

4886

94Ra

sbor

a da

nico

nius

G

urun

g, D

. B.,

Dor

ji, S

., Ts

herin

g, U

. & W

angy

el, J

. T.

2013

Jour

nal o

f Thr

eate

ned

Taxa

5 (1

4): 4

880-

4886

95Sc

hist

ura

cf. b

eava

niTs

herin

g, S

., W

angc

huk,

K.,

Dor

ji, S

. & N

orbu

, P.

2017

Fiel

d gu

ide

to F

ishe

s of W

este

rn B

huta

n, N

RC

RLF

, Bhu

tan

96Sc

hist

ura

cf. k

angj

upkh

ulen

sis

Tshe

ring,

S.,

Wan

gchu

k, K

., D

orji,

S. &

Nor

bu, P

.20

17Fi

eld

guid

e to

Fis

hes o

f Wes

tern

Bhu

tan,

NR

CR

LF, B

huta

n

97Sc

hist

ura

cf. r

etic

ulof

asci

atus

G

urun

g, D

. B.,

Dor

ji, S

., Ts

herin

g, U

. & W

angy

el, J

. T.

2013

Jour

nal o

f Thr

eate

ned

Taxa

5 (1

4): 4

880-

4886

98Sc

hist

ura

cf. s

avon

aTs

herin

g, S

., W

angc

huk,

K.,

Dor

ji, S

. & N

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Fiel

d gu

ide

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ishe

s of W

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RC

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, Bhu

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isi

Gur

ung,

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., D

orji,

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Wan

gyel

, J. T

.20

13Jo

urna

l of T

hrea

tene

d Ta

xa 5

(14)

: 488

0-48

86

100

Schi

stur

a m

ultif

asci

ata

Gur

ung,

D. B

., D

orji,

S.,

Tshe

ring,

U. &

Wan

gyel

, J. T

.20

13Jo

urna

l of T

hrea

tene

d Ta

xa 5

(14)

: 488

0-48

86

101

Schi

stur

a sc

atur

igin

a G

urun

g, D

. B.,

Dor

ji, S

., Ts

herin

g, U

. & W

angy

el, J

. T.

2013

Jour

nal o

f Thr

eate

ned

Taxa

5 (1

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880-

4886

102

Schi

zoth

orax

mol

esw

orth

i G

urun

g, D

. B.,

Dor

ji, S

., Ts

herin

g, U

. & W

angy

el, J

. T.

2013

Jour

nal o

f Thr

eate

ned

Taxa

5 (1

4): 4

880-

4886

103

Schi

zoth

orax

pro

gast

us

Gur

ung,

D. B

., D

orji,

S.,

Tshe

ring,

U. &

Wan

gyel

, J. T

.20

13Jo

urna

l of T

hrea

tene

d Ta

xa 5

(14)

: 488

0-48

86

104

Sem

iplo

tus s

emip

lotu

sG

urun

g, D

. B.,

Dor

ji, S

., Ts

herin

g, U

. & W

angy

el, J

. T.

2013

Jour

nal o

f Thr

eate

ned

Taxa

5 (1

4): 4

880-

4886

105

Tetr

aodo

n cu

tcut

ia

Gur

ung,

D. B

., D

orji,

S.,

Tshe

ring,

U. &

Wan

gyel

, J. T

.20

13Jo

urna

l of T

hrea

tene

d Ta

xa 5

(14)

: 488

0-48

86

106

Tor p

utito

ra

Gur

ung,

D. B

., D

orji,

S.,

Tshe

ring,

U. &

Wan

gyel

, J. T

.20

13Jo

urna

l of T

hrea

tene

d Ta

xa 5

(14)

: 488

0-48

86

107

Tor t

or

Gur

ung,

D. B

., D

orji,

S.,

Tshe

ring,

U. &

Wan

gyel

, J. T

.20

13Jo

urna

l of T

hrea

tene

d Ta

xa 5

(14)

: 488

0-48

86

108

Xene

ntod

on c

anci

la

Gur

ung,

D. B

., D

orji,

S.,

Tshe

ring,

U. &

Wan

gyel

, J. T

.20

13Jo

urna

l of T

hrea

tene

d Ta

xa 5

(14)

: 488

0-48

86

Am

phib

ia a

nd R

eptil

ia

S.N

o.Sp

ecie

sSc

ient

ist (

s)Ye

arR

efer

ence

1Am

olop

s him

alay

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up, T

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or.,

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. J.

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The

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ulle

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3-18

2Am

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s man

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umW

angy

el, J

. T.

2013

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nal o

f Thr

eate

ned

Taxa

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3): 4

774-

4783

3Am

phie

sma

plat

ycep

sW

angy

el, J

. T.

2013

Jour

nal o

f Thr

eate

ned

Taxa

5 (1

3): 4

774-

4783

4C

alot

es m

aria

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, A.,

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ma,

P.,

Sure

ndra

n, H

., N

ath,

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sh, S

., D

utta

, D.,

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dol,

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gdi,

Y.20

16H

erpe

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gy N

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usW

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. T.

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ngs o

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tan

Ecol

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ea o

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aW

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Ham

adry

ad 3

6, 1

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31

7C

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s ass

amen

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ma,

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Sure

ndra

n, H

., N

ath,

A.,

Gho

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., D

utta

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gdi,

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16H

erpe

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8C

uora

am

boin

ensi

sW

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el, J

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gchu

k, D

. & D

as, I

.20

12C

helo

nian

Con

serv

atio

n an

d B

iolo

gy 1

1, N

umbe

r 2

9C

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., W

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, I.

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Che

loni

an C

onse

rvat

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Bio

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Num

ber 2

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, J. T

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an C

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rvat

ion

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11D

endr

elap

his t

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isW

angy

el, J

. T.

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Ham

adry

ad 3

6, 1

: 25-

31

12D

inod

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Wan

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, J. T

.20

13Jo

urna

l of T

hrea

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d Ta

xa 5

(13)

: 477

4-47

83

13Eu

phly

ctis

cya

noph

lyct

isW

angy

el, J

. T.

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Jour

nal o

f Thr

eate

ned

Taxa

5 (1

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774-

4783

14Fe

jerv

arya

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sis

Wan

gyel

, J. T

.20

13Jo

urna

l of T

hrea

tene

d Ta

xa 5

(13)

: 477

4-47

83

15Fe

jerv

arya

pie

rrei

Wan

gyel

, J. T

.20

13Jo

urna

l of T

hrea

tene

d Ta

xa 5

(13)

: 477

4-47

83

82


16Fe

jerv

arya

tera

iens

isW

angy

el, J

. T.

2013

Jour

nal o

f Thr

eate

ned

Taxa

5 (1

3): 4

774-

4783

17G

ekko

gec

koW

angy

el, J

. T.

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Ham

adry

ad 3

6, 1

: 25-

31

18H

ylar

ana

taip

ehen

sis

Wan

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, J. T

.20

13Jo

urna

l of T

hrea

tene

d Ta

xa 5

(13)

: 477

4-47

83

19In

dote

stud

o el

onga

taW

angy

el, J

. T.,

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gchu

k, D

. & D

as, I

.20

12C

helo

nian

Con

serv

atio

n an

d B

iolo

gy 1

1, N

umbe

r 2

20In

gera

na b

orea

lisD

as, A

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arm

a, P

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rend

ran,

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h, A

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ta, D

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l, J.

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angd

i, Y.

2016

Her

peto

logy

Not

es 9

: 261

-278

21Ly

codo

n au

licus

Wan

gyel

, J. T

.20

12H

amad

ryad

36,

1: 2

5-31

22Ly

codo

n fa

scia

tus

Wan

gyel

, J. T

.20

12H

amad

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36,

1: 2

5-31

23Ly

codo

n ja

raW

angy

el, J

. T.

2012

Ham

adry

ad 3

6, 1

: 25-

31

24M

elan

oche

lys t

rica

rina

taW

angy

el, J

. T.,

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gchu

k, D

. & D

as, I

.20

12C

helo

nian

Con

serv

atio

n an

d B

iolo

gy 1

1, N

umbe

r 2

25M

elan

oche

lys t

riju

gaW

angy

el, J

. T.

2013

Jour

nal o

f Thr

eate

ned

Taxa

5 (1

3): 4

774-

4783

26N

aja

naja

Wan

gyel

, J. T

.20

12H

amad

ryad

36,

1: 2

5-31

27N

anor

ana

anna

ndal

iiW

angy

el, J

. T.

2014

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eedi

ngs o

f Bhu

tan

Ecol

ogic

al S

ocie

ty S

ympo

sium

28N

anor

ana

cona

ensi

sW

angy

el, J

. T.

2013

Jour

nal o

f Thr

eate

ned

Taxa

5 (1

3): 4

774-

4783

29N

anor

ana

ples

kei

Wan

gyel

, J. T

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13Jo

urna

l of T

hrea

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d Ta

xa 5

(13)

: 477

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83

30O

ligod

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enio

latu

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f Bhu

tan

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al S

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ty S

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auru

s gra

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Wan

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l of T

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d Ta

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(13)

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83

32Pt

ycto

laem

us g

ular

isW

angy

el, J

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eedi

ngs o

f Bhu

tan

Ecol

ogic

al S

ocie

ty S

ympo

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33Py

thon

biv

ittat

usW

angy

el, J

. T.

2012

Ham

adry

ad 3

6, 1

: 25-

31

34Ra

mph

otyp

hlop

s bra

min

usW

angy

el, J

. T.

2012

Ham

adry

ad 3

6, 1

: 25-

31

35Rh

abdo

phis

subm

inia

tus

Wan

gyel

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.20

12H

amad

ryad

36,

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5-31

36Sp

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mor

phus

mac

ulat

usW

angy

el, J

. T.

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Ham

adry

ad 3

6, 1

: 25-

31

37Sy

lvir

ana

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uent

heri

Wan

gyel

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13Jo

urna

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hrea

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d Ta

xa 5

(13)

: 477

4-47

83

38Sy

lvir

ana

lept

oglo

ssa

Wan

gyel

, J. T

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13Jo

urna

l of T

hrea

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d Ta

xa 5

(13)

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4-47

83

39Tr

imer

esur

us a

lbol

abri

sW

angy

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adry

ad 3

6, 1

: 25-

31

40Tr

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esur

us p

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riru

mD

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arm

a, P

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rend

ran,

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h, A

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hosh

, S.,

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i, Y.

2016

Her

peto

logy

Not

es 9

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41U

pero

don

glob

ulos

usD

as, A

., Sh

arm

a, P

., Su

rend

ran,

H.,

Nat

h, A

., G

hosh

, S.,

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ta, D

., M

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l, J.

& W

angd

i, Y.

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Her

peto

logy

Not

es 9

: 261

-278

42Xe

noph

rys g

land

ulos

aW

angy

el, J

. T.

2013

Jour

nal o

f Thr

eate

ned

Taxa

5 (1

3): 4

774-

4783

43Xe

noph

rys m

ajor

Wan

gyel

, J. T

.20

13Jo

urna

l of T

hrea

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d Ta

xa 5

(13)

: 477

4-47

83

44Xe

noph

rys m

inor

Wan

gyel

, J. T

.20

13Jo

urna

l of T

hrea

tene

d Ta

xa 5

(13)

: 477

4-47

83

Aves

(Bir

ds)

S.N

o.Sp

ecie

sSc

ient

ist (

s)Ye

arR

efer

ence

1At

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tua

Lim

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, A. &

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, F.

2017

Indi

an B

irds 1

3 (6

)

2C

alca

rius

lapp

onic

usC

hoph

el, T

. & S

heru

b20

16In

dian

Bird

s 12

(1)

3C

alla

cant

his b

urto

niB

isho

p, K

. D.

2009

Fork

tail

25

4Pl

uvia

lis fu

lva

Sing

ye, R

., Jo

hnso

n, A

. & H

edem

ark,

M.

2009

Indi

an B

irds 5

(2)

5Sc

hoen

iclu

s rus

ticus

Lilje

, M.

2017

Indi

an B

irds 1

3 (6

): 16

1

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