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Quantitative Ecology Chap 12

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    CHAPTER 12

    Quantitative ecology

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    Quantitative ecology

    Outline

    Population ecology

    Population dispersion

    Biotic potential

    Factors influencing population growth

    Life history patterns

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    Population ecology

    population consists of the individuals of a species that

    occur together at one place at one time

    three important aspects of population

    i. the range throughout which a population occurs

    ii. dispersion of individuals within that range

    iii. size of the population attains

    Population distributions

    most species have relatively limited geographic range

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    Population dispersion

    Randomly spaced

    not common in nature

    individuals do not interact strongly one another

    Uniformly spaced

    always results from competition for resources

    in animals, it occurs because individuals tend to

    defend their territories which provide them resources

    among plants, results of competition for the sunlight,

    nutrients or water

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    Population dispersion

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    Biotic potential

    to understand populations, we must consider how they

    grow and factors in nature limit population growth

    The exponential growth model

    the actual rate of population increase

    r= (b-d) + (i-e) r= difference between the birthrate (b) and death rate (d)e= movement out of area and i= movement into area

    movement of individuals have a major impact onpopulation growth rates

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    Biotic potential

    biotic potential = rate at which a population of a given

    species will increases when no

    limits are placed on its rate of

    growthdN/dt = riN, N = num. of individuals in population,

    dN/dt = rate of change in its num. over time

    ri = innate capacity for growth

    innate capacity of growth of any population isexponential

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    Biotic potential

    Carrying capacity

    how rapidly population grows, they will reach limit

    imposed by several factors such as space, light, water..

    carrying capacity,K = maximum number ofindividuals that environment can support

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    Biotic potential

    Logistic growth model

    as population reaches its carrying capacity, its rate of

    growth slows greatly because lack of resources

    dN/dt = rN(K-N/K)

    as N increases, the fraction by r is multiplied becomes

    smaller, and the rate population increases decline

    in many cases, real population display trendscorresponding to a logistic growth curve

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    Models of population growth

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    Population growth is limited by the

    environment

    population dynamics - study of factors that affectgrowth, stability and decline of population known

    population undergo three distinct phase

    i. growthii. stability

    iii. decline

    population growth occurs when resources exceed thenumber of individuals able to use them

    reproduction is rapid and death rates are low, increasein population size

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    Population growth is limited by the

    environment

    population stability is always the longest phase,

    growing population outstrips its available resources,

    decline occurs because decrease in the individuals

    number in a population, lead to extinctionFactors influencing population growth

    as long resources available, almost all populations will

    ten to grow exponential affect by birth rate and death rate

    intrinsic rate = birth rate death rate

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    The influence of population density

    density-dependent = population size affect populationgrowth rate

    when population approach their carrying capacity,

    competition can be severe, increased risk of mortalityand decrease birthrate

    predators tend to focus on particularly common prey,increased rates of mortality as population increase

    high population led to accumulation of toxic wastes

    behavioral changes also affect population growth rates,results from hormonal actions

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    The influence of population density

    crowded populations may led to decreased in

    population growth rate because of emigration factor

    in some cases, growth rate increase with increase

    population size (Allee effect) individuals are difficult finding mates in population

    that are too sparsely distributed, some species rely on

    large groups to deter predator

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    The influence of population density

    Density-independent effects

    rate of growth of a population at any instant is limited

    by something other that the size of population

    factors such as extremely cold winter, drought, storms,volcanic eruptions may affects populations

    when such events occurs, population extremely

    decrease and will increase rapidly when environmentback to normal

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    The influence of population density

    Survivorship curves

    survivorship is defined as the percentage of an originalpopulation that survives to a given age

    Type I

    - flat at early and middle life and drops suddenlyas death rate increases among the elderly

    - associated with species such as humans and otherlarge mammals that produce few offspring that

    are well cared for.Type II

    - intermediate, mortality more constant over lifespans, can be seen in Hydra and the grey

    squirrels

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    The influence of population density

    Type III

    - very high death rates for the young, followed bylower death rates

    - often associated with organisms, such as oysters,that produce very many offspring but

    provide little or no care.

    some invertebrates show a "stair-stepped" curve withbrief periods of high mortality during molts, followed

    by periods of lower mortality when the exoskeleton ishard.

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    Survivorship curves

    1

    2

    1

    3

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    Life history patterns

    the logistic population model predicts two main lifehistory patterns.

    most populations cannot be characterized as either r-orK-strategists; they have intermediate characteristics.

    r-Selectiona. species that undergo selection to maximize their

    rate of natural increase

    b. often opportunistic species, tend to be colonizers.

    c. strategy for continued existence is based onindividuals having the following traits: (small size,short life span, mature fast, produce manyoffspring, engage in little caring of offspring)

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    Life history patterns

    d. such populations usually exhibit a survivorshipcurve similar to type III.

    e. thus, they rely on rapid dispersal to newunoccupied environments.

    K-Selectiona. species that hold their populations fairly constant

    near the carrying capacity

    b. such populations are equilibrium species, tend to

    be specialists rather than colonizers, and maybecome extinct when their evolved way of life isdisrupted (grizzly bear, Florida panther, etc.).

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    Life history patterns

    c. overall strategy for continued existence is based onhaving the following traits: (large size, long lifespan, slow to mature, produce few offspring,expend considerable energy in care)

    d. such populations usually have a survivorshipcurve similar to type I.

    e. thus, they rely on competitive superiority tosecure limited resources.

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    Life history patterns

    Population pyramids

    at the beginning of twentieth century, humanpopulation grow rapidly

    different countries shows different growth rate (eg.Mexico grow rapidly)

    rate which population can be expected to grow in thefuture can be represent by population pyramids

    human population pyramid displays the age population

    by sex, female in most region have longer lifeexpectancy than male

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    Population pyramids

    1

    2

    1


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