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One of the large, detailed illustrations in Andreas

Vesalius's De humani corporis fabrica, 1543

AnatomyFrom Wikipedia, the free encyclopedia

For the anatomy of humans, see Human anatomy. For the anatomy of plants, see Plant anatomy. For

other uses, see Anatomy (disambiguation).

Anatomy (from the Greek ἀνατέμνω anatemnō "I cut up,cut open" from ἀνά ana "on, upon", and τέμνω temnō "Icut") is the study of the structure of animals and their parts. Insome of its facets, anatomy is closely related to embryology,

comparative anatomy and comparative embryology,[1]

through common roots in evolution. Human anatomy isimportant in medicine.

The discipline of anatomy is subdivided into gross (ormacroscopic) anatomy and microscopic anatomy. Grossanatomy is the study of structures that can, when suitablypresented or dissected, be seen by unaided vision with thenaked eye. Microscopic anatomy is the study of structures ona microscopic scale, including histology (the study of tissues)and cytology (the study of cells).

The history of anatomy is characterized by a continualdevelopment in understanding of the functions of the organsand structures of the human body. Methods have alsoimproved dramatically, advancing from examination ofanimals through dissection of cadavers (dead human bodies)to 20th century techniques including X-ray, ultrasound, andmagnetic resonance imaging.

Contents

1 Overview

2 History

2.1 Ancient

2.2 Early modern

2.3 Late modern

3 Animal tissues

4 Human anatomy

4.1 Research and practice

5 Vertebrate anatomy

5.1 Fish anatomy

5.2 Amphibian anatomy

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5.3 Reptile anatomy

5.4 Bird anatomy

5.5 Mammal anatomy

6 Invertebrate anatomy

6.1 Insect anatomy

7 Other branches of anatomy

8 See also

9 Notes

10 Bibliography

11 External links

Overview

Anatomy is the scientific study of the structure of living things including their systems, organs, and tissues. It includesthe appearance and position of the various parts, the materials from which they are composed, their locations andtheir relationships with other parts. Anatomy is quite distinct from physiology and biochemistry, which deal with thefunctions of those parts. For example, an anatomist is concerned with the shape, size, position, structure, bloodsupply and enervation of an organ such as the liver, while a physiologist is interested in the production of bile and

the role of the liver in nutrition and regulating body functions.[2]

The discipline of anatomy can be subdivided into a number of branches including gross or macroscopic anatomy

and microscopic anatomy.[3] Gross anatomy is the study of structures that are large enough to be seen with thenaked eye. Microscopic anatomy is the study of structures on a microscopic scale, including histology (the study of

tissues), and embryology (the study of the human organism in its immature condition).[1]

Anatomy can be studied using both invasive and non-invasive methods with the goal of obtaining information about

the structure and organization of organs and systems.[1] Methods used include dissection, in which the body isopened and its organs studied, and endoscopy, in which a video camera-equipped instrument is inserted through asmall incision in the body wall and used to explore the internal organs and other structures. Angiography using X-

rays or magnetic resonance angiography are methods to visualize blood vessels.[4][5][6][7]

The term "anatomy" is commonly taken to refer to human anatomy. However, substantially the same structures andtissues are found throughout the animal kingdom and the term also includes the anatomy of other animals. The

structure and tissues of plants are of a dissimilar nature and they are studied in plant anatomy.[2]

History

Main article: History of anatomy

Ancient

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Michiel Jansz van Mierevelt –

Anatomy lesson of Dr. Willem van

der Meer, 1617

In 1600 BCE, the Edwin Smith Papyrus, an Ancient Egyptian medical text, described the heart, its vessels, liver,spleen, kidneys, hypothalamus, uterus and bladder, and showed the blood vessels diverging from the heart. TheEbers Papyrus (c. 1550 BCE) features a "treatise on the heart", with vessels carrying all the body's fluids to or from

every member of the body.[8]

The anatomy of the muscles and skeleton is described in the Hippocratic Corpus, an Ancient Greek medical work

written by unknown authors.[9] Aristotle described vertebrate anatomy based on animal dissection. Praxagorasidentified the difference between arteries and veins. Also in the 4th century BCE, Herophilos and Erasistratusproduced more accurate anatomical descriptions based on vivisection of criminals in Alexandria during the

Ptolemaic dynasty.[10][11]

In the 2nd century Galen, a Roman physician, surgeon and philosopher, wrote the final and highly influential

anatomy treatise of ancient times.[12] He compiled existing knowledge and studied organs through vivisection of

animals.[13] Galen's drawings, based mostly on dog anatomy, became effectively the only anatomical textbook for

the next thousand years.[14] His work was known to Renaissance doctors only through Islamic Golden Age

medicine until it was translated from the Greek some time in the 15th century.[14]

Early modern

Between 1275 and 1326, the anatomists Mondino de Luzzi, AlessandroAchillini and Antonio Benivieni at Bologna carried out the first systematic

human dissections since ancient times.[15][16][17] Mondino's Anatomy of1316 was the first textbook in the mediaeval rediscovery of humananatomy. It describes the body in the order followed in Mondino'sdissections, starting with the abdomen, then the thorax, then the head and

limbs. It was the standard anatomy textbook for the next century.[14]

Andreas Vesalius (1514–1564) (Latinized from Andries van Wezel),professor of anatomy at the University of Padua, is considered the

founder of modern human anatomy.[18] Originally from Brabant, Vesaliuspublished the influential book De humani corporis fabrica ("thestructure of the human body"), a large format book in seven volumes, in

1543.[19] The accurate and intricately detailed illustrations, often in allegorical poses against Italianate landscapes,

are thought to have been made by the artist Jan van Calcar, a pupil of Titian.[14][20]

The artist Leonardo da Vinci (1452–1519) was trained in anatomy by Andrea del Verrocchio.[14] He made use ofhis anatomical knowledge in his artwork, making many sketches of skeletal structures, muscles and organs of

humans and other vertebrates which he dissected.[14][21]

In England, anatomy was the subject of the first public lectures given in any science; these were given by theCompany of Barbers and Surgeons in the 16th century, joined in 1583 by the Lumleian lectures in surgery at the

Royal College of Physicians.[22]

Late modern

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Sagittal section of the head as seen by

a MRI scan

Further information: History of anatomy in the 19th century

In the United States, medical schools began to be set up towards the end of the 18th century. Classes in anatomyneeded a continual stream of cadavers for dissection and these were difficult to obtain. Philadelphia, Baltimore andNew York were all renowned for body snatching activity as criminals raided graveyards at night, removing newly

buried corpses from their coffins.[23] A similar problem existed in Britain where demand for bodies became so great

that grave-raiding and even anatomy murder were practised to obtain cadavers.[24] Some graveyards were in

consequence protected with watchtowers. The practice was halted in Britain by the Anatomy Act of 1832,[25][26]

while in the United States, similar legislation was enacted after the physician William S. Forbes of Jefferson MedicalCollege was found guilty in 1882 of "complicity with resurrectionists in the despoliation of graves in Lebanon

Cemetery".[27]

The teaching of anatomy in Britain was transformed by Sir John Struthers, Regius Professor of Anatomy at theUniversity of Aberdeen from 1863 to 1889. He was responsible for setting up the system of three years of "pre-clinical" academic teaching in the sciences underlying medicine, including especially anatomy. This system lasted untilthe reform of medical training in 1993 and 2003. As well as teaching, he collected many vertebrate skeletons for hismuseum of comparative anatomy, published over 70 research papers, and became famous for his public dissection

of the Tay Whale.[28][29] From 1822 the Royal College of Surgeons regulated the teaching of anatomy in medical

schools.[30] Medical museums provided examples in comparative anatomy, and were often used in teaching.[31]

Ignaz Semmelweis investigated puerperal fever and he discovered how it was caused. He noticed that thefrequently fatal fever occurred more often in mothers examined by medical students than by midwives. The studentswent from the dissecting room to the hospital ward and examined women in childbirth. Semmelweis showed thatwhen the trainees washed their hands in chlorinated lime before each clinical examination, the incidence of puerperal

fever among the mothers could be reduced dramatically.[32]

Before the era of modern medical procedures, the main means forstudying the internal structure of the body were palpation and dissection.It was the advent of microscopy that opened up an understanding of thebuilding blocks that constituted living tissues. Technical advances in thedevelopment of achromatic lenses increased the resolving power of themicroscope and around 1839, Matthias Jakob Schleiden and TheodorSchwann identified that cells were the fundamental unit of organization ofall living things. Study of small structures involved passing light throughthem and the microtome was invented to provide sufficiently thin slices oftissue to examine. Staining techniques using artificial dyes wereestablished to help distinguish between different types of cell. The fieldsof cytology and histology developed from here in the late 19th

century.[33] The invention of the electron microscope brought a greatadvance in resolution power and allowed research into the ultrastructureof cells and the organelles and other structures within them. About thesame time, in the 1950s, the use of X-ray diffraction for studying thecrystal structures of proteins, nucleic acids and other biological molecules

gave rise to a new field of molecular anatomy.[33]

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A diagram of the layers of the human

epidermis

Short wavelength electromagnetic radiation such as X-rays can be passed through the body and are used in medicalradiography to view interior structures which have different degrees of opaqueness. Nowadays, modern techniquessuch as magnetic resonance imaging, computed tomography, fluoroscopy and ultrasound imaging have enabledresearchers and practitioners to examine organs, living or dead, in unprecedented detail. They are used fordiagnostic and therapeutic purposes and provide information on the internal structures and organs of the body to a

degree far beyond the imagination of earlier generations.[34]

Animal tissues

The kingdom Animalia contains multicellular organisms that are heterotrophic and motile (although some havesecondarily adopted a sessile lifestyle). Most animals have bodies differentiated into separate tissues and theseanimals are known as metazoans or eumetazoans. They have an internal digestive chamber, with one or twoopenings, the gametes are produced in multicellular sex organs and the zygotes include a blastula stage in theirembryonic development. Metazoans do not include the sponges, phylum Porifera, or the Placozoans which have

undifferentiated cells.[35]

Animal cells do not have a cell wall and do not contain chloroplasts. If a vacuole is present, it is smaller than thatfound in a plant cell. The body tissues are composed of a great variety of cells such as muscle, nerve and bloodcells. Each typically has a cell membrane formed of phospholipids, cytoplasm and a nucleus. Animal tissues can begrouped into four basic types: connective, epithelial, muscle and nervous. Even the simplest invertebrates contain at

least two types and body structures and organs of higher animals are formed from multiple tissue types.[36]

Connective tissues are fibrous tissues and are made up of cells scattered among non-living material which is calledthe extracellular matrix. Connective tissue gives shape to organs and holds them in place. The main types are looseconnective tissue, adipose tissue, fibrous connective tissue, blood, cartilage and bone. The extracellular matrixcontains proteins, one of the commonest of which is collagen, and plays an important part in organizing andmaintaining tissues. The matrix can be modified to form a skeleton to support or protect the body. An exoskeletonis a thickened, rigid cuticle which is stiffened by mineralization, as in crustaceans or by the cross-linking of its

proteins as in insects. An endoskeleton is internal and is present in all higher animals and many lower ones.[36]

Epithelial tissue is composed of closely packed cells with little intercellularspace. They can be squamous (flat), cuboidal or columnar and rest on abasal lamina, a membrane composed of extracellular matrix secreted bythe epithelial cells. The cells are bound closely to each other by celladhesion molecules. One or more layers of epithelial cells line the cavitiesand surfaces of structures throughout the body including the externalsurface, the respiratory surface and the gut. The epithelial cells on theexternal surface of the body typically secrete an extracellular matrix in theform of a cuticle. In simple animals this may just be a coat of

glycoproteins.[36] In more advanced animals, many glands are formed of

epithelial cells.[37]

Muscle cells (myocytes) form the active contractile tissue of the body.Muscle tissue functions to produce force and cause motion, eitherlocomotion or movement within internal organs. Muscle is formed ofcontractile filaments and is separated into three types; smooth muscle,

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Two facing pages of the 1543

Epitome by Vesalius, with woodcuts

of naked male and female figures

skeletal muscle and obliquely striated muscle. Smooth muscle has no striations when examined microscopically. Itcontracts slowly but maintains contractibility over a wide range of stretch lengths. It is found in such organs as seaanemone tentacles and the body wall of sea cucumbers. Cross-striated muscle contracts rapidly but has a limitedrange of extension. It is found in the movement of appendages and jaws. Obliquely striated muscle is intermediatebetween the other two. The filaments are staggered and this is the type of muscle found in earthworms that can

extend slowly or make rapid contractions.[38] In higher animals striated muscles occur in bundles attached to boneto provide movement and are often arranged in antagonistic sets. Smooth muscle is found in the walls of the uterus,bladder, intestines, stomach, esophagus, respiratory airways, and blood vessels. Cardiac muscle is found only in theheart, allowing it to contract and pump blood round the body.

Nervous tissue is composed of many nerve cells known as neurons which transmit information. In some slow-moving radially symmetrical marine animals such as ctenophores and cnidarians (including sea anemones andjellyfish), the nerves form a nerve net, but in most animals they are organized longitudinally into bundles. In simpleanimals, receptor neurons in the body wall cause a local reaction to a stimulus. In more complex animals,specialised receptor cells such as chemoreceptors and photoreceptors are found in groups and send messages

along neurons to other parts of the organism. Neurons can be connected together in ganglia.[39] In higher animals,specialized receptors are the basis of sense organs and there is a central nervous system (brain and spinal cord) anda peripheral nervous system. The latter consists of sensory nerves that transmit information from sense organs and

motor nerves that influence target organs.[40][41] The peripheral nervous system is divided into the somatic nervoussystem which conveys sensation and controls voluntary muscle, and the autonomic nervous system which

involuntarily controls smooth muscle, certain glands, and internal organs such as the stomach.[42]

Human anatomy

Main article: Human anatomy

Research and practice

Human anatomy, including gross human anatomy and histology, isprimarily the scientific study of the morphology of the adult human body.It differs from physiology in that anatomy is about the morphology ofbiological structures, while physiology is the way those structures actually

work.[1]

Generally, students of certain biological sciences, paramedics,prosthetists and orthotists, physiotherapists, occupational therapists,nurses, and medical students learn gross anatomy and microscopicanatomy from anatomical models, skeletons, textbooks, diagrams,photographs, lectures and tutorials, and in addition, medical studentsgenerally also learn gross anatomy through practical experience ofdissection and inspection of cadavers. The study of microscopic anatomy(or histology) can be aided by practical experience examining histological

preparations (or slides) under a microscope. [43]

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Anatomical chart by

Vesalius, Epitome, 1543

Human anatomy, physiology and biochemistry are complementary basic medical sciences, which are generallytaught to medical students in their first year at medical school. Human anatomycan be taught regionally or systemically; that is, respectively, studying anatomy bybodily regions such as the head and chest, or studying by specific systems, such

as the nervous or respiratory systems.[1] The major anatomy textbook, Gray'sAnatomy, has been reorganized from a systems format to a regional format, in line

with modern teaching methods.[44][45] A thorough working knowledge ofanatomy is required by physicians, especially surgeons and doctors working in

some diagnostic specialties, such as histopathology and radiology. [46]

Academic human anatomists are usually employed by universities, medicalschools or teaching hospitals. They are often involved in teaching anatomy, and

research into certain systems, organs, tissues or cells.[46]

Vertebrate anatomy

All vertebrates are built along the basic chordate body plan: a stiff rod running

through the length of the animal (vertebral column or notochord),[47] with ahollow tube of nervous tissue, the spinal cord above it and the gastrointestinal tract below. Nervous tissue is derivedfrom the ectoderm, connective tissues are derived from mesoderm, and gut is derived from the endoderm. At theposterior end is a tail which continues the spinal cord and vertebrae but not the gut. The mouth is found at the

anterior end of the animal, and the anus at the base of the tail.[48] The defining characteristic of a vertebrate is thevertebral column, in which the notochord has been replaced by a segmented series of stiffer elements (vertebrae)separated by mobile joints (intervertebral discs), which each contain a soft central core (nucleus pulposus), derivedfrom the notochord of the embryo. However, a few vertebrates have secondarily lost this anatomy, retaining the

notochord into adulthood. These include the sturgeon and the coelacanth.[49] Jawed vertebrates are typified bypaired appendages, fins or legs, which may be secondarily lost. The limbs of vertebrates are considered to behomologous because the same underlying skeletal structure was inherited from their last common ancestor. This is

one of the arguments put forward by Charles Darwin to support his theory of evolution.[50]

Fish anatomy

Main article: Fish anatomy

The body of a fish is divided into a head, trunk and tail, although the divisions between the three are not alwaysexternally visible. The skeleton, which forms the support structure inside the fish, is either made of cartilage(cartilaginous fish) or bone (bony fishes). The main skeletal element is the vertebral column, composed ofarticulating vertebrae which are lightweight yet strong. The ribs attach to the spine and there are no limbs or limbgirdles. The main external features of the fish, the fins, are composed of bony spines and soft rays and, with theexception of the caudal fins, have no direct connection with the spine. They are supported by the muscles which

compose the main part of the trunk.[51] The heart has two chambers and pumps the blood through the respiratory

surfaces of the gills and on round the body in a single circulatory loop.[52] The eyes are adapted for seeing

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Skeleton of a butterfly fish showing

the vertebral column and fin rays

Skeleton of Surinam horned frog

(Ceratophrys cornuta)

underwater and have only local vision. There is an inner ear but noexternal or middle ear. Low frequency vibrations are detected by thelateral line system on the flanks which responds to nearby movements

and to changes in water pressure.[51]

Sharks and rays are basal fish with numerous primitive anatomicalfeatures similar to those of ancient fish, including skeletons composed ofcartilage. Their bodies tend to be dorso-ventrally flattened, they usuallyhave five pairs of gill slits and a large mouth set on the underside of thehead. The dermis is covered with separate dermal placoid scales. Theyhave a cloaca into which the urinary and genital passages open, but not aswim bladder. Cartilaginous fish produce a small number of large, yolkyeggs. Some species are ovoviviparous and the young develop internally

but others are oviparous and the larvae develop externally in egg cases.[53]

The bony fish lineage shows more derived anatomical traits, often with major evolutionary changes from the featuresof ancient fish. They have a bony skeleton, are generally laterally flattened, have five pairs of gills protected by anoperculum, and a mouth at or near the tip of the snout. The dermis is covered with overlapping scales. Bony fishhave a swim bladder which helps them maintain a constant depth in the water column, but not a cloaca. They mostly

spawn a large number of small eggs with little yolk which they broadcast into the water column.[53]

Amphibian anatomy

Main article: Amphibian anatomy

Amphibians are a class of animals comprising frogs, salamanders andcaecilians. They are tetrapods, but the caecilians and a few species ofsalamander have either no limbs or their limbs are much reduced in size.Their main bones are hollow and lightweight and are fully ossified and thevertebrae interlock with each other and have articular processes. Theirribs are usually short and may be fused to the vertebrae. Their skulls aremostly broad and short, and are often incompletely ossified. Their skincontains little keratin and lacks scales, but contains many mucous glandsand in some species, poison glands. The hearts of amphibians have threechambers, two atria and one ventricle. They have a urinary bladder andnitrogenous waste products are excreted primarily as urea. Amphibiansbreathe by means of a pump action in which air is first drawn into thebuccopharyngeal region through the nostrils. These are then closed and

the air is forced into the lungs by contraction of the throat.[54] They

supplement this with gas exchange through the skin which needs to be kept moist.[55]

In frogs the pelvic girdle is robust and the hind legs are much longer and stronger than the forelimbs. The feet havefour or five digits and the toes are often webbed for swimming or have suction pads for climbing. Frogs have largeeyes and no tail. Salamanders resemble lizards in appearance; their short legs project sideways, the belly is close toor in contact with the ground and they have a long tail. Caecilians superficially resemble earthworms and arelimbless. They burrow by means of zones of muscle contractions which move along the body and they swim by

undulating their body from side to side.[56]

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Skeleton of a snake, drawn by Richard

Lydekker, 1896

Reptile anatomy

Main article: Reptile anatomy

Reptiles are a class of animals comprising turtles, tuataras, lizards,snakes and crocodiles. They are tetrapods, but the snakes and afew species of lizards either have no limbs or their limbs are muchreduced in size. Their bones are better ossified and their skeletonsstronger than those of amphibians. The teeth are conical andmostly uniform in size. The surface cells of the epidermis aremodified into horny scales which create a waterproof layer.Reptiles are unable to use their skin for respiration as doamphibians and have a more efficient respiratory system drawingair into their lungs by expanding their chest walls. The heartresembles that of the amphibian but there is a septum which morecompletely separates the oxygenated and deoxygenated

bloodstreams. The reproductive system is designed for internal fertilisation, with a copulatory organ present in mostspecies. The eggs are surrounded by amniotic membranes which prevents them from drying out and are laid on

land, or develop internally in some species. The bladder is small as nitrogenous waste is excreted as uric acid.[57]

Turtles are notable for their protective shells. They have an inflexible trunk encased in a horny carapace above anda plastron below. These are formed from bony plates embedded in the dermis which are overlain by horny onesand are partially fused with the ribs and spine. The neck is long and flexible and the head and the legs can be drawnback inside the shell. Turtles are vegetarians and the typical reptile teeth have been replaced by sharp, horny plates.

In aquatic species, the front legs are modified into flippers.[58]

Tuataras superficially resemble lizards but the lineages diverged in the Triassic Period. There is one living species,Sphenodon punctatus. The skull has two openings (fenestrae) on either side and the jaw is rigidly attached to theskull. There is one row of teeth in the lower jaw and this fits between the two rows in the upper jaw when theanimal chews. The teeth are merely projections of bony material from the jaw and eventually wear down. The brainand heart are more primitive than is the case in other reptiles and the lungs have a single chamber and lack bronchi.

The tuatara has a well-developed parietal eye on its forehead.[58]

Lizards have skulls with only one fenestra on each side, the lower bar of bone below the second fenestra havingbeen lost. This results in the jaws being less rigidly attached which allows the mouth to open wider. Lizards aremostly quadrupeds, with the trunk held off the ground by short, sideways-facing legs, but a few species have nolimbs and resemble snakes. Lizards have moveable eyelids, eardrums are present and some species have a central

parietal eye.[58]

Snakes are closely related to lizards, having branched off from a common ancestral lineage during the CretaceousPeriod, and they share many of the same features. The skeleton consists of a skull, a hyoid bone, spine and ribsthough a few species retain a vestige of the pelvis and rear limbs in the form of pelvic spurs. The bar under thesecond fenestra has also been lost and the jaws have extreme flexibility allowing the snake to swallow its preywhole. Snakes lack moveable eyelids, the eyes being covered by transparent "spectacle" scales. They do not haveeardrums but can detect ground vibrations through the bones of their skull. Their forked tongues are used as organsof taste and smell and some species have sensory pits on their heads enabling them to locate warm-blooded

prey.[59]

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Skeleton of the Common Kingfisher

(Alcedo atthis)

Crocodilians are large, low-slung aquatic reptiles with long snouts and large numbers of teeth. The head and trunkare dorso-ventrally flattened and the tail is laterally compressed. It undulates from side to side to force the animalthrough the water when swimming. The tough keratinised scales provide body armour and some are fused to theskull. The nostrils, eyes and ears are elevated above the top of the flat head enabling them to remain above thesurface of the water when the animal is floating. Valves seal the nostrils and ears when it is submerged. Unlike otherreptiles, crocodilians have hearts with four chambers allowing complete separation of oxygenated and

deoxygenated blood.[60]

Bird anatomy

Main article: Bird anatomy

Birds are tetrapods but though their hind limbs are used for walking orhopping, their front limbs are wings covered with feathers and adaptedfor flight. Birds are endothermic, have a high metabolic rate, a lightskeletal system and powerful muscles. The long bones are thin, hollowand very light. Air sac extensions from the lungs occupy the centre ofsome bones. The sternum is wide and usually has a keel and the caudalvertebrae are fused. There are no teeth and the narrow jaws are adaptedinto a horn-covered beak. The eyes are relatively large, particularly innocturnal species such as owls. They face forwards in predators and

sideways in ducks.[61]

The feathers are outgrowths of the epidermis and are found in localizedbands from where they fan out over the skin. Large flight feathers arefound on the wings and tail, contour feathers cover the bird's surface and fine down occurs on young birds andunder the contour feathers of water birds. The only cutaneous gland is the single uropygial gland near the base ofthe tail. This produces an oily secretion that waterproofs the feathers when the bird preens. There are scales on the

legs and feet and claws on the tips of the toes.[61]

Mammal anatomy

Main article: Mammal anatomy

Mammals are a diverse class of animals, mostly terrestrial but some are aquatic and others have evolved flapping orgliding flight. They mostly have four limbs but some aquatic mammals have no limbs or limbs modified into fins andthe forelimbs of bats are modified into wings. The legs of most mammals are situated below the trunk, which is heldwell clear of the ground. The bones of mammals are well ossified and their teeth, which are usually differentiated,are coated in a layer of prismatic enamel. The teeth are shed once (milk teeth) during the animal's lifetime or not atall, as is the case in cetaceans. Mammals have three bones in the middle ear and a cochlea in the inner ear. They areclothed in hair and their skin contains glands which secrete sweat. Some of these glands are specialised asmammary glands, producing milk to feed the young. Mammals breathe with lungs and have a muscular diaphragmseparating the thorax from the abdomen which helps them draw air into the lungs. The mammalian heart has fourchambers and oxygenated and deoxygenated blood are kept entirely separate. Nitrogenous waste is excreted

primarily as urea.[62]

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Skeletons of a Great Dane and a

Chihuahua

Mammals are amniotes, and most are viviparous, giving birth to liveyoung. The exception to this are the egg-laying monotremes, the platypusand the echidnas of Australia. Most other mammals have a placentathrough which the developing foetus obtains nourishment, but inmarsupials, the foetal stage is very short and the immature young is bornand finds its way to its mother's pouch where it latches on to a nipple and

completes its development.[62]

Invertebrate anatomy

Invertebrates constitute a vast array of living organisms ranging from thesimplest unicellular eukaryotes to such complex creatures as the octopus,lobster and dragonfly. By definition, none of them has a backbone. Thecells of single-cell protozoans have the same basic structure as those ofmulticellular animals but some parts are specialised into the equivalent of

tissues and organs. Locomotion is often provided by cilia or flagella or may proceed via the advance ofpseudopodia, food may be gathered by phagocytosis, energy needs may be supplied by photosynthesis and the cell

may be supported by an endoskeleton or an exoskeleton. Some protozoans can form multicellular colonies.[63]

Metazoans are multicellular organism, different groups of cells of which have separate functions. The most basictypes of metazoan tissues are epithelium and connective tissue, both of which are present in nearly all invertebrates.The outer surface of the epidermis is normally formed of epithelial cells and secretes an extracellular matrix whichprovides support to the organism. An endoskeleton derived from the mesoderm is present in echinoderms, spongesand some cephalopods. Exoskeletons are derived from the epidermis and is composed of chitin in arthropods(insects, spiders, ticks, shrimps, crabs, lobsters). Calcium carbonate constitutes the shells of molluscs, brachiopodsand some tube-building polychaete worms and silica forms the exoskeleton of the microscopic diatoms and

radiolaria.[64] Other invertebrates may have no rigid structures but the epidermis may secrete a variety of surfacecoatings such as the pinacoderm of sponges, the gelatinous cuticle of cnidarians (polyps, sea anemones, jellyfish)and the collagenous cuticle of annelids. The outer epithelial layer may include cells of several types including sensorycells, gland cells and stinging cells. There may also be protrusions such as microvilli, cilia, bristles, spines and

tubercles.[65]

Insect anatomy

Main article: Insect anatomy

Insects possess segmented bodies supported by a hard-jointed outer covering made mostly of chitin. The segments

of the body are organized into three distinct parts, a head, a thorax and an abdomen.[66] The head typically bears apair of sensory antennae, a pair of compound eyes, one to three simple eyes (ocelli) and three sets of modifiedappendages that form the mouthparts. The thorax has three pairs of segmented legs, one pair each for the threesegments that compose the thorax and one or two pairs of wings. The abdomen is composed of eleven segments,

some of which may be fused and houses the digestive, respiratory, excretory and reproductive systems.[67] There isconsiderable variations between species and many adaptations to the body parts, especially wings, legs, antennae

and mouthparts.[68]

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Other branches of anatomy

Superficial or surface anatomy is important as the study of anatomical landmarks that can be readily seen

from the exterior contours of the body.[1] It enables physicians or veterinary surgeons to gauge the position

and anatomy of the associated deeper structures. Superficial is a directional term that indicates that structures

are located relatively close to the surface of the body.[69]

Comparative anatomy relates to the comparison of anatomical structures (both gross and microscopic) in

different animals.[1]

Anthropological anatomy or physical anthropology relates to the comparison of the anatomy of different

races of humans.

Artistic anatomy relates to anatomic studies for artistic reasons.

See also

Acland's Video Atlas of Human Anatomy

American Association of Anatomists

Anatomical terminology

Anatomical terms of location

Anatomical terms of motion

Foundational Model of Anatomy

List of human anatomical features

List of human anatomical parts named after people

Outline of human anatomy

Human body

Notes

1. ̂a b c d e f g "Introduction page, "Anatomy of the Human Body". Henry Gray. 20th edition. 1918"

(https://web.archive.org/web/20070316005206/http://www.bartleby.com/107/1.html). Archived from the original

(http://www.bartleby.com/107/1.html) on 16 March 2007. Retrieved 19 March 2007.

2. ̂a b Bozman, E. F. (ed.) (1967). Everyman's Encyclopedia: Anatomy. J. M. Dent & Sons. p. 272.

ASIN B0066E44EC (https://www.amazon.com/dp/B0066E44EC).

3. ^ "Anatomy" (http://medical-dictionary.thefreedictionary.com/microscopic+anatomy). The Free Dictionary. Farlex.

2007. Retrieved 8 July 2013.

4. ^ Gribble, N.; Reynolds, K. (1993). "Use of Angiography to Outline the Cardiovascular Anatomy of the Sand Crab

Portunus pelagicus Linnaeus" (http://www.jstor.org/stable/1549093). Journal of Crustacean Biology 13 (4): 627–

637.

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5. ^ Benson, K. G.; Forrest, L. (1999). "Characterization of the Renal Portal System of the Common Green Iguana

(Iguana iguana) by Digital Subtraction Imaging". Journal of Zoo and Wildlife Medicine 30 (2): 235–241.

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(http://www.hopkinsmedicine.org/healthlibrary/test_procedures/cardiovascular/magnetic_resonance_angiography_

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Present. Harper Collins. pp. 49–50. ISBN 0-00-215173-1.

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pp. 419–427.

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vwUeCzCYKEiAexnoHYBA&redir_esc=y#v=onepage&q=Ptolemaic%20vivisection&f=false). Brill NV. p. 256.

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12. ^ Galen (April 1977). "Galen on the Affected Parts" (http://www.ncbi.nlm.nih.gov/pmc/articles/PMC1081972/).

Medical History 21 (2): 212. PMC 1081972 (https://www.ncbi.nlm.nih.gov/pmc/articles/PMC1081972).

13. ^ Brock, Arthur John Brock (translator), Introduction. Galen. On the Natural Faculties. Edinburgh, 1916.

14. ̂a b c d e f Boas, Marie (1970 (First published by Collins, 1962)). The Scientific Renaissance 1450–1630. Fontana.

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(http://books.google.com/books?id=ABbCI7z4UwMC). Norman. ISBN 978-0-930-40553-3.

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20. ^ O'Malley, C.D. Andreas Vesalius of Brussels, 1514–1564. Berkeley: University of California Press, 1964.

21. ^ Mason, Stephen F. (1962). A History of the Sciences. New York: Collier. p. 550.

22. ^ Boas, Marie (1970 (First published by Collins, 1962)). The Scientific Renaissance 1450–1630. Fontana. p. 229.

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America. Princeton, N.J.: Princeton University Press. ISBN 0-691-05925-X.

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24. ^ Rosner, Lisa. 2010. The Anatomy Murders. Being the True and Spectacular History of Edinburgh's Notorious

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40. ^ Johnston, T.B; Whillis, J, eds. (1944). Grey's Anatomy: Descriptive and Applied (28 ed.). Langmans. p. 1038.

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anatomy/nervous-system). Nervous System (4th ed.). Inkling. Retrieved 30 April 2014.

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Bibliography

Main article: Bibliography of anatomy

"Anatomy of the Human Body". 20th edition. 1918. Henry Gray (http://www.bartleby.com/107/)

External links

Anatomy (http://www.dmoz.org/Health/Medicine/Basic_Sciences/Anatomy) at DMOZ

Anatomy (http://www.bbc.co.uk/podcasts/series/iots/all#playepisode115), In Our Time. BBC Radio 4.

Melvin Bragg with guests Ruth Richardson, Andrew Cunningham and Harold Ellis.

Retrieved from "http://en.wikipedia.org/w/index.php?title=Anatomy&oldid=607067783"

Categories: Anatomy

This page was last modified on 4 May 2014 at 20:06.

Text is available under the Creative Commons Attribution-ShareAlike License; additional terms may apply.

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