+ All Categories
Home > Documents > Zabrus Pelor skoupyi sp. nov. (Coleoptera: Carabidae) from ... · 22 T XONOMIC P RT Zabrus (Pelor)...

Zabrus Pelor skoupyi sp. nov. (Coleoptera: Carabidae) from ... · 22 T XONOMIC P RT Zabrus (Pelor)...

Date post: 14-Sep-2020
Category:
Upload: others
View: 2 times
Download: 0 times
Share this document with a friend
8
21 Klapalekiana, 47: 21–28, 2011 ISSN 1210-6100 Zabrus (Pelor) skoupyi sp. nov. (Coleoptera: Carabidae) from Greece Zabrus (Pelor) skoupyi sp. nov. (Coleoptera: Carabidae) z Řecka Jiř ��J�JWolkerova �11�8, �-�8 01 Kra�li�e, �e�ep�li� e-�ail: a�ara�voln�11�8, �-�8 01 Kra�li�e, �e�ep�li� e-�ail: a�ara�volnTaxonomy, Coleoptera, Carabidae, �abrini, �abrini, Zabrus, new species, distribution, Palaearctic region Abstract. Zabrus (Pelor) skoupyi sp. nov. from northwestern Greece is �escri�e�re�n� comp�re� with is �escri�e�re�n� comp�re� with rel�te� B�lk�n species of the s�en�s Pelor Bonelli, 1810� Te new �pe�ie� live� in te �ra��o�ro�o�ntain�ra��o�ro�o�ntainin te alpine �eadow� at an altit�de of 2000–2200 � a�l� Zabrus (P.) albanicus �pfel�eck 1904 s. l�t. is recor�e� a� new for �ree�e� New data on te di�tri�tion of Z. (P.) aetolus S�a�, 186� lat�, Z. (P.) brevicollis S�a�, 18�� and Z. (P.) peristericus peristericus �pfel�eck 1901 �re presente�. are pre�entedINT��DU�TI�N �i�ht t�x� (species or s�species) of the s�en�s Pelor Bonelli 1810 of the �en�s Zabrus Cl�irville 1806 �re known from continent�l northwestern Greece (Fre�e 1989): Z. aetolus matejkai M�ř�n 1940 (Vermion Oros Mo�nt�ins); Z. aetolus purkynei M�ř�n 1940 (type loc�lity in Mo�nt �jm�kč�l�n on the �or�er �etween Greece �n� the Rep�lic of M�ce�o- nia) Z. brevicollis S�a�, 18� (nortwe�tern and �entral �ree�e) Z. ganglbaueri gan� glbaueri �pfel�eck 1906 (�l�ni� �n� northwestern Greece); Z. graecus subtilis S�a�, 1862 (nortwe�tern and �entral �ree�e) Z. incrassatus (�hrens 1814) (B�lk�n Penins�l�); Z. peristericus �pfel�eck 1901 (�kmos Oros Mo�nt�ins); �n�kmos Oros Mo�nt�ins); �n and Z. spinipes insignis �ül- ler 1932 (Greece). �n ��ition�l species of the s�en�s Pelor fro� te �entioned area i�escri�e�elow. �lso �nother species of this s�en�s is recor�e� from Greece for the rst ti�e in ti� paper M�T�RI�ND M�T�ODS The specimens of new species �re �eposite� in the followin� collections: coll. G. D�lt S�vi�ny-s�r-Or�e Fr�nce; coll. J. �ejk�l �r�slice Czech Rep�lic; coll. P. �r�š� S��rlo�is Germ�ny; coll. �. Orsz�lik Frý�ek-Mstek Czech Rep��lic; coll. V. Sko�pý Žilin� Czech Rep�lic; coll. P. Štěpánek �l�no Czech Rep�lic. Me�s�rements were m�e with � MBS–10 stereo microscope �t m��nic�tions of 8× to 56×. Me�s�rements of �o�y p�rts �n�o�y r�tios �se� in the text were t�ken �s follows: tot�l len�th = len�th me�s�re� from �nterior m�r�in of l�r�m to elytron �pex; len�th of me�i�n lo�e (penis) = len�th of me�i�n lo�e in �ors�l view; wi�th of me�i�n lo�e (penis) = m�xim�l wi�th of me�i�n lo�e in �ors�l view; r�tio of wi�th/len�th of pronot�m = r�tio of m�xim�l wi�th of pronot�m/len�th of pronot�m me�s�re�lon� mi�-line; r�tio of �nterior m�r�in/posterior m�r�in of pronot�m = r�tio of wi�th of pronot�m �etween �nterior �n�les/wi�th of pronot�m �etween posterior �n�les; r�tio of len�th/wi�th of elytr� = r�tio of len�th of lon�er elytron me�s�re� from �s�l �or�er to �pex/m�xim�l wi�th of �oth elytr� com�ine�; r�tio of len�th/wi�th of �pex of me�i�n lo�e (�pex of penis) = r�tio of len�th of �pex me�s�refrom �pic�l �or�er of intern�l s�c to tip of me�i�n lo�e/m�xim�l wi�th of �pex in �ors�l view. P��lishe� M�y 5 2011
Transcript
Page 1: Zabrus Pelor skoupyi sp. nov. (Coleoptera: Carabidae) from ... · 22 T XONOMIC P RT Zabrus (Pelor) skoupyi sp. nov. (Fi s 1–5 10 11) Type material. olotype:: ♂ l elle “Gr. M

21

Klapalekiana, 47: 21–28, 2011ISSN 1210-6100

Zabrus (Pelor) skoupyi sp. nov. (Coleoptera: Carabidae) from Greece

Zabrus (Pelor) skoupyi sp. nov. (Coleoptera: Carabidae) z Řecka

Jiří� ��J��� ��J���

Wolkerova �11��8, ��-��8 01 Kra�li�e, ��e��� �ep��li��� e-�ail: a�ara�voln���� �11��8, ��-��8 01 Kra�li�e, ��e��� �ep��li��� e-�ail: a�ara�voln����

Taxonomy, Coleoptera, Carabidae, �abrini, �abrini,, Zabrus, new species, distribution, Palaearctic region

Abstract. Zabrus (Pelor) skoupyi sp. nov. from northwestern Greece is �escri�e��� ����re��� �n� comp�re� withis �escri�e��� ����re��� �n� comp�re� with rel�te� B�lk�n species of the s���en�s Pelor Bonelli, 1810� T��e new �pe�ie� live� in t��e �ra��o� �ro� �o�ntain��ra��o� �ro� �o�ntain� in t��e alpine �eadow� at an altit�de of 2000–2200 � a���l� Zabrus (P.) albanicus �pfel�eck�� 1904 s. l�t. is recor�e� a� new for �ree�e� New data on t��e di�tri��tion of Z. (P.) aetolus S���a��, 186� �� lat�, Z. (P.) brevicollis S���a��, 18�� and Z. (P.) peristericusperistericus �pfel�eck�� 1901 �re presente�.are pre�ented�

INT��DU�TI�N

�i�ht t�x� (species or s��species) of the s���en�s Pelor Bonelli�� 1810 of the �en�s Zabrus Cl�irville�� 1806 �re known from continent�l northwestern Greece (Fre��e 1989): Z. aetolus matejkai M�ř�n�� 1940 (Vermion Oros Mo�nt�ins); Z. aetolus purkynei M�ř�n�� 1940 (type loc�lity in Mo�nt ��jm�kč�l�n on the �or�er �etween Greece �n� the Rep��lic of M�ce�o-nia)�� Z. brevicollis S���a��, 18�� (nort��we�tern and �entral �ree�e)�� Z. ganglbaueri gan�glbaueri �pfel�eck�� 1906 (�l��ni� �n� northwestern Greece); Z. graecus subtilis S���a��, 1862 (nort��we�tern and �entral �ree�e)�� Z. incrassatus (�hrens�� 1814) (B�lk�n Penins�l�); Z. peristericus �pfel�eck�� 1901 (��kmos Oros Mo�nt�ins); �n���kmos Oros Mo�nt�ins); �n��� and Z. spinipes insignis �� �ül-ler�� 1932 (Greece). �n ���ition�l species of the s���en�s Pelor fro� t��e �entioned area i� �escri�e� �elow. �lso�� �nother species of this s���en�s is recor�e� from Greece for the ��rst ti�e in t��i� paper�

M�T�RI�� �ND M�T�ODS

The specimens of new species �re �eposite� in the followin� collections: coll. G. D����lt�� S�vi�ny-s�r-Or�e�� Fr�nce; coll. J. �ejk�l�� �r�slice�� Czech Rep��lic; coll. P. �r�š��� S��rlo�is�� Germ�ny; coll. �. Orsz�lik�� Frý�ek-Mí�stek�� Czech Rep��lic; coll. V. Sko�pý�� Žilin��� Czech Rep��lic; coll. P. Štěpánek�� �l��no�� Czech Rep��lic.

Me�s�rements were m��e with � MBS–10 stereo microscope �t m��ni��c�tions of 8× to 56×. Me�s�rements of �o�y p�rts �n� �o�y r�tios �se� in the text were t�ken �s follows: tot�l len�th = len�th me�s�re� from �nterior m�r�in of l��r�m to elytron �pex; len�th of me�i�n lo�e (penis) = len�th of me�i�n lo�e in �ors�l view; wi�th of me�i�n lo�e (penis) = m�xim�l wi�th of me�i�n lo�e in �ors�l view; r�tio of wi�th/len�th of pronot�m = r�tio of m�xim�l wi�th of pronot�m/len�th of pronot�m me�s�re� �lon� mi�-line; r�tio of �nterior m�r�in/posterior m�r�in of pronot�m = r�tio of wi�th of pronot�m �etween �nterior �n�les/wi�th of pronot�m �etween posterior �n�les; r�tio of len�th/wi�th of elytr� = r�tio of len�th of lon�er elytron me�s�re� from ��s�l �or�er to �pex/m�xim�l wi�th of �oth elytr� com�ine�; r�tio of len�th/wi�th of �pex of me�i�n lo�e (�pex of penis) = r�tio of len�th of �pex me�s�re� from �pic�l �or�er of intern�l s�c to tip of me�i�n lo�e/m�xim�l wi�th of �pex in �ors�l view.

P��lishe� M�y 5�� 2011

Page 2: Zabrus Pelor skoupyi sp. nov. (Coleoptera: Carabidae) from ... · 22 T XONOMIC P RT Zabrus (Pelor) skoupyi sp. nov. (Fi s 1–5 10 11) Type material. olotype:: ♂ l elle “Gr. M

22

T�XONOMIC P�RT

Zabrus (Pelor) skoupyi sp. nov. (Fi�s 1–5�� 10�� 11)

Type material. �olotype:: ♂�� l��elle� “Gr. M�ce�.�� �etomilits� – Gr�mos mts.�� 23.6.2004�� �lpin�� Sko�pý le�.”�� collecte� in northwestern Greece�� �t the �or�er �etween �pir�s �n� West M�ce�oni� peripheries�� Gr�mmos Oros Mo�nt�ins�� north of �etomilits��� �lpine me��ows�� �n�er stone ne�r snow-��el��� c�. 2100 m �.s.l. (Fi�. 1 �n� V. Sko�pý�� pers. comm.)�� c�. 40�20���20��20���� N�� 20�50�� ��� coll. V. Sko�pý. P�r�types: Nos 1–6�� 4�50�� ��� coll. V. Sko�pý. P�r�types: Nos 1–6�� 450�� ��� coll. V. Sko�pý. P�r�types: Nos 1–6�� 4�� ��� coll. V. Sko�pý. P�r�types: Nos 1–6�� 4 ��� coll. V. Sko�pý. P�r�types: Nos 1–6�� 4 ♂♂, 2 ♀♀�� l��elle� “GR��C� occ.�� Gr�mos“GR��C� occ.�� Gr�mos Mts.�� �etomilits� env.�� 29.v.2001�� �lpin�� 2000–2200 m�� S. Bene�ikt le�.”�� 3 km northwest of29.v.2001�� �lpin�� 2000–2200 m�� S. Bene�ikt le�.”�� 3 km northwest of–2200 m�� S. Bene�ikt le�.”�� 3 km northwest of2200 m�� S. Bene�ikt le�.”�� 3 km northwest of”�� 3 km northwest of �etomilits��� mo�nt�in r�n�e�� e�stern slope�� �n�er stones ne�r sm�ll snow-��el�s (S. Bene�ikt�� pers. comm.)�� coll. J. �ejk�l; Nos �–12�� 4J. �ejk�l; Nos �–12�� 4�� No� �–12, �–12, �12, � ♂♂♂, 2 ♀♀�� s�me ��t� �s Nos 1–6�� coll. V. Sko�pý; Noss�me ��t� �s Nos 1–6�� coll. V. Sko�pý; Nos–6�� coll. V. Sko�pý; Nos6�� coll. V. Sko�pý; Nos 1�–1�, 2–1�, 21�, 2 ♂♂♂, �a�e data a� No� 1–6, �oll� �� D��a�lt�� No� 1�–1�, 2–6, �oll� �� D��a�lt�� No� 1�–1�, 26, �oll� �� D��a�lt�� No� 1�–1�, 2–1�, 21�, 2 ♂♂, 1 ♀, la�elled “�ree�e, Ioánin��� M. Gr�moz�� �etomilits��� 29.5.2001�� 2000 m�� P. �r�s� le�.”�� coll. P. �r�š�; Nos 18–29��–29��29�� 8 ♂♂♂, � ♀♀�� s�me ��t� �s holotype�� coll. V. Sko�pý; Nos 30–31�� 1–�1, 1�1, 1 ♂, 1 ♀, �a�e data a� ��olot�pe,�a�e data a� ��olot�pe, coll. J. �ejk�l; Nos 32–34�� 1–��, 1��, 1 ♂, 2 ♀♀�� l��elle� “Gr. M�ce�.�� �etomilits� – Gr�mos�� 14.6.2006��l��elle� “Gr. M�ce�.�� �etomilits� – Gr�mos�� 14.6.2006��“Gr. M�ce�.�� �etomilits� – Gr�mos�� 14.6.2006��Gr. M�ce�.�� �etomilits� – Gr�mos�� 14.6.2006�� Sko�pý le�.”�� coll. V. Sko�pý; Nos 35–36�� 1”�� coll. V. Sko�pý; Nos 35–36�� 1�� coll. V. Sko�pý; Nos 35–36�� 1–�6, 1�6, 1 ♂, 1 ♀�� l��elle� “Gr. M�ce�.���etomilits� – Gr�mos��l��elle� “Gr. M�ce�.���etomilits� – Gr�mos��“Gr. M�ce�.���etomilits� – Gr�mos��Gr. M�ce�.�� �etomilits� – Gr�mos�� 14.6.2006�� Štěpánek le�.”�� coll. P. Štěpánek; Nos 3�–38�� 1”�� coll. P. Štěpánek; Nos 3�–38�� 1�� coll. P. Štěpánek; Nos 3�–38�� 1–�8, 1�8, 1 ♂, 1 ♀�� l��elle� “GR��C� �or. occ.��l��elle� “GR��C� �or. occ.��“GR��C� �or. occ.��GR��C� �or. occ.�� Gr�mmos mts.�� �etomilis��� 6.6.2008�� 2100 m�� l�t. Orsz�lik”�� coll. �. Orsz�lik.”�� coll. �. Orsz�lik., �oll� K� �r���lik�Description. Win�less species with me�i�m-size� �n� rel�tively n�rrow �o�y (Fi�. 2). Tot�l len�th 12.9–14.5 mm�� mostly 13.1–13.5 mm (holotype 14.0 mm�� �ver��e 13.5 mm��–14.5 mm�� mostly 13.1–13.5 mm (holotype 14.0 mm�� �ver��e 13.5 mm��14.5 mm�� mostly 13.1–13.5 mm (holotype 14.0 mm�� �ver��e 13.5 mm��–13.5 mm (holotype 14.0 mm�� �ver��e 13.5 mm��13.5 mm (holotype 14.0 mm�� �ver��e 13.5 mm�� n = 39). Dors�l si�e ��rk �rown. ���r�m re�-�rown to ��rk �rown. Dors�l si�e shiny in m�les �n� with sli�ht l�stre to ��ll with elytr� less shiny th�n he�� �n� pronot�m in fem�les. Ventr�l si�e re�-�rown to ��rk �rown. M�n�i�les re�-�rown to ��rk �rown. �ntenn�e �n� le�s re�-�rown to �rown. M�xill�ry �n� l��i�l p�lpi p�le re�-�rown to re�-�rown.

�e�� thickene��� sp�rsely to �ensely �n� very ��nely p�nct�te (p�nct�tion visi�le in 30× m��ni��c�tion) with eyes short �n� sli�htly (in holotype) to m�rke�ly convex. �nterior m�r�in of l��r�m with six setifero�s p�nct�res. �nterior p�rt of clype�s with �rc��te c�ll�s. Front�l f�rrows short�� mostly n�rrow �n� sh�llow. �ntenn�e re�chin� ��se of pronot�m when recli-ne�. �ntennomere 1 very �ro��. Ment�m with simple (i.e. not �i���)�� ro�n�e� (exception�lly rect�n��l�r) �n� �ro�� me�i�n tooth.

Pronot�m tr�nsverse (r�tio of wi�th/len�th = 1.56–1.66�� holotype 1.62)�� stron�ly convex��–1.66�� holotype 1.62)�� stron�ly convex��1.66�� holotype 1.62)�� stron�ly convex�� wi�est �ehin� mi��le�� m�rke�ly n�rrowe� tow�r� �nterior �n�les (r�tio of �nterior m�r�in/posterior m�r�in = 0.�4–0.80�� holotype 0.��)�� l�ter�l m�r�ins sli�htly convex (in holotype)–0.80�� holotype 0.��)�� l�ter�l m�r�ins sli�htly convex (in holotype)0.80�� holotype 0.��)�� l�ter�l m�r�ins sli�htly convex (in holotype) to str�i�ht �efore posterior �n�les. �nterior m�r�in of pronot�m very sh�llowly em�r�in�te�� �nterior �n�les sli�htly protr��in��� ro�n�e�. Posterior �n�les m�rke�ly (in holotype) to sli�htly o�t�se�� with �pex shortly ro�n�e�. Pronot�l ��se sh�llowly em�r�in�te (exception�lly �lmost str�i�ht in p�r�type No 30). Inner ��s�l impression sh�llow �n� wi�e (in holotype) or ��sent. O�ter ��s�l impression ��sent. �nterior p�rt of pronot�m sp�rsely to �ensely �n� ��nely p�nct�te. ��ter�l m�r�in of pronot�m n�rrowly fl�ttene� (fl�ttenin� mo�er�tely wi�ene� posteri��)�� very sp�rsely �n� ��nely p�nct�te�� witho�t set�. Pronot�l ��se �ensely to sp�rsely (in holotype) �n� ��nely to co�rsely p�nct�te. ��ter�l �e�� of pronot�m n�rrow�� not thickene�. �piple�ron of pronot�m smooth (mostly) or very sp�rsely �n� ��nely p�nct�te.

�lytr� rel�tively n�rrow (wi�th = 5.3–6.3 mm�� holotype 6.0 mm) �n� elon��te (r�tio of–6.3 mm�� holotype 6.0 mm) �n� elon��te (r�tio of6.3 mm�� holotype 6.0 mm) �n� elon��te (r�tio of len�th/wi�th = 1.48–1.5��� holotype 1.53)�� very stron�ly convex�� �eclivity of elytr� �efore–1.5��� holotype 1.53)�� very stron�ly convex�� �eclivity of elytr� �efore1.5��� holotype 1.53)�� very stron�ly convex�� �eclivity of elytr� �efore

Page 3: Zabrus Pelor skoupyi sp. nov. (Coleoptera: Carabidae) from ... · 22 T XONOMIC P RT Zabrus (Pelor) skoupyi sp. nov. (Fi s 1–5 10 11) Type material. olotype:: ♂ l elle “Gr. M

2�

�pex steep to �lmost vertic�l in l�ter�l view. Interv�ls sli�htly convex (in holotype) or fl�t. Stri�e �istinct�� mo�er�tely �eepene��� very ��nely (in holotype) to ��nely p�nct�te. Row of seti-fero�s p�nct�res in stri� 8 (or in elytr�l interv�l 9) mostly consi�er��ly interr�pte�. Sc�tell�r stri� mostly contin�o�s�� to�chin� (in holotype) or not to�chin� stri� 1. Sc�tell�r setifero�s p�nct�re ��sent. B�se of elytr� sli�htly sin��te to str�i�ht�� ��s�l �e�� sli�htly sin��te. �lytr�l h�meri o�t�se�� ro�n�e� (in holotype) or �n��l�te�� h�mer�l tooth ��sent. ��ter�l �roove of elytron n�rrow. Microsc�lpt�re consistin� of re��l�r meshes�� co�rser in fem�les. �piple�ron of el�tron ��oot��, ���all� wit�� row of ���ort i�pre��ion��

Ventr�l p�rt of thor�x �n� ���omen. Prostern�m in mi��le smooth�� with sh�llow to �eep tr�nsvers�l impression. Prostern�l process not (in holotype) or very sh�llowly to sh�llowly

Fi�. 1. M�p of Greece in�ic�tin� loc�lity of Zabrus skoupyi �p� nov�O�r. 1. M�p� Řeck� s vyzn�čeno� lok�lito� Zabrus skoupyi �p� nov�

Page 4: Zabrus Pelor skoupyi sp. nov. (Coleoptera: Carabidae) from ... · 22 T XONOMIC P RT Zabrus (Pelor) skoupyi sp. nov. (Fi s 1–5 10 11) Type material. olotype:: ♂ l elle “Gr. M

2�

�e��e� (�e�� interr�pte� �pic�lly)�� with 4–8 set�e �nteriorly. Mesothor�x �n� met�thor�x in–8 set�e �nteriorly. Mesothor�x �n� met�thor�x in8 set�e �nteriorly. Mesothor�x �n� met�thor�x in mi��le smooth. �pistern�m of prothor�x very sp�rsely to sp�rsely �n� very ��nely to ��nely p�nct�te. �pistern�m of mesothor�x sp�rsely to �ensely (in holotype) �n� co�rsely to very co�rsely p�nct�te. �pistern�m of met�thor�x very sp�rsely �n� ��nely (in holotype) to co�rsely p�nct�te�� shorter th�n wi�e. Si�es of ���omin�l ventrites 1 �n� 2 sp�rsely �n� co�rsely to ��nely p�nct�te�� si�es of ���omin�l ventrite 3 very sp�rsely to sp�rsely �n� ��nely (in holotype) to �oar�el� p�n�tate, �ide� of a�do�inal ventrite� � and � ��oot�� (���all�) or ver� �par�el� and ��nely p�nct�te�� si�es of ���omin�l ventrite 6 smooth to �lmost smooth. Si�es of ���omin�l ventrites 3 to 5 �n� �lmost entire ���omin�l ventrite 6 wrinkle�. ���omin�l ventrites 3 to 5 wit�� one pair of �etifero�� p�n�t�re�� �ale� wit�� two, fe�ale� wit�� fo�r �etifero�� p�n�t�re� �t posterior m�r�in of l�st ���omin�l ventrite.

�e�s rel�tively lon� �n� slen�er. Ventr�l si�e of fore femor� with 3–5 setifero�s p�nct�-–� �etifero�� p�n�t�-� �etifero�� p�n�t�-res �ehin� �nterior m�r�in. Ventr�l si�e of mi��le femor� with 4–� (mostly 5–6) setifero�s–� (�o�tl� �–6) �etifero��� (�o�tl� �–6) �etifero��–6) �etifero��6) �etifero�� p�nct�res �efore posterior m�r�in. Ventr�l si�e of hin� femor� with 3–5 setifero�s p�nct�res–� �etifero�� p�n�t�re�� �etifero�� p�n�t�re� �efore posterior m�r�in.

�e�e���s. Me�i�n lo�e me�i�m-size� (len�th 4.10–4.31 mm�� wi�th 1.20–1.30 mm; holo-–���1 ��, widt�� 1�20–1��0 ���� ��olo-���1 ��, widt�� 1�20–1��0 ���� ��olo-–1��0 ���� ��olo-1��0 ���� ��olo-type 4.31 �n� 1.30 mm)�� left si�e sh�llowly em�r�in�te �efore �pex �ors�lly (Fi�s 4 �n� 5). �pex of me�i�n lo�e elon��te �n� n�rrow ��ove (r�tio of len�th/wi�th = 1.33–1.44; holotype–1����� ��olot�pe1����� ��olot�pe 1.44)�� sli�htly c�rve� to left. Ri�ht p�r�mere with lon� hook (Fi�s 3 �n� 5).

Styli l�r�e�� l�st stylomere ton��e-sh�pe� (Fi�s 10 �n� 11).Differential diagnosis. Zabrus skoupyi �p� nov� i� a di�tin�t �pe�ie� wit�� a �niq�e �od� ���ape� It is e�sily �istin��ish��le from �ll species of the s���en�s Pelor fro� nort��we�tern �ree�e ��fro� nort��we�tern �ree�e ���� its very �ifferent h��it�s �n� the extremely n�rrow l�ter�l �e�� of pronot�m. The most simil�r �pe�ie�, Z. peristericus �pfel�eck�� 1901�� �iffers in h�vin� the pronot�m more n�rrowe� tow�r� posterior �n�les�� pronot�m wi�est �efore or �t mi��le�� l�ter�l m�r�ins of pronot�m sli�htly sin��te to str�i�ht �efore posterior �n�les. Z. peristericus �iffers in the sh�pe of the �e�e���s (Fi�. 9; �pex of me�i�n lo�e of Z. peristericus is �ro��er)�� �n� its l�r�er �ver��e size (13.8–1�.0 mm��– 1��0 ��,1��0 ��, �ver��e 15.2 mm�� n = 35). The two symp�tric species �iffer �s follows: Z. albanicus �pfel�eck�� 1904 s. l�t. in h�vin� pronot�m �n� elytr� �ro��er�� posterior �n�les of pronot�m more sh�rp�� �n� in the sh�pe of �e�e���s (Fi�. 6; me�i�n lo�e of Z. albanicus i� ���orter and narrower)��)�� Z. brevicollis Sch��m�� 185� in h�vin� pronot�m �n� elytr� �ro��er�� pronot�m more n�rrowe� tow�r� posterior �n�les�� in the sh�pe of �e�e���s (Fi�. �; �pex of me�i�n lo�e of Z. brevicollis is shorter �n� �ro��er)�� �n� l�r�er size (14.�–18.2 mm). The rem�inin� species from north-–18.2 mm). The rem�inin� species from north-18.2 mm). The rem�inin� species from north-nort��-western Greece �iffer especi�lly in the followin� ch�r�cters: Z. aetolus S���a��, 186� �� lat� in h�vin� pronot�m �n� elytr� �ro��er�� posterior �n�les of pronot�m with �istinct �pex�� �n� in the sh�pe of �e�e���s (Fi�. 8; �pex of me�i�n lo�e of Z. aetolus i� �roader)�� Z. ganglbaueri �pfel�eck�� 1906 s. l�t. in h�vin� pronot�m �n� elytr� m�ch �ro��er�� �n� pronot�m wi�est �lo�e to it� �a�e�� Z. graecus Deje�n�� 1828 s. l�t. in h�vin� pronot�m �n� elytr� m�ch �ro��er�� and pronot�� not narrowed po�teriad�� Z. incrassatus (�hrens�� 1814) in h�vin� pronot�m �n� el�tra ����� �roader�� Z. spinipes (F��rici�s�� 1�98) s. l�t. in h�vin� pronot�m �n� elytr� m�ch �ro��er�� elytr� �lmost smooth (witho�t �istinct stri�e)�� �n� l�r�er size (16.8–22.5 mm).–22�� ��)�22�� ��)�Distribution and habitat. Zabrus skoupyi �p� nov� wa� repeatedl� fo�nd in t��e �ra��o��ra��o� Oros Mo�nt�ins�� north or northwest of the vill��e of�etomilits��� �t the �or�er �etween �pir�snorthwest of the vill��e of�etomilits��� �t the �or�er �etween �pir�sof the vill��e of �etomilits��� �t the �or�er �etween �pir�s �n� West M�ce�oni� peripheries (Fi�. 1). The in�ivi���ls were collecte� in �lpine me��ows ne�r snow-��el�s �t �n �ltit��e of c�. 2000–2200 m �.s.l.at an altit�de of �a� 2000–2200 � a���l�–2200 � a���l�2200 � a���l� Zabrus skoupyi �p� nov� �o�t likel� in��a�it� onl� a ��all area�

Page 5: Zabrus Pelor skoupyi sp. nov. (Coleoptera: Carabidae) from ... · 22 T XONOMIC P RT Zabrus (Pelor) skoupyi sp. nov. (Fi s 1–5 10 11) Type material. olotype:: ♂ l elle “Gr. M

2�

Fi�. 2. Zabrus skoupyi �p� nov�, ��olot�pe, ��a�it��, dor�al view���r� 2� Zabrus skoupyi sp. nov.�� holotyp�s�� h��it�s�� �orsální� pohle�.

Page 6: Zabrus Pelor skoupyi sp. nov. (Coleoptera: Carabidae) from ... · 22 T XONOMIC P RT Zabrus (Pelor) skoupyi sp. nov. (Fi s 1–5 10 11) Type material. olotype:: ♂ l elle “Gr. M

26

I i�enti��e� these species of the s���en�s Pelor fro� alpine and ���alpine �one� of t��e �ra��o� �ro� �o�ntain� and near�� �o�ntain area� in nort��we�tern �ree�e:�o�ntain� and near�� �o�ntain area� in nort��we�tern �ree�e: nort��we�tern �ree�e: Z. aetolus s. l�t. (Vor�s Oros Mo�nt�ins�� Vermion Oros Mo�nt�ins�� Vernon Oros Mo�nt�ins�� Trikl�rioVernon Oros Mo�nt�ins�� Trikl�rio Mo�nt�ins�� Smolik�s Oros Mo�nt�ins�� �n� Vori� Pin�os mo�nt�in r�n�e�� ��tár� P�ss); �n� Vori� Pin�os mo�nt�in r�n�e�� ��tár� P�ss); Z. albanicus s. l�t. (Vernon Oros Mo�nt�ins�� Gr�mmos Oros Mo�nt�ins�� Smolik�s Oros�o�ntain�, �ra��o� �ro� �o�ntain�, S�olika� �ro� Mo�nt�ins�� �n� Noti� Pin�os mo�nt�in r�n�e�� west of Trí�k�l�);Noti� Pin�os mo�nt�in r�n�e�� west of Trí�k�l�); Z. brevicollis (�ra��o��ra��o� �ro� �o�ntain�)�� Z. incrassatus (Tym�� Mo�nt�ins�� ��kmos /= Peristeri/ Oros Mo�nt�ins);�� ��kmos /= Peristeri/ Oros Mo�nt�ins);��kmos /= Peristeri/ Oros Mo�nt�ins);)�� Z. peristericus (Tym�� Mo�nt�ins�� ��kmos /= Peristeri/ Oros Mo�nt�ins�� Tso�merk� Mo�n-T�o��erka �o�n-t�ins�� �n� Noti� Pin�os mo�nt�in r�n�e�� west of Trí�k�l�); �n��� and Z. skoupyi �p� nov� (�ra��o��ra��o� Oros Mo�nt�ins). Fre��e (1989) mentione� the occ�rrence of Z. brevicollis in the ��tár� P�ss �n� the Smolik�s Oros Mo�nt�ins. Speciest��e S�olika� �ro� �o�ntain�� Spe�ie� Z. graecus, Z. incrassatus and Z. spinipes are �ore �o��on in t��e �teppe and fore�t �one��Notes. Z. albanicus wa� not �et re�orded fro� �ree�e� It i� a widel� di�tri��ted pol�t�pi� species hitherto known from �l��ni��� �osovo �n� the Rep��lic of M�ce�oni� (�pfel�eck�n� the Rep��lic of M�ce�oni� (�pfel�eck 1904�� G�n�l���er 1915�� M�ř�n 1939�� �ieke 1981�� Fre��e 1989�� G�éor��iev 200��� etc.). Serr�no & �n�új�r (2003) liste� Z. albanicus jablanicensis M�ř�n�� 1939 incorrectly from Bosni� �n� �erze�ovin�. This s��species w�s �escri�e� from the J��l�nic� Mo�nt�ins �t the �or�er �etween �l��ni� �n� the Rep��lic of M�ce�oni� (see M�ř�n 1939 �n� the type series in the N�tion�l M�se�m�� Pr�h�).

Fi�s 3–�� Zabrus skoupyi sp. nov.�� holotype. 3: �e�e���s: me�i�n lo�e �n� ri�ht p�r�mere�� l�ter�l view; 4: �e�e���s: �edian lo�e and para�ere�, dor�o-api�al view� ��r� �–�� Zabrus skoupyi sp. nov.�� holotyp�s. 3: �e�e���s: penis � pr�vá p�r�mer��� �oční� pohle�; 4: �e�e���s: penis � p�r�mery�� �orso�pikální� pohle�.

Page 7: Zabrus Pelor skoupyi sp. nov. (Coleoptera: Carabidae) from ... · 22 T XONOMIC P RT Zabrus (Pelor) skoupyi sp. nov. (Fi s 1–5 10 11) Type material. olotype:: ♂ l elle “Gr. M

2�

The t�xonomy of B�lk�n species of the s���en�s Pelor i� ver� �o�pli�ated� T��e �tat�� of �an� ����pe�ie� i� �n�lear a� t��ere i� not an a���rate de��ription of varia�ilit� a�ro�� t��e di�tri��tion of t��e �pe�ie�� T��at i� w��� I �o�tl� do not ��e t��e ����pe�ie� na�e� of pol�t�pi� �pe�ie� in t��i� �t�d��Name derivation. De�ic�te��� with �re�t ple�s�re�� to Dr. Vl��imí�r Sko�pý�� Žilin� (Czech �ep��li�), one of �olle�tor� of t��e new �pe�ie� and �pe�iali�t in Zabrus�

�C�NOW��DGM�NTS. I �m o�li�e� to S. Bene�ikt (Plzeň) for provi�in� his m�teri�l�� �n� �ll collectors mentione� in M�teri�l �n� metho�s for the kin� lo�n of specimens of the �en�s Zabrus from their collections. I th�nk V. Sko�pý (Žilin�) for � �isc�ssion on some pro�lems of t�xonomy�� �n� J. Růžičk� (Czech University of �ife Sciences�� Pr�h�) for technic�l help with the m�n�script. Speci�l th�nks �o to J. Velc (�l��no) for �r�win� the ����res.

Fi�s 5–11� Zabrus �pp� �–9: �e�e���s: me�i�n lo�e (�orso-�pic�l view) �n� ri�ht p�r�mere (l�ter�l view); 10�� 11: �t�l��, ventral view� �, 10, 11: Z. skoupyi �p� nov� (�: parat�pe No� 1�� 10: parat�pe No� ��� 11: parat�pe No� 6)�� 6: Z. albanicus �pfel�eck�� Greece�� Noti� Pin�os mo�nt�in r�n�e ne�r Pertoúlion (W of Trí�k�l�); �: Z. brevicollis S���a��, Greece�� P�rn�ssos Oros Mo�nt�ins�� Ski centr�m; 8: Z. aetolus Sch��m�� Greece�� Vori� Pin�os mo�nt�in r�n�e�� ��tár� P�ss; 9: Z. peristericus �pfel�eck�� Greece�� Tso�merk� Mo�nt�ins ne�r �th�mánion.��r� �–11� Zabrus �pp� �–9: �e�e���s: penis (�orso�pikální� pohle�) � pr�vá p�r�mer� (�oční� pohle�); 10�� 11: styl�s�� ventrální� pohle�. 5�� 10�� 11: Z. skoupyi sp. nov. (5: p�r�typ�s č. 1; 10: p�r�typ�s č. 5; 11: p�r�typ�s č. 6); 6: Z. alba�alba�nicus �pfel�eck�� Řecko�� horské pásmo Noti� Pin�os pr. Pertoúlion (záp��ně o� Trí�k�ly); �: Z. brevicollis S���a��, Řecko�� pohoří� P�rn�ssos Oros�� Ski centr�m; 8: Z. aetolus Sch��m�� Řecko�� horské pásmo Vori� Pin�os�� průsmyk ��tár�; 9: Z. peristericus �pfel�eck�� Řecko�� pohoří� Tso�merk� pr. �th�mánion.

Page 8: Zabrus Pelor skoupyi sp. nov. (Coleoptera: Carabidae) from ... · 22 T XONOMIC P RT Zabrus (Pelor) skoupyi sp. nov. (Fi s 1–5 10 11) Type material. olotype:: ♂ l elle “Gr. M

28

R�F�R�NC�S

�PF��B�C� V. 1904: Die Käferfauna der Balkanhalbinsel, mit Berücksichtigung Klein�Asiens und der Insel Kreta. Erster Band: Familienreihe Caraboidea. �� Friedländer �nd So��n, Berlin, �22 pp�

FR�UD� �. 1989: Revision �er z�r (?) G�tt�n� “���r�s” Cl�irville 1806 �eh�ren�en�rten mit Bestimm�n�st��ellen“���r�s” Cl�irville 1806 �eh�ren�en �rten mit Bestimm�n�st��ellen (�ol� �ara�idae)� �� Teil��� Teil� Atti dei Museo Civico di Storia Naturale di Trieste 42: �1–1���

G�NG�B�U�R �. 1915: Revision �er G�tt�n� ���r�s Cl�irv. Münchener Koleopterologische Zeitschrift 4: 8�–1���

GUÉORGUI�V B. 200�: 9. Bio�eo�r�phy of the �n�emic C�r��i��e (Coleopter�) in the Centr�l �n� ��stern B�l-Bio�eo�r�phy of the �n�emic C�r��i��e (Coleopter�) in the Centr�l �n� ��stern B�l-k�n Penins�l�. Pp. 29�–356. In: F�T V. & POPOV �. (e�s):& POPOV �. (e�s): Biogeography and Ecology of Bulgaria. Sprin�er�� Dordre���t, 68� pp�

�I��� F. 1981: Die C�r��i��e einer S�mmelreise n�ch M�ze�onien (Insect�: Coleopter�). Acta Musei Macedonici Scientiarum Naturalium 16: �1–101�

M�Ř�N J. 1939: �oo�eo�r���cká � system�tická st��ie o C�r��i�ech Golešnice-pl�niny v jižní� J��osl�vii. Die C�r��i�enf��n� �er Golešnic�-Pl�nin�. Sborník Entomologického Oddělení Národního Musea v Praze 17: 1��–1�0 (in ��e��� and �er�an)�

S�RR�NO J. & �ND�J�R �. 2003: S��tri�e ���rin� Bonelli�� 1810. Pp. 568–5�3. In: ��B� I. & SM�T�N� �.& �ND�J�R �. 2003: S��tri�e ���rin� Bonelli�� 1810. Pp. 568–5�3. In: ��B� I. & SM�T�N� �.–5�3. In: ��B� I. & SM�T�N� �.5�3. In: ��B� I. & SM�T�N� �. (ed�): Catalogue of Palaearctic Coleoptera. Volume 1. Archostemata – Myxophaga – Adephaga� �pollo Books�� Stenstr�p�� 819 pp.

SOU�RN

V práci je popsán nový �r�h střevlí�čk� Zabrus (Pelor) skoupyi sp. nov. ze severozáp��ní�ho Řeck�. Žije v pohoří� Gr�mmos n� �lpinských lo�kách v n��mořské výšce 2000–2200 metrů.–2200 metrů.2200 metrů. Nový �r�h �yl op�kov�ně s�í�rán v �lí�zkosti sněhových polí�.

Dos�� �ylo známo z pevninské části severozáp��ní�ho Řeck� osm t�xonů (�r�hů ne�o po��r�hů) v po�ro�� Pelor Bonelli, 1810 (rod Zabrus �lairville, 1806): Z. aetolus matejkai M�ř�n�� 1940 (pohoří� Vermion Oros); Z. aetolus purkynei M�ř�n�� 1940 (typová lok�lit� n� hoře ��jm�kč�l�n n� společné hr�nici Řeck� � Rep��liky M�ke�onie); Z. brevicollis S���a��, 185� (severozáp��ní� � stře�ní� Řecko); Z. ganglbaueri ganglbaueri �pfel�eck�� 1906 (�l�ánie � severozáp��ní� Řecko); Z. graecus subtilis Sch��m�� 1862 (severozáp��ní� � stře�ní� Řecko); Z. incrassatus (�hrens�� 1814) (B�lkánský poloostrov); Z. peristericus �pfel�eck�� 1901 (pohoří� ��kmos Oros) � a Z. spinipes insignis G. Müller�� 1932 (Řecko).

Z. (P.) skoupyi sp. nov. je ch�r�kteristický �r�h s je�inečným tv�rem těl�. O� všech �r�hů po�ro�� Pelor �os�� známých ze severozáp��ní�ho Řeck� je sn��no rozezn�telný pro svůj velmi o�lišný vzhle� � neo�yčejně úzký postr�nní� lem ští�t�. Nejpo�o�nější� �r�h Z. (P.) peristericus se liší� tv�rem ští�t��� který je pře� z��ní�mi rohy ví�ce zúžený � je nejširší� pře� stře�em ne�o �prostře��� postr�nní�mi okr�ji ští�t��� které jso� pře� z��ní�mi rohy sl��ě vykrojené �ž rovné�� tv�rem �e�e��� (špice penis� Z. peristericus je širší�) � větší� průměrno� velikostí�. Ost�tní� �r�hy m�jí� ští�t � krovky širší��� č�sto velmi výr�zně. D�lší� rozlišov�cí� zn�ky jso� �ve�eny v �n�lickém text�.

T�xonomie ��lkánských �r�hů po�ro�� Pelor je velmi složitá. St�t�s mnoh� po��r�hů je nej�sný�� protože chy�í� �ůkl��né popisy v�ri��ility v rámci rozší�ření� �r�hů. � těchto �ůvo�ů ��tor této st��ie většino� nepo�ží�vá po��r�hová jmén� � polytypických �r�hů.

Práce o�s�h�je nové ú��je o rozší�ření� �r�hů Z. (P.) aetolus S���a��, 186� �� lat�, Z. (P.) albanicus �pfel�eck�� 1904 s. l�t.�� Z. (P.) brevicollis S���a��, 18�� a Z. (P.) peristericusperistericus �pfel-�eck�� 1901 v severozáp��ní�m Řeck�.� Z. (P.) albanicus s. l�t. je nový �r�h pro Řecko.


Recommended